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<document id="11BA92659D78685349A3D7518E4EE4B2" ID-DOI="10.11646/zootaxa.4098.1.1" ID-GBIF-Dataset="61bbe4a2-0c80-4fa7-915e-46cba4581f00" ID-ISSN="1175-5326" ID-Zenodo-Dep="271096" ID-ZooBank="6A79B596-26E0-454B-8830-D69FFCBC4684" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459847611809" checkinUser="plazi" docAuthor="Lonsdale, O. &amp; Marshall, S. A." docDate="2016" docId="03C387D68A23225FEA846A00ED82FB48" docLanguage="en" docName="zootaxa.4098.1.1.pdf.imf" docOrigin="Zootaxa 4098 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nothybus sumatranus Enderlein" docType="treatment" docVersion="12" lastPageNumber="31" masterDocId="FFFAFFAE8A382241EA13686EE977FF8D" masterDocTitle="Revision of the family Nothybidae (Diptera: Schizophora)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="28" updateTime="1698224978163" updateUser="plazi">
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<mods:title id="60141B2573A830EB576A3A2E9EE0FF51">Revision of the family Nothybidae (Diptera: Schizophora)</mods:title>
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<mods:namePart id="D9A27992726BCC01C073649B0C2A0EC0">Lonsdale, O.</mods:namePart>
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<mods:namePart id="710B5088F8715BE62ADD68C0FD598898">Marshall, S. A.</mods:namePart>
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<mods:date id="036DCD48FB59740E42F3CD7D5AD3FA53">2016</mods:date>
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<treatment id="03C387D68A23225FEA846A00ED82FB48" ID-DOI="http://doi.org/10.5281/zenodo.6053704" ID-GBIF-Taxon="119369851" ID-Zenodo-Dep="6053704" LSID="urn:lsid:plazi:treatment:03C387D68A23225FEA846A00ED82FB48" httpUri="http://treatment.plazi.org/id/03C387D68A23225FEA846A00ED82FB48" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28">
<subSubSection id="C370654B8A23225AEA846A00EB36FD27" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BD536C08A23225AEA846A00EB53FD05" blockId="27.[151,577,622,682]" box="[151,548,622,648]" pageId="27" pageNumber="28">
<heading id="D09D81AC8A23225AEA846A00EB53FD05" bold="true" box="[151,548,622,648]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<taxonomicName id="4C6A4D438A23225AEA846A00EB53FD05" ID-CoL="74C5G" authority="Enderlein" authorityName="Enderlein" box="[151,548,622,648]" class="Insecta" family="Nothybidae" genus="Nothybus" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="B91EEAD28A23225AEA846A00EB53FD05" bold="true" box="[151,548,622,648]" pageId="27" pageNumber="28">
<emphasis id="B91EEAD28A23225AEA846A00E8D6FD05" bold="true" box="[151,417,622,648]" italics="true" pageId="27" pageNumber="28">Nothybus sumatranus</emphasis>
Enderlein
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD536C08A23225AEA846AFFEB36FD27" blockId="27.[151,577,622,682]" box="[151,577,657,682]" pageId="27" pageNumber="28">Figures 810, 5563, 77, 78, 114120</paragraph>
</subSubSection>
<subSubSection id="C370654B8A23225AEA846AB7E8A9FC83" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8BD536C08A23225AEA846AB7E8A9FC83" blockId="27.[151,1436,729,782]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB7A11EE8A23225AEA846AB7E8A9FC83" pageId="27" pageNumber="28">
<taxonomicName id="4C6A4D438A23225AEA846AB7EB3EFD62" ID-CoL="74C5G" authority="Enderlein 1922: 176" authorityName="Enderlein" authorityPageNumber="176" authorityYear="1922" box="[151,585,729,751]" class="Insecta" family="Nothybidae" genus="Nothybus" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="B91EEAD28A23225AEA846AB7E805FD62" box="[151,370,729,751]" italics="true" pageId="27" pageNumber="28">Nothybus sumatranus</emphasis>
<treatmentCitation id="0ACB10D18A23225AEB6B6AB7EB3EFD62" author="Enderlein" box="[376,585,729,751]" page="176" pageId="27" pageNumber="28" year="1922">
<bibRefCitation id="EFFB4B318A23225AEB6B6AB7EB61FD62" author="Enderlein" box="[376,534,729,751]" pageId="27" pageNumber="33" refString="Enderlein, G. (1922) Klassifikation der Micropeziden. Archiv fur Naturgeschichte, (A), 88, 140 - 229." type="journal article" year="1922">Enderlein 1922</bibRefCitation>
: 176
</treatmentCitation>
</taxonomicName>
.
<treatmentCitation id="0ACB10D18A23225AE8416AB7EB9BFD62" author="Aczel" box="[594,748,729,751]" page="10" pageId="27" pageNumber="28" year="1955">
<bibRefCitation id="EFFB4B318A23225AE8416AB7EBBEFD62" author="Aczel" box="[594,713,729,751]" pageId="27" pageNumber="32" refString="Aczel, M. L. (1955) Nothybidae, a new family of Diptera. Treubia, 23, 1 - 18." type="journal article" year="1955">Aczél 1955</bibRefCitation>
: 10
</treatmentCitation>
[family revision, key];
<treatmentCitation id="0ACB10D18A23225AE9CB6AB7EDFBFD62" author="Steyskal" box="[984,1164,729,751]" page="31" pageId="27" pageNumber="28" year="1977">
<bibRefCitation id="EFFB4B318A23225AE9CB6AB7ED1EFD62" author="Steyskal" box="[984,1129,729,751]" pageId="27" pageNumber="33" refString="Steyskal, G. C. (1977) Family Nothybidae. In: Delfinado, M. C. &amp; Hardy, D. E. (Eds.), A Catalog of the Diptera of the Oriental Region 3: Suborder Cyclorrhapha (Excluding Division Aschiza). University Press of Hawaii, Honolulu, pp. 30 - 31" type="journal article" year="1977">Steyskal 1977</bibRefCitation>
: 31
</treatmentCitation>
[Oriental catalogue];
<treatmentCitation id="0ACB10D18A23225AEF7A6AB4E811FC83" author="Papp" page="176" pageId="27" pageNumber="28" year="2006">
<bibRefCitation id="EFFB4B318A23225AEF7A6AB4E841FC83" author="Papp" pageId="27" pageNumber="33" refString="Papp, L., Merz, B. &amp; Foldvari, M. (2006) Diptera of Thailand: A summary of the families and genera with references to the species representations. Acta Zoologica Academiae Scientarum Hungaricae, 52 (2), 97 - 269." type="journal article" year="2006">
Papp
<emphasis id="B91EEAD28A23225AEAD46A97E98DFC83" box="[199,250,760,782]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2006
</bibRefCitation>
: 176
</treatmentCitation>
[
<collectingCountry id="F37D76508A23225AEB616A96E8A3FC83" box="[370,468,760,782]" name="Thailand" pageId="27" pageNumber="28">Thailand</collectingCountry>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C370654B8A23225CEA846B54E86BF96B" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="8BD536C08A23225AEA846B54E8F5FCFA" blockId="27.[151,1437,826,2014]" pageId="27" pageNumber="28">
<emphasis id="B91EEAD28A23225AEA846B54E856FCDE" bold="true" box="[151,289,826,851]" pageId="27" pageNumber="28">Description</emphasis>
. Sumatra and Pahang morphotype, including
<typeStatus id="54D188628A23225AE93A6B55EA2EFCDE" box="[809,857,827,851]" pageId="27" pageNumber="28">type</typeStatus>
female and 2 non-type males (also see “Variation” below). (
<figureCitation id="13512A458A23225AEAE96B30E80DFCFA" box="[250,378,862,887]" captionStart="FIGURES 55 57" captionStartId="28.[151,264,1675,1698]" captionTargetBox="[151,1436,193,1654]" captionTargetId="figure@28.[151,1436,193,1655]" captionTargetPageId="28" captionText="FIGURES 55 57. Nothybus sumatranus Enderlein, female syntype, photos; 55: left lateral; 56: head, anterior; 57: head and thorax, dorsal. FIGURES 58 60: N. sumatranus, male, Selangor, photos; 58: left lateral; 59: head, anterior; 60: head and thorax, dorsal. FIGURES 61 63: N. sumatranus, male, Thailand, photos; 61: left lateral; 62: head, anterior; 63: head and thorax, dorsal. FIGURES 64 66: N. triguttatus Bezzi male, Selangor, photos; 64: head and thorax, dorsal; 65: left lateral; 66: head, anterior." httpUri="https://zenodo.org/record/271106/files/figure.png" pageId="27" pageNumber="28">Figs 5557</figureCitation>
)
</paragraph>
<paragraph id="8BD536C08A23225AEAD46BECEA67FBA7" blockId="27.[151,1437,826,2014]" pageId="27" pageNumber="28">
<emphasis id="B91EEAD28A23225AEAD46BECE873FC16" box="[199,260,898,923]" italics="true" pageId="27" pageNumber="28">Head</emphasis>
: Mostly reddish brown with yellowish-orange areas; distal 2/3 of first flagellomere black; frons with anterolateral velvety patches confluent with anterior margin and dark parafacial; ocellar tubercle black; ventral half of face silvery tomentose with glossy medial tubercle dark brown in female and dark yellow in males; midpoint of face sometimes with horizontal brown line evident above silvery region; back of head with transverse band of faint whitish pruinosity above one pair of brown lateral spots.
</paragraph>
<paragraph id="8BD536C08A23225AEAD46C58EB27F9AE" blockId="27.[151,1437,826,2014]" pageId="27" pageNumber="28">
<emphasis id="B91EEAD28A23225AEAD46C58E86FFBC2" box="[199,280,1078,1103]" italics="true" pageId="27" pageNumber="28">Thorax</emphasis>
: Yellowish-orange in base colour with dorsum of thorax darker. Scutum with medial brown stripe reaching anterior margin; stripe bounded by medial 6 rows of setulae anteriorly, and becoming wider postsuturally to nearly reach posterior supra-alar; stripe sometimes confluent with one pair of darker, narrow postsutural stripes originating at dorsocentral seta; one male with darker line along dorsocentral row; AMSA Pahang specimens with lateral margins of stripe expanded to fuse with lateral brown mottling, or with pigment slightly paler overall on thorax and with medial scutal stripe ending shortly before anterior margin (similar to Selangor specimens). Presutural scutum with brown mottling laterally, especially posterolaterally. Dorsal half of postpronotum brown with posterior region iridescent. Notopleuron brown (pigment sometimes slightly irregular) and with posterolateral iridescence, adjoining yellow ovate region on scutum usually bordered with similar irregular brown pigment; tubercle yellow, at least centrally. Scutellum dark brown with margin paler (more so towards venter), apex between and around apical setae orange; subscutellum brown. Metanotum with slight brownish tint, katatergite brown posteriorly or mostly yellow. Proepisternum dirty orange to brown laterally; anterior lobe with distal margin dark brown. Anepisternum brownish and iridescent posterodorsally. Posterodorsal corner of katepisternum and posterior region of anepisternum whitish-yellow.
</paragraph>
<paragraph id="8BD536C08A23225AEAD46E5EEC0AF96D" blockId="27.[151,1437,826,2014]" pageId="27" pageNumber="28">
<emphasis id="B91EEAD28A23225AEAD46E5EE988F9CA" box="[199,255,1584,1607]" italics="true" pageId="27" pageNumber="28">Wing</emphasis>
: Wing length 4.87.0mm. Similar overall to
<taxonomicName id="4C6A4D438A23225AE8ED6E5EEAE2F9CA" box="[766,917,1584,1607]" class="Insecta" family="Nothybidae" genus="Nothybus" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="triguttatus">
<emphasis id="B91EEAD28A23225AE8ED6E5EEAE2F9CA" box="[766,917,1584,1607]" italics="true" pageId="27" pageNumber="28">N. triguttatus</emphasis>
</taxonomicName>
but with pigment paler. Apex with infuscated area enclosing three clear iridescent spots, with second spot in cell r4+5 displaced apically. Transverse brown stripe narrow (more so to CuA1), barely extending into female cell r1 (
<figureCitation id="13512A458A23225AE9716E17EAC3F91F" box="[866,948,1657,1682]" captionStart="FIGURES 67 78" captionStartId="33.[151,264,1695,1718]" captionTargetBox="[151,1428,205,1674]" captionTargetId="figure@33.[151,1436,193,1674]" captionTargetPageId="33" captionText="FIGURES 67 78. wings, photos; 67: Nothybus longicollis (Walker), with relevant veins listed; 68: N. biguttatus Wulp; 69: N. kempi (Brunetti), with relevant cells listed; 70: N. procerus spec. nov., holotype; 71: N. kuznetsovorum Galinskaya &amp; Shatalkin; 72: N. cataractus spec. nov.; 73: N. lineifer Enderlein, syntype; 74: N. decorus Meijere [syn. nov. of N. lineifer], syntype; 75: N. lineifer, CNC specimen; 76: N. triguttatus Bezzi; 77: N. sumatranus Enderlein, syntype; 78: N. sumatranus, Thai specimen." httpUri="https://zenodo.org/record/271107/files/figure.png" pageId="27" pageNumber="28">Fig. 77</figureCitation>
) and filling most of apex of male cell r1 (as in
<figureCitation id="13512A458A23225AEAA66ECEE87CF934" box="[181,267,1696,1721]" captionStart="FIGURES 67 78" captionStartId="33.[151,264,1695,1718]" captionTargetBox="[151,1428,205,1674]" captionTargetId="figure@33.[151,1436,193,1674]" captionTargetPageId="33" captionText="FIGURES 67 78. wings, photos; 67: Nothybus longicollis (Walker), with relevant veins listed; 68: N. biguttatus Wulp; 69: N. kempi (Brunetti), with relevant cells listed; 70: N. procerus spec. nov., holotype; 71: N. kuznetsovorum Galinskaya &amp; Shatalkin; 72: N. cataractus spec. nov.; 73: N. lineifer Enderlein, syntype; 74: N. decorus Meijere [syn. nov. of N. lineifer], syntype; 75: N. lineifer, CNC specimen; 76: N. triguttatus Bezzi; 77: N. sumatranus Enderlein, syntype; 78: N. sumatranus, Thai specimen." httpUri="https://zenodo.org/record/271107/files/figure.png" pageId="27" pageNumber="28">Fig. 78</figureCitation>
). Wing with faint, ill-defined brown spots apically on R4+5 and M1 (leaving at most a thin clear border along margin of veins). Subapical cloud on CuA1 small and indistinct. Vein r-m with narrow medial infuscation.
</paragraph>
<paragraph id="8BD536C08A23225AEAD46E81EA3AF8C3" blockId="27.[151,1437,826,2014]" pageId="27" pageNumber="28">
<emphasis id="B91EEAD28A23225AEAD46E81E98BF88B" box="[199,252,1775,1798]" italics="true" pageId="27" pageNumber="28">Legs</emphasis>
: Coxae and femora light yellow, apices of femora dark brown (with pigment slightly extended along inner surface of fore femur); tibiae and tarsi dark brown except for fore basitarsomere, which is white with base narrowly orange to blackish ventrally (sometimes only setae darkened).
</paragraph>
<paragraph id="8BD536C08A23225AEAD46F34E845F853" blockId="27.[151,1437,826,2014]" pageId="27" pageNumber="28">
<emphasis id="B91EEAD28A23225AEAD46F34E846F8FE" box="[199,305,1882,1907]" italics="true" pageId="27" pageNumber="28">Abdomen</emphasis>
: Tergites 24 dark brown except for anterolateral corner of tergite 2; male tergite 4 with broad orange-tinted region sublaterally. Tergite 5 black and velvety. Tergite 6 brown to brownish, darker laterally and posterolaterally. Some Pahang males with tergite 4 apparently entirely dark brown (possibly due to poor preservation).
</paragraph>
<caption id="DF1566488A24225DEA846EE5E85FF890" httpUri="https://zenodo.org/record/271106/files/figure.png" pageId="28" pageNumber="29" targetBox="[151,1436,193,1654]" targetPageId="28">
<paragraph id="8BD536C08A24225DEA846EE5E85FF890" blockId="28.[151,1436,1675,1821]" pageId="28" pageNumber="29">
<emphasis id="B91EEAD28A24225DEA846EE5E821F92C" bold="true" box="[151,342,1675,1698]" pageId="28" pageNumber="29">FIGURES 5557.</emphasis>
<taxonomicName id="4C6A4D438A24225DEB4E6EE2EBDFF92F" authority="Enderlein" authorityName="Enderlein" box="[349,680,1676,1698]" class="Insecta" family="Nothybidae" genus="Nothybus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="B91EEAD28A24225DEB4E6EE2EB4CF92F" box="[349,571,1676,1698]" italics="true" pageId="28" pageNumber="29">Nothybus sumatranus</emphasis>
Enderlein
</taxonomicName>
, female syntype, photos; 55: left lateral; 56: head, anterior; 57: head and thorax, dorsal.
<emphasis id="B91EEAD28A24225DEB266EC4E880F94D" bold="true" box="[309,503,1706,1729]" pageId="28" pageNumber="29">FIGURES 5860:</emphasis>
<taxonomicName id="4C6A4D438A24225DE8126EC2EBEFF94C" box="[513,664,1708,1729]" class="Insecta" family="Nothybidae" genus="Nothybus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="B91EEAD28A24225DE8126EC2EBEFF94C" box="[513,664,1708,1729]" italics="true" pageId="28" pageNumber="29">N. sumatranus</emphasis>
</taxonomicName>
, male, Selangor, photos; 58: left lateral; 59: head, anterior; 60: head and thorax, dorsal.
<emphasis id="B91EEAD28A24225DEB266EA7E880F953" bold="true" box="[309,503,1737,1760]" pageId="28" pageNumber="29">FIGURES 6163:</emphasis>
<taxonomicName id="4C6A4D438A24225DE8136EA4EBE0F952" box="[512,663,1738,1759]" class="Insecta" family="Nothybidae" genus="Nothybus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="B91EEAD28A24225DE8136EA4EBE0F952" box="[512,663,1738,1759]" italics="true" pageId="28" pageNumber="29">N. sumatranus</emphasis>
</taxonomicName>
, male, Thailand, photos; 61: left lateral; 62: head, anterior; 63: head and thorax, dorsal.
<emphasis id="B91EEAD28A24225DEB236E89E898F970" bold="true" box="[304,495,1767,1790]" pageId="28" pageNumber="29">FIGURES 6466:</emphasis>
<taxonomicName id="4C6A4D438A24225DEBE56E87EBC9F973" authority="Bezzi" authorityName="Bezzi" box="[502,702,1768,1790]" class="Insecta" family="Nothybidae" genus="Nothybus" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="triguttatus">
<emphasis id="B91EEAD28A24225DEBE56E87EB0AF973" box="[502,637,1769,1790]" italics="true" pageId="28" pageNumber="29">N. triguttatus</emphasis>
Bezzi
</taxonomicName>
male, Selangor, photos; 64: head and thorax, dorsal; 65: left lateral; 66: head, anterior.
</paragraph>
</caption>
<paragraph id="8BD536C08A24225CEAD46F27ED64FE20" blockId="28.[151,1436,1865,2034]" lastBlockId="29.[151,1437,151,2021]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="B91EEAD28A24225DEAD46F27E80BF8EF" box="[199,380,1865,1890]" italics="true" pageId="28" pageNumber="29">Male terminalia</emphasis>
: (
<figureCitation id="13512A458A24225DEB876F27EB47F8EF" box="[404,560,1865,1890]" captionStart="FIGURES 114 120" captionStartId="40.[151,264,1915,1938]" captionTargetBox="[188,1428,210,1883]" captionTargetId="figure@40.[151,1436,193,1894]" captionTargetPageId="40" captionText="FIGURES 114 120. Nothybus sumatranus Enderlein, male abdomen; 114: external components, segment 6 to cercus, left lateral; 115: external components past sternite 8, posterior; 116: external components, segment 6 to cercus, ventral; 117: external components past sternite 8, anterior; 118: ejaculatory apodeme; 119: internal components, ventral; 120: same, left lateral." httpUri="https://zenodo.org/record/271114/files/figure.png" pageId="28" pageNumber="29">Figs 114120</figureCitation>
) Tergite 6 band-like with several setae along posterolateral margin. Sternite 6 longer than wide, subtriangular with rounded anterior margin, setose on distal half with several longer marginal setae. Sternite 7 not evident in dissected material. Sternite 8 large and dome-like with several posterolateral setae marginally. Epandrium setose, fused to posterior half of surstylus. Surstylus approximately as high as epandrium, angled anteriorly but with slight posterior curve, minute setulae mostly clustered on distal half and inner-distal margin beside cluster of tubercle-like setae, inner surface confluent with subepandrial sclerite. Cercus flat, fingerlike. Hypandrium broadly fused to phallapodeme, setose medially and on narrow, prominent process with one longer apical seta; anteromedial margin broadly rounded, ventral lobe with ventromedial swelling with numerous setae anteriorly. Pregonite with slightly widened base adjoining hypandrium, outer surface minutely spinulose and textured, and with clear, gradually tapered apex; with one long anteroventral seta and several minute ventrolateral setae. Basiphallus triangular, sclerotized laterally, terminating in narrow, clear, sac-like process. Distiphallus long and unusually broad with one pair of thick ribbon-like bands divided on distal 1/3 into two pairs of narrower bands (apically fused into W-shaped structure with medial process elongate and bent).
</paragraph>
<paragraph id="8BD536C08A25225CEAD469D7E8EFFD25" blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
<emphasis id="B91EEAD28A25225CEAD469D7E8C7FE5D" box="[199,432,439,464]" italics="true" pageId="29" pageNumber="30">Variation—Selangor</emphasis>
: (Figs 5860) Scutum with medial stripe not reaching anterior margin, with stripe either atrophied or broken subapically, leaving a floating spot anteriorly; stripe usually at least partially confluent with one pair of darker, narrow postsutural stripes originating at dorsocentral seta; notal pigmentation difficult to discern in BMNH specimens due to state of preservation. Metanotum yellowish-orange with brown mottling. Anepisternum with brownish mottling, especially anteriorly. Wing pigment strong, as in
<figureCitation id="13512A458A25225CEE846A29ED9BFDED" box="[1175,1260,583,608]" captionStart="FIGURES 67 78" captionStartId="33.[151,264,1695,1718]" captionTargetBox="[151,1428,205,1674]" captionTargetId="figure@33.[151,1436,193,1674]" captionTargetPageId="33" captionText="FIGURES 67 78. wings, photos; 67: Nothybus longicollis (Walker), with relevant veins listed; 68: N. biguttatus Wulp; 69: N. kempi (Brunetti), with relevant cells listed; 70: N. procerus spec. nov., holotype; 71: N. kuznetsovorum Galinskaya &amp; Shatalkin; 72: N. cataractus spec. nov.; 73: N. lineifer Enderlein, syntype; 74: N. decorus Meijere [syn. nov. of N. lineifer], syntype; 75: N. lineifer, CNC specimen; 76: N. triguttatus Bezzi; 77: N. sumatranus Enderlein, syntype; 78: N. sumatranus, Thai specimen." httpUri="https://zenodo.org/record/271107/files/figure.png" pageId="29" pageNumber="30">Fig. 78</figureCitation>
. Tergite 4 with orange sublateral spots sometimes atrophied (mostly black); female tergite 6 brown along anterior, lateral and posterolateral margins.
</paragraph>
<paragraph id="8BD536C08A25225CEAD46ADBE990FBDA" blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
<emphasis id="B91EEAD28A25225CEAD46ADBE8D3FD41" box="[199,420,693,716]" italics="true" pageId="29" pageNumber="30">
Variation—
<collectingCountry id="F37D76508A25225CEB586ADBE8D3FD41" box="[331,420,693,716]" name="Vietnam" pageId="29" pageNumber="30">Vietnam</collectingCountry>
</emphasis>
: Velvety anterolateral patch on frons sometimes narrowly separated from anterior margin; ocellar spot often extending anteriorly as narrow stripe that is longer in females; parafacial sometimes only brownish medially, along eye margin, or entirely yellow. Some Vietnamese specimens with scutal pattern as for Sumatra specimens (notal stripe wider, unbroken and reaching anterior margin), but some with presutural section of medial stripe fading before margin, or stripe faded anteriorly and near transverse suture to leave darker floating presutural spot (as in Selangor material); medial stripe slightly to strongly fused to posterolateral stripe that is itself faded to darkly pigmented; anepisternum with more extensive brownish mottling, especially anteriorly (similar to Selangor material), sometimes covering entire sclerite. Scutellum sometimes more extensively black along outer margins. Tergite 3 and sometimes also 2 sometimes orange sublaterally; paired orange regions on tergite 4 large, sometimes confluent, sometimes enveloping sclerite excluding posterior margin. Wing pigment strong, as in
<figureCitation id="13512A458A25225CEF636B99E9C1FBB9" captionStart="FIGURES 67 78" captionStartId="33.[151,264,1695,1718]" captionTargetBox="[151,1428,205,1674]" captionTargetId="figure@33.[151,1436,193,1674]" captionTargetPageId="33" captionText="FIGURES 67 78. wings, photos; 67: Nothybus longicollis (Walker), with relevant veins listed; 68: N. biguttatus Wulp; 69: N. kempi (Brunetti), with relevant cells listed; 70: N. procerus spec. nov., holotype; 71: N. kuznetsovorum Galinskaya &amp; Shatalkin; 72: N. cataractus spec. nov.; 73: N. lineifer Enderlein, syntype; 74: N. decorus Meijere [syn. nov. of N. lineifer], syntype; 75: N. lineifer, CNC specimen; 76: N. triguttatus Bezzi; 77: N. sumatranus Enderlein, syntype; 78: N. sumatranus, Thai specimen." httpUri="https://zenodo.org/record/271107/files/figure.png" pageId="29" pageNumber="30">Fig. 78</figureCitation>
. One female with tergite 6 brown and transverse stripe on wing broader anteriorly, expanding to surround apex of R2+3.
</paragraph>
<paragraph id="8BD536C08A25225CEAD46C06EC3FFA67" blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
<emphasis id="B91EEAD28A25225CEAD46C06E8B0FBF2" box="[199,455,1126,1151]" italics="true" pageId="29" pageNumber="30">
Variation—
<collectingCountry id="F37D76508A25225CEB586C08E808FBF2" box="[331,383,1126,1151]" name="Thailand" pageId="29" pageNumber="30">Thai</collectingCountry>
males
</emphasis>
: (Figs 6163) Velvety anterolateral patch on frons sometimes narrowly separated from anterior margin of frons; facial tubercle light brown or with slightly brownish tint; sometimes light brown to dark yellow. Frons with one pair of small white pruinescent patches anterior to posterior fronto-orbital. Thoracic and wing pattern sometimes as described for Selangor specimens, but sometimes paler or with medial scutal stripe strongly narrowed to midpoint with presutural section absent, and with brownish apical spots on R4+5 and M1 darker. Subscutellum and lateral surface of scutellum paler; scutellum sometimes with dorsal margin yellowishorange and venter of subscutellum yellow. Anepisternum with light whitish-grey pruinosity and with brownish mottling dorsally and anteromedially. Only dorsal margin of proepisternum brownish. Base of fore basitarsomere very narrowly brown. Abdomen with orange spots on tergite 4 larger, sometimes partially confluent, at least anteriorly, or widely confluent with only tergite margins brown (as in most
<collectingCountry id="F37D76508A25225CE9FF6DC3ED68FA4B" box="[1004,1055,1453,1478]" name="Thailand" pageId="29" pageNumber="30">Thai</collectingCountry>
females). Terminalia of dissected
<collectingCountry id="F37D76508A25225CEA846DBFE9BCFA67" box="[151,203,1489,1514]" name="Thailand" pageId="29" pageNumber="30">Thai</collectingCountry>
male as above except apex of distiphallus more slender with margins slightly converging, not parallel.
</paragraph>
<paragraph id="8BD536C08A25225CEAD46D98E86BF96B" blockId="29.[151,1437,151,2021]" pageId="29" pageNumber="30">
<emphasis id="B91EEAD28A25225CEAD46D98E8AAF983" box="[199,477,1525,1550]" italics="true" pageId="29" pageNumber="30">
Variation—
<collectingCountry id="F37D76508A25225CEB586D9BE808F983" box="[331,383,1525,1550]" name="Thailand" pageId="29" pageNumber="30">Thai</collectingCountry>
females
</emphasis>
: As for
<collectingCountry id="F37D76508A25225CE82E6D9BEB07F983" box="[573,624,1525,1550]" name="Thailand" pageId="29" pageNumber="30">Thai</collectingCountry>
males except as follows. Face with very faint brown medial transverse line and glossy ventral tubercle brown to dark brown. Notal pattern sometimes faded, being reduced to smaller stripes and spots, including one large spot on postpronotum and adjacent spot on scutum; remnant of medial stripe often strong presuturally. Anepisternum with markings restricted to posterodorsal margin and one pale vertical anteromedial stripe. Fore basitarsomere with brown basal spot reduced to small vestige ventrally. Tergite 4 with dark pigment reduced, sometimes with only margins dark brown. Tergite 6 with lateral margin and anteromedial spot brown.
</paragraph>
</subSubSection>
<subSubSection id="C370654B8A25225CEAD46E9FEDF2F8A3" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BD536C08A25225CEAD46E9FED85F887" blockId="29.[151,1437,151,2021]" box="[199,1266,1777,1802]" pageId="29" pageNumber="30">
<emphasis id="B91EEAD28A25225CEAD46E9FE817F887" bold="true" box="[199,352,1777,1802]" pageId="29" pageNumber="30">Distribution.</emphasis>
<collectingCountry id="F37D76508A25225CEB746E9FE8A3F887" box="[359,468,1777,1802]" name="Indonesia" pageId="29" pageNumber="30">Indonesia</collectingCountry>
(Sumatra),
<collectingCountry id="F37D76508A25225CE84A6E9FEBC8F887" box="[601,703,1777,1802]" name="Malaysia" pageId="29" pageNumber="30">Malaysia</collectingCountry>
(Pahang,
<collectingCountry id="F37D76508A25225CE9236E9FEAE4F887" box="[816,915,1777,1802]" name="Malaysia" pageId="29" pageNumber="30">Sarawak</collectingCountry>
, Selangor),
<collectingCountry id="F37D76508A25225CEE0A6E9FED08F887" box="[1049,1151,1777,1802]" name="Thailand" pageId="29" pageNumber="30">Thailand</collectingCountry>
,
<collectingCountry id="F37D76508A25225CEE986E9FED99F887" box="[1163,1262,1777,1802]" name="Vietnam" pageId="29" pageNumber="30">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8BD536C08A25225CEAD46F7BEDF2F8A3" blockId="29.[151,1437,151,2021]" box="[199,1157,1813,1838]" pageId="29" pageNumber="30">
<emphasis id="B91EEAD28A25225CEAD46F7BE8ACF8A3" bold="true" box="[199,475,1813,1838]" pageId="29" pageNumber="30">
<typeStatus id="54D188628A25225CEAD46F7BE85EF8A3" box="[199,297,1813,1838]" pageId="29" pageNumber="30" type="syntype">Syntype</typeStatus>
.
<collectingCountry id="F37D76508A25225CEB266F7BE8A1F8A3" box="[309,470,1813,1838]" name="Indonesia" pageId="29" pageNumber="30">INDONESIA</collectingCountry>
.
</emphasis>
Sumatra: N.O. Sumatra, Deli, Dr. Martin S.G. (1♀, ZMHU).
</paragraph>
</subSubSection>
<subSubSection id="C370654B8A25225FEA846F57EB73FC61" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BD536C08A25225FEA846F57EB73FC61" blockId="29.[151,1437,151,2021]" lastBlockId="30.[151,1437,151,1221]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<emphasis id="B91EEAD28A25225CEA846F57EBC3F8DF" bold="true" box="[151,692,1849,1874]" pageId="29" pageNumber="30">
Additional material examined.
<collectingCountry id="F37D76508A25225CE81D6F57EBD8F8DF" box="[526,687,1849,1874]" name="Indonesia" pageId="29" pageNumber="30">INDONESIA</collectingCountry>
.
</emphasis>
Sumatra: Sibolangit, forest,
<date id="FFD410008A25225CE9EA6F57ED11F8DF" box="[1017,1126,1849,1874]" pageId="29" pageNumber="30" value="1985-06-03">3.vi.1985</date>
, J.W. Ismay, BM 1986-283 (
<specimenCount id="9D6CFD498A25225CEAB36F30E9B2F8FB" box="[160,197,1886,1910]" pageId="29" pageNumber="30" type="generic">2♂</specimenCount>
,
<collectionCode id="ED7BAE058A25225CEACA6F30E846F8FB" box="[217,305,1886,1910]" pageId="29" pageNumber="30">BMNH</collectionCode>
),
<date id="FFD410008A25225CEB5F6F33E8B3F8FB" box="[332,452,1885,1910]" pageId="29" pageNumber="30" value="1983-11-12">12.xi.1983</date>
(
<specimenCount id="9D6CFD498A25225CEBCE6F30EB76F8FB" box="[477,513,1886,1910]" pageId="29" pageNumber="30" type="generic">1♂</specimenCount>
,
<collectionCode id="ED7BAE058A25225CE8066F30EB19F8FB" box="[533,622,1886,1910]" pageId="29" pageNumber="30">BMNH</collectionCode>
), (
<specimenCount id="9D6CFD498A25225CE8816F30EBC1F8FB" box="[658,694,1886,1910]" pageId="29" pageNumber="30" type="generic">1♂</specimenCount>
,
<collectionCode id="ED7BAE058A25225CE8D96F30EA54F8FB" box="[714,803,1886,1910]" pageId="29" pageNumber="30">BMNH</collectionCode>
).
<emphasis id="B91EEAD28A25225CE92F6F33EAADF8FB" bold="true" box="[828,986,1885,1910]" pageId="29" pageNumber="30">
<collectingCountry id="F37D76508A25225CE92F6F33EAA3F8FB" box="[828,980,1885,1910]" name="Malaysia" pageId="29" pageNumber="30">MALAYSIA</collectingCountry>
.
</emphasis>
<collectingCountry id="F37D76508A25225CE9FB6F33ED48F8FB" box="[1000,1087,1885,1910]" name="Malaysia" pageId="29" pageNumber="30">Malaya</collectingCountry>
, Pahang: Gua Che Yatim to Terenggan,
<date id="FFD410008A25225CEB096FEFE8EBF817" box="[282,412,1921,1946]" pageId="29" pageNumber="30" value="1958-12-17">17.xii.1958</date>
, L.W. Quate (
<specimenCount id="9D6CFD498A25225CE8246FECEB2BF817" box="[567,604,1922,1946]" pageId="29" pageNumber="30" type="generic">2♂</specimenCount>
,
<collectionCode id="ED7BAE058A25225CE8746FECEBCEF817" LSID="urn:lsid:biocol.org:col:34697" box="[615,697,1922,1946]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="29" pageNumber="30">BPBM</collectionCode>
), Pahang: W.
<collectingCountry id="F37D76508A25225CE9416FEFEACCF817" box="[850,955,1921,1946]" name="Malaysia" pageId="29" pageNumber="30">Malaysia</collectingCountry>
, Old Bentong Pass nr Gombak,
<date id="FFD410008A25225CEF0C6FEFECE0F817" box="[1311,1431,1921,1946]" pageId="29" pageNumber="30" value="1985-04-16">16.iv.1985</date>
, D.K. McAlpine (
<specimenCount id="9D6CFD498A25225CEB456FC7E80DF84C" box="[342,378,1961,1985]" pageId="29" pageNumber="30" type="generic">1♂</specimenCount>
,
<collectionCode id="ED7BAE058A25225CEB956FC7E8ACF84C" box="[390,475,1961,1985]" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="29" pageNumber="30">AMSA</collectionCode>
),
<collectingCountry id="F37D76508A25225CEBFD6FC6EB24F84C" box="[494,595,1960,1985]" name="Malaysia" pageId="29" pageNumber="30">Sarawak</collectingCountry>
: 4th Div, R.G.S. Mulu Expedition, BM 1977-543, forest path to camp 1,
<date id="FFD410008A25225CEF616FC7E9C1F868" pageId="29" pageNumber="30" value="1977-09-19" valueMax="1977-09-22" valueMin="1977-09-19">19 22</date>
.ix.1977, D. Hollis (2♀,
<collectionCode id="ED7BAE058A25225CEBC16FA3EB5DF868" box="[466,554,1997,2021]" pageId="29" pageNumber="30">BMNH</collectionCode>
), Selangor:
<collectingCountry id="F37D76508A25225CE8A56FA2EA7AF868" box="[694,781,1996,2021]" name="Malaysia" pageId="29" pageNumber="30">Malaya</collectingCountry>
, Gombak Field Stn.,
<date id="FFD410008A25225CEE156FA2EDF6F868" box="[1030,1153,1996,2021]" pageId="29" pageNumber="30" value="1977-11-14">14.xi.1977</date>
, B. Bendell (
<specimenCount id="9D6CFD498A25225CEF0C6FA3EC34F868" box="[1311,1347,1997,2021]" pageId="29" pageNumber="30" type="generic">1♂</specimenCount>
,
<collectionCode id="ED7BAE058A25225CEF416FA3ECE7F868" box="[1362,1424,1997,2021]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="29" pageNumber="30">CNC</collectionCode>
),
<collectingCountry id="F37D76508A26225FEA8468F9E999FF3D" box="[151,238,151,176]" name="Malaysia" pageId="30" pageNumber="31">Malaya</collectingCountry>
, Selangor, F.M.S., Gombak Valley,
<date id="FFD410008A26225FE89468F9EA71FF3D" box="[647,774,151,176]" pageId="30" pageNumber="31" value="1930-12-26">26.xii.1930</date>
(
<specimenCount id="9D6CFD498A26225FE90A68F6EA4AFF3D" box="[793,829,152,176]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
,
<collectionCode id="ED7BAE058A26225FE95868F6EAD4FF3D" box="[843,931,152,176]" pageId="30" pageNumber="31">BMNH</collectionCode>
), Selangor, Ulu Gombak,
<quantity id="4C929B258A26225FEEDD68F9EC7BFF3D" box="[1230,1292,151,176]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="30" pageNumber="31" unit="ft" value="800.0">800ft</quantity>
, J.W. Ismay, BM 1986-283,
<date id="FFD410008A26225FEB5568D2E814FF58" box="[326,355,188,213]" pageId="30" pageNumber="31" value="1983-11-10">10</date>
.xi.1983 (
<specimenCount id="9D6CFD498A26225FEBC368D2E883FF59" box="[464,500,188,212]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
,
<collectionCode id="ED7BAE058A26225FE81168D2EB2DFF59" box="[514,602,188,212]" pageId="30" pageNumber="31">BMNH</collectionCode>
),
<date id="FFD410008A26225FE87C68D2EB90FF58" box="[623,743,188,213]" pageId="30" pageNumber="31" value="1983-11-12">12.xi.1983</date>
(
<specimenCount id="9D6CFD498A26225FE8EA68D2EA6BFF59" box="[761,796,188,212]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
1♀,
<collectionCode id="ED7BAE058A26225FE94468D2EAD8FF59" box="[855,943,188,212]" pageId="30" pageNumber="31">BMNH</collectionCode>
), Selangor: Hulu, Langat,
<date id="FFD410008A26225FEECF68D2EC28FF58" box="[1244,1375,188,213]" pageId="30" pageNumber="31" value="1988-06" valueMax="1988-07-31" valueMin="1988-06-00">vivii.1988</date>
, C.S. Doi, bamboo thicket, CIE
<accessionNumber id="9439AB238A26225FEBD6688EEB68FF75" box="[453,543,224,248]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A19844" pageId="30" pageNumber="31" type="EnaNcbi">A19844</accessionNumber>
(1♀,
<collectionCode id="ED7BAE058A26225FE870688EEBCCFF75" box="[611,699,224,248]" pageId="30" pageNumber="31">BMNH</collectionCode>
),
<collectingCountry id="F37D76508A26225FE8C368B1EA42FF75" box="[720,821,223,248]" name="Malaysia" pageId="30" pageNumber="31">Sarawak</collectingCountry>
: Mt. Kalujong,
<quantity id="4C929B258A26225FE9FA68B1ED4CFF75" box="[1001,1083,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="5.4864" pageId="30" pageNumber="31" unit="ft" value="1.8">1,800ft</quantity>
,
<date id="FFD410008A26225FEE5A68B1EDC2FF75" box="[1097,1205,223,248]" pageId="30" pageNumber="31" value="1932-11-04">4.xi.1932</date>
, Oxford Univ. Exp., B.M Hobby &amp; A.W. Moore, B.M.1933-254, primitive forest, undergrowth (1♀,
<collectionCode id="ED7BAE058A26225FEE38696AEDF4FE91" box="[1067,1155,260,284]" pageId="30" pageNumber="31">BMNH</collectionCode>
), Selangor, Old Bentong Pass nr Genting turnoff,
<date id="FFD410008A26225FEBBF6949EB54FECD" box="[428,547,295,320]" pageId="30" pageNumber="31" value="1985-04-11">11.iv.1985</date>
, D.K. McAlpine (1♀,
<collectionCode id="ED7BAE058A26225FE9326946EA01FECD" box="[801,886,296,320]" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="30" pageNumber="31">AMSA</collectionCode>
), Genting Highlands, Road Junction,
<date id="FFD410008A26225FEF0E6949ECE0FECD" box="[1309,1431,295,320]" pageId="30" pageNumber="31" value="1973-06-15">15.vi.1973</date>
, D.K. McAlpine (
<specimenCount id="9D6CFD498A26225FEB736922E8F4FEE9" box="[352,387,332,356]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
1♀,
<collectionCode id="ED7BAE058A26225FEBD76922EB6EFEE9" box="[452,537,332,356]" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="30" pageNumber="31">AMSA</collectionCode>
), Bentong Rd.,
<quantity id="4C929B258A26225FE8C56922EA61FEE8" box="[726,790,332,357]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="km" value="30.0">30km</quantity>
ex Kuala Lumpur, Gombak, K. Khoe,
<date id="FFD410008A26225FEEFF6922EC2AFEE8" box="[1260,1373,332,357]" pageId="30" pageNumber="31" value="1987-12-01">1.xii.1987</date>
(1♀,
<collectionCode id="ED7BAE058A26225FEA84691EE99BFE05" box="[151,236,368,392]" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="30" pageNumber="31">AMSA</collectionCode>
),
<date id="FFD410008A26225FEB106901E8F4FE05" box="[259,387,367,392]" pageId="30" pageNumber="31" value="1987-12-30">30.xii.1987</date>
(
<specimenCount id="9D6CFD498A26225FEB84691EE8CDFE05" box="[407,442,368,392]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
1♀,
<collectionCode id="ED7BAE058A26225FEBE9691EEB38FE05" box="[506,591,368,392]" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="30" pageNumber="31">AMSA</collectionCode>
), Genting Highlands, Bentong Pass, D.K. McAlpine,
<date id="FFD410008A26225FEED06901EC4FFE05" box="[1219,1336,367,392]" pageId="30" pageNumber="31" value="1985-04-10">10.iv.1985</date>
(1♀ 1?,
<collectionCode id="ED7BAE058A26225FEA8469FAE99BFE21" box="[151,236,404,428]" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="30" pageNumber="31">AMSA</collectionCode>
),
<date id="FFD410008A26225FEB1169FAE80FFE20" box="[258,376,404,429]" pageId="30" pageNumber="31" value="1985-04-12">12.iv.1985</date>
(
<specimenCount id="9D6CFD498A26225FEB9F69FAE8C6FE21" box="[396,433,404,428]" pageId="30" pageNumber="31" type="generic">2♂</specimenCount>
,
<collectionCode id="ED7BAE058A26225FEBD369FAEB61FE21" box="[448,534,404,428]" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="30" pageNumber="31">AMSA</collectionCode>
).
<emphasis id="B91EEAD28A26225FE83F69FAEBBDFE20" bold="true" box="[556,714,404,429]" pageId="30" pageNumber="31">
<collectingCountry id="F37D76508A26225FE83F69FAEBB2FE20" box="[556,709,404,429]" name="Thailand" pageId="30" pageNumber="31">THAILAND</collectingCountry>
.
</emphasis>
“Peninsular
<collectingCountry id="F37D76508A26225FE97269FAEAE8FE20" box="[865,927,404,429]" name="Thailand" pageId="30" pageNumber="31">Siam</collectingCountry>
”, Nakon Sri Tamarat, Khao duang,
<quantity id="4C929B258A26225FEF5169FAECEFFE20" box="[1346,1432,404,429]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="30" pageNumber="31" unit="ft" value="2000.0">2000 ft</quantity>
,
<date id="FFD410008A26225FEA8469D9E864FE5D" box="[151,275,439,464]" pageId="30" pageNumber="31" value="1922-03-21">21.iii.1922</date>
, H.M. Pendlebury (1♀,
<collectionCode id="ED7BAE058A26225FE82569D6EBF9FE5D" box="[566,654,440,464]" pageId="30" pageNumber="31">BMNH</collectionCode>
), S Khao Lak Nat. Par., Tone Chong Fah Fall,
<quantity id="4C929B258A26225FEEDE69D9EC6CFE5D" box="[1229,1307,439,464]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="100.0">100km</quantity>
N Phuket,
<date id="FFD410008A26225FEA8469B2E87EFE79" box="[151,265,476,500]" pageId="30" pageNumber="31" value="1993-10-20">20.x.1993</date>
, F. Kaplan &amp; A. Freidberg (
<specimenCount id="9D6CFD498A26225FE85C69B2EB04FE79" box="[591,627,476,500]" pageId="30" pageNumber="31" type="generic">2♂</specimenCount>
1♀,
<collectionCode id="ED7BAE058A26225FE8BC69B2EB9CFE79" box="[687,747,476,500]" httpUri="http://grbio.org/cool/ze71-842p" name="Tel-Aviv University" pageId="30" pageNumber="31">
<collectingCountry id="F37D76508A26225FE8BC69B2EB9CFE79" box="[687,747,476,500]" name="China" pageId="30" pageNumber="31">TAU</collectingCountry>
</collectionCode>
), S Khao Sok Nat. Per., Rt. 401,
<date id="FFD410008A26225FEE7869B2EDFDFE79" box="[1131,1162,476,500]" pageId="30" pageNumber="31" value="1993-10-22">22</date>
.x.1993, F. Kaplan &amp; A. Freidberg (2♀,
<collectionCode id="ED7BAE058A26225FEB416A6EE8F9FD95" box="[338,398,512,536]" httpUri="http://grbio.org/cool/ze71-842p" name="Tel-Aviv University" pageId="30" pageNumber="31">
<collectingCountry id="F37D76508A26225FEB416A6EE8F9FD95" box="[338,398,512,536]" name="China" pageId="30" pageNumber="31">TAU</collectingCountry>
</collectionCode>
), Banna, Chawang, nr Nabon,
<quantity id="4C929B258A26225FE9156A6EEA4DFD95" box="[774,826,512,536]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="31" unit="m" value="70.0">70m</quantity>
,
<date id="FFD410008A26225FE95F6991EACFFD95" box="[844,952,511,536]" pageId="30" pageNumber="31" value="1958-09-06">6.ix.1958</date>
, J.L. Gressitt (
<specimenCount id="9D6CFD498A26225FEE626A6EEDE2FD95" box="[1137,1173,512,536]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
,
<collectionCode id="ED7BAE058A26225FEEBB6A6EED92FD95" box="[1192,1253,512,536]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="30" pageNumber="31">CNC</collectionCode>
), Khorat Prov.: Sakaerat,
<quantity id="4C929B258A26225FEB1B6A4AE821FDB0" box="[264,342,548,573]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="km" value="300.0">300km</quantity>
NE of Bangkok,
<date id="FFD410008A26225FE8376A4AEBE5FDB0" box="[548,658,548,573]" pageId="30" pageNumber="31" value="1968-03-01">1.iii.1968</date>
, dry diplocot forest, D.E. Hardy (2♀,
<collectionCode id="ED7BAE058A26225FEE5D6A4AEDD7FDB1" LSID="urn:lsid:biocol.org:col:34697" box="[1102,1184,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="30" pageNumber="31">BPBM</collectionCode>
), Doi Suthep,
<quantity id="4C929B258A26225FEF546A4AECEFFDB1" box="[1351,1432,548,572]" metricMagnitude="3" metricUnit="m" metricValue="1.278" pageId="30" pageNumber="31" unit="m" value="1278.0">1278m</quantity>
,
<date id="FFD410008A26225FEA846A29E835FDED" box="[151,322,583,608]" pageId="30" pageNumber="31" value="1958-03-29" valueMax="1958-05-04" valueMin="1958-03-29">29.iii4.v.1958</date>
, water margin, T.C. Maa (1♀,
<collectionCode id="ED7BAE058A26225FE8AD6A26EA78FDD2" LSID="urn:lsid:biocol.org:col:34697" box="[702,783,584,607]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="30" pageNumber="31">BPBM</collectionCode>
; 1♀,
<collectionCode id="ED7BAE058A26225FE9446A26EAE2FDED" box="[855,917,584,608]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="30" pageNumber="31">CNC</collectionCode>
), Chantiburi Prov.: Kwad Soi Dao Wildlife Sanctuary, breeding centre,
<date id="FFD410008A26225FEBC76A02EB38FD08" box="[468,591,620,645]" pageId="30" pageNumber="31" value="1987-08-09">9.viii.1987</date>
, R. deKeyzer (
<specimenCount id="9D6CFD498A26225FE8EE6A02EA57FD09" box="[765,800,620,644]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
1♀,
<collectionCode id="ED7BAE058A26225FE9486A02EAC7FD09" box="[859,944,620,644]" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="30" pageNumber="31">AMSA</collectionCode>
), Chiang Rai Prov.: Pong Pah,
<date id="FFD410008A26225FEF056A02ECEFFD08" box="[1302,1432,620,645]" pageId="30" pageNumber="31" value="2013-07-15">
15.vii.
<quantity id="4C929B258A26225FEF4A6A02ECEFFD08" box="[1369,1432,620,645]" metricMagnitude="3" metricUnit="m" metricValue="2.0134499999999997" pageId="30" pageNumber="31" unit="m" value="2013.45">2013</quantity>
</date>
, 450m, 20.3812˚N, 99.8670˚E, M. Hauser (
<specimenCount id="9D6CFD498A26225FE86D6AFEEBD5FD25" box="[638,674,656,680]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
,
<collectionCode id="ED7BAE058A26225FE8BC6AFEEB8AFD25" LSID="urn:lsid:biocol.org:col:33156" box="[687,765,656,680]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="30" pageNumber="31">CSCA</collectionCode>
).
<emphasis id="B91EEAD28A26225FE9026AE1EAD7FD25" bold="true" box="[785,928,655,680]" pageId="30" pageNumber="31">
<collectingCountry id="F37D76508A26225FE9026AE1EAECFD25" box="[785,923,655,680]" name="Vietnam" pageId="30" pageNumber="31">VIETNAM</collectingCountry>
.
</emphasis>
<quantity id="4C929B258A26225FE9BB6AE1EA9FFD25" box="[936,1000,655,680]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="30" pageNumber="31" unit="km" value="22.0">22km</quantity>
S of Nha Trang,
<date id="FFD410008A26225FEEA26AE1EC22FD25" box="[1201,1365,655,680]" pageId="30" pageNumber="31" value="1960-11-20" valueMax="1960-11-26" valueMin="1960-11-20">2026.xi.1960</date>
, C.M. Yoshimoto (
<specimenCount id="9D6CFD498A26225FEB326ADAE833FD41" box="[289,324,692,716]" pageId="30" pageNumber="31" type="generic">2♂</specimenCount>
1♀,
<collectionCode id="ED7BAE058A26225FEB6E6ADAE8B8FD41" LSID="urn:lsid:biocol.org:col:34697" box="[381,463,692,716]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="30" pageNumber="31">BPBM</collectionCode>
), Ban Me Thuot,
<quantity id="4C929B258A26225FE8856ADAEBAFFD40" box="[662,728,692,717]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="m" value="500.0">500m</quantity>
,
<date id="FFD410008A26225FE8F76ADAEA08FD40" box="[740,895,692,717]" pageId="30" pageNumber="31" value="1960-05-16" valueMax="1960-05-18" valueMin="1960-05-16">1618.v.1960</date>
, L.W. Quate (
<specimenCount id="9D6CFD498A26225FEE0E6ADAED37FD41" box="[1053,1088,692,716]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
3♀,
<collectionCode id="ED7BAE058A26225FEE6A6ADAEDBCFD41" LSID="urn:lsid:biocol.org:col:34697" box="[1145,1227,692,716]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="30" pageNumber="31">BPBM</collectionCode>
), Dak Song,
<quantity id="4C929B258A26225FEF4F6ADAECEAFD40" box="[1372,1437,692,717]" metricMagnitude="4" metricUnit="m" metricValue="7.6" pageId="30" pageNumber="31" unit="km" value="76.0">76km</quantity>
SW of Ban Me Thuot,
<quantity id="4C929B258A26225FEB896AB6E8ABFD7D" box="[410,476,728,752]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="30" pageNumber="31" unit="m" value="870.0">870m</quantity>
,
<date id="FFD410008A26225FEBFB6AB9EBF4FD7D" box="[488,643,727,752]" pageId="30" pageNumber="31" value="1960-05-19" valueMax="1960-05-21" valueMin="1960-05-19">1921.v.1960</date>
, L.W. Quate (1♀,
<collectionCode id="ED7BAE058A26225FE9416AB6EAD3FD7D" LSID="urn:lsid:biocol.org:col:34697" box="[850,932,728,752]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="30" pageNumber="31">BPBM</collectionCode>
), Fyan,
<quantity id="4C929B258A26225FE9ED6AB6EDF0FD7D" box="[1022,1159,728,752]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="30" pageNumber="31" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">9001000m</quantity>
,
<date id="FFD410008A26225FEE816AB9EC2CFD7D" box="[1170,1371,727,752]" pageId="30" pageNumber="31" value="1961-07-11" valueMax="1961-08-09" valueMin="1961-07-11">11.vii9.viii.1961</date>
, N.R. Spencer (1♀,
<collectionCode id="ED7BAE058A26225FEB2C6A92E8E6FC99" LSID="urn:lsid:biocol.org:col:34697" box="[319,401,764,788]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="30" pageNumber="31">BPBM</collectionCode>
), Ninh Binh, Cuc Phuong National Park, 20°2113ʺN, 105°3514ʺE,
<quantity id="4C929B258A26225FEEDA6A92EC33FC98" box="[1225,1348,764,789]" metricMagnitude="2" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="30" pageNumber="31" unit="m" value="425.0" valueMax="500.0" valueMin="350.0">350500m</quantity>
,
<date id="FFD410008A26225FEF466A92E980FCB5" pageId="30" pageNumber="31" value="2013-04-30" valueMax="2013-05-08" valueMin="2013-04-30">30.iv 8.v.2013</date>
S.A.
<collectingCountry id="F37D76508A26225FEB246B71E8ECFCB5" box="[311,411,799,824]" name="Marshall Islands" pageId="30" pageNumber="31">Marshall</collectingCountry>
&amp; S.M. Paiero (
<specimenCount id="9D6CFD498A26225FE8416B4EEB00FCB5" box="[594,631,800,824]" pageId="30" pageNumber="31" type="generic">4♂</specimenCount>
1♀,
<collectionCode id="ED7BAE058A26225FE8BD6B4EEB89FCB5" box="[686,766,800,824]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="30" pageNumber="31">DEBU</collectionCode>
),
<date id="FFD410008A26225FE9016B4EEA03FCB5" box="[786,884,800,824]" pageId="30" pageNumber="31" value="2013-05-02">2.v.2013</date>
,
<collectingCountry id="F37D76508A26225FE9936B71EA93FCB5" box="[896,996,799,824]" name="Marshall Islands" pageId="30" pageNumber="31">Marshall</collectingCountry>
&amp; Paiero (2♀,
<collectionCode id="ED7BAE058A26225FEE836B4EED97FCBA" box="[1168,1248,800,823]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="30" pageNumber="31">DEBU</collectionCode>
;
<specimenCount id="9D6CFD498A26225FEEF86B4EEC79FCB5" box="[1259,1294,800,824]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
1♀,
<collectionCode id="ED7BAE058A26225FEF556B4EECE7FCB5" LSID="urn:lsid:biocol.org:col:12905" box="[1350,1424,800,824]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12905" name="Cuc Phuong National Park" pageId="30" pageNumber="31">CPNP</collectionCode>
), Ninh Binh, Cuc Phuong National Park, 20°1835ʺN, 105°3706ʺE,
<date id="FFD410008A26225FE99D6B2AED63FCD0" box="[910,1044,836,861]" pageId="30" pageNumber="31" value="2013-08-11">11.viii.2013</date>
S.A.
<collectingCountry id="F37D76508A26225FEE446B2AEDCCFCD0" box="[1111,1211,836,861]" name="Marshall Islands" pageId="30" pageNumber="31">Marshall</collectingCountry>
(
<specimenCount id="9D6CFD498A26225FEEDD6B2AED85FCD1" box="[1230,1266,836,860]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
,
<collectionCode id="ED7BAE058A26225FEEEC6B2AEC27FCD1" box="[1279,1360,836,860]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="30" pageNumber="31">DEBU</collectionCode>
), Ninh Binh Province: Cuc Phuong National Park, 20˚2101ʺN, 105˚3535ʺE, ex hand collected,
<date id="FFD410008A26225FEE646B09EC6CFC0D" box="[1143,1307,871,896]" pageId="30" pageNumber="31" value="2014-06-13" valueMax="2014-06-17" valueMin="2014-06-13">1317.vi.2014</date>
, M. Hauser &amp; N. vonEllenreider (2♀,
<collectionCode id="ED7BAE058A26225FEBC56BE2EB53FC29" LSID="urn:lsid:biocol.org:col:33156" box="[470,548,908,932]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="30" pageNumber="31">CSCA</collectionCode>
), Guang Ninh, Cat Ba Island, 20˚4736ʺN, 106˚5955ʺE,
<date id="FFD410008A26225FEEC36BE2EC2FFC28" box="[1232,1368,908,933]" pageId="30" pageNumber="31" value="2011-08-12">12.viii.2011</date>
, S.A.
<collectingCountry id="F37D76508A26225FEA846BC1E98CFC45" box="[151,251,943,968]" name="Marshall Islands" pageId="30" pageNumber="31">Marshall</collectingCountry>
(
<specimenCount id="9D6CFD498A26225FEB1D6BDEE845FC45" box="[270,306,944,968]" pageId="30" pageNumber="31" type="generic">1♂</specimenCount>
,
<collectionCode id="ED7BAE058A26225FEB526BDEE8E6FC45" box="[321,401,944,968]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="30" pageNumber="31">DEBU</collectionCode>
), Bac Giang, An Lac, Khe Ro Forest, 21˚2026ʺN, 106˚5800ʺE,
<quantity id="4C929B258A26225FEE916BC1ED98FC45" box="[1154,1263,943,968]" metricMagnitude="2" metricUnit="m" metricValue="1.125" metricValueMax="1.5" metricValueMin="0.75" pageId="30" pageNumber="31" unit="m" value="112.5" valueMax="150.0" valueMin="75.0">75150m</quantity>
,
<date id="FFD410008A26225FEEEE6BC1ECEFFC45" box="[1277,1432,943,968]" pageId="30" pageNumber="31" value="2013-05-16" valueMax="2013-05-18" valueMin="2013-05-16">1618.v.2013</date>
,
<collectingCountry id="F37D76508A26225FEA846BBAE98CFC60" box="[151,251,980,1005]" name="Marshall Islands" pageId="30" pageNumber="31">Marshall</collectingCountry>
&amp; Paiero (2♀,
<collectionCode id="ED7BAE058A26225FEBB46BBAE88FFC61" box="[423,504,980,1004]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="30" pageNumber="31">DEBU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C370654B8A26225FEAD46B99ED82FB48" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BD536C08A26225FEAD46B99ED82FB48" blockId="30.[151,1437,151,1221]" pageId="30" pageNumber="31">
<emphasis id="B91EEAD28A26225FEAD46B99E838FB9D" bold="true" box="[199,335,1015,1040]" pageId="30" pageNumber="31">Comments.</emphasis>
<taxonomicName id="4C6A4D438A26225FEB776B99EB12FB9D" box="[356,613,1015,1040]" class="Insecta" family="Nothybidae" genus="Nothybus" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="B91EEAD28A26225FEB776B99EB12FB9D" box="[356,613,1015,1040]" italics="true" pageId="30" pageNumber="31">Nothybus sumatranus</emphasis>
</taxonomicName>
is relatively widespread in the Oriental Region, and the most morphologically variable of all
<taxonomicName id="4C6A4D438A26225FEBE56C72EB17FBB8" box="[502,608,1052,1077]" class="Insecta" family="Nothybidae" genus="Nothybus" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEAD28A26225FEBE56C72EB17FBB8" box="[502,608,1052,1077]" italics="true" pageId="30" pageNumber="31">Nothybus</emphasis>
</taxonomicName>
species. Specimens vary in width and length of the transverse brown band on the wing, strength of the scutal stripe, size (or absence) of the orange spots on tergite 4, and colour of the facial tubercle and female tergite 6, but there are no clear patterns of correlated characters supporting the division of
<taxonomicName id="4C6A4D438A26225FEF916C0BE86DFB2D" class="Insecta" family="Nothybidae" genus="Nothybus" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="B91EEAD28A26225FEF916C0BE86DFB2D" italics="true" pageId="30" pageNumber="31">N. sumatranus</emphasis>
</taxonomicName>
into additional taxa. Furthermore, the male terminalia of non-Sumatran specimens, except for one
<collectingCountry id="F37D76508A26225FEF7A6CE9ECEAFB2D" box="[1385,1437,1159,1184]" name="Thailand" pageId="30" pageNumber="31">Thai</collectingCountry>
male with the apex of the phallus slightly narrowed, are identical with the Sumatran male dissected.
</paragraph>
</subSubSection>
</treatment>
</document>