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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1021.63147" ID-GBIF-Dataset="5b6e22ed-ee96-4fa0-990b-0e8f78961670" ID-PMC="PMC7940320" ID-Pensoft-Pub="1313-2970-1021-1" ID-Pensoft-UUID="6CABF438D29F5681A0D771275C5791FF" ID-PubMed="33716539" ID-ZooBank="E24182958A9F425D87285686C41FF59B" ModsDocID="1313-2970-1021-1" checkinTime="1614718828288" checkinUser="pensoft" docAuthor="Skarzynski, Dariusz, Smolis, Adrian, Kovac, Ľubomir &amp; Porco, David" docDate="2021" docId="3048909794415CAFAF7712BC8C024C10" docLanguage="en" docName="ZooKeys 1021: 1-18" docOrigin="ZooKeys 1021" docPubDate="2021-03-01" docSource="http://dx.doi.org/10.3897/zookeys.1021.63147" docTitle="Ceratophysella stachi Skarzynski, Smolis &amp; Porco 2021, sp. nov." docType="treatment" docUuid="40BF0CC2-AF12-43DB-9F35-368E6EC004F8" docUuidSource="ZooBank" docVersion="6" id="6CABF438D29F5681A0D771275C5791FF" lastPageNumber="1" masterDocId="6CABF438D29F5681A0D771275C5791FF" masterDocTitle="A new European species of Ceratophysella (Collembola, Hypogastruridae) revealed by morphological data and DNA barcodes" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1668149838477" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new European species of Ceratophysella (Collembola, Hypogastruridae) revealed by morphological data and DNA barcodes</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Skarzynski, Dariusz</mods:namePart>
<mods:affiliation>Institute of Environmental Biology, Department of Invertebrate Biology, Evolution and Conservation, University of Wroclaw, Przybyszewskiego 65, 51 - 148, Wroclaw, Poland</mods:affiliation>
<mods:nameIdentifier type="email">dariusz.skarzynski@uwr.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smolis, Adrian</mods:namePart>
<mods:affiliation>Institute of Environmental Biology, Department of Invertebrate Biology, Evolution and Conservation, University of Wroclaw, Przybyszewskiego 65, 51 - 148, Wroclaw, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kovac, Ľubomir</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8194-2128</mods:nameIdentifier>
<mods:affiliation>Department of Zoology, Institute of Biology and Ecology, Faculty of Science, Pavol Jozef Safarik University, Moyzesova 11, 041 54, Kosice, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Porco, David</mods:namePart>
<mods:affiliation>Musee National d'Histoire Naturelle, 25 rue Munster, 2160, Luxembourg, Luxembourg</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1021</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>18</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1021.63147</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1021.63147</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1021-1</mods:identifier>
<mods:identifier type="ZooBank">E24182958A9F425D87285686C41FF59B</mods:identifier>
<mods:identifier type="Pensoft-UUID">6CABF438D29F5681A0D771275C5791FF</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="179985158" LSID="urn:lsid:zoobank.org:act:40BF0CC2-AF12-43DB-9F35-368E6EC004F8" httpUri="http://treatment.plazi.org/id/3048909794415CAFAF7712BC8C024C10" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/40BF0CC2-AF12-43DB-9F35-368E6EC004F8" authority="Skarżyński, Smolis &amp; Porco" authorityName="Skarzynski, Smolis &amp; Porco" authorityYear="2021" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella stachi" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stachi" status="sp. nov.">
Ceratophysella stachi
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
, Smolis &amp; Porco
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Ceratophysella stachi sp. nov. 3 chaetotaxy of thoracic tergum II 4 chaetotaxy of abdominal terga III-VI." figureDoi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1">Figures 3-4</figureCitation>
<figureCitation captionStart="Figures 59" captionStartId="F4" captionText="Figures 5 - 9. Ceratophysella stachi sp. nov. 5 chaetotaxy of dorsal side of antennal segments III-IV 6 chaetotaxy of ventral side of antennal segment IV 7 postantennal organ and neighbor ocelli 8 claw I 9 dens and mucro." figureDoi="10.3897/zookeys.1021.63147.figures5-9" httpUri="https://binary.pensoft.net/fig/515256" pageId="0" pageNumber="1">, 5-9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="Skarżyński &amp; Smolis &amp; Kováč &amp; Porco, 2021" authorityName="Skarżyński &amp; Smolis &amp; Kováč &amp; Porco" authorityYear="2021" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella granulata" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="granulata" status="sp. nov.">Ceratophysella granulata</taxonomicName>
: Fjellberg, 1998: 41.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">
<typeStatus>Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
(
<collectionCode>DIBEC</collectionCode>
)
</emphasis>
:
<materialsCitation ID-GBIF-Occurrence="3063647306" collectingDate="2009-10-20" collectionCode="DIBEC" collectorName="M. Furgol. Paratypes" country="Poland" county="Tylawa village" elevation="500" location="Carpathians" municipality="Ostra Gora" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
female,
<collectingCountry name="Poland">Poland</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:86655269F2F9314ECCF849DE29A50ACE" country="Poland" county="Tylawa village" municipality="Ostra Gora" name="Carpathians">Carpathians</location>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:081D983A0B33D1D24C47D8FBDBD211BE" country="Poland" county="Tylawa village" municipality="Ostra Gora" name="Beskid Niski Mts">Beskid Niski Mts</location>
, litter of the
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:A47C4058248C2996AD6E5A18C0D1B870" country="Poland" county="Tylawa village" municipality="Ostra Gora" name="Carpathian">Carpathian</location>
beech forest on the slopes of
<collectingMunicipality>
Ostra
<normalizedToken originalValue="Góra">Gora</normalizedToken>
</collectingMunicipality>
near
<collectingCounty>Tylawa village</collectingCounty>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</quantity>
a.s.l.
</elevation>
,
<date value="2009-10-20">
<collectingDate value="2009-10-20">20.X.2009</collectingDate>
</date>
, leg.
<collectorName>
M.
<normalizedToken originalValue="Furgoł">Furgol</normalizedToken>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
</collectorName>
(
<emphasis bold="true" pageId="0" pageNumber="1">
<collectionCode>DIBEC</collectionCode>
)
</emphasis>
:
<specimenCount type="male">2 males</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063647312" collectingDate="2001-05-14" collectionCode="DIBEC" collectorName="A. Smolis &amp; Smolis, D" country="Poland" county="Tylawa village" elevation="500" location="Carpathians" municipality="Ostra Gora" specimenCount="1" typeStatus="holotype">
male, juv.,
<date value="2001-05-14">
<collectingDate value="2001-05-14">14.V.2001</collectingDate>
</date>
, leg.
<collectorName>
A.
<collectorName>Smolis, D</collectorName>
</collectorName>
.
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
, other data same as holotype
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063647304" collectingDate="2002-05-14" collectionCode="DIBEC" collectorName="A. Smolis &amp; Smolis, D" country="Poland" county="Tylawa village" elevation="500" location="Carpathians" municipality="Ostra Gora" specimenCount="8" specimenCount-female="3" specimenCount-male="5" typeStatus="holotype">
<specimenCount type="female">3 females</specimenCount>
,
<specimenCount type="male">5 males</specimenCount>
, juv.,
<date value="2002-05-14">
<collectingDate value="2002-05-14">14.V.2002</collectingDate>
</date>
, leg.
<collectorName>
A.
<collectorName>Smolis, D</collectorName>
</collectorName>
.
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
, other data same as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3063647317" collectingDate="1994-03-20" collectingDateMax="1994-03-24" collectingDateMin="1994-03-20" collectionCode="AF" collectorName="A. Fjellberg &amp; Funen &amp; Syltemade Adal" country="Denmark" county="Jutland" location="Rold Skov" municipality="Himmerland" specimenCount="8" specimenCount-female="6" specimenCount-male="2">
<collectingCountry name="Denmark">Denmark</collectingCountry>
(
<collectionCode>AF</collectionCode>
, leg.
<collectorName>A. Fjellberg</collectorName>
): female,
<collectingCounty>Jutland</collectingCounty>
,
<collectingMunicipality>Himmerland</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:3EBF1D3C7C0A1D09D56F661CE2E878B4" country="Denmark" county="Jutland" municipality="Himmerland" name="Rold Skov">Rold Skov</location>
,
<taxonomicName class="Dicotyledoneae" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Fagus</emphasis>
</taxonomicName>
liter,
<date value="1994-03-20">
<collectingDate value="1994-03-20">20.III.1994</collectingDate>
</date>
, 94.024;
<collectorName>Funen</collectorName>
:
<specimenCount type="female">6 females</specimenCount>
,
<specimenCount type="male">2 males</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:15B5C4FB0CDCE67CE9C1BDB5EF7E8E60" country="Denmark" county="Jutland" municipality="Himmerland" name="Fiskerup Skov">Fiskerup Skov</location>
, forest stream,
<date value="1994-03-24">
<collectingDate value="1994-03-24">24.III.1994</collectingDate>
</date>
, 94.076; female,
<collectorName>Syltemade Adal</collectorName>
,
<taxonomicName class="Dicotyledoneae" family="Oleaceae" genus="Fraxinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fraxinus" order="Oleales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Fraxinus</emphasis>
</taxonomicName>
/
<taxonomicName class="Insecta" family="Tingidae" genus="Ulmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Ulmus" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Ulmus</emphasis>
</taxonomicName>
/
<taxonomicName class="Dicotyledoneae" family="Caprifoliaceae" genus="Viburnum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Viburnum" order="Dipsacales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Viburnum</emphasis>
</taxonomicName>
litter,
<date value="1994-03-23">
<collectingDate value="1994-03-23">23.III.1994</collectingDate>
</date>
, 94.070
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063647314" collectingDate="1972-05-01" collectionCode="SMNG, V" collectorName="Hiebsch" country="Germany" location="Wanninchen" specimenCount="12" specimenCount-female="6" specimenCount-male="3" stateProvince="Brandenburg">
<collectingCountry name="Germany">Germany</collectingCountry>
(
<collectionCode country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" type="">SMNG</collectionCode>
):
<collectingRegion country="Germany" name="Brandenburg">Brandenburg</collectingRegion>
:
<specimenCount type="female">6 females</specimenCount>
,
<specimenCount type="male">3 males</specimenCount>
,
<specimenCount type="generic">3 juv.</specimenCount>
,
<normalizedToken originalValue="“Wanninchen”">
&quot;
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:DF27A5EC579138028761B884FAB38999" country="Germany" name="Wanninchen" stateProvince="Brandenburg">Wanninchen</location>
&quot;
</normalizedToken>
nature reserve, bog,
<date value="1972-05-01">
<collectingDate value="1972-05-01">
1.
<collectionCode country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.1972
</collectingDate>
</date>
, leg.
<collectorName>Hiebsch</collectorName>
</materialsCitation>
, 11911;
<specimenCount type="female">9 females</specimenCount>
, male,
<normalizedToken originalValue="“Wanninchen”">&quot;Wanninchen&quot;</normalizedToken>
nature reserve, wet heather, 1972, leg. Hiebsch,
<materialsCitation ID-GBIF-Occurrence="3063647308" collectingDate="1972-05-01" collectionCode="V" collectorName="Hiebsch" country="Germany" location="Bergener Moor" specimenCount="2">
11911; male,
<specimenCount type="generic">2 juv.</specimenCount>
, &quot;
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:8210A8F00DB43BCB8F25E9344CE0A682" country="Germany" name="Bergener Moor">Bergener Moor</location>
&quot; nature reserve, moor,
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Bryopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="0" pageNumber="1" phylum="Bryophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sphagnum</emphasis>
</taxonomicName>
, heather, pine forest,
<date value="1972-05-01">
<collectingDate value="1972-05-01">
1.
<collectionCode country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.1972
</collectingDate>
</date>
, leg.
<collectorName>Hiebsch</collectorName>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3063647302" collectingDate="1985" collectingDateMax="1995" collectingDateMin="1985" collectorName="W. Boehle. Mecklenburg-West Pomerania" country="Germany" location="Vogelsberg" specimenCount="5" specimenCount-female="5" stateProvince="Hessen">
11912.
<collectingRegion country="Germany" name="Hessen">Hesse</collectingRegion>
:
<specimenCount type="female">5 females</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:C89627ECBC5296CB65890DBD5F79F89D" country="Germany" name="Vogelsberg" stateProvince="Hessen">Vogelsberg</location>
,
<date value="1985" valueMax="1995">
<collectingDate value="1985" valueMax="1995">1985-1995</collectingDate>
</date>
, leg.
<collectorName>
W.
<normalizedToken originalValue="Böhle">Boehle</normalizedToken>
. Mecklenburg-West Pomerania
</collectorName>
</materialsCitation>
:
<materialsCitation ID-GBIF-Occurrence="3063647318" collectingDate="2013-10-16" collectorName="U. Burkhardt" country="Germany" location="Neustrelitz" municipality="Mueritz National Park" specimenCount="1">
female,
<collectingMunicipality>
<normalizedToken originalValue="Müritz">Mueritz</normalizedToken>
National Park
</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:6A30EFE34EB00DB2350D0B93F106FB7D" country="Germany" municipality="Mueritz National Park" name="Neustrelitz">Neustrelitz</location>
, soil
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" unit="cm" value="2.5" valueMax="5.0" valueMin="0.0">0-5 cm</quantity>
, alder swamp woods not on acid peat,
<date value="2013-10-16">
<collectingDate value="2013-10-16">16.X.2013</collectingDate>
</date>
, leg.
<collectorName>U. Burkhardt</collectorName>
</materialsCitation>
, 264-F865;
<specimenCount type="female">4 females</specimenCount>
,
<specimenCount type="male">2 males</specimenCount>
, Serrahn, Hauptmannsberg near Feldberg, sandy-gravelly moraine, largely unforested since the Middle Ages, 1973, leg. Hiebsch/
<materialsCitation ID-GBIF-Occurrence="3063647313" collectingDate="2013-10-16" collectingDateMax="2014-11-05" collectingDateMin="2013-10-16" collectionCode="R" collectorName="U. Burkardt &amp; U. Burkhardt" country="Germany" county="Greifswald" location="Klockenbruch" municipality="Serrahn" specimenCount="1">
ILN
<collectingCounty>Greifswald</collectingCounty>
, 11915; female,
<collectingMunicipality>Serrahn</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:88F1EDF36464C550C286DC74F4328485" country="Germany" county="Greifswald" municipality="Serrahn" name="Klockenbruch">Klockenbruch</location>
, active, relatively undamaged raised bogs, moss/
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Bryopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="0" pageNumber="1" phylum="Bryophyta" rank="genus">Sphagnum</taxonomicName>
,
<date value="2013-10-16">
<collectingDate value="2013-10-16">16.X.2013</collectingDate>
</date>
, leg.
<collectionCode country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:277124ED57256F15946554E4410F3E77" country="Germany" county="Greifswald" municipality="Serrahn" name="Lehmitz">Lehmitz</location>
and
<collectorName>U. Burkardt</collectorName>
, 261-01; male,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:903487B02C97F0B1DCDDCCF7FA6000EC" country="Germany" county="Greifswald" municipality="Serrahn" name="Serrahn">Serrahn</location>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:47702EC0F9B6142B0A0651AD46C1EA28" country="Germany" county="Greifswald" municipality="Serrahn" name="Klockenbruch">Klockenbruch</location>
, active, relatively undamaged raised bogs, moss/
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Bryopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="0" pageNumber="1" phylum="Bryophyta" rank="genus">Sphagnum</taxonomicName>
,
<date value="2014-11-05">
<collectingDate value="2014-11-05">5.XI.2014</collectingDate>
</date>
, leg.
<collectionCode country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:471E04700BB8324A4AEF3C0E8BEBDA05" country="Germany" county="Greifswald" municipality="Serrahn" name="Lehmitz">Lehmitz</location>
and
<collectorName>U. Burkardt</collectorName>
, 401-12; female,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:F2CCF6A01A8B94C9B64B7B6A28A4145A" country="Germany" county="Greifswald" municipality="Serrahn" name="Serrahn">Serrahn</location>
,
<normalizedToken originalValue="“Kesselmoor”">&quot;Kesselmoor&quot;</normalizedToken>
nature reserve, active, relatively undamaged raised bogs, moss/
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Bryopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="0" pageNumber="1" phylum="Bryophyta" rank="genus">Sphagnum</taxonomicName>
,
<date value="2013-10-16">
<collectingDate value="2013-10-16">16.X.2013</collectingDate>
</date>
, leg.
<collectionCode country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:B8E4B60A30590B530F13206868F2D295" country="Germany" county="Greifswald" municipality="Serrahn" name="Lehmitz">Lehmitz</location>
, 49077-08; male,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:5CD718760581D5D0A3EE9411446BDFE3" country="Germany" county="Greifswald" municipality="Serrahn" name="Serrahn">Serrahn</location>
, soil
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" unit="cm" value="2.5" valueMax="5.0" valueMin="0.0">0-5 cm</quantity>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:84F3813F61A19E9D1971D8231B9D3767" country="Germany" county="Greifswald" municipality="Serrahn" name="Medio-European">Medio-European</location>
collinear woodrush beech forest,
<date value="2013-10-16">
<collectingDate value="2013-10-16">16.X.2013</collectingDate>
</date>
, leg.
<collectorName>U. Burkhardt</collectorName>
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="3063647311" collectingDate="1971-07-22" collectorName="W. Dunger" country="Germany" location="Kleiner Kranichsee" municipality="Erzgebirge" specimenCount="1" stateProvince="Sachsen">
263-03.
<collectingRegion country="Germany" name="Sachsen">Saxony</collectingRegion>
: female and juv.,
<collectingMunicipality>Erzgebirge</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:D9D30B60B493B61DDC5DA10515D5E2FB" country="Germany" municipality="Erzgebirge" name="Kleiner Kranichsee" stateProvince="Sachsen">Kleiner Kranichsee</location>
,
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Bryopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="0" pageNumber="1" phylum="Bryophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sphagnum</emphasis>
</taxonomicName>
,
<date value="1971-07-22">
<collectingDate value="1971-07-22">22.VII. 1971</collectingDate>
</date>
, leg.
<collectorName>W. Dunger</collectorName>
</materialsCitation>
, 7871.
<materialsCitation ID-GBIF-Occurrence="3063647305" collectingDate="1991-07-21" collectorName="M. Ursone" country="Luxembourg" location="Tintesmuehle" municipality="Vallee d'Our" specimenCount="1">
<collectingCountry name="Luxembourg">Luxembourg</collectingCountry>
, subadult male,
<collectingMunicipality>
<normalizedToken originalValue="Vallée">Vallee</normalizedToken>
<normalizedToken originalValue="dOur">d'Our</normalizedToken>
</collectingMunicipality>
:
<normalizedToken originalValue="Tintesmühle">
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:B2E6E6B4F21082029E02D323C6524177" country="Luxembourg" municipality="Vallee d'Our" name="Tintesmuehle">Tintesmuehle</location>
</normalizedToken>
, near the river,
<date value="1991-07-21">
<collectingDate value="1991-07-21">21.VII.1991</collectingDate>
</date>
, leg.
<collectorName>M. Ursone</collectorName>
</materialsCitation>
, L-91-101 (
<collectionCode>ISEZ</collectionCode>
).
<materialsCitation ID-GBIF-Occurrence="3063647315" collectingDate="1993-11-14" collectingDateMax="2009-04-22" collectingDateMin="1993-11-14" collectionCode="AF" collectorName="A. Fjellberg &amp; Barum &amp; Dalivannet &amp; Larvik, N &amp; Hvarnes &amp; Larvik &amp; Granasnekollen &amp; Brunlanes &amp; Hummerbakken &amp; Camp. &amp; Tjome &amp; Sando" country="Norway" location="Tappenbergvann" specimenCount="9" specimenCount-female="9" stateProvince="Akershus">
<collectingCountry name="Norway">Norway</collectingCountry>
(
<collectionCode>AF</collectionCode>
, leg.
<collectorName>A. Fjellberg</collectorName>
):
<collectingRegion country="Norway" name="Akershus">Akershus</collectingRegion>
:
<specimenCount type="female">4 females</specimenCount>
,
<normalizedToken originalValue="“Östmarka”">&quot;Oestmarka&quot;</normalizedToken>
nature reserve,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:5241BFAF409827CBEF350DCA092F2FF4" country="Norway" name="Tappenbergvann" stateProvince="Akershus">Tappenbergvann</location>
, old spruce cones,
<date value="1995-05-18">
<collectingDate value="1995-05-18">18.V.1995</collectingDate>
</date>
, 95.163; female,
<collectorName>Barum</collectorName>
,
<collectorName>Dalivannet</collectorName>
,
<date value="1997-11-16">
<collectingDate value="1997-11-16">16.XI.1997</collectingDate>
</date>
.
<collectingRegion country="Norway" name="Vestfold">Vestfold</collectingRegion>
: male, 32/93;
<specimenCount type="female">2 females</specimenCount>
,
<collectorName>Larvik, N</collectorName>
. Holtesetra,
<collectorName>Hvarnes</collectorName>
, lush
<taxonomicName class="Dicotyledoneae" family="Betulaceae" genus="Alnus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Alnus" order="Fagales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Alnus</emphasis>
</taxonomicName>
and
<taxonomicName class="Dicotyledoneae" family="Oleaceae" genus="Fraxinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fraxinus" order="Oleales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Fraxinus</emphasis>
</taxonomicName>
forest,
<date value="2007-11-24">
<collectingDate value="2007-11-24">24.XI.2007</collectingDate>
</date>
, 7.290; female and male,
<collectorName>Larvik</collectorName>
,
<collectorName>Granasnekollen</collectorName>
,
<collectorName>Hvarnes</collectorName>
, litter, oak/beech,
<date value="2004-09-25">
<collectingDate value="2004-09-25">25.IX.2004</collectingDate>
</date>
, 04.086;
<specimenCount type="female">3 females</specimenCount>
,
<collectorName>Brunlanes</collectorName>
,
<collectorName>Hummerbakken</collectorName>
,
<collectingRegion country="Norway" name="Telemark">Telemark</collectingRegion>
<collectorName>Camp.</collectorName>
, plant debris, beach,
<date value="1993-11-14">
<collectingDate value="1993-11-14">14.XI.1993</collectingDate>
</date>
, 93.077; female,
<normalizedToken originalValue="Tjøme">
<collectorName>Tjome</collectorName>
</normalizedToken>
,
<normalizedToken originalValue="Sandø">
<collectorName>Sando</collectorName>
</normalizedToken>
, N-stranda, spongy
<taxonomicName class="Insecta" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinus" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pinus</emphasis>
</taxonomicName>
litter behind beach,
<date value="2009-04-22">
<collectingDate value="2009-04-22">22.IV.2009</collectingDate>
</date>
, 9.086
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063647301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="DIBEC" collectorName="A. Smolis &amp; D. Skarzynski" country="Poland" county="Carpathians" elevation="600" location="Muszyna" municipality="Beskid Sadecki Mts" specimenCount="5" specimenCount-male="3">
<collectingCountry name="Poland">Poland</collectingCountry>
(
<collectingCounty>Carpathians</collectingCounty>
,
<collectionCode>DIBEC</collectionCode>
): female,
<specimenCount type="male">3 males</specimenCount>
,
<specimenCount type="generic">2 juv.</specimenCount>
,
<collectingMunicipality>
Beskid
<normalizedToken originalValue="Sądecki">Sadecki</normalizedToken>
Mts
</collectingMunicipality>
: &quot;Las Lipowy
<normalizedToken originalValue="Obrożyska”">Obrozyska&quot;</normalizedToken>
nature reserve near
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:62609E95EABC2A0FC344EC3A8A9C73E6" country="Poland" county="Carpathians" municipality="Beskid Sadecki Mts" name="Muszyna">Muszyna</location>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
a.s.l.
</elevation>
, mosses on rocks and trees,
<collectingDate isEnumeration="true">
<date value="2004-05-01">1.V.2004</date>
,
<date value="2005-06-25">25.VI.2005</date>
</collectingDate>
, leg.
<collectorName>A. Smolis</collectorName>
,
<collectorName>
D.
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063647309" collectingDate="2004-05-03" collectionCode="V" collectorName="A. Smolis &amp; D. Skarzynski" country="Poland" elevation="600" location="Roztoka Ryterska" specimenCount="1">
female,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:51D2FFDC02B73EF089F1D1FF90ECB96F" country="Poland" name="Roztoka Ryterska">Roztoka Ryterska</location>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
a.s.l.
</elevation>
, litter and mosses near stream,
<date value="2004-05-03">
<collectingDate value="2004-05-03">
3.
<collectionCode country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.2004
</collectingDate>
</date>
, leg.
<collectorName>A. Smolis</collectorName>
,
<collectorName>
D.
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063647316" collectingDate="2000-05-03" collectionCode="V" collectorName="A. Smolis" country="Poland" elevation="850" location="Uhryn" specimenCount="5" specimenCount-female="5">
<specimenCount type="female">5 females</specimenCount>
,
<normalizedToken originalValue="“Uhryń”">
&quot;
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:32107697CA6BA1D9CF9D46FAD58C8CD8" country="Poland" name="Uhryn">Uhryn</location>
&quot;
</normalizedToken>
nature reserve,
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">850 m</quantity>
a.s.l.
</elevation>
, litter of fir-beech forest,
<date value="2000-05-03">
<collectingDate value="2000-05-03">
3.
<collectionCode country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.2000
</collectingDate>
</date>
, leg.
<collectorName>A. Smolis</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063647322" collectingDate="2003-05-10" collectionCode="V" collectorName="A. Smolis &amp; D. Skarzynski" country="Poland" elevation="850" location="Barnowiec" specimenCount="5" specimenCount-male="3">
female,
<specimenCount type="male">3 males</specimenCount>
,
<specimenCount type="generic">2 juv.</specimenCount>
,
<normalizedToken originalValue="“Barnowiec”">
&quot;
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:D2FF9880EE21ABBF82C0044DA4F17E24" country="Poland" name="Barnowiec">Barnowiec</location>
&quot;
</normalizedToken>
nature reserve,
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">850 m</quantity>
a.s.l.
</elevation>
, mosses on rocks in an old beech forest,
<date value="2003-05-10">
<collectingDate value="2003-05-10">
10.
<collectionCode country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.2003
</collectingDate>
</date>
, leg.
<collectorName>A. Smolis</collectorName>
,
<collectorName>
D.
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
</collectorName>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063647319" collectingDate="2004-05-02" collectionCode="V" collectorName="A. Smolis &amp; D. Skarzynski. Bieszczady Mts" country="Poland" elevation="800" location="Jaworzyna Krynicka" specimenCount="6" specimenCount-female="4" specimenCount-male="2">
<specimenCount type="female">4 females</specimenCount>
,
<specimenCount type="male">2 males</specimenCount>
, SE slopes of
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:73D302FBE8E32B351C9C095BCE16F039" country="Poland" name="Jaworzyna Krynicka">Jaworzyna Krynicka</location>
, litter in a beech forest, ca
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
</quantity>
a.s.l,
<date value="2004-05-02">
<collectingDate value="2004-05-02">
2.
<collectionCode country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.2004
</collectingDate>
</date>
, leg.
<collectorName>A. Smolis</collectorName>
,
<collectorName>
D.
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
. Bieszczady Mts
</collectorName>
</materialsCitation>
:
<materialsCitation ID-GBIF-Occurrence="3063647303" collectingDate="2000-05-19" collectionCode="V" collectorName="A. Smolis." country="Poland" elevation="1000" location="Krzemieniec" specimenCount="2" specimenCount-female="2">
<specimenCount type="female">2 females</specimenCount>
, N slopes of
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:322E4B72A141C932DEC8E55E5A5022E5" country="Poland" name="Krzemieniec">Krzemieniec</location>
,
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
, litter in a stream valley,
<date value="2000-05-19">
<collectingDate value="2000-05-19">
19.
<collectionCode country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.2000
</collectingDate>
</date>
, leg.
<collectorName>A. Smolis.</collectorName>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3063647320" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="S. Bakaeva" country="Ukraine" location="Ivano-Frankove village" specimenCount="1" stateProvince="L'viv">
<collectingCountry name="Ukraine">Ukraine</collectingCountry>
: male,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:469E17613A087BF2BB88B08CC62AB96D" country="Ukraine" name="Ivano-Frankove village" stateProvince="L'viv">Ivano-Frankove village</location>
,
<collectingRegion country="Ukraine" name="L'viv">Lviv District</collectingRegion>
, beech and elm forest, leaf litter and soil, leg.
<collectorName>S. Bakaeva</collectorName>
</materialsCitation>
, 2.2.4.8 (
<collectionCode>SMNHL</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Dedicated to Jan Stach, the excellent specialist in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="order">Collembola</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Body length
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1-2 mm</quantity>
. Colour (in alcohol) bluish-gray to bluish-black. Tegumental granulation strong, with fields of especially coarse granules on head (large uniform field covering whole dorsal side except antennal bases), thoracic terga II-III (two large subaxial fields and two lateral ones of medium size, Fig.
<figureCitation captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Ceratophysella stachi sp. nov. 3 chaetotaxy of thoracic tergum II 4 chaetotaxy of abdominal terga III-VI." figureDoi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1">3</figureCitation>
), abdominal terga I-III (variable distribution: from four - Fig.
<figureCitation captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Ceratophysella stachi sp. nov. 3 chaetotaxy of thoracic tergum II 4 chaetotaxy of abdominal terga III-VI." figureDoi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1">4</figureCitation>
to seven fields of medium size as in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. granulata" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="granulata">
<emphasis italics="true" pageId="0" pageNumber="1">C. granulata</emphasis>
</taxonomicName>
; see
<bibRefCitation DOI="https://doi.org/10.3409/173491504783995825" author="Skarzynski, D" journalOrPublisher="Acta Zoologica Cracoviensia" pageId="0" pageNumber="1" pagination="147 - 153" refId="B45" refString="Skarzynski, D, 2004a. Taxonomic position of Ceratophysella granulata STACH, 1949 and Ceratophysella silvatica (RUSEK, 1964) in the light of morphological and laboratory hybridization studies. Acta Zoologica Cracoviensia 47 (3-4): 147 - 153, DOI: https://doi.org/10.3409/173491504783995825" title="Taxonomic position of Ceratophysella granulata STACH, 1949 and Ceratophysella silvatica (RUSEK, 1964) in the light of morphological and laboratory hybridization studies." url="https://doi.org/10.3409/173491504783995825" volume="47" year="2004 a">
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
2004a
</bibRefCitation>
: fig. 8), abdominal tergum IV (medium axial field and two lateral large ones, Fig.
<figureCitation captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Ceratophysella stachi sp. nov. 3 chaetotaxy of thoracic tergum II 4 chaetotaxy of abdominal terga III-VI." figureDoi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1">4</figureCitation>
) and abdominal terga V-VI (large uniform fields covering almost whole dorsal side, Fig.
<figureCitation captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Ceratophysella stachi sp. nov. 3 chaetotaxy of thoracic tergum II 4 chaetotaxy of abdominal terga III-VI." figureDoi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1">4</figureCitation>
). 6-9 granules between macrosetae p1 on abdominal tergum
<collectionCode country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.
</paragraph>
<caption doi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1" start="Figures 3, 4" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 3, 4.</emphasis>
<taxonomicName authorityName="Skarzynski, Smolis &amp; Porco" authorityYear="2021" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella stachi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stachi">
<emphasis italics="true" pageId="0" pageNumber="1">Ceratophysella stachi</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">3</emphasis>
chaetotaxy of thoracic tergum II
<emphasis bold="true" pageId="0" pageNumber="1">4</emphasis>
chaetotaxy of abdominal terga III-VI.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Arrangement of setae on head typical for the genus, spine-like setae absent. Dorsal chaetotaxy of
<typeStatus>type</typeStatus>
B (Figs
<figureCitation captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Ceratophysella stachi sp. nov. 3 chaetotaxy of thoracic tergum II 4 chaetotaxy of abdominal terga III-VI." figureDoi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1">3</figureCitation>
,
<figureCitation captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Ceratophysella stachi sp. nov. 3 chaetotaxy of thoracic tergum II 4 chaetotaxy of abdominal terga III-VI." figureDoi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1">4</figureCitation>
). Thoracic terga II-III with setae m6 present and one additional seta outside lateral sensillum m7 present or absent. Setae p1 on abdominal tergum IV developed as macrosetae, p2 as microsetae, setae p3 present. Differentiation of dorsal setae into micro- and macrosetae distinct. Setae long (ratio p1 microsetae and p2 macrosetae on thoracic tergum II/inner edge of claws III = 1-1.3 and 1.8-2.8, respectively), thick, curved, pointed at tips and only slightly serrate. Body sensilla (s) short (ratio sensillum p4 and m7 on thoracic tergum II/inner edge of claws III = 0.6-1 and 0.4-0.6, respectively), thin and smooth. Microsensilla (ms) on thoracic tergum II present (Fig.
<figureCitation captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Ceratophysella stachi sp. nov. 3 chaetotaxy of thoracic tergum II 4 chaetotaxy of abdominal terga III-VI." figureDoi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1">3</figureCitation>
). Subcoxae I, II, III with 1, 2, 3 setae, respectively.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Antennal segment IV with simple or lobed apical vesicle, subapical organite (or), microsensillum (ms), 7 (2 lateral, 5 dorsal) cylindrical, subequal sensilla and 15-25 slightly curved blunt-tipped sensilla in ventral field (Figs
<figureCitation captionStart="Figures 59" captionStartId="F4" captionText="Figures 5 - 9. Ceratophysella stachi sp. nov. 5 chaetotaxy of dorsal side of antennal segments III-IV 6 chaetotaxy of ventral side of antennal segment IV 7 postantennal organ and neighbor ocelli 8 claw I 9 dens and mucro." figureDoi="10.3897/zookeys.1021.63147.figures5-9" httpUri="https://binary.pensoft.net/fig/515256" pageId="0" pageNumber="1">5</figureCitation>
,
<figureCitation captionStart="Figures 59" captionStartId="F4" captionText="Figures 5 - 9. Ceratophysella stachi sp. nov. 5 chaetotaxy of dorsal side of antennal segments III-IV 6 chaetotaxy of ventral side of antennal segment IV 7 postantennal organ and neighbor ocelli 8 claw I 9 dens and mucro." figureDoi="10.3897/zookeys.1021.63147.figures5-9" httpUri="https://binary.pensoft.net/fig/515256" pageId="0" pageNumber="1">6</figureCitation>
). Antennal segment III organ with two long (lateral) and two short (internal), curved sensilla (Fig.
<figureCitation captionStart="Figures 59" captionStartId="F4" captionText="Figures 5 - 9. Ceratophysella stachi sp. nov. 5 chaetotaxy of dorsal side of antennal segments III-IV 6 chaetotaxy of ventral side of antennal segment IV 7 postantennal organ and neighbor ocelli 8 claw I 9 dens and mucro." figureDoi="10.3897/zookeys.1021.63147.figures5-9" httpUri="https://binary.pensoft.net/fig/515256" pageId="0" pageNumber="1">5</figureCitation>
). Microsensillum on antennal segment III present. Eversible sac between antennal segments III and IV present. Antennal segment I with 7 setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Ocelli: 8 + 8. Postantennal organ 1.8-2.3 times as large as single ocellus; the former with four lobes, its anterior pair larger than posterior pair. Accessory boss large (equal to or only slightly smaller than posterior lobes of postantennal organ), often granulated (Fig.
<figureCitation captionStart="Figures 59" captionStartId="F4" captionText="Figures 5 - 9. Ceratophysella stachi sp. nov. 5 chaetotaxy of dorsal side of antennal segments III-IV 6 chaetotaxy of ventral side of antennal segment IV 7 postantennal organ and neighbor ocelli 8 claw I 9 dens and mucro." figureDoi="10.3897/zookeys.1021.63147.figures5-9" httpUri="https://binary.pensoft.net/fig/515256" pageId="0" pageNumber="1">7</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Labrum with 5, 5, 4 setae; 4 prelabrals present. Maxillary head of
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. armata" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="armata">
<emphasis italics="true" pageId="0" pageNumber="1">C. armata</emphasis>
</taxonomicName>
<typeStatus>type</typeStatus>
(
<bibRefCitation author="Fjellberg, A" journalOrPublisher="Annales de la Societe Royale Zoologique de Belgique" pageId="0" pageNumber="1" pagination="89 - 99" refId="B14" refString="Fjellberg, A, 1984. Maxillary structures in Hypogastruridae (Collembola). Annales de la Societe Royale Zoologique de Belgique 114: 89 - 99" title="Maxillary structures in Hypogastruridae (Collembola)." volume="114" year="1984">Fjellberg 1984</bibRefCitation>
: fig. 18). Labial palp as shown in
<bibRefCitation author="Fjellberg, A" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="1" pagination="309 - 330" refId="B16" refString="Fjellberg, A, 1999. The labial palp in Collembola. Zoologischer Anzeiger 237: 309 - 330" title="The labial palp in Collembola." volume="237" year="1999">Fjellberg (1999</bibRefCitation>
: Fig.
<figureCitation captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Ceratophysella stachi sp. nov. 3 chaetotaxy of thoracic tergum II 4 chaetotaxy of abdominal terga III-VI." figureDoi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1">4</figureCitation>
), but with 6 proximal setae. Outer maxillary lobe with one sublobal hair.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Tibiotarsi I, II, III with 19, 19, 18 setae, respectively, clavate setae absent. Claws with inner tooth and a pair of lateral teeth. Empodial appendage with broad lamelliform base and filiform apex reaching inner tooth or slightly beyond, ratio empodial appendage/ inner edge of claws = 0.4-0.7 (Fig.
<figureCitation captionStart="Figures 59" captionStartId="F4" captionText="Figures 5 - 9. Ceratophysella stachi sp. nov. 5 chaetotaxy of dorsal side of antennal segments III-IV 6 chaetotaxy of ventral side of antennal segment IV 7 postantennal organ and neighbor ocelli 8 claw I 9 dens and mucro." figureDoi="10.3897/zookeys.1021.63147.figures5-9" httpUri="https://binary.pensoft.net/fig/515256" pageId="0" pageNumber="1">8</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Ventral tube with 4 + 4 setae. Furca well developed. Ratio dens + mucro/inner edge of claw III = 1.8-2.2, ratio dens/mucro = 1.7-2.2. Dens with uniform fine granules and 7 dorsal setae (2-4 inner setae modified) (Fig.
<figureCitation captionStart="Figures 59" captionStartId="F4" captionText="Figures 5 - 9. Ceratophysella stachi sp. nov. 5 chaetotaxy of dorsal side of antennal segments III-IV 6 chaetotaxy of ventral side of antennal segment IV 7 postantennal organ and neighbor ocelli 8 claw I 9 dens and mucro." figureDoi="10.3897/zookeys.1021.63147.figures5-9" httpUri="https://binary.pensoft.net/fig/515256" pageId="0" pageNumber="1">9</figureCitation>
). Mucro wide at tip (ratio width of apical part/length of mucro = 0.4-0.6, usually 0.5), boat-like, with large outer lamella, (Fig.
<figureCitation captionStart="Figures 59" captionStartId="F4" captionText="Figures 5 - 9. Ceratophysella stachi sp. nov. 5 chaetotaxy of dorsal side of antennal segments III-IV 6 chaetotaxy of ventral side of antennal segment IV 7 postantennal organ and neighbor ocelli 8 claw I 9 dens and mucro." figureDoi="10.3897/zookeys.1021.63147.figures5-9" httpUri="https://binary.pensoft.net/fig/515256" pageId="0" pageNumber="1">9</figureCitation>
). Retinaculum with 4 + 4 teeth.
</paragraph>
<caption doi="10.3897/zookeys.1021.63147.figures5-9" httpUri="https://binary.pensoft.net/fig/515256" pageId="0" pageNumber="1" start="Figures 59" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 5-9.</emphasis>
<taxonomicName authorityName="Skarzynski, Smolis &amp; Porco" authorityYear="2021" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella stachi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stachi">
<emphasis italics="true" pageId="0" pageNumber="1">Ceratophysella stachi</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">5</emphasis>
chaetotaxy of dorsal side of antennal segments III-IV
<emphasis bold="true" pageId="0" pageNumber="1">6</emphasis>
chaetotaxy of ventral side of antennal segment IV
<emphasis bold="true" pageId="0" pageNumber="1">7</emphasis>
postantennal organ and neighbor ocelli
<emphasis bold="true" pageId="0" pageNumber="1">8</emphasis>
claw I
<emphasis bold="true" pageId="0" pageNumber="1">9</emphasis>
dens and mucro.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Anal spines yellowish, slightly curved, situated on high basal papillae, 1.1-1.7 times as long as inner edge of claws III (Fig.
<figureCitation captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Ceratophysella stachi sp. nov. 3 chaetotaxy of thoracic tergum II 4 chaetotaxy of abdominal terga III-VI." figureDoi="10.3897/zookeys.1021.63147.figures3-4" httpUri="https://binary.pensoft.net/fig/515255" pageId="0" pageNumber="1">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3063647323" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Denmark" county="The" elevation="1000" location="Jutland" municipality="It" specimenCount="1" stateProvince="L'viv">
<collectingCounty>The</collectingCounty>
range of distribution of
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. stachi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stachi">
<emphasis italics="true" pageId="0" pageNumber="1">C. stachi</emphasis>
</taxonomicName>
sp. nov. appears to be relatively wide.
<collectingMunicipality>It</collectingMunicipality>
is known from
<collectingCountry name="Denmark">Denmark</collectingCountry>
(
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:DC460D093C896F99FBF61FA4BCBA985C" country="Denmark" county="The" municipality="It" name="Jutland" stateProvince="L'viv">Jutland</location>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:54D9EFB7EEB466C71194AED26F9FA2F3" country="Denmark" county="The" municipality="It" name="Funen" stateProvince="L'viv">Funen</location>
),
<collectingCountry name="Germany">Germany</collectingCountry>
(Brandenburg, Hesse,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:DC12E9C8F8FB512A863B7AF74BC80328" country="Denmark" county="The" municipality="It" name="Mecklenburg-West Pomerania" stateProvince="L'viv">Mecklenburg-West Pomerania</location>
, Saxony),
<collectingCountry name="Luxembourg">Luxembourg</collectingCountry>
, southern
<collectingCountry name="Norway">Norway</collectingCountry>
(Akershus, Vestfold),
<collectingCountry name="Poland">Poland</collectingCountry>
(
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:121D6205B746FD37BE12D1FA4EA04611" country="Denmark" county="The" municipality="It" name="Carpathians" stateProvince="L'viv">Carpathians</location>
:
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:7040F73EAB74D9EB4A6819011790684F" country="Denmark" county="The" municipality="It" name="Beskid Niski" stateProvince="L'viv">Beskid Niski</location>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:971CD894536421E36FF07860183963C8" country="Denmark" county="The" municipality="It" name="Beskid Sadecki" stateProvince="L'viv">
Beskid
<normalizedToken originalValue="Sądecki">Sadecki</normalizedToken>
</location>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:7ED2388C577C70361BEFC59A201D20A9" country="Denmark" county="The" municipality="It" name="Bieszczady mountains" stateProvince="L'viv">Bieszczady mountains</location>
) and
<collectingCountry name="Ukraine">Ukraine</collectingCountry>
(
<collectingRegion country="Ukraine" name="L'viv">Lviv District</collectingRegion>
) (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Ceratophysella stachi sp. nov. (black circles) and C. granulata (empty circles). Abbreviations: AKE - Akershus, B - Bieszczady Mts, BM - Beskid Maly Mts, BN - Beskid Niski Mts, BRA - Brandenburg, BS - Beskid Sadecki Mts, BW - Beskid Wyspowy Mts, BZ - Beskid Zywiecki Mts, CH - Cierna hora Mts, FUN - Funen, HES - Hesse, JUT - Jutland, L - Levocske vrchy Mts, LD - Lviv District, LT - Low Tatra Mts, LUX - Luxembourg, M - Muranska planina Plateau, MEC - Mecklenburg-West Pomerania, P - Pieniny Mts, S - Slovak Paradise, SAX - Saxony, SK - Slovak Karst, T - Tatra Mts, VES - Vestfold, VF - Veľka Fatra Mts." figureDoi="10.3897/zookeys.1021.63147.figure1" httpUri="https://binary.pensoft.net/fig/515253" pageId="0" pageNumber="1">1</figureCitation>
).
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:493B35D803D8FD4515FFF85F722FFCC8" country="Denmark" county="The" municipality="It" name="Probably" stateProvince="L'viv">Probably</location>
this species is distributed much more widely in
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:3DA9EB1D378EA1F2373AF1942020C0A1" country="Denmark" county="The" municipality="It" name="Europe" stateProvince="L'viv">Europe</location>
, but additional research is needed to prove this thesis.
<taxonomicName authorityName="Skarzynski, Smolis &amp; Porco" authorityYear="2021" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella stachi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stachi">
<emphasis italics="true" pageId="0" pageNumber="1">Ceratophysella stachi</emphasis>
</taxonomicName>
sp. nov. lives in lowlands and in the mountains (up to ca.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</elevation>
</quantity>
a.s.l) where it inhabits litter and mosses in different
<typeStatus>types</typeStatus>
of forests, and also heathlands and bogs
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Skarzynski, Smolis &amp; Porco" authorityYear="2021" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella stachi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stachi">
<emphasis italics="true" pageId="0" pageNumber="1">Ceratophysella stachi</emphasis>
</taxonomicName>
sp. nov. belongs to a small European branch of species of the
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. armata" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="armata">
<emphasis italics="true" pageId="0" pageNumber="1">C. armata</emphasis>
</taxonomicName>
-group, which have strong tegumentary granulation, with distinct fields of coarse granules:
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. granulata" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="granulata">
<emphasis italics="true" pageId="0" pageNumber="1">C. granulata</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. lawrencei" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis italics="true" pageId="0" pageNumber="1">C. lawrencei</emphasis>
</taxonomicName>
(Gisin, 1963),
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. neomeridionalis" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="neomeridionalis">
<emphasis italics="true" pageId="0" pageNumber="1">C. neomeridionalis</emphasis>
</taxonomicName>
(Nosek &amp;
<normalizedToken originalValue="Červek">Cervek</normalizedToken>
, 1970),
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. scotica" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scotica">
<emphasis italics="true" pageId="0" pageNumber="1">C. scotica</emphasis>
</taxonomicName>
(Carpenter &amp; Evans, 1899), and
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. silvatica" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="silvatica">
<emphasis italics="true" pageId="0" pageNumber="1">C. silvatica</emphasis>
</taxonomicName>
. It differs from all of them in the chaetotaxy of the lateral parts of the thoracic terga II-III (setae m6 present and one additional seta outside lateral sensillum m7 present or absent vs setae m6 and additional setae absent) which is exceptional within the whole
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. armata" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="armata">
<emphasis italics="true" pageId="0" pageNumber="1">C. armata</emphasis>
</taxonomicName>
-group (both characters are found in the genus, but in other groups of species: formerly classified as
<taxonomicName authorityName="Wray" authorityYear="1953" class="Collembola" family="Hypogastruridae" genus="Mitchellania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitchellania" order="Poduromorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Mitchellania</emphasis>
</taxonomicName>
Wray, 1953 and
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. denticulata" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis italics="true" pageId="0" pageNumber="1">C. denticulata</emphasis>
</taxonomicName>
). The remaining differences between
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. stachi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stachi">
<emphasis italics="true" pageId="0" pageNumber="1">C. stachi</emphasis>
</taxonomicName>
sp. nov. and related species mentioned above are summarized in Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Characteristics of Ceratophysella stachi sp. nov., C. granulata and related species. Based on Carpenter and Evans (1899), Gisin (1963), Rusek (1964), Nosek and Cervek (1967, 1970), Christian (1987), Babenko et al. (1994), Fjellberg (1998), Skarzynski (2004 a, 2006), Thibaud et al. (2004), Kaprus' et al. (2006), Danyi and Traser (2008), Kovac et al. (2016), and our own data. Abbreviations: d 2 - spine-like setae d 2 on head, oc 2 - spine-like setae oc 2 on head, m 6 - setae m 6 on thoracic terga II-III, p 3 - setae p 3 on abdominal tergum IV, e / cl - ratio empodium / claw." pageId="0" pageNumber="1">4</tableCitation>
.
</paragraph>
<caption pageId="0" pageNumber="1" start="Table 4" startId="T4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 4.</emphasis>
Characteristics of
<taxonomicName authorityName="Skarzynski, Smolis &amp; Porco" authorityYear="2021" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella stachi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stachi">
<emphasis italics="true" pageId="0" pageNumber="1">Ceratophysella stachi</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. granulata" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="granulata">
<emphasis italics="true" pageId="0" pageNumber="1">C. granulata</emphasis>
</taxonomicName>
and related species. Based on
<bibRefCitation author="Carpenter, GH" journalOrPublisher="Proceedings of the Royal Society of Edinburgh" pageId="0" pageNumber="1" pagination="221 - 266" refId="B7" refString="Carpenter, GH, Evans, W, 1899. The Collembola and Thysanura of the Edinburgh district. Proceedings of the Royal Society of Edinburgh 14: 221 - 266" title="The Collembola and Thysanura of the Edinburgh district." volume="14" year="1899">Carpenter and Evans (1899)</bibRefCitation>
,
<bibRefCitation author="Gisin, H" journalOrPublisher="Revue suisse de Zoologie" pageId="0" pageNumber="1" pagination="77 - 101" refId="B19" refString="Gisin, H, 1963. Collemboles d'Europe V. Revue suisse de Zoologie 70: 77 - 101" title="Collemboles d'Europe V." volume="70" year="1963">Gisin (1963)</bibRefCitation>
,
<bibRefCitation author="Rusek, J" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="1" pagination="432 - 440" refId="B43" refString="Rusek, J, 1964. Zwei neue Collembolenarten aus der Mittel-Slowakei (CSSR). Zoologischer Anzeiger 173: 432 - 440" title="Zwei neue Collembolenarten aus der Mittel-Slowakei (CSSR)." volume="173" year="1964">Rusek (1964)</bibRefCitation>
,
<bibRefCitation author="Nosek, J" journalOrPublisher="Acta Societatis Zoologicae Bohemoslovenicae" pageId="0" pageNumber="1" pagination="245 - 249" refId="B36" refString="Nosek, J, Cervek, S, 1967. A new species of springtail from the mediterranean region Hypogastrura (Ceratophysella) meridionalis sp. n. (Collembola: Hypogastruridae). Acta Societatis Zoologicae Bohemoslovenicae 31: 245 - 249" title="A new species of springtail from the mediterranean region Hypogastrura (Ceratophysella) meridionalis sp. n. (Collembola: Hypogastruridae)." volume="31" year="1967">
Nosek and
<normalizedToken originalValue="Červek">Cervek</normalizedToken>
(1967
</bibRefCitation>
,
<bibRefCitation author="Nosek, J" journalOrPublisher="Acta Societatis Zoologicae Bohemoslovenicae" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/40807834" refId="B37" refString="Nosek, J, Cervek, S, 1970. Beseitigung eines Homonyms in der Gattung Hypogastrura (Insecta: Collembola). Revue suisse de Zoologie 77: 207. https://www.biodiversitylibrary.org/page/40807834" title="Beseitigung eines Homonyms in der Gattung Hypogastrura (Insecta: Collembola). Revue suisse de Zoologie 77: 207." url="https://www.biodiversitylibrary.org/page/40807834" year="1970">1970</bibRefCitation>
),
<bibRefCitation author="Christian, E" journalOrPublisher="Verlag der Oesterreichischen Akademie der Wissenschaften, Wien" pageId="0" pageNumber="1" refId="B9" refString="Christian, E, 1987. Catalogus Faunae Austriae. XIIa, U. Kl.: Collembola (Springschwaenze). Verlag der Oesterreichischen Akademie der Wissenschaften, Wien" title="Catalogus Faunae Austriae. XIIa, U. Kl.: Collembola (Springschwaenze)." year="1987">Christian (1987)</bibRefCitation>
,
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, and our own data. Abbreviations: d2 - spine-like setae d2 on head, oc2 - spine-like setae oc2 on head, m6 - setae m6 on thoracic terga II-III, p3 - setae p3 on abdominal tergum IV, e/cl - ratio empodium/claw.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3063647321" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" elevation="2" location="Distribution Habitat" municipality="Litter" specimenCount="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Species</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">d2</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
oc
<quantity metricMagnitude="0" metricUnit="m" metricValue="2.0" unit="m" value="2.0">
<elevation metricMagnitude="0" metricUnit="m" metricValue="2.0" unit="m" value="2.0">
2
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">m6</th>
</elevation>
</quantity>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">p3</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">e/cl</th>
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:CDCB0DBB472F25D6FA40DD5A59DAD6BA" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Distribution Habitat">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Distribution</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Habitat preferences</th>
</location>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. granulata" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="granulata">
<emphasis italics="true" pageId="0" pageNumber="1">C. granulata</emphasis>
</taxonomicName>
1
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.5-0.7</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:E287210D9BDDBE7EF16EB2F6F6C39576" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Polish">Polish</location>
and
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:9ABE52BDFC85740407A1AE376AA67762" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Slovak Carpathians">Slovak Carpathians</location>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Ceratophysella stachi sp. nov. (black circles) and C. granulata (empty circles). Abbreviations: AKE - Akershus, B - Bieszczady Mts, BM - Beskid Maly Mts, BN - Beskid Niski Mts, BRA - Brandenburg, BS - Beskid Sadecki Mts, BW - Beskid Wyspowy Mts, BZ - Beskid Zywiecki Mts, CH - Cierna hora Mts, FUN - Funen, HES - Hesse, JUT - Jutland, L - Levocske vrchy Mts, LD - Lviv District, LT - Low Tatra Mts, LUX - Luxembourg, M - Muranska planina Plateau, MEC - Mecklenburg-West Pomerania, P - Pieniny Mts, S - Slovak Paradise, SAX - Saxony, SK - Slovak Karst, T - Tatra Mts, VES - Vestfold, VF - Veľka Fatra Mts." figureDoi="10.3897/zookeys.1021.63147.figure1" httpUri="https://binary.pensoft.net/fig/515253" pageId="0" pageNumber="1">1</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:5A52120327CEC87B669E0A63C3457DF9" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Cold">Cold</location>
and humid places in the mountains: mosses in alpine zone, litter and mosses in dwarf mountain pine zone and deep gorges and caves in montane forests zone
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. lawrencei" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis italics="true" pageId="0" pageNumber="1">C. lawrencei</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.5-1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:38A75ACDB169AEC3DD53E5039F81914F" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Austrian">Austrian</location>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:C765FD1324CB1DEAC45F8116845C0372" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Italian">Italian</location>
and
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:9472FFA7EBDB06CBEC3A9157BC1D11B5" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Swiss Alps">Swiss Alps</location>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:6CA37650A3373E5D290B5B8128C74DA6" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Apennines">Apennines</location>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:CBFB99255EDDEF9FABA820BBA05F9B8B" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Polish Tatra Mts.">
Polish Tatra
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:F771E975235FF01B1A61D979FEC180AA" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Mts. Litter">
Mts.
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Litter, mosses on rocks in upper montane zone and above, caves</td>
</location>
</location>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. neomeridionalis" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="neomeridionalis">
<emphasis italics="true" pageId="0" pageNumber="1">C. neomeridionalis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.2-0.3</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:F39CB6B880F3E2694781FD8F15F4CFC5" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Slovenian Dinaric Mts">Slovenian Dinaric Mts</location>
,
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:EABB644DCF006A3776B9C3380B445E7C" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Polish">Polish</location>
and
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:9BF16705FFF7D5A05DCDFA597C88528E" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Ukrainian Carpathians Litter">
Ukrainian Carpathians
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Litter, mosses on rocks in montane zone</td>
</location>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. scotica" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scotica">
<emphasis italics="true" pageId="0" pageNumber="1">C. scotica</emphasis>
</taxonomicName>
2
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.7-1.1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<collectingCountry isEnumeration="true">Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden</collectingCountry>
,
<collectingCountry name="Ukraine">Ukraine</collectingCountry>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<location LSID="urn:lsid:plazi:treatment:3048909794415CAFAF7712BC8C024C10:1CB60FC266A387FD703D6B39FE68C220" country="Belarus, Denmark, Finland, Germany, Great Britain, Ireland, Norway, Poland, Russia, Sweden" county="Litter" municipality="Litter" name="Hygrophilous">Hygrophilous</location>
and tyrphophilous species living in lowlands and mountains
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. silvatica" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="silvatica">
<emphasis italics="true" pageId="0" pageNumber="1">C. silvatica</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.3-0.4</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<collectingCountry isEnumeration="true">Hungary, Italy, Poland, Romania, Slovakia</collectingCountry>
,
<collectingCountry name="Ukraine">Ukraine</collectingCountry>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<collectingMunicipality>Litter</collectingMunicipality>
, mosses on rocks in upland and mountain forests
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. stachi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stachi">
<emphasis italics="true" pageId="0" pageNumber="1">C. stachi</emphasis>
</taxonomicName>
sp. nov.3
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">+</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.4-0.7</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<collectingCountry isEnumeration="true">Denmark, Germany, Luxembourg, Norway, Poland</collectingCountry>
,
<collectingCountry name="Ukraine">Ukraine</collectingCountry>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Ceratophysella stachi sp. nov. (black circles) and C. granulata (empty circles). Abbreviations: AKE - Akershus, B - Bieszczady Mts, BM - Beskid Maly Mts, BN - Beskid Niski Mts, BRA - Brandenburg, BS - Beskid Sadecki Mts, BW - Beskid Wyspowy Mts, BZ - Beskid Zywiecki Mts, CH - Cierna hora Mts, FUN - Funen, HES - Hesse, JUT - Jutland, L - Levocske vrchy Mts, LD - Lviv District, LT - Low Tatra Mts, LUX - Luxembourg, M - Muranska planina Plateau, MEC - Mecklenburg-West Pomerania, P - Pieniny Mts, S - Slovak Paradise, SAX - Saxony, SK - Slovak Karst, T - Tatra Mts, VES - Vestfold, VF - Veľka Fatra Mts." figureDoi="10.3897/zookeys.1021.63147.figure1" httpUri="https://binary.pensoft.net/fig/515253" pageId="0" pageNumber="1">1</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<collectingCounty>Litter</collectingCounty>
and mosses in different
<typeStatus>types</typeStatus>
of forests in lowlands and mountains, also heathlands and bogs
</td>
</tr>
</materialsCitation>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<tableNote pageId="0" pageNumber="1">1 Accessory boss near post-antennal organ small (about half the size of posterior lobes of post-antennal organ), mucro narrow at tip (ratio width of apical part/length of mucro = 0.22-0.52 (mean 39). 2 Fields of especially coarse granules only on abdominal terga IV-VI. 3 Thoracic terga II-III with one additional seta outside lateral sensillum m7 present or absent.</tableNote>
</paragraph>
</subSubSection>
</treatment>
</document>