treatments-xml/data/24/2F/0B/242F0B25FFA7FFE46ACBF88AFC73FA1B.xml

<document ID-DOI="10.26107/RBZ-2021-0069" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174477" ID-ZooBank="4D1086F5-6FC4-4FB7-9A83-A13769F4BF2B" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1665290878726" checkinUser="felipe" docAuthor="Kottelat, Maurice &amp; Lim, Kelvin K. P." docDate="2021" docId="242F0B25FFA7FFE46ACBF88AFC73FA1B" docLanguage="en" docName="RafflesBZool.69.522–540.pdf" docOrigin="Raffles Bulletin of Zoology 69" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.4:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="4" docTitle="Barbodes zakariaismaili Kottelat &amp; Lim 2021, new species" docType="treatment" docVersion="3" lastPageNumber="536" masterDocId="D816735DFFADFFEA6957FFB3FF81FF8E" masterDocTitle="Two new species of Barbodes from the Malay Peninsula and comments on ‘ cryptic species’ in the B. binotatus group (Teleostei: Cyprinidae)" masterLastPageNumber="540" masterPageNumber="522" pageNumber="532" updateTime="1665307045777" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Two new species of Barbodes from the Malay Peninsula and comments on ‘ cryptic species’ in the B. binotatus group (Teleostei: Cyprinidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kottelat, Maurice</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lim, Kelvin K. P.</mods:namePart>
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<mods:title>Raffles Bulletin of Zoology</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-12-10</mods:number>
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<mods:detail type="volume">
<mods:number>69</mods:number>
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<mods:start>522</mods:start>
<mods:end>540</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.26107/RBZ-2021-0069</mods:identifier>
<mods:identifier type="ISSN">2345-7600</mods:identifier>
<mods:identifier type="Zenodo-Dep">7174477</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7171142" ID-Zenodo-Dep="7171142" LSID="urn:lsid:plazi:treatment:242F0B25FFA7FFE46ACBF88AFC73FA1B" httpUri="http://treatment.plazi.org/id/242F0B25FFA7FFE46ACBF88AFC73FA1B" lastPageId="14" lastPageNumber="536" pageId="10" pageNumber="532">
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<paragraph blockId="10.[924,1341,1849,1905]" box="[924,1341,1849,1873]" pageId="10" pageNumber="532">
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<emphasis bold="true" box="[924,1341,1849,1873]" pageId="10" pageNumber="532">
<taxonomicName authority="Kottelat &amp; Lim, 2021" authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[924,1192,1849,1873]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="532" phylum="Chordata" rank="species" species="zakariaismaili" status="new species">
<emphasis bold="true" box="[924,1192,1849,1873]" italics="true" pageId="10" pageNumber="532">Barbodes zakariaismaili</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1207,1341,1849,1873]" pageId="10" pageNumber="532" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1058,1206,1881,1905]" pageId="10" pageNumber="532" type="description">
<paragraph blockId="10.[924,1341,1849,1905]" box="[1058,1206,1881,1905]" pageId="10" pageNumber="532">
(
<figureCitation box="[1066,1157,1881,1905]" captionStart="Fig" captionStartId="12.[113,148,1102,1124]" captionTargetBox="[113,1451,189,1086]" captionTargetId="figure-258@12.[113,1451,189,1086]" captionTargetPageId="12" captionText="Fig. 13. Barbodes zakariaismaili, new species, Malaysia: Pahang: Cameron Highlands; a, ZRC 52347, holotype, 66.3 mm SL; b, ZRC 62315, paratype, 58.3 mm SL. (Photographs by M. Kottelat)." figureDoi="http://doi.org/10.5281/zenodo.7174509" httpUri="https://zenodo.org/record/7174509/files/figure.png" pageId="10" pageNumber="532">Figs. 13</figureCitation>
, 
<figureCitation box="[1170,1197,1881,1905]" captionStart="Fig" captionStartId="13.[136,171,1993,2015]" captionTargetBox="[136,1458,268,1984]" captionTargetId="figure-55@13.[136,1474,258,1985]" captionTargetPageId="13" captionText="Fig. 14. Barbodes zakariaismaili, new species, Malaysia: Pahang: Cameron Highlands; a, b, CMK 27384, paratypes: 27.8 mm SL (a) and 33.7 mm SL (b); c, d, ZRC 62315, paratypes: 30.1 mm SL (c) and 35.5 mm SL (d). (Photographs by M. Kottelat)." figureDoi="http://doi.org/10.5281/zenodo.7174511" httpUri="https://zenodo.org/record/7174511/files/figure.png" pageId="10" pageNumber="532">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="533" pageId="10" pageNumber="532" type="materials_examined">
<paragraph blockId="10.[813,1452,1945,2065]" pageId="10" pageNumber="532">
<materialsCitation accessionNumber="ZRC 52347" collectingDate="1992-10-26" collectionCode="ZRC" collectorName="M. Zakaria-Ismail" country="Malaysia" county="Cameron Highlands" elevation="200" latitude="4.436111" location="Sungai" longLatPrecision="21" longitude="101.49444" municipality="Sungai Boh" pageId="10" pageNumber="532" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<emphasis bold="true" box="[813,926,1945,1969]" pageId="10" pageNumber="532">
<typeStatus box="[813,921,1945,1969]" pageId="10" pageNumber="532">Holotype</typeStatus>
.
</emphasis>
<accessionNumber box="[936,1070,1945,1969]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC52347" pageId="10" pageNumber="532">
<collectionCode box="[936,989,1945,1969]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="532">ZRC</collectionCode>
52347
</accessionNumber>
, 
<quantity box="[1085,1185,1945,1969]" metricMagnitude="-2" metricUnit="m" metricValue="6.609999999999999" pageId="10" pageNumber="532" unit="mm" value="66.1">66.1 mm</quantity>
SL; 
<collectingCountry box="[1245,1349,1945,1969]" name="Malaysia" pageId="10" pageNumber="532">Malaysia</collectingCountry>
: 
<collectingRegion box="[1364,1448,1945,1969]" country="Malaysia" name="Pahang" pageId="10" pageNumber="532">Pahang</collectingRegion>
: 
<collectingRegion box="[813,894,1977,2001]" country="Malaysia" name="Pahang" pageId="10" pageNumber="532">Pahang</collectingRegion>
drainage: 
<collectingCounty box="[1018,1247,1977,2001]" pageId="10" pageNumber="532">Cameron Highlands</collectingCounty>
: 
<collectingMunicipality box="[1260,1396,1977,2001]" pageId="10" pageNumber="532">
<location LSID="urn:lsid:plazi:treatment:242F0B25FFA7FFE46ACBF88AFC73FA1B:A959ECE8FFA7FFE06DBBF80AFAB9F85F" box="[1260,1336,1977,2001]" country="Malaysia" county="Cameron Highlands" latitude="4.436111" longLatPrecision="21" longitude="101.49444" municipality="Sungai Boh" name="Sungai" pageId="10" pageNumber="532" stateProvince="Pahang">Sungai</location>
Boh
</collectingMunicipality>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="532" unit="m" value="200.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="532" unit="m" value="200.0">200 m</elevation>
</quantity>
upstream from 
<location LSID="urn:lsid:plazi:treatment:242F0B25FFA7FFE46ACBF88AFC73FA1B:A959ECE8FFA7FFE06AA7F86AFB1DF87F" box="[1008,1180,2009,2033]" country="Malaysia" county="Cameron Highlands" latitude="4.436111" longLatPrecision="21" longitude="101.49444" municipality="Sungai Boh" name="Sungai Menson" pageId="10" pageNumber="532" stateProvince="Pahang">Sungai Menson</location>
; 
<geoCoordinate box="[1193,1307,2008,2033]" degrees="4" direction="north" minutes="26" orientation="latitude" pageId="10" pageNumber="532" precision="15" seconds="10" value="4.436111">4°26′10″N</geoCoordinate>
<geoCoordinate box="[1315,1452,2008,2033]" degrees="101" direction="east" minutes="29" orientation="longitude" pageId="10" pageNumber="532" precision="15" seconds="40" value="101.49444">101°29′40″E</geoCoordinate>
[about 
<quantity box="[889,994,2041,2065]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="532" unit="m" value="700.0">
<elevation box="[889,994,2041,2065]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="532" unit="m" value="700.0">700 masl</elevation>
</quantity>
]; 
<collectorName box="[1013,1212,2041,2065]" pageId="10" pageNumber="532">M. Zakaria-Ismail</collectorName>
, 
<date box="[1225,1412,2041,2065]" pageId="10" pageNumber="532" value="1992-10-26">
<collectingDate box="[1225,1412,2041,2065]" pageId="10" pageNumber="532" value="1992-10-26">26 October 1992</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7174507" ID-Zenodo-Dep="7174507" httpUri="https://zenodo.org/record/7174507/files/figure.png" pageId="11" pageNumber="533" startId="11.[136,171,1678,1700]" targetBox="[136,1474,189,1663]" targetPageId="11">
<paragraph blockId="11.[136,1474,1678,1729]" pageId="11" pageNumber="533">
Fig. 12. 
<taxonomicName authority=", Borneo" authorityName="Borneo" box="[219,464,1678,1700]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="533" phylum="Chordata" rank="species" species="bunau">
<emphasis box="[219,377,1678,1699]" italics="true" pageId="11" pageNumber="533">Barbodes bunau</emphasis>
, Borneo
</taxonomicName>
: Indonesia: Kalimantan Timur: 
<emphasis bold="true" box="[775,787,1678,1699]" pageId="11" pageNumber="533">a</emphasis>
, 
<emphasis bold="true" box="[800,813,1678,1699]" pageId="11" pageNumber="533">b</emphasis>
, CMK 9535, 20.2 (a) and 28.3 mm SL (b), Sungai Sebuku; 
<emphasis bold="true" box="[1405,1415,1678,1699]" pageId="11" pageNumber="533">c</emphasis>
, ZRC 49867, paratype, 67.5 mm, Sungai Sesayap. (Photographs by M. Kottelat (a, b) and Tan H. H. (c)).
</paragraph>
</caption>
<paragraph blockId="11.[135,775,1784,1840]" pageId="11" pageNumber="533">
<materialsCitation accessionNumber="ZRC 62315, CMK 27384" collectingDate="1992-10-26" collectionCode="ZRC" collectorName="M. Zakaria-Ismail" country="Malaysia" county="Cameron Highlands" elevation="200" latitude="4.436111" location="Sungai" longLatPrecision="21" longitude="101.49444" municipality="Sungai Boh" pageId="11" pageNumber="533" specimenCode="18" specimenCount="1" stateProvince="Pahang" typeStatus="paratype">
<emphasis bold="true" box="[136,256,1784,1808]" pageId="11" pageNumber="533">
<typeStatus box="[136,251,1784,1808]" pageId="11" pageNumber="533">Paratypes</typeStatus>
.
</emphasis>
<accessionNumber box="[261,387,1784,1808]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC62315" pageId="11" pageNumber="533">
<collectionCode box="[261,313,1784,1808]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
62315
</accessionNumber>
, 
<specimenCode box="[396,424,1784,1808]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">18</specimenCode>
, 30.0– 
<quantity box="[494,587,1784,1808]" metricMagnitude="-2" metricUnit="m" metricValue="6.74" pageId="11" pageNumber="533" unit="mm" value="67.4">67.4 mm</quantity>
SL; 
<accessionNumber box="[636,771,1784,1808]" httpUri="http://www.ncbi.nlm.nih.gov/protein/CMK27384" pageId="11" pageNumber="533">CMK 27384</accessionNumber>
, 9, 
<quantity box="[163,320,1816,1840]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" metricValueMax="6.79" metricValueMin="2.41" pageId="11" pageNumber="533" unit="mm" value="46.0" valueMax="67.9" valueMin="24.1">24.1–67.9 mm</quantity>
SL; same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph blockId="11.[134,776,1880,2096]" lastBlockId="11.[834,1474,1784,2064]" pageId="11" pageNumber="533">
<materialsCitation location="All" pageId="11" pageNumber="533" specimenCount="2" stateProvince="Pahang, Pahang">
<emphasis bold="true" box="[136,361,1880,1904]" pageId="11" pageNumber="533">
<typeStatus box="[136,361,1880,1904]" isOtherTypeStatus="true" pageId="11" pageNumber="533">Additional material</typeStatus>
</emphasis>
(non 
<typeStatus box="[424,485,1880,1904]" pageId="11" pageNumber="533">types</typeStatus>
). 
<location LSID="urn:lsid:plazi:treatment:242F0B25FFA7FFE46ACBF88AFC73FA1B:A959ECE8FFA6FFE168A0F8EBFD98F8FE" box="[503,537,1880,1904]" name="All" pageId="11" pageNumber="533" stateProvince="Pahang, Pahang">All</location>
from 
<collectingRegion box="[604,775,1880,1904]" isEnumeration="true" pageId="11" pageNumber="533">Pahang, Pahang</collectingRegion>
drainage
</materialsCitation>
. 
<materialsCitation accessionNumber="ZRC 60583" collectingDate="1990-08-19" collectionCode="ZRC" latitude="4.4245" location="Sungai" longLatPrecision="7" longitude="101.5083" municipality="Sungai Menson" pageId="11" pageNumber="533" specimenCode="4" specimenCount="1" stateProvince="Pahang, Pahang">
<accessionNumber box="[240,366,1912,1936]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC60583" pageId="11" pageNumber="533">
<collectionCode box="[240,292,1912,1936]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
60583
</accessionNumber>
, 
<specimenCode box="[376,391,1912,1936]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">4</specimenCode>
, 
<quantity box="[401,554,1912,1936]" metricMagnitude="-2" metricUnit="m" metricValue="5.26" metricValueMax="8.129999999999999" metricValueMin="2.3899999999999997" pageId="11" pageNumber="533" unit="mm" value="52.599999999999994" valueMax="81.3" valueMin="23.9">23.9–81.3 mm</quantity>
SL; 
<collectingMunicipality box="[603,771,1912,1936]" pageId="11" pageNumber="533">Sungai Menson</collectingMunicipality>
, tributary of 
<location LSID="urn:lsid:plazi:treatment:242F0B25FFA7FFE46ACBF88AFC73FA1B:A959ECE8FFA6FFE16843F82BFEE0F83E" box="[276,353,1944,1968]" latitude="4.4245" longLatPrecision="7" longitude="101.5083" municipality="Sungai Menson" name="Sungai" pageId="11" pageNumber="533" stateProvince="Pahang, Pahang">Sungai</location>
Bertam (
<geoCoordinate box="[465,574,1944,1968]" degrees="4.4245" direction="north" orientation="latitude" pageId="11" pageNumber="533" precision="5" value="4.4245">4.4245°N</geoCoordinate>
<geoCoordinate box="[585,723,1944,1968]" degrees="101.5083" direction="east" orientation="longitude" pageId="11" pageNumber="533" precision="5" value="101.5083">101.5083°E</geoCoordinate>
); 
<date pageId="11" pageNumber="533" value="1990-08-19">
<collectingDate pageId="11" pageNumber="533" value="1990-08-19">19 August 1990</collectingDate>
</date>
</materialsCitation>
. — 
<materialsCitation accessionNumber="ZRC 60590" collectingDate="1992-10-26" collectionCode="ZRC" latitude="4.45" location="Sungai" longLatPrecision="7" longitude="101.5057" municipality="Sungai Menson" pageId="11" pageNumber="533" specimenCode="1" specimenCount="1" stateProvince="Pahang, Pahang">
<accessionNumber box="[312,435,1976,2000]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC60590" pageId="11" pageNumber="533">
<collectionCode box="[312,363,1976,2000]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
60590
</accessionNumber>
, 
<specimenCode box="[444,457,1976,2000]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">1</specimenCode>
, 
<quantity box="[468,560,1976,2000]" metricMagnitude="-2" metricUnit="m" metricValue="6.05" pageId="11" pageNumber="533" unit="mm" value="60.5">60.5 mm</quantity>
SL; 
<collectingMunicipality box="[607,771,1976,2000]" pageId="11" pageNumber="533">Sungai Menson</collectingMunicipality>
, tributary of 
<location LSID="urn:lsid:plazi:treatment:242F0B25FFA7FFE46ACBF88AFC73FA1B:A959ECE8FFA6FFE16843F86BFEE0F87E" box="[276,353,2008,2032]" latitude="4.45" longLatPrecision="7" longitude="101.5057" municipality="Sungai Menson" name="Sungai" pageId="11" pageNumber="533" stateProvince="Pahang, Pahang">Sungai</location>
Bertam (
<geoCoordinate box="[464,574,2008,2032]" degrees="4.4500" direction="north" orientation="latitude" pageId="11" pageNumber="533" precision="5" value="4.45">4.4500°N</geoCoordinate>
<geoCoordinate box="[585,722,2008,2032]" degrees="101.5057" direction="east" orientation="longitude" pageId="11" pageNumber="533" precision="5" value="101.5057">101.5057°E</geoCoordinate>
); 
<date pageId="11" pageNumber="533" value="1992-10-26">
<collectingDate pageId="11" pageNumber="533" value="1992-10-26">26 October 1992</collectingDate>
</date>
</materialsCitation>
. — 
<materialsCitation accessionNumber="ZRC 60584" collectingDate="1992-09-28" collectionCode="ZRC" location="Kuala Kernip" municipality="Sungai" pageId="11" pageNumber="533" specimenCode="6" specimenCount="1" stateProvince="Pahang, Pahang">
<accessionNumber box="[329,457,2040,2064]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC60584" pageId="11" pageNumber="533">
<collectionCode box="[329,381,2040,2064]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
60584
</accessionNumber>
, 
<specimenCode box="[468,483,2040,2064]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">6</specimenCode>
, 
<quantity box="[495,649,2040,2064]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" metricValueMax="6.65" metricValueMin="2.65" pageId="11" pageNumber="533" unit="mm" value="46.5" valueMax="66.5" valueMin="26.5">26.5–66.5 mm</quantity>
SL; 
<collectingMunicipality box="[701,775,2040,2064]" pageId="11" pageNumber="533">Sungai</collectingMunicipality>
So’ok near 
<location LSID="urn:lsid:plazi:treatment:242F0B25FFA7FFE46ACBF88AFC73FA1B:A959ECE8FFA6FFE169AAF7ABFE0DF7BE" box="[253,396,2072,2096]" municipality="Sungai" name="Kuala Kernip" pageId="11" pageNumber="533" stateProvince="Pahang, Pahang">Kuala Kernip</location>
; 
<date box="[405,608,2072,2096]" pageId="11" pageNumber="533" value="1992-09-28">
<collectingDate box="[405,608,2072,2096]" pageId="11" pageNumber="533" value="1992-09-28">28 September 1992</collectingDate>
</date>
</materialsCitation>
. — 
<materialsCitation accessionNumber="ZRC 60576" box="[647,1065,1784,2096]" collectionCode="ZRC" pageId="11" pageNumber="533" specimenCode="4, 43.5" specimenCount="1" stateProvince="Pahang, Pahang">
<accessionNumber box="[647,771,2072,2096]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC60576" pageId="11" pageNumber="533">
<collectionCode box="[647,698,2072,2096]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
60576
</accessionNumber>
, 
<specimenCode box="[834,849,1784,1808]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">4</specimenCode>
, 
<specimenCode box="[863,909,1784,1808]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">43.5</specimenCode>
–69.0 mm SL
</materialsCitation>
; 
<materialsCitation accessionNumber="ZRC 60577" box="[1079,1468,1784,1808]" collectionCode="ZRC" pageId="11" pageNumber="533" specimenCode="11" specimenCount="1" stateProvince="Pahang, Pahang">
<accessionNumber box="[1079,1210,1784,1808]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC60577" pageId="11" pageNumber="533">
<collectionCode box="[1079,1131,1784,1808]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
60577
</accessionNumber>
, 
<specimenCode box="[1224,1252,1784,1808]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">11</specimenCode>
, 
<quantity box="[1267,1424,1784,1808]" metricMagnitude="-2" metricUnit="m" metricValue="4.59" metricValueMax="6.89" metricValueMin="2.29" pageId="11" pageNumber="533" unit="mm" value="45.9" valueMax="68.9" valueMin="22.9">22.9–68.9 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation accessionNumber="ZRC 60578" collectingDate="1993-02-16" collectionCode="ZRC" latitude="4.061" location="Sungai" longLatPrecision="7" longitude="101.761" pageId="11" pageNumber="533" specimenCode="3" specimenCount="1" stateProvince="Pahang, Pahang">
<accessionNumber box="[835,964,1816,1840]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC60578" pageId="11" pageNumber="533">
<collectionCode box="[835,887,1816,1840]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
60578
</accessionNumber>
, 
<specimenCode box="[975,990,1816,1840]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">3</specimenCode>
, 
<quantity box="[1002,1157,1816,1840]" metricMagnitude="-2" metricUnit="m" metricValue="4.995" metricValueMax="6.279999999999999" metricValueMin="3.71" pageId="11" pageNumber="533" unit="mm" value="49.95" valueMax="62.8" valueMin="37.1">37.1–62.8 mm</quantity>
SL; 
<location LSID="urn:lsid:plazi:treatment:242F0B25FFA7FFE46ACBF88AFC73FA1B:A959ECE8FFA6FFE16DECF8ABFA84F8BE" box="[1211,1285,1816,1840]" latitude="4.061" longLatPrecision="7" longitude="101.761" name="Sungai" pageId="11" pageNumber="533" stateProvince="Pahang, Pahang">Sungai</location>
Tersang 
<locationDeviation firstLocation="Bukit Talom" pageId="11" pageNumber="533" secondLocation="Sungai Koyan">between Bukit Talom and Sungai Koyan</locationDeviation>
(
<geoCoordinate box="[1187,1292,1848,1872]" degrees="4.0610" direction="north" orientation="latitude" pageId="11" pageNumber="533" precision="5" value="4.061">4.0610°N</geoCoordinate>
<geoCoordinate box="[1298,1429,1848,1872]" degrees="101.7610" direction="east" orientation="longitude" pageId="11" pageNumber="533" precision="5" value="101.761">101.7610°E</geoCoordinate>
); 
<date pageId="11" pageNumber="533" value="1993-02-16">
<collectingDate pageId="11" pageNumber="533" value="1993-02-16">16 February 1993</collectingDate>
</date>
</materialsCitation>
. — 
<materialsCitation accessionNumber="ZRC 60581" collectingDate="1992-11-04" collectionCode="ZRC" latitude="4.6307" location="Sungai" longLatPrecision="7" longitude="102.064" pageId="11" pageNumber="533" specimenCode="22, 13.5" specimenCount="1" stateProvince="Pahang, Pahang">
<accessionNumber box="[1036,1163,1880,1904]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC60581" pageId="11" pageNumber="533">
<collectionCode box="[1036,1088,1880,1904]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
60581
</accessionNumber>
, 
<specimenCode box="[1173,1202,1880,1904]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">22</specimenCode>
, 
<specimenCode box="[1212,1257,1880,1904]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">13.5</specimenCode>
–26.0 mm SL; small stream draining to 
<location LSID="urn:lsid:plazi:treatment:242F0B25FFA7FFE46ACBF88AFC73FA1B:A959ECE8FFA6FFE16D45F8CBFBDDF81E" box="[1042,1116,1912,1936]" latitude="4.6307" longLatPrecision="7" longitude="102.064" name="Sungai" pageId="11" pageNumber="533" stateProvince="Pahang, Pahang">Sungai</location>
Tanom (
<geoCoordinate box="[1216,1321,1912,1936]" degrees="4.6307" direction="north" orientation="latitude" pageId="11" pageNumber="533" precision="5" value="4.6307">4.6307°N</geoCoordinate>
<geoCoordinate box="[1330,1462,1912,1936]" degrees="102.0640" direction="east" orientation="longitude" pageId="11" pageNumber="533" precision="5" value="102.064">102.0640°E</geoCoordinate>
); 
<date box="[835,1039,1944,1968]" pageId="11" pageNumber="533" value="1992-11-04">
<collectingDate box="[835,1039,1944,1968]" pageId="11" pageNumber="533" value="1992-11-04">4 November 1992</collectingDate>
</date>
</materialsCitation>
. — 
<materialsCitation accessionNumber="ZRC 52348" box="[1089,1469,1944,1968]" collectionCode="ZRC" pageId="11" pageNumber="533" specimenCode="9" specimenCount="1" stateProvince="Pahang, Pahang">
<accessionNumber box="[1089,1222,1944,1968]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC52348" pageId="11" pageNumber="533">
<collectionCode box="[1089,1141,1944,1968]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
52348
</accessionNumber>
, 
<specimenCode box="[1236,1251,1944,1968]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">9</specimenCode>
, 
<quantity box="[1266,1425,1944,1968]" metricMagnitude="-2" metricUnit="m" metricValue="6.495" metricValueMax="7.83" metricValueMin="5.16" pageId="11" pageNumber="533" unit="mm" value="64.95" valueMax="78.3" valueMin="51.6">51.6–78.3 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation accessionNumber="ZRC 60587" box="[835,1228,1976,2000]" collectionCode="ZRC" pageId="11" pageNumber="533" specimenCode="6" specimenCount="1" stateProvince="Pahang, Pahang">
<accessionNumber box="[835,971,1976,2000]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC60587" pageId="11" pageNumber="533">
<collectionCode box="[835,888,1976,2000]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
60587
</accessionNumber>
, 
<specimenCode box="[986,1002,1976,2000]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">6</specimenCode>
, 
<quantity box="[1018,1182,1976,2000]" metricMagnitude="-2" metricUnit="m" metricValue="5.01" metricValueMax="6.609999999999999" metricValueMin="3.41" pageId="11" pageNumber="533" unit="mm" value="50.099999999999994" valueMax="66.1" valueMin="34.1">34.1–66.1 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation accessionNumber="ZRC 60588" collectingDate="1993-02-15" collectionCode="ZRC" latitude="3.9833" location="Sungai" longLatPrecision="7" longitude="101.6258" municipality="Sungai Kenor" pageId="11" pageNumber="533" specimenCode="1" specimenCount="1" stateProvince="Pahang, Pahang">
<accessionNumber box="[1244,1379,1976,2000]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZRC60588" pageId="11" pageNumber="533">
<collectionCode box="[1244,1297,1976,2000]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">ZRC</collectionCode>
60588
</accessionNumber>
, 
<specimenCode box="[1394,1408,1976,2000]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="533">1</specimenCode>
, 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="5.71" pageId="11" pageNumber="533" unit="mm" value="57.1">57.1 mm</quantity>
SL; 
<collectingMunicipality box="[929,1080,2008,2032]" pageId="11" pageNumber="533">Sungai Kenor</collectingMunicipality>
, tributary of 
<location LSID="urn:lsid:plazi:treatment:242F0B25FFA7FFE46ACBF88AFC73FA1B:A959ECE8FFA6FFE16D94F86BFA8CF87E" box="[1219,1293,2008,2032]" latitude="3.9833" longLatPrecision="7" longitude="101.6258" municipality="Sungai Kenor" name="Sungai" pageId="11" pageNumber="533" stateProvince="Pahang, Pahang">Sungai</location>
Lipis (
<geoCoordinate box="[1369,1474,2008,2032]" degrees="3.9833" direction="north" orientation="latitude" pageId="11" pageNumber="533" precision="5" value="3.9833">3.9833°N</geoCoordinate>
<geoCoordinate box="[836,967,2040,2064]" degrees="101.6258" direction="east" orientation="longitude" pageId="11" pageNumber="533" precision="5" value="101.6258">101.6258°E</geoCoordinate>
); 
<date box="[988,1185,2040,2064]" pageId="11" pageNumber="533" value="1993-02-15">
<collectingDate box="[988,1185,2040,2064]" pageId="11" pageNumber="533" value="1993-02-15">15 February 1993</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7174509" ID-Zenodo-Dep="7174509" httpUri="https://zenodo.org/record/7174509/files/figure.png" pageId="12" pageNumber="534" startId="12.[113,148,1102,1124]" targetBox="[113,1451,189,1086]" targetPageId="12">
<paragraph blockId="12.[113,1452,1101,1152]" pageId="12" pageNumber="534">
Fig. 13. 
<taxonomicName authority="Kottelat &amp; Lim, 2021" authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[198,436,1102,1123]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="534" phylum="Chordata" rank="species" species="zakariaismaili" status="new species">
<emphasis box="[198,436,1102,1123]" italics="true" pageId="12" pageNumber="534">Barbodes zakariaismaili</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[451,571,1102,1124]" pageId="12" pageNumber="534" rank="species">new species</taxonomicNameLabel>
, Malaysia: Pahang: Cameron Highlands; 
<emphasis bold="true" box="[983,995,1102,1123]" pageId="12" pageNumber="534">a</emphasis>
, ZRC 52347, holotype, 66.3 mm SL; 
<emphasis bold="true" box="[1377,1390,1102,1123]" pageId="12" pageNumber="534">b</emphasis>
, ZRC 62315, paratype, 58.3 mm SL. (Photographs by M. Kottelat).
</paragraph>
</caption>
<subSubSection pageId="12" pageNumber="534" type="diagnosis">
<paragraph blockId="12.[112,754,1207,1648]" pageId="12" pageNumber="534">
<emphasis bold="true" box="[113,241,1207,1231]" pageId="12" pageNumber="534">Diagnosis.</emphasis>
<taxonomicName authority="Kottelat &amp; Lim, 2021" authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[254,548,1208,1232]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="534" phylum="Chordata" rank="species" species="zakariaismaili" status="new species">
<emphasis box="[254,548,1208,1232]" italics="true" pageId="12" pageNumber="534">Barbodes zakariaismaili</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[568,716,1208,1232]" pageId="12" pageNumber="534" rank="species">new species</taxonomicNameLabel>
, is distinguished from all other species of the 
<taxonomicName box="[614,753,1240,1264]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="534" phylum="Chordata" rank="species" species="binotatus">
<emphasis box="[614,753,1240,1264]" italics="true" pageId="12" pageNumber="534">B. binotatus</emphasis>
</taxonomicName>
group by its unique colour pattern in adults, including a faint longitudinally elongate blackish midlateral mark from the upper extremity of the gill opening to below the dorsal-fin origin; a black spot below the anterior part of the dorsal-fin base, extending downwards to the midlateral row as a narrow triangular mark; and a blackish spot at the end of the caudal peduncle. Other characters useful for identification, but not unique to the species, are: slender body (depth 2.9–3.1 times in SL); interobital area convex; eye not flush with dorsal profile, relatively small (4–5 times in head length, 1.5–1.9 times in interorbital distance); juveniles with a conspicuous reticulate pattern made of black pigments on scale pockets.
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="536" pageId="12" pageNumber="534" type="description">
<paragraph blockId="12.[112,755,1687,2096]" lastBlockId="12.[813,1451,1208,1264]" pageId="12" pageNumber="534">
<emphasis bold="true" box="[113,252,1687,1711]" pageId="12" pageNumber="534">Description.</emphasis>
General appearance is shown in 
<figureCitation box="[617,705,1688,1712]" captionStart="Fig" captionStartId="12.[113,148,1102,1124]" captionTargetBox="[113,1451,189,1086]" captionTargetId="figure-258@12.[113,1451,189,1086]" captionTargetPageId="12" captionText="Fig. 13. Barbodes zakariaismaili, new species, Malaysia: Pahang: Cameron Highlands; a, ZRC 52347, holotype, 66.3 mm SL; b, ZRC 62315, paratype, 58.3 mm SL. (Photographs by M. Kottelat)." figureDoi="http://doi.org/10.5281/zenodo.7174509" httpUri="https://zenodo.org/record/7174509/files/figure.png" pageId="12" pageNumber="534">Figs. 13</figureCitation>
and 
<figureCitation box="[115,145,1720,1744]" captionStart="Fig" captionStartId="13.[136,171,1993,2015]" captionTargetBox="[136,1458,268,1984]" captionTargetId="figure-55@13.[136,1474,258,1985]" captionTargetPageId="13" captionText="Fig. 14. Barbodes zakariaismaili, new species, Malaysia: Pahang: Cameron Highlands; a, b, CMK 27384, paratypes: 27.8 mm SL (a) and 33.7 mm SL (b); c, d, ZRC 62315, paratypes: 30.1 mm SL (c) and 35.5 mm SL (d). (Photographs by M. Kottelat)." figureDoi="http://doi.org/10.5281/zenodo.7174511" httpUri="https://zenodo.org/record/7174511/files/figure.png" pageId="12" pageNumber="534">14</figureCitation>
. Morphometric data of 
<typeStatus box="[413,507,1720,1744]" pageId="12" pageNumber="534">holotype</typeStatus>
and 
<specimenCount box="[566,711,1720,1744]" pageId="12" pageNumber="534" type="generic" typeStatus="paratypes">six paratypes</specimenCount>
are given in 
<tableCitation box="[210,326,1752,1776]" captionStart="Table 1" captionStartId="5.[137,192,189,211]" captionText="Table 1. Morphometric data of Barbodes sellifer, new species, holotype, ZRC 12354, and eight paratypes (ZRC 12355, 17754, 50405, CMK 6027) and B. zakariaismaili, new species, holotype, ZRC 52347, and six paratypes, ZRC 62315. Values of holotype (H) included in ranges and means." pageId="12" pageNumber="534">Table 1. A</tableCitation>
relatively slender species of 
<taxonomicName authorityName="Bleeker" authorityYear="1859" box="[643,745,1752,1776]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="534" phylum="Chordata" rank="genus">
<emphasis box="[643,745,1752,1776]" italics="true" pageId="12" pageNumber="534">Barbodes</emphasis>
</taxonomicName>
, with body depth at dorsal-fin origin about 2.9–3.1 times in standard length, and depth of caudal peduncle 2.0–2.3 times in body depth. Dorsal profile of head and body arched, with a shallow concavity at nape, and a low post-occipital hump in specimens above about 
<quantity box="[467,542,1911,1935]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="534" unit="mm" value="50.0">50 mm</quantity>
SL. Snout pointed, directed forwards. Ventral profile slightly less arched than dorsal profile. Interorbital area convex; eye not flush with dorsal profile of head, diameter 1.5–1.9 times in interorbital distance (
<figureCitation box="[211,288,2039,2064]" captionStart="Fig" captionStartId="6.[113,148,743,765]" captionTargetBox="[113,1451,189,728]" captionTargetId="figure-688@6.[113,1451,189,728]" captionTargetPageId="6" captionText="Fig. 5. a, Barbodes sellifer, new species, CMK 6027, paratype, 59.5 mm SL; b, B. zakariaismaili, new species, ZRC 52347, holotype, 66.3 mm SL. Note differences in snout shape, and in size and position of eye. (Photographs by M. Kottelat)." figureDoi="http://doi.org/10.5281/zenodo.7174487" httpUri="https://zenodo.org/record/7174487/files/figure.png" pageId="12" pageNumber="534">Fig. 5b</figureCitation>
). Mouth subterminal, postlabial groove deep, narrowly interrupted in middle; snout slightly projecting, rostral fold covering upper part of upper lip; lower jaw not enclosed in upper lip.
</paragraph>
<paragraph blockId="12.[812,1453,1304,1808]" pageId="12" pageNumber="534">Dorsal fin with 3 (3) or 4 (4*) visible unbranched and 8½ branched rays; origin above lateral line scale 7 (1), 8 (5*), or 9 (1); distal edge straight to slightly convex; last unbranched ray moderately thick, with 17–25 serrae along most of posterior edge of stiff portion, space between serrae narrower than width of serrae. Pectoral fin rounded, with 15 (4*) or 16 (3) rays (including anterior unbranched ray); reaching almost to pelvic-fin base. Pelvic fin with slightly rounded posterior edge, with 9 (6*) or 10 (1) rays (including anterior unbranched ray); reaching to about 2 scales in front of anus; pelvic axillary scale present, about ¼ of fin length. Anal fin with 3 unbranched and 5½ branched rays; posterior edge straight. Caudal fin with 10–11 + 9 principal rays (of which 8+8 (1) or 9+8 (6*) branched), forked, lobes rounded, tip slightly pointed. Depth of caudal peduncle 1.2–1.4 times in its length.</paragraph>
<paragraph blockId="12.[813,1452,1847,2064]" lastBlockId="14.[113,752,187,275]" lastPageId="14" lastPageNumber="536" pageId="12" pageNumber="534">
21+3 (1), 22+2 (5*), or 23+2 (1) scales along lateral line, 8 (1), 9 (5*), or 10 (1) predorsal scales, ½3/1/4½ (1) or ½4/1/4½ (6*) rows of scales from dorsal-fin origin to about 2 scales in front of pelvic-fin base, ½2/1/2½ scales rows in transverse line on caudal peduncle, 3 scale rows between lateral line and pelvic-fin origin. Posteriormost scale on each lobe of caudal fin larger than preceding ones (
<figureCitation box="[1359,1440,2039,2064]" captionStart="Fig" captionStartId="7.[136,171,955,977]" captionTargetBox="[136,1474,189,940]" captionTargetId="figure-7@7.[136,1474,189,940]" captionTargetPageId="7" captionText="Fig. 6. Scales at caudal-fin base in: a, Barbodes sellifer, new species, ZRC 12355, 69.2 mm SL; and b, B. zakariaismaili, new species, ZRC 62315, 67.4 mm SL. Edge of posteriormost scales enhanced. (Photographs by M. Kottelat)." figureDoi="http://doi.org/10.5281/zenodo.7174489" httpUri="https://zenodo.org/record/7174489/files/figure.png" pageId="12" pageNumber="534">Fig. 6b</figureCitation>
). Two pairs of maxillary barbels; anterior one reaching about middle of postorbital area of head; posterior one reaching posterior edge of opercle.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7174511" ID-Zenodo-Dep="7174511" httpUri="https://zenodo.org/record/7174511/files/figure.png" pageId="13" pageNumber="535" startId="13.[136,171,1993,2015]" targetBox="[136,1458,268,1984]" targetPageId="13">
<paragraph blockId="13.[136,1474,1993,2044]" pageId="13" pageNumber="535">
Fig. 14. 
<taxonomicName authority="Kottelat &amp; Lim, 2021" authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[218,454,1993,2014]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="535" phylum="Chordata" rank="species" species="zakariaismaili" status="new species">
<emphasis box="[218,454,1993,2014]" italics="true" pageId="13" pageNumber="535">Barbodes zakariaismaili</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[466,585,1993,2015]" pageId="13" pageNumber="535" rank="species">new species</taxonomicNameLabel>
, Malaysia: Pahang: Cameron Highlands; 
<emphasis bold="true" box="[988,1000,1993,2014]" pageId="13" pageNumber="535">a</emphasis>
, 
<emphasis bold="true" box="[1012,1025,1993,2014]" pageId="13" pageNumber="535">b</emphasis>
, CMK 27384, paratypes: 27.8 mm SL (a) and 33.7 mm SL (b); 
<emphasis bold="true" box="[311,320,2022,2043]" pageId="13" pageNumber="535">c</emphasis>
, 
<emphasis bold="true" box="[333,346,2022,2043]" pageId="13" pageNumber="535">d</emphasis>
, ZRC 62315, paratypes: 30.1 mm SL (c) and 35.5 mm SL (d). (Photographs by M. Kottelat).
</paragraph>
</caption>
<paragraph blockId="14.[113,753,315,947]" pageId="14" pageNumber="536">
<emphasis bold="true" box="[113,262,315,339]" pageId="14" pageNumber="536">Colouration.</emphasis>
Preserved specimens: specimens above 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="536" unit="mm" value="35.0">35 mm</quantity>
SL: body and head yellowish brown, darker on back, yellowish on belly. No reticulate pattern on body. On scale row +1 and on anterior 3–5 scales of row 0, scale pockets covered by dark pigments, centre of scales dark brown, posterior fourth of scales paler brown and with sparse black pigments; sometimes appearing as a midlateral row of dark spots. In anterior third, these scales superimposed on a faint longitudinally elongate blackish midlateral mark (made of deeper pigments) from upper extremity of gill opening to below dorsal-fin origin. On row +2, centre of scales less dark and mark on scale pocket narrower. On lateral line row and rows –1 and –2, only dark scale pocket pigmentation. A faint dark grey spot at end of caudal peduncle. A black spot on sheath scales of dorsal fin or on upper part of row +4, at base of branched dorsal-fin rays 1–2; area of spot equal to about one scale; a faint triangular extension, about 1–1.5 scale wide, reaching to row +3 or +2 (then in contact with midlateral row of dark scales). All fins hyaline; in a few specimens, distal edge of anal fin blackish.
</paragraph>
<paragraph blockId="14.[112,754,987,1364]" pageId="14" pageNumber="536">
Juveniles. Only three small specimens available, 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="3.075" metricValueMax="3.37" metricValueMin="2.78" pageId="14" pageNumber="536" unit="mm" value="30.75" valueMax="33.7" valueMin="27.8">27.8–33.7 mm</quantity>
SL (
<figureCitation box="[204,317,1019,1044]" captionStart="Fig" captionStartId="13.[136,171,1993,2015]" captionTargetBox="[136,1458,268,1984]" captionTargetId="figure-55@13.[136,1474,258,1985]" captionTargetPageId="13" captionText="Fig. 14. Barbodes zakariaismaili, new species, Malaysia: Pahang: Cameron Highlands; a, b, CMK 27384, paratypes: 27.8 mm SL (a) and 33.7 mm SL (b); c, d, ZRC 62315, paratypes: 30.1 mm SL (c) and 35.5 mm SL (d). (Photographs by M. Kottelat)." figureDoi="http://doi.org/10.5281/zenodo.7174511" httpUri="https://zenodo.org/record/7174511/files/figure.png" pageId="14" pageNumber="536">Fig. 14a–c</figureCitation>
). Body with conspicuous reticulate pattern made of black pigments on scale pocket, on rows +4 to –2. Elongated blotch on anterior third of body present, made of pigments in deeper layer. An elongated patch of pigments under scales on row +1, extending from below end of dorsalfin base to end of anal-fin base, vaguely forming two or three spots. Roundish black spot at posterior extremity of caudal peduncle. Black spot at dorsal-fin base as in adults. Black spot on body, at base of anal fin. Distal margin of anal fin black. Dorsal fin: black pigments along edges of branched rays near branching point.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="536" type="distribution">
<paragraph blockId="14.[113,753,1403,1620]" pageId="14" pageNumber="536">
<emphasis bold="true" box="[113,260,1403,1427]" pageId="14" pageNumber="536">Distribution.</emphasis>
<taxonomicName authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[265,526,1403,1427]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="zakariaismaili">
<emphasis box="[265,526,1403,1427]" italics="true" pageId="14" pageNumber="536">Barbodes zakariaismaili</emphasis>
</taxonomicName>
is presently recorded only from Peninsular 
<collectingCountry box="[349,449,1435,1459]" name="Malaysia" pageId="14" pageNumber="536">Malaysia</collectingCountry>
, in tributaries of the Sungai Jelai of the 
<collectingRegion box="[243,322,1467,1491]" country="Malaysia" name="Pahang" pageId="14" pageNumber="536">Pahang</collectingRegion>
drainage in north-western 
<collectingRegion box="[614,753,1467,1492]" country="Malaysia" name="Pahang" pageId="14" pageNumber="536">Pahang State</collectingRegion>
(
<figureCitation box="[123,196,1499,1524]" captionStart="Fig" captionStartId="7.[136,171,1877,1899]" captionTargetBox="[143,1471,1072,1857]" captionTargetId="figure-55@7.[143,998,1072,1857]" captionTargetPageId="7" captionText="Fig. 7. Distribution of Barbodes sellifer, new species (green marks) and B. zakariaismaili, new species (orange marks) on Malay Peninsula and Sumatra, based on examined material. Stars are type localities. Each mark may consist of more than one locality. (K. K. P. Lim)." figureDoi="http://doi.org/10.5281/zenodo.7174491" httpUri="https://zenodo.org/record/7174491/files/figure.png" pageId="14" pageNumber="536">Fig. 7</figureCitation>
). The 
<typeStatus box="[275,323,1500,1524]" pageId="14" pageNumber="536">type</typeStatus>
locality, Sungai Boh, is a small and shallow hill stream about 4–6 metres wide, under forest canopy. It has clear and cool water flowing over rock and sand substrate (Khaironizam M. Z., pers. comm.).
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="536" type="etymology">
<paragraph blockId="14.[113,753,1659,1748]" pageId="14" pageNumber="536">
<emphasis bold="true" box="[113,243,1660,1684]" pageId="14" pageNumber="536">Etymology.</emphasis>
The species is named for Mohd. Zakaria-Ismail in appreciation for his work on the fish fauna of 
<collectingCountry box="[648,749,1691,1715]" name="Malaysia" pageId="14" pageNumber="536">Malaysia</collectingCountry>
. A noun in the genitive, indeclinable.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="536" type="discussion">
<paragraph blockId="14.[112,752,1787,1971]" pageId="14" pageNumber="536">
<emphasis bold="true" box="[113,228,1788,1812]" pageId="14" pageNumber="536">Remarks.</emphasis>
<taxonomicName authority="Kottelat &amp; Lim, 2021" authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[239,518,1787,1811]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="zakariaismaili" status="new species">
<emphasis box="[239,518,1787,1811]" italics="true" pageId="14" pageNumber="536">Barbodes zakariaismaili</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[536,677,1787,1811]" pageId="14" pageNumber="536" rank="species">new species</taxonomicNameLabel>
, has a relatively slender body, with the ventral profile only slightly less curved than the dorsal one, and the tip of the snout about level with the axis of the body. This body shape is unusual in the 
<taxonomicName box="[175,302,1915,1939]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="binotatus">
<emphasis box="[175,302,1915,1939]" italics="true" pageId="14" pageNumber="536">B. binotatus</emphasis>
</taxonomicName>
group; among the named species presently recognised as valid, we have seen it only in 
<taxonomicName box="[603,732,1947,1971]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="binotatus">
<emphasis box="[603,732,1947,1971]" italics="true" pageId="14" pageNumber="536">B. binotatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="14.[113,752,2011,2099]" lastBlockId="14.[812,1452,189,405]" pageId="14" pageNumber="536">
<taxonomicName box="[113,321,2011,2035]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="binotatus">
<emphasis box="[113,321,2011,2035]" italics="true" pageId="14" pageNumber="536">Barbodes binotatus</emphasis>
</taxonomicName>
(
<figureCitation box="[336,412,2011,2036]" captionStart="Fig" captionStartId="10.[113,148,1292,1314]" captionTargetBox="[113,1451,714,1277]" captionTargetId="figure-395@10.[113,1451,714,1277]" captionTargetPageId="10" captionText="Fig. 11. Barbodes binotatus, CMK 8976, 43.0 mm SL; Java: Ciliwung at Cilebut. (Photograph by M. Kottelat)." figureDoi="http://doi.org/10.5281/zenodo.7174505" httpUri="https://zenodo.org/record/7174505/files/figure.png" pageId="14" pageNumber="536">Fig. 11</figureCitation>
) is distinguished from the other species of the group by the presence of a midlateral stripe from the upper extremity of the gill opening to the spot at the end of the caudal peduncle, in both juveniles and adults; 
<taxonomicName authority="Kottelat &amp; Lim, 2021" authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[813,915,221,245]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="sellifer" status="new species">
<emphasis box="[813,915,221,245]" italics="true" pageId="14" pageNumber="536">B. sellifer</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[927,1057,221,245]" pageId="14" pageNumber="536" rank="species">new species</taxonomicNameLabel>
, by the large triangular to rectangular blotch below the dorsal-fin base, 
<taxonomicName authorityName="Herre" authorityYear="1940" box="[1184,1287,253,277]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="banksi">
<emphasis box="[1184,1287,253,277]" italics="true" pageId="14" pageNumber="536">B. banksi</emphasis>
</taxonomicName>
by the narrow slanted bar under the dorsal-fin origin, and 
<taxonomicName box="[1310,1451,285,309]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="rhombeus">
<emphasis box="[1310,1451,285,309]" italics="true" pageId="14" pageNumber="536">B. rhombeus</emphasis>
</taxonomicName>
has only a small black spot at the origin of the dorsal fin, at the end of the caudal peduncle, and a midlateral row of 3–5 spots sometimes connected by a thin stripe.
</paragraph>
<paragraph blockId="14.[811,1453,445,1109]" pageId="14" pageNumber="536">
<taxonomicName authority="Kottelat &amp; Lim, 2021" authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[813,1094,445,469]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="zakariaismaili" status="new species">
<emphasis box="[813,1094,445,469]" italics="true" pageId="14" pageNumber="536">Barbodes zakariaismaili</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1113,1255,445,469]" pageId="14" pageNumber="536" rank="species">new species</taxonomicNameLabel>
, is distinguished from 
<taxonomicName authority="Kottelat &amp; Lim, 2021" authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[875,982,477,501]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="sellifer" status="new species">
<emphasis box="[875,982,477,501]" italics="true" pageId="14" pageNumber="536">B. sellifer</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[999,1134,477,501]" pageId="14" pageNumber="536" rank="species">new species</taxonomicNameLabel>
, in having, in adults, a faint longitudinally elongate blackish midlateral mark from the upper extremity of the gill opening to below the dorsal-fin origin (vs. absence), a black spot below the anterior part of the dorsal-fin base, extending downwards to the midlateral row as a narrow triangular mark (vs., in adults, a large triangular to rectangular blotch extending downwards from in front of and beneath the base of the dorsal fin, sometimes incomplete or narrower), a more pointed snout (
<figureCitation box="[1214,1281,733,757]" captionStart="Fig" captionStartId="6.[113,148,743,765]" captionTargetBox="[113,1451,189,728]" captionTargetId="figure-688@6.[113,1451,189,728]" captionTargetPageId="6" captionText="Fig. 5. a, Barbodes sellifer, new species, CMK 6027, paratype, 59.5 mm SL; b, B. zakariaismaili, new species, ZRC 52347, holotype, 66.3 mm SL. Note differences in snout shape, and in size and position of eye. (Photographs by M. Kottelat)." figureDoi="http://doi.org/10.5281/zenodo.7174487" httpUri="https://zenodo.org/record/7174487/files/figure.png" pageId="14" pageNumber="536">Fig. 5</figureCitation>
), and a smaller eye (diameter 21–25% HL vs. 26–30). Our morphometric data, based on a few specimens of both species in the size range 
<quantity box="[882,999,829,853]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" metricValueMax="9.8" metricValueMin="6.6" pageId="14" pageNumber="536" unit="mm" value="82.0" valueMax="98.0" valueMin="66.0">66–98 mm</quantity>
SL, suggest that 
<taxonomicName authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[1191,1375,829,853]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="zakariaismaili">
<emphasis box="[1191,1375,829,853]" italics="true" pageId="14" pageNumber="536">B. zakariaismaili</emphasis>
</taxonomicName>
differs from 
<taxonomicName authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[872,977,861,885]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="sellifer">
<emphasis box="[872,977,861,885]" italics="true" pageId="14" pageNumber="536">B. sellifer</emphasis>
</taxonomicName>
in having a shorter (27–28% SL vs. 28–32) and shallower head (depth 20–22% SL vs. 22–24), a smaller predorsal length (53–58% SL vs. 58–63) and a more slender body (depth at dorsal-fin origin 32–35% SL vs. 37–42; depth of caudal peduncle 15–16% SL vs. 16–17). However, we have not measured a sufficient number of specimens from enough localities to be certain that all these differences would hold for all populations.
</paragraph>
<paragraph blockId="14.[813,1452,1149,1429]" pageId="14" pageNumber="536">
Presently, we include in 
<taxonomicName authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[1077,1260,1149,1173]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="zakariaismaili">
<emphasis box="[1077,1260,1149,1173]" italics="true" pageId="14" pageNumber="536">B. zakariaismaili</emphasis>
</taxonomicName>
material from the Sungai Jelai watershed only. Its presence in the headwaters of adjacent drainages is not impossible. We are aware of 
<taxonomicName authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[813,920,1245,1269]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="sellifer">
<emphasis box="[813,920,1245,1269]" italics="true" pageId="14" pageNumber="536">B. sellifer</emphasis>
</taxonomicName>
-like fish from the 
<collectingRegion box="[1132,1228,1245,1269]" country="Malaysia" name="Kelantan" pageId="14" pageNumber="536">Kelantan</collectingRegion>
drainages, but have not seen fresh or well-preserved specimens to be able to determine their identity; they may be 
<taxonomicName authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[1251,1445,1309,1333]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="zakariaismaili">
<emphasis box="[1251,1445,1309,1333]" italics="true" pageId="14" pageNumber="536">B. zakariaismaili</emphasis>
</taxonomicName>
. It is noteworthy that 
<taxonomicName authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[1047,1232,1341,1365]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="zakariaismaili">
<emphasis box="[1047,1232,1341,1365]" italics="true" pageId="14" pageNumber="536">B. zakariaismaili</emphasis>
</taxonomicName>
is recorded in hilly areas, while 
<taxonomicName authorityName="Kottelat &amp; Lim" authorityYear="2021" box="[949,1053,1373,1397]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="536" phylum="Chordata" rank="species" species="sellifer">
<emphasis box="[949,1053,1373,1397]" italics="true" pageId="14" pageNumber="536">B. sellifer</emphasis>
</taxonomicName>
occurs in the lowlands, in areas then covered by forest.
</paragraph>
</subSubSection>
</treatment>
</document>