624 lines
111 KiB
XML
624 lines
111 KiB
XML
<document id="8DD8BC8B8812B4144AF7F6B434ACE965" ID-DOI="10.11646/zootaxa.3682.1.6" ID-GBIF-Dataset="e8a0fb91-227c-4b35-92f4-a3b409035052" ID-ISSN="1175-5326" ID-Zenodo-Dep="218044" ID-ZooBank="1B811565-C78A-4CF2-9B89-723A20524280" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460641999739" checkinUser="plazi" docAuthor="Thomsen, Vibe G., Rho, Hyun Soo, Kim, Dongsung & Sørensen, Martin V." docDate="2013" docId="6F11A30112442231FF538EB8F488FBC2" docLanguage="en" docName="zt03682p142.pdf" docOrigin="Zootaxa 3682 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dracoderes nidhug Thomsen, Rho, Kim & Sørensen, 2013, new species" docType="treatment" docVersion="6" lastPageNumber="140" masterDocId="9328DB7912452236FFC48B29F60CFF95" masterDocTitle="A new species of Dracoderes (Kinorhyncha: Dracoderidae) from Korea provides further support for a dracoderid-homalorhagid relationship" masterLastPageNumber="142" masterPageNumber="133" pageNumber="134" updateTime="1698336042082" updateUser="plazi">
|
||
<mods:mods id="7455A41A0DFA91831379CF95E4F70E57" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="D26D7FA1EC59A7E02C70C63192797498">
|
||
<mods:title id="36C4FF69FC91E94F7BC36AE8B067E9F0">A new species of Dracoderes (Kinorhyncha: Dracoderidae) from Korea provides further support for a dracoderid-homalorhagid relationship</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="377366FE905CBF53C44036CAB22A7434" type="personal">
|
||
<mods:role id="0F1F628A69039B333D9D059670BEDD5E">
|
||
<mods:roleTerm id="1ED057A6D45DDD7F2FA94B7435D3CF9B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A4BD9BA202B496D41C5434B1D0C2AD86">Thomsen, Vibe G.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F524BF55393D8D573D4F8E9FA9AAA61E" type="personal">
|
||
<mods:role id="4B8B50D190193C6B37324D1646363223">
|
||
<mods:roleTerm id="FBFC6B7E1DFAD6A26970BC47DADC5EF9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="65272F7CCED9ABF72F7FF2CD529B0938">Rho, Hyun Soo</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="856FC156A672B8BBBD09E436AE8A6F7B" type="personal">
|
||
<mods:role id="7E4DC35ADA5F8BD8FC078870BDAE542E">
|
||
<mods:roleTerm id="A2995228030B2544F90BE8CEF5013B54">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E59A6E499DDFE7CFF73A87999E902F92">Kim, Dongsung</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A1ED64E53E543BAD3AA18358AE1BE000" type="personal">
|
||
<mods:role id="B8A739E116D50958F4E50B5048A7D674">
|
||
<mods:roleTerm id="FA4D4E00F59EC6AF859336D573DCFC61">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A46D7BD26CF56A766C74157F7134A271">Sørensen, Martin V.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="8006AE89A1813C9BE1B5DA520040C574">text</mods:typeOfResource>
|
||
<mods:relatedItem id="D6088602248923D804D0586A22995958" type="host">
|
||
<mods:titleInfo id="0095F390535EC39DF86FA3C4033BF92D">
|
||
<mods:title id="D8CADB6DF04E77660ED08D295258797D">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="A245AEC60CCCFAD45F53DC63BDF38078">
|
||
<mods:date id="5EA1A2D0E7BA5835BB3BA02B2EC175AB">2013</mods:date>
|
||
<mods:detail id="A4A1A16715FE895BFCCFA78F1827D15C" type="volume">
|
||
<mods:number id="2FD1500E26C210DB19613CB4795611B4">3682</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="5EB79267F820B42B33EC853124D16246" type="issue">
|
||
<mods:number id="BF02BD641C0769252B46806B82E850B3">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="983148E200A07BEAC42478AA4AF4BF70" unit="page">
|
||
<mods:start id="1EA82D580A81EC389C62DFDF9E5BA5E5">133</mods:start>
|
||
<mods:end id="9A4243D2727103988A4FFD3C87A51375">142</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="54B3EE708397300FE7F5851E1F30592B">journal article</mods:classification>
|
||
<mods:identifier id="95670806CA4141238F9B30FC7E463405" type="DOI">10.11646/zootaxa.3682.1.6</mods:identifier>
|
||
<mods:identifier id="499F99AFDEB33F64FBEFB14138B05033" type="GBIF-Dataset">e8a0fb91-227c-4b35-92f4-a3b409035052</mods:identifier>
|
||
<mods:identifier id="38FD1D5FBAA038DC13A4DF4B4BF183BA" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="CFCE24CFF34186F6A56CEA0852DBAD41" type="Zenodo-Dep">218044</mods:identifier>
|
||
<mods:identifier id="99A8144FC52244D67E9653CD531F3A17" type="ZooBank">1B811565-C78A-4CF2-9B89-723A20524280</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="6F11A30112442231FF538EB8F488FBC2" ID-DOI="http://doi.org/10.5281/zenodo.5620879" ID-GBIF-Taxon="119571390" ID-Zenodo-Dep="5620879" LSID="urn:lsid:plazi:treatment:6F11A30112442231FF538EB8F488FBC2" httpUri="http://treatment.plazi.org/id/6F11A30112442231FF538EB8F488FBC2" lastPageId="7" lastPageNumber="140" pageId="1" pageNumber="134">
|
||
<subSubSection id="AFA2419C12442237FF538EB8F7BAFA59" pageId="1" pageNumber="134" type="nomenclature">
|
||
<paragraph id="E707121712442237FF538EB8F41AFA3E" blockId="1.[151,534,1425,1484]" box="[151,534,1425,1451]" pageId="1" pageNumber="134">
|
||
<heading id="BC4FA57B12442237FF538EB8F41AFA3E" bold="true" box="[151,534,1425,1451]" fontSize="11" level="1" pageId="1" pageNumber="134" reason="1">
|
||
<emphasis id="D5CCCE0512442237FF538EB8F41AFA3E" bold="true" box="[151,534,1425,1451]" pageId="1" pageNumber="134">
|
||
<taxonomicName id="20B8699412442237FF538EB8F772FA3E" box="[151,382,1425,1451]" class="Kinorhyncha" family="Dracoderidae" genus="Dracoderes" kingdom="Animalia" order="Cyclorhagida" pageId="1" pageNumber="134" phylum="Cephalorhyncha" rank="species" species="nidhug" status="sp. nov.">
|
||
<emphasis id="D5CCCE0512442237FF538EB8F772FA3E" bold="true" box="[151,382,1425,1451]" italics="true" pageId="1" pageNumber="134">Dracoderes nidhug</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CEFF737E12442237FE428EB8F41AFA3E" box="[390,534,1425,1451]" pageId="1" pageNumber="134" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E707121712442237FF538E9DF7BAFA59" blockId="1.[151,534,1425,1484]" box="[151,438,1460,1484]" pageId="1" pageNumber="134">
|
||
(
|
||
<figureCitation id="7F830E9212442237FF5A8E9DF72AFA59" box="[158,294,1460,1484]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[151,250,1861,1884]" captionStartId-1="5.[151,250,1440,1463]" captionStartId-2="6.[151,250,1577,1600]" captionTargetBox-0="[171,1414,289,1845]" captionTargetBox-1="[167,1419,196,1404]" captionTargetBox-2="[171,1422,198,1541]" captionTargetId-0="figure@4.[170,1429,254,1845]" captionTargetId-1="figure@5.[167,1419,194,1411]" captionTargetId-2="figure@6.[171,1422,194,1568]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." captionText-1="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." captionText-2="FIGURE 3. Scanning electron micrographs showing overviews and details in head and trunk morphology of Dracoderes nidhug nov. sp. A. Dorsal overview of whole specimen. B. Mouth cone with pharynx extended. C. lateroventral overview of whole specimen. D. Segment 2, ventral view. E. Detail of mid- and subdorsal areas of segment 2. F. Detail of lateroventral area of segment 2. G. Segments 7 to 9, lateral view. H. Segment 11 in male, ventral view. I. Segment 11 in male, lateral view. Abbreviations: fs, fringed spine; hs, helioscalid; ld, laterodorsal sensory spot; loo, large outer oral styles; lts, lateral terminal spine; lvt, lateroventral tubule; mds, midadorsal spine; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; scl, subcuticular line; sdt, subdorsal tubule; sl, sublateral sensory spot; soo, small outer oral styles; te, tergal extension; tr, trichoscalid; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri-0="https://zenodo.org/record/218045/files/figure.png" httpUri-1="https://zenodo.org/record/218046/files/figure.png" httpUri-2="https://zenodo.org/record/218047/files/figure.png" pageId="1" pageNumber="134">Figures 1–3</figureCitation>
|
||
,
|
||
<tableCitation id="AA3A27AC12442237FEF78E9DF7A1FA59" box="[307,429,1460,1484]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="2.[151,239,254,278]" captionStartId-1="3.[151,239,290,314]" captionTargetBox-0="[159,1260,433,1386]" captionTargetBox-1="[160,1380,533,1010]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="TABLE 1. Measurements of two adult specimens of Dracoderes nidhug nov. sp. from the East Sea, Korea, including number of measured specimens (n) and standard deviation (S. D.). Abbreviations: (ac): acicular spine; LTS: lateral terminal spine; LV: lateroventral; MD: middorsal spine; MSW- 5: Maximum sternal width, measured on segment 5 in this species; PD: dorsal spine displaced to paradorsal position; S: segment length; SW- 10, standard width, always measured on segment 10; TL: trunk length." captionText-1="TABLE 2. Summary of nature and location of sensory spots, glandular cell outlets, spines and tubules arranged by series in Dracoderes nidhug nov. sp. Abbreviations: LA: lateral accessory; LD: laterodorsal; LV: lateroventral; MD: middorsal; ML: midlateral; PD: paradorsal; PV: paraventral; SD: subdorsal; SL: sublateral; VL: ventrolateral; VM: ventromedial; ac: acicular spine; fs: fringed spine; gc, glandular cell outlet; lts: lateral terminal spine; (m): male condition of sexually dimorphic character; ne, nephridiopore; (pe): marks that the structure is perispinal and its exact position along the longitudinal series may have been obscured by eventual lateral spine displacement; ps, penile spine; ss, sensory spot; tu, tubule." httpUri-0="http://table.plazi.org/id/B3C7429F12472234FF538BD7F7F4FE00" httpUri-1="http://table.plazi.org/id/B3C7429F12462235FF538A0BF6EDFE6C" pageId="1" pageNumber="134" tableUuid-0="B3C7429F12472234FF538BD7F7F4FE00" tableUuid-1="B3C7429F12462235FF538A0BF6EDFE6C">Tables 1–2</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AFA2419C12442237FF538ED5F42EF915" pageId="1" pageNumber="134" type="diagnosis">
|
||
<paragraph id="E707121712442237FF538ED5F42EF915" blockId="1.[151,1436,1532,2033]" pageId="1" pageNumber="134">
|
||
<emphasis id="D5CCCE0512442237FF538ED5F71DF980" bold="true" box="[151,273,1532,1557]" pageId="1" pageNumber="134">Diagnosis.</emphasis>
|
||
<taxonomicName id="20B8699412442237FEDC8ED5F797F981" box="[280,411,1532,1556]" class="Kinorhyncha" family="Dracoderidae" genus="Dracoderes" kingdom="Animalia" order="Cyclorhagida" pageId="1" pageNumber="134" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="D5CCCE0512442237FEDC8ED5F797F981" box="[280,411,1532,1556]" italics="true" pageId="1" pageNumber="134">Dracoderes</emphasis>
|
||
</taxonomicName>
|
||
with dorsal spines on segments 3 to 9, alternatingly offset to lateral sides on segments 3 to 8, subdorsal tubules on segment 2, lateroventral tubules on segments 2, 5 and 10, and lateral accessory tubules on segment 8. Tergal extensions of terminal segment with thin, flexible tip. Males with two large penile spines and a minute, lateroventral fringed spine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AFA2419C12442237FF038DA4F216F811" pageId="1" pageNumber="134" type="materials_examined">
|
||
<paragraph id="E707121712442237FF038DA4F216F811" blockId="1.[151,1436,1532,2033]" pageId="1" pageNumber="134">
|
||
<emphasis id="D5CCCE0512442237FF038DA4F779F930" bold="true" box="[199,373,1676,1701]" pageId="1" pageNumber="134">
|
||
<typeStatus id="3803ACB512442237FF038DA4F70CF930" box="[199,256,1677,1701]" pageId="1" pageNumber="134">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="3803ACB512442237FE448DA5F7E7F931" box="[384,491,1676,1700]" pageId="1" pageNumber="134" type="holotype">Holotype</typeStatus>
|
||
, adult female, collected on
|
||
<date id="930634D712442237FCEF8DA5F201F931" box="[811,1037,1676,1700]" pageId="1" pageNumber="134" value="2011-11-11">November 11, 2011</date>
|
||
in the East Sea, off Uljin, locality MAP37: 37°01ʹ26ʹ N, 129°30ʹ55ʹ E, from mud at
|
||
<quantity id="2040BFF212442237FD198D9AF524F95E" box="[733,808,1715,1739]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="134" unit="m" value="160.0">160 m</quantity>
|
||
depth; mounted in Fluoromount G®, deposited at the
|
||
<collectionCode id="81A98AD212442237FF538DF1F6E3F97A" box="[151,239,1752,1775]" pageId="1" pageNumber="134">NHMD</collectionCode>
|
||
, under accession number:
|
||
<collectionCode id="81A98AD212442237FDF58DFEF48EF97A" box="[561,642,1751,1775]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="134">ZMUC</collectionCode>
|
||
<collectionCode id="81A98AD212442237FD548DFEF4C2F97A" box="[656,718,1751,1775]" pageId="1" pageNumber="134">KIN-</collectionCode>
|
||
649. Allotype, adult male, same collecting data as
|
||
<typeStatus id="3803ACB512442237FAF78DFEF394F97A" box="[1331,1432,1751,1775]" pageId="1" pageNumber="134" type="holotype">holotype</typeStatus>
|
||
, mounted in Fluoromount G®, deposited at
|
||
<collectionCode id="81A98AD212442237FDB48DD6F4CAF883" box="[624,710,1791,1814]" pageId="1" pageNumber="134">NHMD</collectionCode>
|
||
under accession number:
|
||
<collectionCode id="81A98AD212442237FC2D8DD7F236F883" box="[1001,1082,1790,1814]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="134">ZMUC</collectionCode>
|
||
<collectionCode id="81A98AD212442237FB858DD7F272F883" box="[1089,1150,1790,1814]" pageId="1" pageNumber="134">KIN-</collectionCode>
|
||
650.
|
||
<typeStatus id="3803ACB512442237FB708DD6F328F882" box="[1204,1316,1791,1815]" pageId="1" pageNumber="134" type="paratype">Paratypes</typeStatus>
|
||
, four adult females and three adult males, collected on same date and locality as
|
||
<typeStatus id="3803ACB512442237FC118C0DF236F8A9" box="[981,1082,1828,1852]" pageId="1" pageNumber="134" type="holotype">holotype</typeStatus>
|
||
, mounted in Fluoromount G®, deposited at
|
||
<collectionCode id="81A98AD212442237FEEE8C60F773F8F5" box="[298,383,1865,1888]" pageId="1" pageNumber="134">NHMD</collectionCode>
|
||
under accession numbers:
|
||
<collectionCode id="81A98AD212442237FD788C61F501F8F5" box="[700,781,1864,1888]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="134">ZMUC</collectionCode>
|
||
<collectionCode id="81A98AD212442237FCD38C61F558F8F5" box="[791,852,1864,1888]" pageId="1" pageNumber="134">KIN-</collectionCode>
|
||
651 to
|
||
<collectionCode id="81A98AD212442237FC628C61F5E8F8F5" box="[934,996,1864,1888]" pageId="1" pageNumber="134">KIN-</collectionCode>
|
||
657. Additional material includes nine specimens mounted for SEM, stored in the personal collection of the last author.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AFA2419C12442237FF038CB8F5FEF859" pageId="1" pageNumber="134" type="etymology">
|
||
<paragraph id="E707121712442237FF038CB8F5FEF859" blockId="1.[151,1436,1532,2033]" pageId="1" pageNumber="134">
|
||
<emphasis id="D5CCCE0512442237FF038CB8F742F83C" bold="true" box="[199,334,1937,1961]" pageId="1" pageNumber="134">Etymology.</emphasis>
|
||
The species is named after the dragon “
|
||
<taxonomicName id="20B8699412442237FCDF8CB9F57AF83D" box="[795,886,1936,1960]" class="Kinorhyncha" family="Dracoderidae" genus="Dracoderes" kingdom="Animalia" order="Cyclorhagida" pageId="1" pageNumber="142" phylum="Cephalorhyncha" rank="species" species="nidhug">Nidhug</taxonomicName>
|
||
”, known from Nordic mythology. At the
|
||
<collectingCountry id="9FAF528712442237FA8B8CB9F370F83D" box="[1359,1404,1936,1960]" name="India" pageId="1" pageNumber="134">End</collectingCountry>
|
||
of Days,
|
||
<taxonomicName id="20B8699412442237FF1B8C9DF73FF859" box="[223,307,1972,1996]" class="Kinorhyncha" family="Dracoderidae" genus="Dracoderes" kingdom="Animalia" order="Cyclorhagida" pageId="1" pageNumber="142" phylum="Cephalorhyncha" rank="species" species="nidhug">Nidhug</taxonomicName>
|
||
will bite and cut the roots of Ash Yggdrasil – the Tree of Life.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AFA2419C12442231FF038CF1F488FBC2" lastPageId="7" lastPageNumber="140" pageId="1" pageNumber="134" type="description">
|
||
<paragraph id="E707121712442237FF038CF1F390F865" blockId="1.[151,1436,1532,2033]" box="[199,1436,2008,2033]" pageId="1" pageNumber="134">
|
||
<emphasis id="D5CCCE0512442237FF038CF1F755F864" bold="true" box="[199,345,2008,2033]" pageId="1" pageNumber="134">Description.</emphasis>
|
||
Adult specimens with of a head, a neck and eleven trunk segments (
|
||
<figureCitation id="7F830E9212442237FB478CF1F2C5F865" box="[1155,1225,2008,2032]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="1" pageNumber="134">Figs 1</figureCitation>
|
||
A–B, 2A, 3A, C).
|
||
</paragraph>
|
||
<paragraph id="E707121712472234FF538BB1F478FF41" blockId="2.[151,1436,151,212]" pageId="2" pageNumber="135">
|
||
Measurements and dimensions are given in
|
||
<tableCitation id="AA3A27AC12472234FD4A8BBEF4EEFF3A" box="[654,738,151,175]" captionStart="TABLE 1" captionStartId="2.[151,239,254,278]" captionTargetBox="[159,1260,433,1386]" captionTargetPageId="2" captionText="TABLE 1. Measurements of two adult specimens of Dracoderes nidhug nov. sp. from the East Sea, Korea, including number of measured specimens (n) and standard deviation (S. D.). Abbreviations: (ac): acicular spine; LTS: lateral terminal spine; LV: lateroventral; MD: middorsal spine; MSW- 5: Maximum sternal width, measured on segment 5 in this species; PD: dorsal spine displaced to paradorsal position; S: segment length; SW- 10, standard width, always measured on segment 10; TL: trunk length." httpUri="http://table.plazi.org/id/B3C7429F12472234FF538BD7F7F4FE00" pageId="2" pageNumber="135" tableUuid="B3C7429F12472234FF538BD7F7F4FE00">Table 1</tableCitation>
|
||
, and a summary of sensory spot, spine, tubule and glandular cell outlet positions is provided in
|
||
<tableCitation id="AA3A27AC12472234FDDE8B95F47CFF41" box="[538,624,188,212]" captionStart="TABLE 2" captionStartId="3.[151,239,290,314]" captionTargetBox="[160,1380,533,1010]" captionTargetPageId="3" captionText="TABLE 2. Summary of nature and location of sensory spots, glandular cell outlets, spines and tubules arranged by series in Dracoderes nidhug nov. sp. Abbreviations: LA: lateral accessory; LD: laterodorsal; LV: lateroventral; MD: middorsal; ML: midlateral; PD: paradorsal; PV: paraventral; SD: subdorsal; SL: sublateral; VL: ventrolateral; VM: ventromedial; ac: acicular spine; fs: fringed spine; gc, glandular cell outlet; lts: lateral terminal spine; (m): male condition of sexually dimorphic character; ne, nephridiopore; (pe): marks that the structure is perispinal and its exact position along the longitudinal series may have been obscured by eventual lateral spine displacement; ps, penile spine; ss, sensory spot; tu, tubule." httpUri="http://table.plazi.org/id/B3C7429F12462235FF538A0BF6EDFE6C" pageId="2" pageNumber="135" tableUuid="B3C7429F12462235FF538A0BF6EDFE6C">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="B3C7429F12472234FF538BD7F7F4FE00" ID-Table-UUID="B3C7429F12472234FF538BD7F7F4FE00" httpUri="http://table.plazi.org/id/B3C7429F12472234FF538BD7F7F4FE00" pageId="2" pageNumber="135" targetBox="[159,1260,433,1386]" targetIsTable="true" targetPageId="2">
|
||
<paragraph id="E707121712472234FF538BD7F7F4FE00" blockId="2.[151,1436,254,405]" pageId="2" pageNumber="135">
|
||
<emphasis id="D5CCCE0512472234FF538BD7F706FE83" bold="true" box="[151,266,254,278]" pageId="2" pageNumber="135">TABLE 1.</emphasis>
|
||
Measurements of two adult specimens of
|
||
<taxonomicName id="20B8699412472234FD188BD7F5A0FE80" box="[732,940,254,277]" class="Kinorhyncha" family="Dracoderidae" genus="Dracoderes" kingdom="Animalia" order="Cyclorhagida" pageId="2" pageNumber="135" phylum="Cephalorhyncha" rank="species" species="nidhug">
|
||
<emphasis id="D5CCCE0512472234FD188BD7F5A0FE80" box="[732,940,254,277]" italics="true" pageId="2" pageNumber="135">Dracoderes nidhug</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D5CCCE0512472234FC718BD7F200FE83" bold="true" box="[949,1036,254,278]" pageId="2" pageNumber="135">nov. sp.</emphasis>
|
||
from the East Sea, Korea, including number of measured specimens (n) and standard deviation (S.D.). Abbreviations: (ac): acicular spine; LTS: lateral terminal spine; LV: lateroventral; MD: middorsal spine; MSW-5: Maximum sternal width, measured on segment 5 in this species; PD: dorsal spine displaced to paradorsal position; S: segment length; SW-10, standard width, always measured on segment 10; TL: trunk length.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E707121712472234FF5B8A98F2E6FAFF" pageId="2" pageNumber="135">
|
||
<table id="95B8E0B71247DDC9FF5B8A98F2E0FAFF" box="[159,1260,433,1386]" gridcols="4" gridrows="27" pageId="2" pageNumber="135">
|
||
<tr id="598810551247DDC9FF5B8A98F2E0FE52" box="[159,1260,433,455]" gridrow="0" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8A98F719FE52" box="[159,277,433,455]" gridcol="0" gridrow="0" pageId="2" pageNumber="135">Character</th>
|
||
<th id="1A5979291247DDC9FE6D8A98F4A9FE52" box="[425,677,433,455]" gridcol="1" gridrow="0" pageId="2" pageNumber="135">n Range</th>
|
||
<th id="1A5979291247DDC9FC508A98F5EEFE52" box="[916,994,433,455]" gridcol="2" gridrow="0" pageId="2" pageNumber="135">Mean</th>
|
||
<th id="1A5979291247DDC9FB4D8A98F2E0FE52" box="[1161,1260,433,455]" gridcol="3" gridrow="0" pageId="2" pageNumber="135">S.D.</th>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8AF3F2E0FE65" box="[159,1260,474,496]" gridrow="1" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8AF3F719FE65" box="[159,277,474,496]" gridcol="0" gridrow="1" pageId="2" pageNumber="135">TL</th>
|
||
<td id="1A5979291247DDC9FE6D8AF3F4A9FE65" box="[425,677,474,496]" gridcol="1" gridrow="1" pageId="2" pageNumber="135">9 374–427 µm</td>
|
||
<td id="1A5979291247DDC9FC508AF3F5EEFE65" box="[916,994,474,496]" gridcol="2" gridrow="1" pageId="2" pageNumber="135">403 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8AF3F2E0FE65" box="[1161,1260,474,496]" gridcol="3" gridrow="1" pageId="2" pageNumber="135">15,85 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8AD0F2E0FD9A" box="[159,1260,505,527]" gridrow="2" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8AD0F719FD9A" box="[159,277,505,527]" gridcol="0" gridrow="2" pageId="2" pageNumber="135">MSW-5</th>
|
||
<td id="1A5979291247DDC9FE6D8AD0F4A9FD9A" box="[425,677,505,527]" gridcol="1" gridrow="2" pageId="2" pageNumber="135">9 87–95 µm</td>
|
||
<td id="1A5979291247DDC9FC508AD0F5EEFD9A" box="[916,994,505,527]" gridcol="2" gridrow="2" pageId="2" pageNumber="135">90 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8AD0F2E0FD9A" box="[1161,1260,505,527]" gridcol="3" gridrow="2" pageId="2" pageNumber="135">3,14 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8931F2E0FDBB" box="[159,1260,536,558]" gridrow="3" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8931F719FDBB" box="[159,277,536,558]" gridcol="0" gridrow="3" pageId="2" pageNumber="135">MSW-5/TL</th>
|
||
<td id="1A5979291247DDC9FE6D8931F4A9FDBB" box="[425,677,536,558]" gridcol="1" gridrow="3" pageId="2" pageNumber="135">9 21,0–23,3 %</td>
|
||
<td id="1A5979291247DDC9FC508931F5EEFDBB" box="[916,994,536,558]" gridcol="2" gridrow="3" pageId="2" pageNumber="135">22.3 %</td>
|
||
<td id="1A5979291247DDC9FB4D8931F2E0FDBB" box="[1161,1260,536,558]" gridcol="3" gridrow="3" pageId="2" pageNumber="135">0,68 %</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B891FF2E0FDD9" box="[159,1260,566,588]" gridrow="4" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B891FF719FDD9" box="[159,277,566,588]" gridcol="0" gridrow="4" pageId="2" pageNumber="135">SW-10</th>
|
||
<td id="1A5979291247DDC9FE6D891FF4A9FDD9" box="[425,677,566,588]" gridcol="1" gridrow="4" pageId="2" pageNumber="135">9 59–68 µm</td>
|
||
<td id="1A5979291247DDC9FC50891FF5EEFDD9" box="[916,994,566,588]" gridcol="2" gridrow="4" pageId="2" pageNumber="135">65 µm</td>
|
||
<td id="1A5979291247DDC9FB4D891FF2E0FDD9" box="[1161,1260,566,588]" gridcol="3" gridrow="4" pageId="2" pageNumber="135">3,21 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B897CF2E0FDFE" box="[159,1260,597,619]" gridrow="5" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B897CF719FDFE" box="[159,277,597,619]" gridcol="0" gridrow="5" pageId="2" pageNumber="135">SW-10/TL</th>
|
||
<td id="1A5979291247DDC9FE6D897CF4A9FDFE" box="[425,677,597,619]" gridcol="1" gridrow="5" pageId="2" pageNumber="135">9 14,9–17,0 %</td>
|
||
<td id="1A5979291247DDC9FC50897CF5EEFDFE" box="[916,994,597,619]" gridcol="2" gridrow="5" pageId="2" pageNumber="135">16.0 %</td>
|
||
<td id="1A5979291247DDC9FB4D897CF2E0FDFE" box="[1161,1260,597,619]" gridcol="3" gridrow="5" pageId="2" pageNumber="135">0,84 %</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B895AF2E0FD1F" box="[159,1260,627,650]" gridrow="6" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B895AF719FD1F" box="[159,277,627,650]" gridcol="0" gridrow="6" pageId="2" pageNumber="135">S1</th>
|
||
<td id="1A5979291247DDC9FE6D895AF4A9FD1F" box="[425,677,627,650]" gridcol="1" gridrow="6" pageId="2" pageNumber="135">9 43–48 µm</td>
|
||
<td id="1A5979291247DDC9FC50895AF5EEFD1F" box="[916,994,627,650]" gridcol="2" gridrow="6" pageId="2" pageNumber="135">46 µm</td>
|
||
<td id="1A5979291247DDC9FB4D895AF2E0FD1F" box="[1161,1260,627,650]" gridcol="3" gridrow="6" pageId="2" pageNumber="135">1,58 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B89BBF2E0FD3D" box="[159,1260,658,680]" gridrow="7" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B89BBF719FD3D" box="[159,277,658,680]" gridcol="0" gridrow="7" pageId="2" pageNumber="135">S2</th>
|
||
<td id="1A5979291247DDC9FE6D89BBF4A9FD3D" box="[425,677,658,680]" gridcol="1" gridrow="7" pageId="2" pageNumber="135">9 36–40 µm</td>
|
||
<td id="1A5979291247DDC9FC5089BBF5EEFD3D" box="[916,994,658,680]" gridcol="2" gridrow="7" pageId="2" pageNumber="135">38 µm</td>
|
||
<td id="1A5979291247DDC9FB4D89BBF2E0FD3D" box="[1161,1260,658,680]" gridcol="3" gridrow="7" pageId="2" pageNumber="135">1,32 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B899BF2E0FD52" box="[159,1260,690,711]" gridrow="8" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B899BF719FD52" box="[159,277,690,711]" gridcol="0" gridrow="8" pageId="2" pageNumber="135">S3</th>
|
||
<td id="1A5979291247DDC9FE6D899BF4A9FD52" box="[425,677,690,711]" gridcol="1" gridrow="8" pageId="2" pageNumber="135">9 37–41 µm</td>
|
||
<td id="1A5979291247DDC9FC50899BF5EEFD52" box="[916,994,690,711]" gridcol="2" gridrow="8" pageId="2" pageNumber="135">38 µm</td>
|
||
<td id="1A5979291247DDC9FB4D899BF2E0FD52" box="[1161,1260,690,711]" gridcol="3" gridrow="8" pageId="2" pageNumber="135">1,22 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B89F9F2E0FD73" box="[159,1260,720,742]" gridrow="9" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B89F9F719FD73" box="[159,277,720,742]" gridcol="0" gridrow="9" pageId="2" pageNumber="135">S4</th>
|
||
<td id="1A5979291247DDC9FE6D89F9F4A9FD73" box="[425,677,720,742]" gridcol="1" gridrow="9" pageId="2" pageNumber="135">9 39–44 µm</td>
|
||
<td id="1A5979291247DDC9FC5089F9F5EEFD73" box="[916,994,720,742]" gridcol="2" gridrow="9" pageId="2" pageNumber="135">41 µm</td>
|
||
<td id="1A5979291247DDC9FB4D89F9F2E0FD73" box="[1161,1260,720,742]" gridcol="3" gridrow="9" pageId="2" pageNumber="135">1,88 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B89C7F2E0FC91" box="[159,1260,750,772]" gridrow="10" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B89C7F719FC91" box="[159,277,750,772]" gridcol="0" gridrow="10" pageId="2" pageNumber="135">S5</th>
|
||
<td id="1A5979291247DDC9FE6D89C7F4A9FC91" box="[425,677,750,772]" gridcol="1" gridrow="10" pageId="2" pageNumber="135">9 39–47 µm</td>
|
||
<td id="1A5979291247DDC9FC5089C7F5EEFC91" box="[916,994,750,772]" gridcol="2" gridrow="10" pageId="2" pageNumber="135">43 µm</td>
|
||
<td id="1A5979291247DDC9FB4D89C7F2E0FC91" box="[1161,1260,750,772]" gridcol="3" gridrow="10" pageId="2" pageNumber="135">2,33 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8824F2E0FCB6" box="[159,1260,781,803]" gridrow="11" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8824F719FCB6" box="[159,277,781,803]" gridcol="0" gridrow="11" pageId="2" pageNumber="135">S6</th>
|
||
<td id="1A5979291247DDC9FE6D8824F4A9FCB6" box="[425,677,781,803]" gridcol="1" gridrow="11" pageId="2" pageNumber="135">9 40–46 µm</td>
|
||
<td id="1A5979291247DDC9FC508824F5EEFCB6" box="[916,994,781,803]" gridcol="2" gridrow="11" pageId="2" pageNumber="135">43 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8824F2E0FCB6" box="[1161,1260,781,803]" gridcol="3" gridrow="11" pageId="2" pageNumber="135">2,09 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8802F2E0FCD7" box="[159,1260,811,834]" gridrow="12" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8802F719FCD7" box="[159,277,811,834]" gridcol="0" gridrow="12" pageId="2" pageNumber="135">S7</th>
|
||
<td id="1A5979291247DDC9FE6D8802F4A9FCD7" box="[425,677,811,834]" gridcol="1" gridrow="12" pageId="2" pageNumber="135">9 39–45 µm</td>
|
||
<td id="1A5979291247DDC9FC508802F5EEFCD7" box="[916,994,811,834]" gridcol="2" gridrow="12" pageId="2" pageNumber="135">42 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8802F2E0FCD7" box="[1161,1260,811,834]" gridcol="3" gridrow="12" pageId="2" pageNumber="135">2,24 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8863F2E0FCF5" box="[159,1260,842,864]" gridrow="13" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8863F719FCF5" box="[159,277,842,864]" gridcol="0" gridrow="13" pageId="2" pageNumber="135">S8</th>
|
||
<td id="1A5979291247DDC9FE6D8863F4A9FCF5" box="[425,677,842,864]" gridcol="1" gridrow="13" pageId="2" pageNumber="135">9 36–43 µm</td>
|
||
<td id="1A5979291247DDC9FC508863F5EEFCF5" box="[916,994,842,864]" gridcol="2" gridrow="13" pageId="2" pageNumber="135">41 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8863F2E0FCF5" box="[1161,1260,842,864]" gridcol="3" gridrow="13" pageId="2" pageNumber="135">2,30 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8840F2E0FCEA" box="[159,1260,873,895]" gridrow="14" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8840F719FCEA" box="[159,277,873,895]" gridcol="0" gridrow="14" pageId="2" pageNumber="135">S9</th>
|
||
<td id="1A5979291247DDC9FE6D8840F4A9FCEA" box="[425,677,873,895]" gridcol="1" gridrow="14" pageId="2" pageNumber="135">9 36–43 µm</td>
|
||
<td id="1A5979291247DDC9FC508840F5EEFCEA" box="[916,994,873,895]" gridcol="2" gridrow="14" pageId="2" pageNumber="135">40 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8840F2E0FCEA" box="[1161,1260,873,895]" gridcol="3" gridrow="14" pageId="2" pageNumber="135">2,24 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B88A1F2E0FC0B" box="[159,1260,904,926]" gridrow="15" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B88A1F719FC0B" box="[159,277,904,926]" gridcol="0" gridrow="15" pageId="2" pageNumber="135">S10</th>
|
||
<td id="1A5979291247DDC9FE6D88A1F4A9FC0B" box="[425,677,904,926]" gridcol="1" gridrow="15" pageId="2" pageNumber="135">9 29–33 µm</td>
|
||
<td id="1A5979291247DDC9FC5088A1F5EEFC0B" box="[916,994,904,926]" gridcol="2" gridrow="15" pageId="2" pageNumber="135">32 µm</td>
|
||
<td id="1A5979291247DDC9FB4D88A1F2E0FC0B" box="[1161,1260,904,926]" gridcol="3" gridrow="15" pageId="2" pageNumber="135">1,33 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B888EF2E0FC29" box="[159,1260,935,956]" gridrow="16" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B888EF719FC29" box="[159,277,935,956]" gridcol="0" gridrow="16" pageId="2" pageNumber="135">S11</th>
|
||
<td id="1A5979291247DDC9FE6D888EF4A9FC29" box="[425,677,935,956]" gridcol="1" gridrow="16" pageId="2" pageNumber="135">9 39–47 µm</td>
|
||
<td id="1A5979291247DDC9FC50888EF5EEFC29" box="[916,994,935,956]" gridcol="2" gridrow="16" pageId="2" pageNumber="135">43 µm</td>
|
||
<td id="1A5979291247DDC9FB4D888EF2E0FC29" box="[1161,1260,935,956]" gridcol="3" gridrow="16" pageId="2" pageNumber="135">2,42 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B88CAF2E0FC6F" box="[159,1260,995,1018]" gridrow="17" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B88CAF719FC6F" box="[159,277,995,1018]" gridcol="0" gridrow="17" pageId="2" pageNumber="135">MD 3 (ac)</th>
|
||
<td id="1A5979291247DDC9FE6D88CAF4A9FC6F" box="[425,677,995,1018]" gridcol="1" gridrow="17" pageId="2" pageNumber="135">9 43–57 µm</td>
|
||
<td id="1A5979291247DDC9FC5088CAF5EEFC6F" box="[916,994,995,1018]" gridcol="2" gridrow="17" pageId="2" pageNumber="135">50 µm</td>
|
||
<td id="1A5979291247DDC9FB4D88CAF2E0FC6F" box="[1161,1260,995,1018]" gridcol="3" gridrow="17" pageId="2" pageNumber="135">4,85 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8F2BF2E0FB8C" box="[159,1260,1026,1049]" gridrow="18" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8F2BF719FB8C" box="[159,277,1026,1049]" gridcol="0" gridrow="18" pageId="2" pageNumber="135">PD 4 (ac)</th>
|
||
<td id="1A5979291247DDC9FE6D8F2BF4A9FB8C" box="[425,677,1026,1049]" gridcol="1" gridrow="18" pageId="2" pageNumber="135">9 40–62 µm</td>
|
||
<td id="1A5979291247DDC9FC508F2BF5EEFB8C" box="[916,994,1026,1049]" gridcol="2" gridrow="18" pageId="2" pageNumber="135">55 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8F2BF2E0FB8C" box="[1161,1260,1026,1049]" gridcol="3" gridrow="18" pageId="2" pageNumber="135">7,64 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8F08F2E0FBAD" box="[159,1260,1057,1080]" gridrow="19" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8F08F719FBAD" box="[159,277,1057,1080]" gridcol="0" gridrow="19" pageId="2" pageNumber="135">PD 5 (ac)</th>
|
||
<td id="1A5979291247DDC9FE6D8F08F4A9FBAD" box="[425,677,1057,1080]" gridcol="1" gridrow="19" pageId="2" pageNumber="135">9 40–70 µm</td>
|
||
<td id="1A5979291247DDC9FC508F08F5EEFBAD" box="[916,994,1057,1080]" gridcol="2" gridrow="19" pageId="2" pageNumber="135">58 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8F08F2E0FBAD" box="[1161,1260,1057,1080]" gridcol="3" gridrow="19" pageId="2" pageNumber="135">9,55 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8F69F2E0FBC3" box="[159,1260,1088,1110]" gridrow="20" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8F69F719FBC3" box="[159,277,1088,1110]" gridcol="0" gridrow="20" pageId="2" pageNumber="135">PD 6 (ac)</th>
|
||
<td id="1A5979291247DDC9FE6D8F69F4A9FBC3" box="[425,677,1088,1110]" gridcol="1" gridrow="20" pageId="2" pageNumber="135">8 55–63 µm</td>
|
||
<td id="1A5979291247DDC9FC508F69F5EEFBC3" box="[916,994,1088,1110]" gridcol="2" gridrow="20" pageId="2" pageNumber="135">59 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8F69F2E0FBC3" box="[1161,1260,1088,1110]" gridcol="3" gridrow="20" pageId="2" pageNumber="135">2,75 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8F77F2E0FBE0" box="[159,1260,1118,1141]" gridrow="21" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8F77F719FBE0" box="[159,277,1118,1141]" gridcol="0" gridrow="21" pageId="2" pageNumber="135">PD 7 (ac)</th>
|
||
<td id="1A5979291247DDC9FE6D8F77F4A9FBE0" box="[425,677,1118,1141]" gridcol="1" gridrow="21" pageId="2" pageNumber="135">9 44–57 µm</td>
|
||
<td id="1A5979291247DDC9FC508F77F5EEFBE0" box="[916,994,1118,1141]" gridcol="2" gridrow="21" pageId="2" pageNumber="135">51 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8F77F2E0FBE0" box="[1161,1260,1118,1141]" gridcol="3" gridrow="21" pageId="2" pageNumber="135">4,40 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8F54F2E0FB27" box="[159,1260,1149,1202]" gridrow="22" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8F54F719FB27" box="[159,277,1149,1202]" gridcol="0" gridrow="22" pageId="2" pageNumber="135">PD 8 (ac) MD 9 (ac)</th>
|
||
<td id="1A5979291247DDC9FE6D8F54F4A9FB27" box="[425,677,1149,1202]" gridcol="1" gridrow="22" pageId="2" pageNumber="135">8 35–47 µm 9 48–57 µm</td>
|
||
<td id="1A5979291247DDC9FC508F54F5EEFB27" box="[916,994,1149,1202]" gridcol="2" gridrow="22" pageId="2" pageNumber="135">43 µm 53 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8F54F2E0FB27" box="[1161,1260,1149,1202]" gridcol="3" gridrow="22" pageId="2" pageNumber="135">4,34 µm 3,35 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8FF0F2E0FB65" box="[159,1260,1241,1264]" gridrow="23" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8FF0F719FB65" box="[159,277,1241,1264]" gridcol="0" gridrow="23" pageId="2" pageNumber="135">LV 6 (ac)</th>
|
||
<td id="1A5979291247DDC9FE6D8FF0F4A9FB65" box="[425,677,1241,1264]" gridcol="1" gridrow="23" pageId="2" pageNumber="135">9 30–37 µm</td>
|
||
<td id="1A5979291247DDC9FC508FF0F5EEFB65" box="[916,994,1241,1264]" gridcol="2" gridrow="23" pageId="2" pageNumber="135">34 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8FF0F2E0FB65" box="[1161,1260,1241,1264]" gridcol="3" gridrow="23" pageId="2" pageNumber="135">2,12 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8FD1F2E0FA9B" box="[159,1260,1272,1294]" gridrow="24" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8FD1F719FA9B" box="[159,277,1272,1294]" gridcol="0" gridrow="24" pageId="2" pageNumber="135">LV 7 (ac)</th>
|
||
<td id="1A5979291247DDC9FE6D8FD1F4A9FA9B" box="[425,677,1272,1294]" gridcol="1" gridrow="24" pageId="2" pageNumber="135">9 30–36 µm</td>
|
||
<td id="1A5979291247DDC9FC508FD1F5EEFA9B" box="[916,994,1272,1294]" gridcol="2" gridrow="24" pageId="2" pageNumber="135">34 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8FD1F2E0FA9B" box="[1161,1260,1272,1294]" gridcol="3" gridrow="24" pageId="2" pageNumber="135">2,35 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8E3EF2E0FAB8" box="[159,1260,1303,1325]" gridrow="25" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8E3EF719FAB8" box="[159,277,1303,1325]" gridcol="0" gridrow="25" pageId="2" pageNumber="135">LV 8 (ac)</th>
|
||
<td id="1A5979291247DDC9FE6D8E3EF4A9FAB8" box="[425,677,1303,1325]" gridcol="1" gridrow="25" pageId="2" pageNumber="135">9 27–33 µm</td>
|
||
<td id="1A5979291247DDC9FC508E3EF5EEFAB8" box="[916,994,1303,1325]" gridcol="2" gridrow="25" pageId="2" pageNumber="135">31 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8E3EF2E0FAB8" box="[1161,1260,1303,1325]" gridcol="3" gridrow="25" pageId="2" pageNumber="135">1,80 µm</td>
|
||
</tr>
|
||
<tr id="598810551247DDC9FF5B8E1CF2E0FAFF" box="[159,1260,1333,1386]" gridrow="26" pageId="2" pageNumber="135">
|
||
<th id="1A5979291247DDC9FF5B8E1CF719FAFF" box="[159,277,1333,1386]" gridcol="0" gridrow="26" pageId="2" pageNumber="135">LV9 (ac) LTS</th>
|
||
<td id="1A5979291247DDC9FE6D8E1CF4A9FAFF" box="[425,677,1333,1386]" gridcol="1" gridrow="26" pageId="2" pageNumber="135">9 26–31 µm 8 200–229 µm</td>
|
||
<td id="1A5979291247DDC9FC508E1CF5EEFAFF" box="[916,994,1333,1386]" gridcol="2" gridrow="26" pageId="2" pageNumber="135">28 µm 215 µm</td>
|
||
<td id="1A5979291247DDC9FB4D8E1CF2E0FAFF" box="[1161,1260,1333,1386]" gridcol="3" gridrow="26" pageId="2" pageNumber="135">2,19 µm 11,41 µm</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="E707121712472234FF038E87F4C9F973" blockId="2.[151,1437,1454,2018]" pageId="2" pageNumber="135">
|
||
The head consists of a retractable mouth cone and an introvert. Inner armature of mouth cone with pharyngeal crown, one ring of helioscalids, and two rings of inner oral styles. The exact number of helioscalids and inner oral styles could not be determined. Outer mouth cone armature with nine outer oral styles that alternate in size between large well-developed ones in positions anterior to odd numbered introvert sections, and much smaller ones anterior to even numbered sections (
|
||
<figureCitation id="7F830E9212472234FE168D17F419F9C3" box="[466,533,1598,1622]" captionStart="FIGURE 3" captionStartId="6.[151,250,1577,1600]" captionTargetBox="[171,1422,198,1541]" captionTargetId="figure@6.[171,1422,194,1568]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs showing overviews and details in head and trunk morphology of Dracoderes nidhug nov. sp. A. Dorsal overview of whole specimen. B. Mouth cone with pharynx extended. C. lateroventral overview of whole specimen. D. Segment 2, ventral view. E. Detail of mid- and subdorsal areas of segment 2. F. Detail of lateroventral area of segment 2. G. Segments 7 to 9, lateral view. H. Segment 11 in male, ventral view. I. Segment 11 in male, lateral view. Abbreviations: fs, fringed spine; hs, helioscalid; ld, laterodorsal sensory spot; loo, large outer oral styles; lts, lateral terminal spine; lvt, lateroventral tubule; mds, midadorsal spine; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; scl, subcuticular line; sdt, subdorsal tubule; sl, sublateral sensory spot; soo, small outer oral styles; te, tergal extension; tr, trichoscalid; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218047/files/figure.png" pageId="2" pageNumber="135">Fig. 3</figureCitation>
|
||
B). An outer oral style is missing anterior to introvert section 6. Large styles are composed of two joined units; distal unit terminates into hook-like structure (
|
||
<figureCitation id="7F830E9212472234FBC78D4BF24AF9EF" box="[1027,1094,1634,1658]" captionStart="FIGURE 3" captionStartId="6.[151,250,1577,1600]" captionTargetBox="[171,1422,198,1541]" captionTargetId="figure@6.[171,1422,194,1568]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs showing overviews and details in head and trunk morphology of Dracoderes nidhug nov. sp. A. Dorsal overview of whole specimen. B. Mouth cone with pharynx extended. C. lateroventral overview of whole specimen. D. Segment 2, ventral view. E. Detail of mid- and subdorsal areas of segment 2. F. Detail of lateroventral area of segment 2. G. Segments 7 to 9, lateral view. H. Segment 11 in male, ventral view. I. Segment 11 in male, lateral view. Abbreviations: fs, fringed spine; hs, helioscalid; ld, laterodorsal sensory spot; loo, large outer oral styles; lts, lateral terminal spine; lvt, lateroventral tubule; mds, midadorsal spine; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; scl, subcuticular line; sdt, subdorsal tubule; sl, sublateral sensory spot; soo, small outer oral styles; te, tergal extension; tr, trichoscalid; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218047/files/figure.png" pageId="2" pageNumber="135">Fig. 3</figureCitation>
|
||
B). Small styles are extremely under-developed, and resemble the soft inner oral styles. The introvert carries rings of scalids, but even though all specimens mounted for SEM had their introverts extended, scalids always tended to be collapsed and smacked together, so further examination was not possible.
|
||
</paragraph>
|
||
<paragraph id="E707121712472234FF038DDBF26FF8E3" blockId="2.[151,1437,1454,2018]" pageId="2" pageNumber="135">
|
||
Neck with nine placids, with a joint between the placids and segment 1 (
|
||
<figureCitation id="7F830E9212472234FBC58DDBF24AF89F" box="[1025,1094,1778,1802]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="2" pageNumber="135">Figs 1</figureCitation>
|
||
A–B, 2B). The widest placids are the midventral and midlateral ones, ca. 19 µm at base, whereas the widths of the remaining ones vary between 16 µm and 18 µm. The 5 most ventral placids are situated closely together, whereas the midlateral, laterodorsal and subdorsal placids have cuticular foldings occupying the space between them (
|
||
<figureCitation id="7F830E9212472234FC388C77F230F8E3" box="[1020,1084,1886,1910]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="2" pageNumber="135">Fig. 1</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="E707121712472235FF038CABF254FF62" blockId="2.[151,1437,1454,2018]" lastBlockId="3.[151,1436,151,247]" lastPageId="3" lastPageNumber="136" pageId="2" pageNumber="135">
|
||
The first segment forms a closed cuticular ring, whereas the following 10 are composed of a tergal and two sternal plates (
|
||
<figureCitation id="7F830E9212472234FEFC8C8FF772F82B" box="[312,382,1958,1982]" captionStart="FIGURE 2" captionStartId="5.[151,250,1440,1463]" captionTargetBox="[167,1419,196,1404]" captionTargetId="figure@5.[167,1419,194,1411]" captionTargetPageId="5" captionText="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="2" pageNumber="135">Figs 2</figureCitation>
|
||
A, 3C). Junctions between tergal and sternal plates are easily observed with LM (e.g.
|
||
<figureCitation id="7F830E9212472234FAFC8C8FF376F82B" box="[1336,1402,1958,1982]" captionStart="FIGURE 2" captionStartId="5.[151,250,1440,1463]" captionTargetBox="[167,1419,196,1404]" captionTargetId="figure@5.[167,1419,194,1411]" captionTargetPageId="5" captionText="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="2" pageNumber="135">Fig. 2</figureCitation>
|
||
C), but in SEM only the midsternal junction is marked by a visible line, whereas a continuous outer cuticle is present in the position of the tergosternal junctions, indicating that the plate junctions are intracuticular. Maximum sternal width is reached at segment 5, and the sternal widths taper gradually from this segment and towards the posterior end. The overall trunk outline appears cigar-shaped, rather than rectangular (
|
||
<figureCitation id="7F830E9212462235FC358BF6F238FF62" box="[1009,1076,223,247]" captionStart="FIGURE 2" captionStartId="5.[151,250,1440,1463]" captionTargetBox="[167,1419,196,1404]" captionTargetId="figure@5.[167,1419,194,1411]" captionTargetPageId="5" captionText="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="3" pageNumber="136">Fig. 2</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<caption id="B3C7429F12462235FF538A0BF6EDFE6C" ID-Table-UUID="B3C7429F12462235FF538A0BF6EDFE6C" httpUri="http://table.plazi.org/id/B3C7429F12462235FF538A0BF6EDFE6C" pageId="3" pageNumber="136" targetBox="[160,1380,533,1010]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="E707121712462235FF538A0BF6EDFE6C" blockId="3.[151,1436,290,505]" pageId="3" pageNumber="136">
|
||
<emphasis id="D5CCCE0512462235FF538A0BF704FEAF" bold="true" box="[151,264,290,314]" pageId="3" pageNumber="136">TABLE 2.</emphasis>
|
||
Summary of nature and location of sensory spots, glandular cell outlets, spines and tubules arranged by series in
|
||
<taxonomicName id="20B8699412462235FF768A6BF773FECC" box="[178,383,322,345]" class="Kinorhyncha" family="Dracoderidae" genus="Dracoderes" kingdom="Animalia" order="Cyclorhagida" pageId="3" pageNumber="136" phylum="Cephalorhyncha" rank="species" species="nidhug">
|
||
<emphasis id="D5CCCE0512462235FF768A6BF773FECC" box="[178,383,322,345]" italics="true" pageId="3" pageNumber="136">Dracoderes nidhug</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D5CCCE0512462235FE418A6BF7D5FECF" bold="true" box="[389,473,322,346]" pageId="3" pageNumber="136">nov. sp.</emphasis>
|
||
Abbreviations: LA: lateral accessory; LD: laterodorsal; LV: lateroventral; MD: middorsal; ML: midlateral; PD: paradorsal; PV: paraventral; SD: subdorsal; SL: sublateral; VL: ventrolateral; VM: ventromedial; ac: acicular spine; fs: fringed spine; gc, glandular cell outlet; lts: lateral terminal spine; (m): male condition of sexually dimorphic character; ne, nephridiopore; (pe): marks that the structure is perispinal and its exact position along the longitudinal series may have been obscured by eventual lateral spine displacement; ps, penile spine; ss, sensory spot; tu, tubule.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E707121712462235FF64893CF299FC67" pageId="3" pageNumber="136">
|
||
<table id="95B8E0B71246DDC9FF64893CF368FC67" box="[160,1380,533,1010]" gridcols="8" gridrows="12" pageId="3" pageNumber="136">
|
||
<tr id="598810551246DDC9FF64893CF368FDBE" box="[160,1380,533,555]" gridrow="0" pageId="3" pageNumber="136">
|
||
<th id="1A5979291246DDC9FF64893CF79FFDBE" box="[160,403,533,555]" gridcol="0" gridrow="0" pageId="3" pageNumber="136">Position Segment MD</th>
|
||
<th id="1A5979291246DDC9FE7C893CF412FDBE" box="[440,542,533,555]" gridcol="1" gridrow="0" pageId="3" pageNumber="136">PD</th>
|
||
<th id="1A5979291246DDC9FDA5893CF4D7FDBE" box="[609,731,533,555]" gridcol="2" gridrow="0" pageId="3" pageNumber="136">SD LD</th>
|
||
<th id="1A5979291246DDC9FCD4893CF5F1FDBE" box="[784,1021,533,555]" gridcol="3" gridrow="0" pageId="3" pageNumber="136">ML LA SL</th>
|
||
<th id="1A5979291246DDC9FBE4893CF251FDBE" box="[1056,1117,533,555]" gridcol="4" gridrow="0" pageId="3" pageNumber="136">LV</th>
|
||
<th id="1A5979291246DDC9FB45893CF2AFFDBE" box="[1153,1187,533,555]" gridcol="5" gridrow="0" pageId="3" pageNumber="136">VL</th>
|
||
<th id="1A5979291246DDC9FB20893CF301FDBE" box="[1252,1293,533,555]" gridcol="6" gridrow="0" pageId="3" pageNumber="136">VM</th>
|
||
<th id="1A5979291246DDC9FA80893CF368FDBE" box="[1348,1380,533,555]" gridcol="7" gridrow="0" pageId="3" pageNumber="136">PV</th>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF648916F368FDC1" box="[160,1380,575,596]" gridrow="1" pageId="3" pageNumber="136" rowspan-1="1" rowspan-4="1" rowspan-7="1">
|
||
<th id="1A5979291246DDC9FF648916F79FFDC1" box="[160,403,575,596]" gridcol="0" gridrow="1" pageId="3" pageNumber="136">1</th>
|
||
<td id="1A5979291246DDC9FDA58916F4D7FDC1" box="[609,731,575,596]" gridcol="2" gridrow="1" pageId="3" pageNumber="136">ss ss</td>
|
||
<td id="1A5979291246DDC9FCD48916F5F1FDC1" box="[784,1021,575,596]" gridcol="3" gridrow="1" pageId="3" pageNumber="136">ss ss</td>
|
||
<td id="1A5979291246DDC9FB458916F2AFFDC1" box="[1153,1187,575,596]" gridcol="5" gridrow="1" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FB208916F301FDC1" box="[1252,1293,575,596]" gridcol="6" gridrow="1" pageId="3" pageNumber="136">ss</td>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF648941F368FDEB" box="[160,1380,616,638]" gridrow="2" pageId="3" pageNumber="136" rowspan-1="1" rowspan-3="1">
|
||
<th id="1A5979291246DDC9FF648941F79FFDEB" box="[160,403,616,638]" gridcol="0" gridrow="2" pageId="3" pageNumber="136">2</th>
|
||
<td id="1A5979291246DDC9FDA58941F4D7FDEB" box="[609,731,616,638]" gridcol="2" gridrow="2" pageId="3" pageNumber="136">tu ss,ss</td>
|
||
<td id="1A5979291246DDC9FBE48941F251FDEB" box="[1056,1117,616,638]" gridcol="4" gridrow="2" pageId="3" pageNumber="136">tu</td>
|
||
<td id="1A5979291246DDC9FB458941F2AFFDEB" box="[1153,1187,616,638]" gridcol="5" gridrow="2" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FB208941F301FDEB" box="[1252,1293,616,638]" gridcol="6" gridrow="2" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FA808941F368FDEB" box="[1348,1380,616,638]" gridcol="7" gridrow="2" pageId="3" pageNumber="136">gc</td>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF6489BBF368FD32" box="[160,1380,658,679]" gridrow="3" pageId="3" pageNumber="136" rowspan-4="1">
|
||
<th id="1A5979291246DDC9FF6489BBF79FFD32" box="[160,403,658,679]" gridcol="0" gridrow="3" pageId="3" pageNumber="136">3 ac</th>
|
||
<td id="1A5979291246DDC9FE7C89BBF412FD32" box="[440,542,658,679]" gridcol="1" gridrow="3" pageId="3" pageNumber="136">ss,</td>
|
||
<td id="1A5979291246DDC9FDA589BBF4D7FD32" box="[609,731,658,679]" gridcol="2" gridrow="3" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FCD489BBF5F1FD32" box="[784,1021,658,679]" gridcol="3" gridrow="3" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FB4589BBF2AFFD32" box="[1153,1187,658,679]" gridcol="5" gridrow="3" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FB2089BBF301FD32" box="[1252,1293,658,679]" gridcol="6" gridrow="3" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FA8089BBF368FD32" box="[1348,1380,658,679]" gridcol="7" gridrow="3" pageId="3" pageNumber="136">gc</td>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF648992F368FD44" box="[160,1380,699,721]" gridrow="4" pageId="3" pageNumber="136" rowspan-4="1">
|
||
<th id="1A5979291246DDC9FF648992F79FFD44" box="[160,403,699,721]" gridcol="0" gridrow="4" pageId="3" pageNumber="136">4</th>
|
||
<td id="1A5979291246DDC9FE7C8992F412FD44" box="[440,542,699,721]" gridcol="1" gridrow="4" pageId="3" pageNumber="136">ac, ss (pe)</td>
|
||
<td id="1A5979291246DDC9FDA58992F4D7FD44" box="[609,731,699,721]" gridcol="2" gridrow="4" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FCD48992F5F1FD44" box="[784,1021,699,721]" gridcol="3" gridrow="4" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FB458992F2AFFD44" box="[1153,1187,699,721]" gridcol="5" gridrow="4" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FB208992F301FD44" box="[1252,1293,699,721]" gridcol="6" gridrow="4" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FA808992F368FD44" box="[1348,1380,699,721]" gridcol="7" gridrow="4" pageId="3" pageNumber="136">gc</td>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF6489CAF368FD6F" box="[160,1380,739,762]" gridrow="5" pageId="3" pageNumber="136">
|
||
<th id="1A5979291246DDC9FF6489CAF79FFD6F" box="[160,403,739,762]" gridcol="0" gridrow="5" pageId="3" pageNumber="136">5</th>
|
||
<td id="1A5979291246DDC9FE7C89CAF412FD6F" box="[440,542,739,762]" gridcol="1" gridrow="5" pageId="3" pageNumber="136">ac, ss (pe)</td>
|
||
<td id="1A5979291246DDC9FDA589CAF4D7FD6F" box="[609,731,739,762]" gridcol="2" gridrow="5" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FCD489CAF5F1FD6F" box="[784,1021,739,762]" gridcol="3" gridrow="5" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FBE489CAF251FD6F" box="[1056,1117,739,762]" gridcol="4" gridrow="5" pageId="3" pageNumber="136">tu</td>
|
||
<td id="1A5979291246DDC9FB4589CAF2AFFD6F" box="[1153,1187,739,762]" gridcol="5" gridrow="5" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FB2089CAF301FD6F" box="[1252,1293,739,762]" gridcol="6" gridrow="5" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FA8089CAF368FD6F" box="[1348,1380,739,762]" gridcol="7" gridrow="5" pageId="3" pageNumber="136">gc</td>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF648824F368FCB6" box="[160,1380,781,803]" gridrow="6" pageId="3" pageNumber="136">
|
||
<th id="1A5979291246DDC9FF648824F79FFCB6" box="[160,403,781,803]" gridcol="0" gridrow="6" pageId="3" pageNumber="136">6</th>
|
||
<td id="1A5979291246DDC9FE7C8824F412FCB6" box="[440,542,781,803]" gridcol="1" gridrow="6" pageId="3" pageNumber="136">ac, ss (pe)</td>
|
||
<td id="1A5979291246DDC9FDA58824F4D7FCB6" box="[609,731,781,803]" gridcol="2" gridrow="6" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FCD48824F5F1FCB6" box="[784,1021,781,803]" gridcol="3" gridrow="6" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FBE48824F251FCB6" box="[1056,1117,781,803]" gridcol="4" gridrow="6" pageId="3" pageNumber="136">ac</td>
|
||
<td id="1A5979291246DDC9FB458824F2AFFCB6" box="[1153,1187,781,803]" gridcol="5" gridrow="6" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FB208824F301FCB6" box="[1252,1293,781,803]" gridcol="6" gridrow="6" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FA808824F368FCB6" box="[1348,1380,781,803]" gridcol="7" gridrow="6" pageId="3" pageNumber="136">gc</td>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF64881EF368FCD8" box="[160,1380,823,845]" gridrow="7" pageId="3" pageNumber="136" rowspan-6="1">
|
||
<th id="1A5979291246DDC9FF64881EF79FFCD8" box="[160,403,823,845]" gridcol="0" gridrow="7" pageId="3" pageNumber="136">7</th>
|
||
<td id="1A5979291246DDC9FE7C881EF412FCD8" box="[440,542,823,845]" gridcol="1" gridrow="7" pageId="3" pageNumber="136">ac, ss (pe)</td>
|
||
<td id="1A5979291246DDC9FDA5881EF4D7FCD8" box="[609,731,823,845]" gridcol="2" gridrow="7" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FCD4881EF5F1FCD8" box="[784,1021,823,845]" gridcol="3" gridrow="7" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FBE4881EF251FCD8" box="[1056,1117,823,845]" gridcol="4" gridrow="7" pageId="3" pageNumber="136">ac</td>
|
||
<td id="1A5979291246DDC9FB45881EF2AFFCD8" box="[1153,1187,823,845]" gridcol="5" gridrow="7" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FA80881EF368FCD8" box="[1348,1380,823,845]" gridcol="7" gridrow="7" pageId="3" pageNumber="136">gc</td>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF648849F368FCE3" box="[160,1380,864,886]" gridrow="8" pageId="3" pageNumber="136">
|
||
<th id="1A5979291246DDC9FF648849F79FFCE3" box="[160,403,864,886]" gridcol="0" gridrow="8" pageId="3" pageNumber="136">8</th>
|
||
<td id="1A5979291246DDC9FE7C8849F412FCE3" box="[440,542,864,886]" gridcol="1" gridrow="8" pageId="3" pageNumber="136">ac, ss (pe)</td>
|
||
<td id="1A5979291246DDC9FDA58849F4D7FCE3" box="[609,731,864,886]" gridcol="2" gridrow="8" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FCD48849F5F1FCE3" box="[784,1021,864,886]" gridcol="3" gridrow="8" pageId="3" pageNumber="136">tu ss</td>
|
||
<td id="1A5979291246DDC9FBE48849F251FCE3" box="[1056,1117,864,886]" gridcol="4" gridrow="8" pageId="3" pageNumber="136">ac</td>
|
||
<td id="1A5979291246DDC9FB458849F2AFFCE3" box="[1153,1187,864,886]" gridcol="5" gridrow="8" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FB208849F301FCE3" box="[1252,1293,864,886]" gridcol="6" gridrow="8" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FA808849F368FCE3" box="[1348,1380,864,886]" gridcol="7" gridrow="8" pageId="3" pageNumber="136">gc</td>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF6488A3F368FC0A" box="[160,1380,906,927]" gridrow="9" pageId="3" pageNumber="136">
|
||
<th id="1A5979291246DDC9FF6488A3F79FFC0A" box="[160,403,906,927]" gridcol="0" gridrow="9" pageId="3" pageNumber="136">9 ac</th>
|
||
<td id="1A5979291246DDC9FE7C88A3F412FC0A" box="[440,542,906,927]" gridcol="1" gridrow="9" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FDA588A3F4D7FC0A" box="[609,731,906,927]" gridcol="2" gridrow="9" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FCD488A3F5F1FC0A" box="[784,1021,906,927]" gridcol="3" gridrow="9" pageId="3" pageNumber="136">ne ss</td>
|
||
<td id="1A5979291246DDC9FBE488A3F251FC0A" box="[1056,1117,906,927]" gridcol="4" gridrow="9" pageId="3" pageNumber="136">ac</td>
|
||
<td id="1A5979291246DDC9FB4588A3F2AFFC0A" box="[1153,1187,906,927]" gridcol="5" gridrow="9" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FB2088A3F301FC0A" box="[1252,1293,906,927]" gridcol="6" gridrow="9" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FA8088A3F368FC0A" box="[1348,1380,906,927]" gridcol="7" gridrow="9" pageId="3" pageNumber="136">gc</td>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF64889AF368FC5C" box="[160,1380,947,969]" gridrow="10" pageId="3" pageNumber="136" rowspan-1="1" rowspan-6="1">
|
||
<th id="1A5979291246DDC9FF64889AF79FFC5C" box="[160,403,947,969]" gridcol="0" gridrow="10" pageId="3" pageNumber="136">10</th>
|
||
<td id="1A5979291246DDC9FDA5889AF4D7FC5C" box="[609,731,947,969]" gridcol="2" gridrow="10" pageId="3" pageNumber="136">ss ss</td>
|
||
<td id="1A5979291246DDC9FCD4889AF5F1FC5C" box="[784,1021,947,969]" gridcol="3" gridrow="10" pageId="3" pageNumber="136">ss (m)</td>
|
||
<td id="1A5979291246DDC9FBE4889AF251FC5C" box="[1056,1117,947,969]" gridcol="4" gridrow="10" pageId="3" pageNumber="136">tu</td>
|
||
<td id="1A5979291246DDC9FB45889AF2AFFC5C" box="[1153,1187,947,969]" gridcol="5" gridrow="10" pageId="3" pageNumber="136">ss</td>
|
||
<td id="1A5979291246DDC9FA80889AF368FC5C" box="[1348,1380,947,969]" gridcol="7" gridrow="10" pageId="3" pageNumber="136">gc</td>
|
||
</tr>
|
||
<tr id="598810551246DDC9FF6488F2F368FC67" box="[160,1380,987,1010]" gridrow="11" pageId="3" pageNumber="136" rowspan-1="1" rowspan-6="1" rowspan-7="1">
|
||
<th id="1A5979291246DDC9FF6488F2F79FFC67" box="[160,403,987,1010]" gridcol="0" gridrow="11" pageId="3" pageNumber="136">11</th>
|
||
<td id="1A5979291246DDC9FDA588F2F4D7FC67" box="[609,731,987,1010]" gridcol="2" gridrow="11" pageId="3" pageNumber="136">ss,ss</td>
|
||
<td id="1A5979291246DDC9FCD488F2F5F1FC67" box="[784,1021,987,1010]" gridcol="3" gridrow="11" pageId="3" pageNumber="136">ps,ps (m) lts</td>
|
||
<td id="1A5979291246DDC9FBE488F2F251FC67" box="[1056,1117,987,1010]" gridcol="4" gridrow="11" pageId="3" pageNumber="136">fs (m)</td>
|
||
<td id="1A5979291246DDC9FB4588F2F2AFFC67" box="[1153,1187,987,1010]" gridcol="5" gridrow="11" pageId="3" pageNumber="136">ss</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="E707121712462235FF038F18F5A1FA24" blockId="3.[151,1436,1073,2034]" pageId="3" pageNumber="136">
|
||
Segment 1 without spines or hair. Sensory spots are present in subdorsal, sublateral and ventrolateral positions near the IJ-line, and more medially on the segment in laterodorsal, midlateral and ventromedial positions (
|
||
<figureCitation id="7F830E9212462235FAF78F7CF379FBF8" box="[1331,1397,1109,1133]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="3" pageNumber="136">Figs 1</figureCitation>
|
||
A– B, 3D). Sensory spots on this and all following segments are rather small and rounded, with cuticular papillae. The cuticle appears smooth, except on the ventral side where two indistinct markings form a line with presumably thinner cuticle (
|
||
<figureCitation id="7F830E9212462235FE838FE8F783FB4C" box="[327,399,1217,1241]" captionStart="FIGURE 2" captionStartId="5.[151,250,1440,1463]" captionTargetBox="[167,1419,196,1404]" captionTargetId="figure@5.[167,1419,194,1411]" captionTargetPageId="5" captionText="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="3" pageNumber="136">Figs 2</figureCitation>
|
||
B, 3D). The lines run from ventrolateral positions at the anterior segment margin and extend posteriorly and obliquely to a more ventromedial position. At the level of the IJ-line, the folds bend towards each other, and meet midventrally. The folds are extremely weak in LM (but see
|
||
<figureCitation id="7F830E9212462235FBE68E20F265FAB4" box="[1058,1129,1289,1313]" captionStart="FIGURE 2" captionStartId="5.[151,250,1440,1463]" captionTargetBox="[167,1419,196,1404]" captionTargetId="figure@5.[167,1419,194,1411]" captionTargetPageId="5" captionText="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="3" pageNumber="136">Fig. 2</figureCitation>
|
||
C), but were visible in all specimens prepared for SEM. The posterior segment margin is straight along the dorsal and lateral sides, but forms a broad, midventrally pointed extension (
|
||
<figureCitation id="7F830E9212462235FDB28E78F4CDFAFC" box="[630,705,1361,1385]" captionStart="FIGURE 2" captionStartId="5.[151,250,1440,1463]" captionTargetBox="[167,1419,196,1404]" captionTargetId="figure@5.[167,1419,194,1411]" captionTargetPageId="5" captionText="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="3" pageNumber="136">Figs 2</figureCitation>
|
||
B, 3D). A pair of small indentations in the posterior margin is present in line with the tergosternal junctions of the following segment. Except at the indentations, the segment edge forms a finely serrated fringe around the segment (
|
||
<figureCitation id="7F830E9212462235FCCF8EB0F540FA24" box="[779,844,1433,1457]" captionStart="FIGURE 3" captionStartId="6.[151,250,1577,1600]" captionTargetBox="[171,1422,198,1541]" captionTargetId="figure@6.[171,1422,194,1568]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs showing overviews and details in head and trunk morphology of Dracoderes nidhug nov. sp. A. Dorsal overview of whole specimen. B. Mouth cone with pharynx extended. C. lateroventral overview of whole specimen. D. Segment 2, ventral view. E. Detail of mid- and subdorsal areas of segment 2. F. Detail of lateroventral area of segment 2. G. Segments 7 to 9, lateral view. H. Segment 11 in male, ventral view. I. Segment 11 in male, lateral view. Abbreviations: fs, fringed spine; hs, helioscalid; ld, laterodorsal sensory spot; loo, large outer oral styles; lts, lateral terminal spine; lvt, lateroventral tubule; mds, midadorsal spine; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; scl, subcuticular line; sdt, subdorsal tubule; sl, sublateral sensory spot; soo, small outer oral styles; te, tergal extension; tr, trichoscalid; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218047/files/figure.png" pageId="3" pageNumber="136">Fig. 3</figureCitation>
|
||
D, E–F).
|
||
</paragraph>
|
||
<paragraph id="E707121712462235FF038E94F2D6F88C" blockId="3.[151,1436,1073,2034]" pageId="3" pageNumber="136">
|
||
Segment 2 without spines, but with small tubules in subdorsal and lateroventral positions (
|
||
<figureCitation id="7F830E9212462235FB0C8E94F300FA40" box="[1224,1292,1469,1493]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="3" pageNumber="136">Figs 1</figureCitation>
|
||
A–B, 2C–D, 3E–F). Sensory spots are present as twin pairs in laterodorsal position, and single pairs in ventrolateral and ventromedial positions. The ventrolateral pair is placed towards the anterior end of the segment, whereas the rest are located medially on the segment. Glandular cells are present in paraventral positions on this and all following segments, down to segment 10. Indentations are present in the posterior segment margin in same position as on the preceding segments, but on these and the following segments, the indentations are not as deep as those in segment 1. Cuticular hairs are arranged in two rows that extend from subdorsal to ventromedial positions, with the shortest hairs in the anterior row (
|
||
<figureCitation id="7F830E9212462235FE7E8D90F7F2F944" box="[442,510,1721,1745]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="3" pageNumber="136">Figs 1</figureCitation>
|
||
A, 3E). The rows are interrupted midlaterally by hairless spaces. Anterior part of segment, near overlapping free flap from preceding segment, with secondary fringe resembling a curtain with fine, posterior serration. Posterior segment margin serrated, with alternatingly larger and smaller teeth.
|
||
</paragraph>
|
||
<paragraph id="E707121712462232FF038C0CF2C4FF6D" blockId="3.[151,1436,1073,2034]" lastBlockId="4.[151,1437,151,248]" lastPageId="4" lastPageNumber="137" pageId="3" pageNumber="136">
|
||
Segment 3 with a single dorsal spine, slightly displaced to the left or right, into a more lateral position (
|
||
<figureCitation id="7F830E9212462235FAAE8C0CF6ADF8F4" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="3" pageNumber="136">Figs 1</figureCitation>
|
||
A, 2D, 3A); dorsal spines on this and following six segments are thin and acicular, composed of a basal sheath, and an acicular end piece with finely serrated margins. Trunk cuticle around the attachment points of dorsal spines with conspicuous subcuticular lateral extensions that are best observed with LM (
|
||
<figureCitation id="7F830E9212462235FBE48CB8F26FF83C" box="[1056,1123,1937,1961]" captionStart="FIGURE 2" captionStartId="5.[151,250,1440,1463]" captionTargetBox="[167,1419,196,1404]" captionTargetId="figure@5.[167,1419,194,1411]" captionTargetPageId="5" captionText="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="3" pageNumber="136">Fig. 2</figureCitation>
|
||
D). The proximal part of the spine attaches to the trunk cuticle through a spherical, condyle-like structure, suggesting a ball-and-socket articulation. Spine with perispinal paradorsal sensory spots (
|
||
<figureCitation id="7F830E9212462235FC9F8CF0F593F864" box="[859,927,2009,2033]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="3" pageNumber="136">Fig. 1</figureCitation>
|
||
A). Additional sensory spots are located in laterodorsal, sublateral, ventrolateral and ventromedial positions. Hairs are located as on the preceding segment, but with an additional short row placed above the other two from subdorsal position and across the hairless gap. Glandular cell outlets, secondary fringe and posterior segment margin as on preceding segment.
|
||
</paragraph>
|
||
<caption id="B3C7429F12412232FF538C6CF32CF860" httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="4" pageNumber="137" targetBox="[171,1414,289,1845]" targetPageId="4">
|
||
<paragraph id="E707121712412232FF538C6CF32CF860" blockId="4.[151,1436,1861,2037]" pageId="4" pageNumber="137">
|
||
<emphasis id="D5CCCE0512412232FF538C6CF718F8CE" bold="true" box="[151,276,1861,1884]" pageId="4" pageNumber="137">FIGURE 1.</emphasis>
|
||
Line art illustrations showing general habitus in
|
||
<taxonomicName id="20B8699412412232FCC88C6FF5DDF8C9" box="[780,977,1862,1884]" class="Kinorhyncha" family="Dracoderidae" genus="Dracoderes" kingdom="Animalia" order="Cyclorhagida" pageId="4" pageNumber="137" phylum="Cephalorhyncha" rank="species" species="nidhug">
|
||
<emphasis id="D5CCCE0512412232FCC88C6FF5DDF8C9" box="[780,977,1862,1884]" italics="true" pageId="4" pageNumber="137">Dracoderes nidhug</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D5CCCE0512412232FC1C8C6FF246F8CE" bold="true" box="[984,1098,1862,1883]" pageId="4" pageNumber="137">nov. sp. A.</emphasis>
|
||
Female, dorsal view.
|
||
<emphasis id="D5CCCE0512412232FAE88C6FF348F8CE" bold="true" box="[1324,1348,1862,1883]" pageId="4" pageNumber="137">B.</emphasis>
|
||
Female, ventral view.
|
||
<emphasis id="D5CCCE0512412232FEE08C4DF730F8EE" bold="true" box="[292,316,1892,1915]" pageId="4" pageNumber="137">C.</emphasis>
|
||
Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B3C7429F12402233FF538E89F4EDF918" httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="5" pageNumber="138" targetBox="[167,1419,196,1404]" targetPageId="5">
|
||
<paragraph id="E707121712402233FF538E89F4EDF918" blockId="5.[151,1436,1440,1677]" pageId="5" pageNumber="138">
|
||
<emphasis id="D5CCCE0512402233FF538E89F719FA20" bold="true" box="[151,277,1440,1463]" pageId="5" pageNumber="138">FIGURE 2.</emphasis>
|
||
Light micrographs showing overviews and details in
|
||
<taxonomicName id="20B8699412402233FC8F8E89F21CFA23" box="[843,1040,1440,1462]" class="Kinorhyncha" family="Dracoderidae" genus="Dracoderes" kingdom="Animalia" order="Cyclorhagida" pageId="5" pageNumber="138" phylum="Cephalorhyncha" rank="species" species="nidhug">
|
||
<emphasis id="D5CCCE0512402233FC8F8E89F21CFA23" box="[843,1040,1440,1462]" italics="true" pageId="5" pageNumber="138">Dracoderes nidhug</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D5CCCE0512402233FBDE8E88F261FA23" bold="true" box="[1050,1133,1441,1462]" pageId="5" pageNumber="138">nov. sp.</emphasis>
|
||
, displaying female holotype, ZMUC KIN-649 (
|
||
<emphasis id="D5CCCE0512402233FE918E96F7CCFA41" bold="true" box="[341,448,1471,1492]" pageId="5" pageNumber="138">A–B, E–F</emphasis>
|
||
) and male allotype, ZMUC KIN-550 (
|
||
<emphasis id="D5CCCE0512402233FC938E97F5C0FA40" bold="true" box="[855,972,1470,1493]" pageId="5" pageNumber="138">C–D, G–H</emphasis>
|
||
).
|
||
<emphasis id="D5CCCE0512402233FC278E96F5F0FA41" bold="true" box="[995,1020,1471,1492]" pageId="5" pageNumber="138">A.</emphasis>
|
||
Ventral overview of trunk and head.
|
||
<emphasis id="D5CCCE0512402233FA408E96F390FA41" bold="true" box="[1412,1436,1471,1492]" pageId="5" pageNumber="138">B.</emphasis>
|
||
Neck and segment 1, dorsal view.
|
||
<emphasis id="D5CCCE0512402233FE368EF4F406FA61" bold="true" box="[498,522,1501,1524]" pageId="5" pageNumber="138">C.</emphasis>
|
||
Neck and segments 1 and 2, ventral view.
|
||
<emphasis id="D5CCCE0512402233FC7F8EF7F5D8FA66" bold="true" box="[955,980,1502,1523]" pageId="5" pageNumber="138">D.</emphasis>
|
||
Segments 2 to 4, dorsal view.
|
||
<emphasis id="D5CCCE0512402233FACD8EF7F32CFA66" bold="true" box="[1289,1312,1502,1523]" pageId="5" pageNumber="138">E.</emphasis>
|
||
Segments 7 to 9, ventral view.
|
||
<emphasis id="D5CCCE0512402233FE908ED5F769F984" bold="true" box="[340,357,1532,1553]" pageId="5" pageNumber="138">F.</emphasis>
|
||
Detail showing areas around tergosternal junctions on segments 5 and 6.
|
||
<emphasis id="D5CCCE0512402233FB968ED2F26AF987" bold="true" box="[1106,1126,1531,1554]" pageId="5" pageNumber="138">G.</emphasis>
|
||
Segment 11 and penile spines in male.
|
||
<emphasis id="D5CCCE0512402233FF378D32F701F9A5" bold="true" box="[243,269,1563,1584]" pageId="5" pageNumber="138">H.</emphasis>
|
||
Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E707121712402233FF038D90F5C5F8AB" blockId="5.[151,1437,1721,1998]" pageId="5" pageNumber="138">Segment 4 with single dorsal spine, slightly displaced to a more lateral position, opposite to the side of the dorsal spine on the preceding segment. Hairs are arranged in three rows, all interrupted by the same hairless areas as on segment 2; however with a small laterodorsal patch of hair in each row. Sensory spots, glandular cell outlets, secondary fringe and posterior segment margin as on preceding segment.</paragraph>
|
||
<paragraph id="E707121712402233FF038C60F6F0F85B" blockId="5.[151,1437,1721,1998]" pageId="5" pageNumber="138">
|
||
Segment 5 with single dorsal spine, slightly displaced to a more lateral position, opposite to the side of the dorsal spine on the preceding segment. A pair of short tubules is present in the lateroventral positions (
|
||
<figureCitation id="7F830E9212402233FAFE8C44F373F810" box="[1338,1407,1901,1925]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="5" pageNumber="138">Figs 1</figureCitation>
|
||
B, 2F). Sensory spots, glandular cells, cuticular hairs, secondary fringe and posterior segment margin as on preceding segment.
|
||
</paragraph>
|
||
<caption id="B3C7429F12432230FF538D00F586F883" httpUri="https://zenodo.org/record/218047/files/figure.png" pageId="6" pageNumber="139" targetBox="[171,1422,198,1541]" targetPageId="6">
|
||
<paragraph id="E707121712432230FF538D00F586F883" blockId="6.[151,1437,1577,1814]" pageId="6" pageNumber="139">
|
||
<emphasis id="D5CCCE0512432230FF538D00F71AF9AA" bold="true" box="[151,278,1577,1600]" pageId="6" pageNumber="139">FIGURE 3.</emphasis>
|
||
Scanning electron micrographs showing overviews and details in head and trunk morphology of
|
||
<taxonomicName id="20B8699412432230FAE18D03F6D1F9CB" class="Kinorhyncha" family="Dracoderidae" genus="Dracoderes" kingdom="Animalia" order="Cyclorhagida" pageId="6" pageNumber="139" phylum="Cephalorhyncha" rank="species" species="nidhug">
|
||
<emphasis id="D5CCCE0512432230FAE18D03F6D1F9CB" italics="true" pageId="6" pageNumber="139">Dracoderes nidhug</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D5CCCE0512432230FF218D60F75BF9CB" bold="true" box="[229,343,1609,1630]" pageId="6" pageNumber="139">nov. sp. A.</emphasis>
|
||
Dorsal overview of whole specimen.
|
||
<emphasis id="D5CCCE0512432230FD1B8D60F4FBF9CB" bold="true" box="[735,759,1609,1630]" pageId="6" pageNumber="139">B.</emphasis>
|
||
Mouth cone with pharynx extended.
|
||
<emphasis id="D5CCCE0512432230FBBD8D61F29EF9CA" bold="true" box="[1145,1170,1608,1631]" pageId="6" pageNumber="139">C.</emphasis>
|
||
lateroventral overview of whole specimen.
|
||
<emphasis id="D5CCCE0512432230FE838D41F753F9E8" bold="true" box="[327,351,1640,1661]" pageId="6" pageNumber="139">D.</emphasis>
|
||
Segment 2, ventral view.
|
||
<emphasis id="D5CCCE0512432230FDA28D41F471F9E8" bold="true" box="[614,637,1640,1661]" pageId="6" pageNumber="139">E.</emphasis>
|
||
Detail of mid- and subdorsal areas of segment 2.
|
||
<emphasis id="D5CCCE0512432230FBB78D41F288F9E8" bold="true" box="[1139,1156,1640,1661]" pageId="6" pageNumber="139">F.</emphasis>
|
||
Detail of lateroventral area of segment 2.
|
||
<emphasis id="D5CCCE0512432230FEEB8DACF74FF909" bold="true" box="[303,323,1669,1692]" pageId="6" pageNumber="139">G.</emphasis>
|
||
Segments 7 to 9, lateral view.
|
||
<emphasis id="D5CCCE0512432230FD558DAFF4A7F90E" bold="true" box="[657,683,1670,1691]" pageId="6" pageNumber="139">H.</emphasis>
|
||
Segment 11 in male, ventral view.
|
||
<emphasis id="D5CCCE0512432230FBE78DAFF23FF90E" bold="true" box="[1059,1075,1670,1691]" pageId="6" pageNumber="139">I.</emphasis>
|
||
Segment 11 in male, lateral view. Abbreviations: fs, fringed spine; hs, helioscalid; ld, laterodorsal sensory spot; loo, large outer oral styles; lts, lateral terminal spine; lvt, lateroventral tubule; mds, midadorsal spine; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; scl, subcuticular line; sdt, subdorsal tubule; sl, sublateral sensory spot; soo, small outer oral styles; te, tergal extension; tr, trichoscalid; vl, ventrolateral sensory spot; vm, ventromedial sensory spot.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E707121712432231FF038C6AF2C1FF6D" blockId="6.[151,1437,1859,2027]" lastBlockId="7.[151,1437,151,1111]" lastPageId="7" lastPageNumber="140" pageId="6" pageNumber="139">
|
||
Segment 6 with single dorsal spine, slightly displaced to a more lateral position, opposite to the side of the dorsal spine on the preceding segment. A pair of acicular spines present in the lateroventral positions (
|
||
<figureCitation id="7F830E9212432230FAF08C4EF379F8EA" box="[1332,1397,1895,1919]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="6" pageNumber="139">Fig. 1</figureCitation>
|
||
B); lateroventral spines are composed as the dorsal ones, with proximal ball, basal sheath and acicular end piece, but appear somewhat shorter and stouter. Trunk cuticle strongly reinforced around the attachment point of the lateroventral spines, creating conspicuous subcuticular thickenings and ventrally directed extensions (see similar from segment 8 on
|
||
<figureCitation id="7F830E9212422231FE438BBEF7C3FF3A" box="[391,463,151,175]" captionStart="FIGURE 2" captionStartId="5.[151,250,1440,1463]" captionTargetBox="[167,1419,196,1404]" captionTargetId="figure@5.[167,1419,194,1411]" captionTargetPageId="5" captionText="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="7" pageNumber="140">Fig. 2</figureCitation>
|
||
E). Similar extensions are present around the lateroventral spines on all following segments. Cuticular hairs appear as on the preceding segment, but are sparser and in only two rows. Sensory spots, glandular cells outlets, secondary fringe and posterior segment margin as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="E707121712422231FF038A2DF541FEF1" blockId="7.[151,1437,151,1111]" pageId="7" pageNumber="140">Segment 7 is identical with preceding segment, except for the dorsal spine that is laterally displaced to the opposite side, the ventromedial sensory spots that are absent and cuticular hairs being reduced to a patch of two rows placed laterodorsally, and a few single hairs subdorsally.</paragraph>
|
||
<paragraph id="E707121712422231FF038A46F5E0FE39" blockId="7.[151,1437,151,1111]" pageId="7" pageNumber="140">
|
||
Segment 8 identical with segment 6, except for the presence of short lateral accessory tubules (
|
||
<figureCitation id="7F830E9212422231FACE8A46F35CFE12" box="[1290,1360,367,391]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="7" pageNumber="140">Figs 1</figureCitation>
|
||
B, 2E, 3G), and the cuticular hairs now only being present as a few subdorsal ones.
|
||
</paragraph>
|
||
<paragraph id="E707121712422231FF038A9EF318FD8D" blockId="7.[151,1437,151,1111]" pageId="7" pageNumber="140">
|
||
Segment 9 with dorsal and lateroventral spines (
|
||
<figureCitation id="7F830E9212422231FD358A9EF53AFE5A" box="[753,822,439,463]" captionStart="FIGURE 3" captionStartId="6.[151,250,1577,1600]" captionTargetBox="[171,1422,198,1541]" captionTargetId="figure@6.[171,1422,194,1568]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs showing overviews and details in head and trunk morphology of Dracoderes nidhug nov. sp. A. Dorsal overview of whole specimen. B. Mouth cone with pharynx extended. C. lateroventral overview of whole specimen. D. Segment 2, ventral view. E. Detail of mid- and subdorsal areas of segment 2. F. Detail of lateroventral area of segment 2. G. Segments 7 to 9, lateral view. H. Segment 11 in male, ventral view. I. Segment 11 in male, lateral view. Abbreviations: fs, fringed spine; hs, helioscalid; ld, laterodorsal sensory spot; loo, large outer oral styles; lts, lateral terminal spine; lvt, lateroventral tubule; mds, midadorsal spine; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; scl, subcuticular line; sdt, subdorsal tubule; sl, sublateral sensory spot; soo, small outer oral styles; te, tergal extension; tr, trichoscalid; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218047/files/figure.png" pageId="7" pageNumber="140">Fig. 3</figureCitation>
|
||
G). The dorsal spine is located in a perfect middorsal position. Nephridial pores present in lateral accessory positions (
|
||
<figureCitation id="7F830E9212422231FCBB8AF5F5C9FE61" box="[895,965,476,500]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="7" pageNumber="140">Figs 1</figureCitation>
|
||
B, 2E). The segment is hairless. Sensory spots, glandular cells outlets, secondary fringe and posterior segment margin as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="E707121712422231FF03890DF5C5FD10" blockId="7.[151,1437,151,1111]" pageId="7" pageNumber="140">
|
||
Segment 10 without spines and cuticular hairs, but with pair of lateroventral tubules (
|
||
<figureCitation id="7F830E9212422231FB6A890DF2F9FDA9" box="[1198,1269,548,572]" captionStart="FIGURE 2" captionStartId="5.[151,250,1440,1463]" captionTargetBox="[167,1419,196,1404]" captionTargetId="figure@5.[167,1419,194,1411]" captionTargetPageId="5" captionText="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="7" pageNumber="140">Fig. 2</figureCitation>
|
||
B–C). Sensory spots present in subdorsal, laterodorsal, sublateral (males only) and ventrolateral positions. Glandular cells outlets, secondary fringe and posterior segment margin as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="E707121712422231FF0389A6F488FBC2" blockId="7.[151,1437,151,1111]" pageId="7" pageNumber="140">
|
||
Segment 11 with lateral terminal spines. Males with two flexible and long (60–70 µm) penile spines emerging laterally from the intersegmental joint between segment 10 and 11 (
|
||
<figureCitation id="7F830E9212422231FC56899DF5DBFD59" box="[914,983,692,716]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="7" pageNumber="140">Figs 1</figureCitation>
|
||
C, 2G–H, 3H–I). In most specimens the two penile spines tended to be stuck together, appearing as one spine with a grove, but one specimen clearly showed that the spines are separated (Fig,
|
||
<date id="930634D712422231FDA889D5F48AFC81" box="[620,646,764,788]" pageId="7" pageNumber="140">3I</date>
|
||
). Males furthermore with a pair of short (14 µm), lateroventral flexible spines, with tiny fringes arranged around them as rings (
|
||
<figureCitation id="7F830E9212422231FCE98836F575FCA2" box="[813,889,799,823]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="7" pageNumber="140">Figs. 1</figureCitation>
|
||
C–2H, 3H–I). Females do not show any outer, sexual dimorphic characters (
|
||
<figureCitation id="7F830E9212422231FE26886DF42CFCC9" box="[482,544,836,860]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="7" pageNumber="140">Fig. 1</figureCitation>
|
||
A–B). Two pairs of subdorsal sensory spots present, one on the anterior part of segment, and the second one towards the tips of the tergal extensions. Sensory spots furthermore present in ventrolateral positions near the lateral, posterior corners of the sternal plates (
|
||
<figureCitation id="7F830E9212422231FBF888A5F288FC31" box="[1084,1156,908,932]" captionStart="FIGURE 3" captionStartId="6.[151,250,1577,1600]" captionTargetBox="[171,1422,198,1541]" captionTargetId="figure@6.[171,1422,194,1568]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs showing overviews and details in head and trunk morphology of Dracoderes nidhug nov. sp. A. Dorsal overview of whole specimen. B. Mouth cone with pharynx extended. C. lateroventral overview of whole specimen. D. Segment 2, ventral view. E. Detail of mid- and subdorsal areas of segment 2. F. Detail of lateroventral area of segment 2. G. Segments 7 to 9, lateral view. H. Segment 11 in male, ventral view. I. Segment 11 in male, lateral view. Abbreviations: fs, fringed spine; hs, helioscalid; ld, laterodorsal sensory spot; loo, large outer oral styles; lts, lateral terminal spine; lvt, lateroventral tubule; mds, midadorsal spine; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; scl, subcuticular line; sdt, subdorsal tubule; sl, sublateral sensory spot; soo, small outer oral styles; te, tergal extension; tr, trichoscalid; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218047/files/figure.png" pageId="7" pageNumber="140">Fig. 3</figureCitation>
|
||
H). Glandular cells and cuticular hairs are not present. Secondary fringe present dorsally, on some specimens stretching nearly halfway down the segment. Posterior margin of tergal plate bulges before extending into triangular tergal extensions, extending well beyond the truncate sternal plates (
|
||
<figureCitation id="7F830E9212422231FD1D88DEF511FB9A" box="[729,797,1015,1039]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="7" pageNumber="140">Figs 1</figureCitation>
|
||
A–C, 2H, 3H). Dorsal side of tergal extensions smooth ventral side densely covered with short, papillar hairs (
|
||
<figureCitation id="7F830E9212422231FCCC8F35F54BFBA1" box="[776,839,1052,1076]" captionStart="FIGURE 1" captionStartId="4.[151,250,1861,1884]" captionTargetBox="[171,1414,289,1845]" captionTargetId="figure@4.[170,1429,254,1845]" captionTargetPageId="4" captionText="FIGURE 1. Line art illustrations showing general habitus in Dracoderes nidhug nov. sp. A. Female, dorsal view. B. Female, ventral view. C. Segments 10 and 11 in male, ventral view. Abbreviations: cf, cuticular folding between placids; fs, fringed spine; gco, glandular cell outlet; icl, intracuticular line; lat, lateral accessory tubule; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; mds, middorsal spine; ml, midlateral sensory spot; ne, nephridiopore; pd, paradorsal sensory spot; pds, paradorsal spine; pe, penile spine; pl, placid; sd, subdorsal sensory spot; sdt, subdorsal tubule; sl, sublateral sensory spot; sus, subcuticular structure; te, tergal extension; vl, ventrolateral sensory spot; vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218045/files/figure.png" pageId="7" pageNumber="140">Figs1</figureCitation>
|
||
C, 3H). Each tergal extension has a flexible, spinous elongation extending from the tip (
|
||
<figureCitation id="7F830E9212422231FDD98F16F46CFBC2" box="[541,608,1087,1111]" captionStart="FIGURE 2" captionStartId="5.[151,250,1440,1463]" captionTargetBox="[167,1419,196,1404]" captionTargetId="figure@5.[167,1419,194,1411]" captionTargetPageId="5" captionText="FIGURE 2. Light micrographs showing overviews and details in Dracoderes nidhug nov. sp., displaying female holotype, ZMUC KIN- 649 (A – B, E – F) and male allotype, ZMUC KIN- 550 (C – D, G – H). A. Ventral overview of trunk and head. B. Neck and segment 1, dorsal view. C. Neck and segments 1 and 2, ventral view. D. Segments 2 to 4, dorsal view. E. Segments 7 to 9, ventral view. F. Detail showing areas around tergosternal junctions on segments 5 and 6. G. Segment 11 and penile spines in male. H. Segment 11 in male, ventral view. Abbreviations: fs, fringed spine; icl, intracuticular line; lat, lateral accessory tubule; ldp, laterodorsal placid; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tubule; ne, nephridiopore; pds, paradorsal spine; pe, penile spine; sdp, subdorsal placid; sdt, subdorsal tubule; sus, subcuticular structure; te, tergal extension; tr, trichoscalid, vm, ventromedial sensory spot." httpUri="https://zenodo.org/record/218046/files/figure.png" pageId="7" pageNumber="140">Fig. 2</figureCitation>
|
||
H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |