171 lines
21 KiB
XML
171 lines
21 KiB
XML
<document id="5091336D60327E78A1AD79A54AE85050" ID-DOI="10.11646/palaeoentomology.6.3.13" ID-ISSN="2624-2834" ID-Zenodo-Dep="8209295" ID-ZooBank="12D99FD9-32AF-4F30-AF15-35CD80E7A586" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1691005736286" checkinUser="plazi" docAuthor="PANKOWSKI, MAXIMILIAN G. & Fanti, FABRIZIO FANTI" docDate="2023" docId="03C387DBD029FFAF87B2FB95E332F7FC" docLanguage="en" docName="palaeoentomology.6.3.13.pdf" docOrigin="Palaeoentomology 6 (3)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.3.13" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Malthinus (Malthinus) pauljohnsoni PANKOWSKI & Fanti 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="306" masterDocId="FFFAFFA3D02FFFA9873AFFE3E744FFF4" masterDocTitle="Six new species of fossil soldier beetles (Coleoptera: Cantharidae) from Eocene Baltic amber" masterLastPageNumber="312" masterPageNumber="300" pageNumber="306" updateTime="1691482849966" updateUser="tatiana">
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<mods:title id="E0EA418D63C1123C85F34903C01E26DC">Six new species of fossil soldier beetles (Coleoptera: Cantharidae) from Eocene Baltic amber</mods:title>
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<mods:namePart id="283CBA5E047319454A9FB50CF6D17827">PANKOWSKI, MAXIMILIAN G.</mods:namePart>
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<mods:namePart id="23A73F24F1EEC699B3CEA077921F83E7">Fanti, FABRIZIO FANTI</mods:namePart>
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<mods:title id="F5AF01BC9735563D8F83C6062FD4D924">Palaeoentomology</mods:title>
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<mods:part id="502F97966A3B456F9677937F307E7D9C">
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<mods:date id="D324B5F3235D8BE72AABB9244E5D5B19">2023</mods:date>
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<mods:number id="0842308D52808B4ADADD8FD769F41CB4">2023-06-29</mods:number>
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<subSubSection id="C3706546D029FFAF87B2FB95E5D5FB64" box="[136,657,1142,1169]" pageId="6" pageNumber="306" type="nomenclature">
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<paragraph id="8BD536CDD029FFAF87B2FB95E5D5FB64" blockId="6.[136,657,1142,1204]" box="[136,657,1142,1169]" pageId="6" pageNumber="306">
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<heading id="D09D81A1D029FFAF87B2FB95E5D5FB64" bold="true" box="[136,657,1142,1169]" fontSize="10" level="2" pageId="6" pageNumber="306" reason="3">
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<emphasis id="B91EEADFD029FFAF87B2FB95E5D5FB64" bold="true" box="[136,657,1142,1169]" pageId="6" pageNumber="306">
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<taxonomicName id="4C6A4D4ED029FFAF87B2FB95E575FB65" authority="PANKOWSKI & Fanti, 2023" authorityName="PANKOWSKI & Fanti" authorityYear="2023" box="[136,561,1142,1169]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="306" phylum="Arthropoda" rank="species" species="pauljohnsoni" status="sp. nov." subGenus="Malthinus">
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<emphasis id="B91EEADFD029FFAF87B2FB95E644FB65" bold="true" box="[136,256,1142,1169]" italics="true" pageId="6" pageNumber="306">Malthinus</emphasis>
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(
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<emphasis id="B91EEADFD029FFAF862AFB95E6CCFB65" bold="true" box="[272,392,1142,1169]" italics="true" pageId="6" pageNumber="306">Malthinus</emphasis>
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)
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<emphasis id="B91EEADFD029FFAF86A2FB95E575FB65" bold="true" box="[408,561,1142,1169]" italics="true" pageId="6" pageNumber="306">pauljohnsoni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22D57A4D029FFAF8502FB95E5D5FB64" box="[568,657,1142,1168]" pageId="6" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3706546D029FFAF87B2FB7AE798FB40" box="[136,220,1177,1204]" pageId="6" pageNumber="306" type="description">
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||
<paragraph id="8BD536CDD029FFAF87B2FB7AE798FB40" blockId="6.[136,657,1142,1204]" box="[136,220,1177,1204]" pageId="6" pageNumber="306">
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||
(
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<figureCitation id="13512A48D029FFAF87AAFB7AE797FB40" box="[144,211,1177,1204]" captionStart="FIGURE 5" captionStartId="7.[136,235,1446,1470]" captionTargetBox="[280,1308,189,1423]" captionTargetId="figure-203@7.[280,1308,189,1423]" captionTargetPageId="7" captionText="FIGURE 5. Malthinus (Malthinus) pauljohnsoni sp. nov. in Baltic amber. Holotype (USNM PAL 787826). A, Dorsal view. B, Ventral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8209305" httpUri="https://zenodo.org/record/8209305/files/figure.png" pageId="6" pageNumber="306">Fig. 5</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3706546D029FFAF87B2FB3CE536FA71" pageId="6" pageNumber="306" type="materials_examined">
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<paragraph id="8BD536CDD029FFAF87B2FB3CE630FACB" blockId="6.[136,778,1247,2077]" pageId="6" pageNumber="306">
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<materialsCitation id="3B023C90D029FFAF87B2FB3CE630FACB" collectionCode="USNM" county="Smithsonian National Museum of Natural History" location="Female" municipality="Baltic" pageId="6" pageNumber="306" specimenCode="PAL 787826" specimenCount="1" typeStatus="holotype">
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<emphasis id="B91EEADFD029FFAF87B2FB3CE7B8FB0D" bold="true" box="[136,252,1247,1273]" pageId="6" pageNumber="306">
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<typeStatus id="54D1886FD029FFAF87B2FB3CE7B3FB0D" box="[136,247,1247,1273]" pageId="6" pageNumber="306">Holotype</typeStatus>
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.
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</emphasis>
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||
<location id="8EB56016D029FFAF863CFB3CE61FFB0D" LSID="urn:lsid:plazi:treatment:03C387DBD029FFAF87B2FB95E332F7FC:8EB56016D029FFAF863CFB3CE61FFB0D" box="[262,347,1247,1273]" county="Smithsonian National Museum of Natural History" municipality="Baltic" name="Female" pageId="6" pageNumber="306">Female</location>
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, inclusion in
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<collectingMunicipality id="6BB1ACB7D029FFAF86C0FB3CE579FB0D" box="[506,573,1247,1273]" pageId="6" pageNumber="306">Baltic</collectingMunicipality>
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amber, housed at the
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<collectingCounty id="62B44E41D029FFAF878DFAE1E442FAE8" box="[183,774,1282,1308]" pageId="6" pageNumber="306">Smithsonian National Museum of Natural History</collectingCounty>
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,
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<collectionCode id="ED7BAE08D029FFAF87B2FAC6E79DFACB" box="[136,217,1317,1343]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="306" type="Museum">USNM</collectionCode>
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<specimenCode id="DBCC9EB6D029FFAF87DAFAC6E634FACB" box="[224,368,1317,1343]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="306" type="Museum">PAL 787826</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD536CDD029FFAF8787FAABE536FA71" blockId="6.[136,778,1247,2077]" pageId="6" pageNumber="306">Syninclusions: Botanical fragments, stellate hairs, small air bubbles and a Diptera (with eggs).</paragraph>
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</subSubSection>
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<subSubSection id="C3706546D029FFAF8787FA6DE62FF9E5" pageId="6" pageNumber="306" type="etymology">
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<paragraph id="8BD536CDD029FFAF8787FA6DE62FF9E5" blockId="6.[136,778,1247,2077]" pageId="6" pageNumber="306">
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<emphasis id="B91EEADFD029FFAF8787FA6DE600FA5C" bold="true" box="[189,324,1422,1448]" pageId="6" pageNumber="306">Etymology.</emphasis>
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Species named in honor of Dr. Paul Johnson, a professor at
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<collectingRegion id="49AEF82FD029FFAF869FFA52E5C3FA3F" box="[421,647,1457,1483]" country="United States of America" name="South Dakota" pageId="6" pageNumber="306">South Dakota State</collectingRegion>
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University who is always happy to provide his invaluable expertise on beetle taxonomy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3706546D029FFAF8787F9FAE388FED2" pageId="6" pageNumber="306" type="diagnosis">
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<paragraph id="8BD536CDD029FFAF8787F9FAE650F94B" blockId="6.[136,778,1247,2077]" pageId="6" pageNumber="306">
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<emphasis id="B91EEADFD029FFAF8787F9FAE67CF9C7" bold="true" box="[189,312,1561,1587]" pageId="6" pageNumber="306">Diagnosis.</emphasis>
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This species belongs to the genus
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<taxonomicName id="4C6A4D4ED029FFAF87B2F9DFE7BEF9A2" authorityName="Latreille" authorityYear="1806" box="[136,250,1596,1622]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="306" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EEADFD029FFAF87B2F9DFE7BEF9A2" box="[136,250,1596,1622]" italics="true" pageId="6" pageNumber="306">Malthinus</emphasis>
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</taxonomicName>
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based on its globular and distally pointed last maxillary palpomere, the short elytra, head restricted behind the eyes and the elytra with impressed punctuation.
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</paragraph>
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<paragraph id="8BD536CDD029FFAF8787F92BE680F7E9" blockId="6.[136,778,1247,2077]" pageId="6" pageNumber="306">
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Only three fossil species of
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<taxonomicName id="4C6A4D4ED029FFAF852EF92BE5C2F916" authorityName="Latreille" authorityYear="1806" box="[532,646,1736,1762]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="306" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EEADFD029FFAF852EF92BE5C2F916" box="[532,646,1736,1762]" italics="true" pageId="6" pageNumber="306">Malthinus</emphasis>
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</taxonomicName>
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have been described so far, all from Baltic amber (
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<bibRefCitation id="EFFB4B3CD029FFAF856AF908E780F8DC" author="Kuska, A. & Kania, I." pageId="6" pageNumber="306" pagination="49 - 56" refId="ref8113" refString="Kuska, A. & Kania, I. (2010) New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa, 2400 (1), 49 - 56. https: // doi. org / 10.11646 / zootaxa. 2400.1.5" type="journal article" year="2010">Kuśka & Kania, 2010</bibRefCitation>
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||
;
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<bibRefCitation id="EFFB4B3CD029FFAF87ECF8EDE54CF8DC" author="Fanti, F. & Damgaard, A. L." box="[214,520,1806,1832]" pageId="6" pageNumber="306" pagination="1 - 32" refId="ref7214" refString="Fanti, F. & Damgaard, A. L. (2018) Fossil soldier beetles from Baltic amber of the Anders Damgaard amber collection (Coleoptera Cantharidae). Baltic Journal of Coleopterology, 18 (1), 1 - 32." type="journal article" year="2018">Fanti & Damgaard, 2018</bibRefCitation>
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||
;
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<bibRefCitation id="EFFB4B3CD029FFAF8521F8EDE781F8BF" author="Pankowski, M. G. & Fanti, F." pageId="6" pageNumber="306" pagination="548 - 556" refId="ref8344" refString="Pankowski, M. G. & Fanti, F. (2022) Two new fossil species of soldier beetles (Coleoptera, Cantharidae, Malthininae) from Baltic amber. Zootaxa, 5165 (4), 548 - 556. https: // doi. org / 10.11646 / zootaxa. 5165.4.6" type="journal article" year="2022">Pankowski & Fanti, 2022</bibRefCitation>
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||
).
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<taxonomicName id="4C6A4D4ED029FFAF87E0F8D2E59CF8BF" authority="Fanti & Damgaard, 2018" authorityName="Fanti & Damgaard" authorityYear="2018" box="[218,728,1841,1867]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="306" phylum="Arthropoda" rank="species" species="rifbjergi">
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<emphasis id="B91EEADFD029FFAF87E0F8D2E6F4F8BF" box="[218,432,1841,1867]" italics="true" pageId="6" pageNumber="306">Malthinus rifbjergi</emphasis>
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<bibRefCitation id="EFFB4B3CD029FFAF8680F8D2E59CF8BF" author="Fanti, F. & Damgaard, A. L." box="[442,728,1841,1867]" pageId="6" pageNumber="306" pagination="1 - 32" refId="ref7214" refString="Fanti, F. & Damgaard, A. L. (2018) Fossil soldier beetles from Baltic amber of the Anders Damgaard amber collection (Coleoptera Cantharidae). Baltic Journal of Coleopterology, 18 (1), 1 - 32." type="journal article" year="2018">Fanti & Damgaard, 2018</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C6A4D4ED029FFAF87B2F8B7E52CF89A" authority="Kuska & Kania, 2010" authorityName="Kuska & Kania" authorityYear="2010" box="[136,616,1876,1902]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="306" phylum="Arthropoda" rank="species" species="danieli">
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<emphasis id="B91EEADFD029FFAF87B2F8B7E611F89A" box="[136,341,1876,1902]" italics="true" pageId="6" pageNumber="306">Malthinus danieli</emphasis>
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<bibRefCitation id="EFFB4B3CD029FFAF865BF8B7E52CF89A" author="Kuska, A. & Kania, I." box="[353,616,1876,1902]" pageId="6" pageNumber="306" pagination="49 - 56" refId="ref8113" refString="Kuska, A. & Kania, I. (2010) New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa, 2400 (1), 49 - 56. https: // doi. org / 10.11646 / zootaxa. 2400.1.5" type="journal article" year="2010">Kuśka & Kania, 2010</bibRefCitation>
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||
</taxonomicName>
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are the most similar to the new species described here, but they have the pronotum only narrower anteriorly (
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<bibRefCitation id="EFFB4B3CD029FFAF8575F879E780F823" author="Kuska, A. & Kania, I." pageId="6" pageNumber="306" pagination="49 - 56" refId="ref8113" refString="Kuska, A. & Kania, I. (2010) New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa, 2400 (1), 49 - 56. https: // doi. org / 10.11646 / zootaxa. 2400.1.5" type="journal article" year="2010">Kuśka & Kania, 2010</bibRefCitation>
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||
;
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<bibRefCitation id="EFFB4B3CD029FFAF87ECF85EE543F823" author="Fanti, F. & Damgaard, A. L." box="[214,519,1981,2007]" pageId="6" pageNumber="306" pagination="1 - 32" refId="ref7214" refString="Fanti, F. & Damgaard, A. L. (2018) Fossil soldier beetles from Baltic amber of the Anders Damgaard amber collection (Coleoptera Cantharidae). Baltic Journal of Coleopterology, 18 (1), 1 - 32." type="journal article" year="2018">Fanti & Damgaard, 2018</bibRefCitation>
|
||
), where in
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<taxonomicName id="4C6A4D4ED029FFAF85A2F85EE65EF80D" authority="PANKOWSKI & Fanti, 2023" authorityName="PANKOWSKI & Fanti" authorityYear="2023" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="306" phylum="Arthropoda" rank="species" species="pauljohnsoni" status="sp. nov.">
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<emphasis id="B91EEADFD029FFAF85A2F85EE65EF80D" italics="true" pageId="6" pageNumber="306">Malthinus pauljohnsoni</emphasis>
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</taxonomicName>
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<emphasis id="B91EEADFD029FFAF861EF803E6C4F80E" bold="true" box="[292,384,2016,2042]" pageId="6" pageNumber="306">
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<taxonomicNameLabel id="A22D57A4D029FFAF861EF803E6C4F80E" box="[292,384,2016,2042]" pageId="6" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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the pronotum is widely expanded before the posterior corners.
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</paragraph>
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<paragraph id="8BD536CDD029FFAF8465FF7DE388FED2" blockId="6.[810,1452,158,2056]" pageId="6" pageNumber="306">
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<emphasis id="B91EEADFD029FFAF8465FF7DE325FF4C" bold="true" box="[863,1121,158,184]" pageId="6" pageNumber="306">Locality and horizon.</emphasis>
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Amber mine in the Yantarny settlement, Sambian Peninsula,
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<collectingRegion id="49AEF82FD029FFAF83FAFF20E2E3FF29" box="[1216,1447,195,221]" country="Russia" name="Kaliningrad" pageId="6" pageNumber="306">Kaliningrad region</collectingRegion>
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,
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<collectingCountry id="F37D765DD029FFAF8410FF0BE43CFEF6" box="[810,888,232,258]" name="Russia" pageId="6" pageNumber="306">Russia</collectingCountry>
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. Middle Eocene (Lutetian) (47.8–41.2 Ma) to late Eocene (Priabonian) (37.8–33.9 Ma).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3706546D029FFAF8465FED2E253F881" pageId="6" pageNumber="306" type="description">
|
||
<paragraph id="8BD536CDD029FFAF8465FED2E36CFE4D" blockId="6.[810,1452,158,2056]" pageId="6" pageNumber="306">
|
||
<emphasis id="B91EEADFD029FFAF8465FED2E4B5FEBF" bold="true" box="[863,1009,305,331]" pageId="6" pageNumber="306">Description.</emphasis>
|
||
Female, defined on the basis of the short antennae and wide and rounded last sternite. Body length: about
|
||
<quantity id="4C929B28D029FFAF84FFFE98E35BFE61" box="[965,1055,379,405]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="6" pageNumber="306" unit="mm" value="4.9">4.9 mm</quantity>
|
||
. Body entirely dark brown without yellow spots on elytra.
|
||
</paragraph>
|
||
<paragraph id="8BD536CDD029FFAF8465FE27E253F881" blockId="6.[810,1452,158,2056]" pageId="6" pageNumber="306">Adult, alate, slender. Head completely exposed, strongly narrowed behind eyes, very strongly wrinkled, with deep and wide punctation, interocular dorsal distance about 2.5–2.6 times greater than eye diameter. Eyes large, convex, rounded, located in the upper lateral part of head. Mandibles falciform, elongate, thin. Maxillary palpi 4- segmented, with last palpomere globular and apically pointed. Labial palpi 3-segmented, with last palpomere globular and apically pointed. Antennae filiform, 11- segmented, short, surpassing half of elytra but not reaching the elytral apex; antennomere I elongate, slightly club-shaped; antennomere II about 2.1 times shorter than antennomere I; antennomeres III–IX subequal, very slightly longer and slender than antennomere II; antennomere X very slightly shorter than previous ones; antennomere XI elongate, very slightly pointed; all antennomeres covered by short setae. Pronotum longer than wide, narrower than head, slightly punctate, equipped with short setae, anterior margin straight and strongly bordered, posterior margin undulate, sides with a wide expansion before the posterior corners.Scutellum wide with truncate apex, enlarged in the middle. Elytra short, revealing three and a half abdominal segments uncovered, wider than pronotum, covered with deep punctation (smaller punctation close to the elytral apex) in rows and some setae, parallel-sided, rounded at apex. Hind wings slightly infuscate, clearly exceeding elytra and slightly exceeding last abdominal segment. Metasternum with straight posterior margin, covered with dispersed and short setae and very shallow punctation. Sternites transverse with shallow punctation. Last tergite and last sternite wide and rounded. Legs slender, pubescent; coxae short, stout; trochanters elongate with rounded apex; femora enlarged, rather straight; tibiae cylindrical and thin, equipped with apical spurs, pro- and mesotibiae shorter than pro- and mesofemora, metatibiae longer than metafemora. Tarsomere I thin, elongate; tarsomere II about 1.8 times shorter than tarsomere I; tarsomere III very short, rounded; tarsomere IV strongly bilobed; tarsomere V elongate, slightly curved, slender; claws simple without tooth. Male unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3706546D029FFAF8465F863E332F7FC" pageId="6" pageNumber="306" type="discussion">
|
||
<paragraph id="8BD536CDD029FFAF8465F863E332F7FC" blockId="6.[810,1452,158,2056]" pageId="6" pageNumber="306">
|
||
<emphasis id="B91EEADFD029FFAF8465F863E497F86E" bold="true" box="[863,979,1920,1946]" pageId="6" pageNumber="306">Remarks.</emphasis>
|
||
The yellow amber piece measures approximately 18 × 13 ×
|
||
<quantity id="4C929B28D029FFAF834CF846E396F84B" box="[1142,1234,1957,1983]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="306" unit="mm" value="10.0">10 mm</quantity>
|
||
. The inclusion is complete. The last tergites and ventral part of the head are covered with white emulsion.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |