335 lines
47 KiB
XML
335 lines
47 KiB
XML
<document id="72A788B189D948BF3611925A9355FE46" ID-CLB-Dataset="254440" ID-DOI="10.11646/zootaxa.5270.3.4" ID-GBIF-Dataset="ec485338-4b66-4557-be6c-b53ac23a4678" ID-ISSN="1175-5326" ID-Zenodo-Dep="7860353" ID-ZooBank="D3EE46BD-0755-4730-AF3C-ACD92085311A" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1682061938543" checkinUser="plazi" docAuthor="Valdés, Ángel, Feliciano, Kendall & Malaquias, Manuel A. E." docDate="2023" docId="03C38798FF9646738ED72DC74231FC47" docLanguage="en" docName="zootaxa.5270.3.4.pdf" docOrigin="Zootaxa 5270 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5270.3.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pupa charlesi Valdés & Feliciano & Malaquias 2023, sp. nov." docType="treatment" docVersion="6" lastPageNumber="499" masterDocId="FFFAFFE0FF8D466F8E402C1C413AFFB6" masterDocTitle="The genus Pupa Röding, 1798 (Mollusca, Gastropoda, Acteonidae) in New Caledonia with notes on Recent species" masterLastPageNumber="506" masterPageNumber="471" pageNumber="498" updateTime="1699514729445" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D02F7519AFDEBA7D3240264E262560F2">The genus Pupa Röding, 1798 (Mollusca, Gastropoda, Acteonidae) in New Caledonia with notes on Recent species</mods:title>
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<mods:namePart id="8B6094CC5CFB24EDC80E90C0FBA5E22B">Valdés, Ángel</mods:namePart>
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<mods:affiliation id="2BFA56D7D206104235490C4BD477D86C">Department of Biological Sciences, California State Polytechnic University, 3801 West Temple Avenue, Pomona, California 91768, USA. & aavaldes @ cpp. edu; https: // orcid. org / 0000 - 0002 - 2347 - 4896</mods:affiliation>
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<mods:nameIdentifier id="7C3E8DBAE79AFFD61DB101C919CB1873" type="email">aavaldes@cpp.edu</mods:nameIdentifier>
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<mods:namePart id="C3A44436B62677E8B0D53DA479827CEB">Feliciano, Kendall</mods:namePart>
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<mods:nameIdentifier id="D1C97B8ED87473F9C40EE3A9F9C534BC" type="ORCID">0000-0003-0536-2628</mods:nameIdentifier>
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<mods:affiliation id="DF97CEC8A88D7C51EBB509DC77BA8236">Department of Biological Sciences, California State Polytechnic University, 3801 West Temple Avenue, Pomona, California 91768, USA. & https: // orcid. org / 0000 - 0003 - 0536 - 2628</mods:affiliation>
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<mods:namePart id="49F449EB34276BA0D243D4D0DEE66436">Malaquias, Manuel A. E.</mods:namePart>
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<mods:affiliation id="FBAC5B98F3D56D97E9EC3F8D5609AA82">Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, N- 5020 Bergen, Norway. https: // orcid. org / 0000 - 0002 - 9668 - 945 X</mods:affiliation>
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<mods:title id="8383435C9CA810B9B850718758CAF483">Zootaxa</mods:title>
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<mods:date id="48C30927555944F670F5CBB7E1EE8993">2023</mods:date>
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<mods:number id="EC825C512963D648CFE245C3DDED7B2B">2023-04-21</mods:number>
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<treatment id="03C38798FF9646738ED72DC74231FC47" ID-DOI="http://doi.org/10.5281/zenodo.7860412" ID-GBIF-Taxon="209569253" ID-Zenodo-Dep="7860412" LSID="urn:lsid:plazi:treatment:03C38798FF9646738ED72DC74231FC47" httpUri="http://treatment.plazi.org/id/03C38798FF9646738ED72DC74231FC47" lastPageId="28" lastPageNumber="499" pageId="27" pageNumber="498">
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<heading id="D09D81E2FF9646748ED72DC74099FE43" bold="true" box="[151,419,474,501]" fontSize="11" level="1" pageId="27" pageNumber="498" reason="1">
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<emphasis id="B91EEA9CFF9646748ED72DC74099FE43" bold="true" box="[151,419,474,501]" pageId="27" pageNumber="498">
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<taxonomicName id="4C6A4D0DFF9646748ED72DC74004FE43" ID-CoL="6c8a7099-4f33-439a-b985-858fe91a5f14" authority="Valdés & Feliciano & Malaquias, 2023" authorityName="Valdés & Feliciano & Malaquias" authorityYear="2023" box="[151,318,474,501]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="27" pageNumber="498" phylum="Mollusca" rank="species" species="charlesi" status="sp. nov.">
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<emphasis id="B91EEA9CFF9646748ED72DC74004FE43" bold="true" box="[151,318,474,501]" italics="true" pageId="27" pageNumber="498">Pupa charlesi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22D57E7FF9646748F062DC74099FE43" box="[326,419,475,501]" pageId="27" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3706505FF9646748ED72DE34314FDAF" box="[151,558,510,537]" pageId="27" pageNumber="498" type="description">
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<paragraph id="8BD5368EFF9646748ED72DE34314FDAF" blockId="27.[151,558,474,537]" box="[151,558,510,537]" pageId="27" pageNumber="498">
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(
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<figureCitation id="13512A0BFF9646748EDF2DE341C6FDAF" box="[159,252,511,537]" captionStart="FIGURE 2" captionStartId="6.[151,250,1786,1810]" captionTargetBox="[151,1436,181,1762]" captionTargetId="figure-24@6.[151,1436,181,1762]" captionTargetPageId="6" captionText="FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159." figureDoi="http://doi.org/10.5281/zenodo.7860362" httpUri="https://zenodo.org/record/7860362/files/figure.png" pageId="27" pageNumber="498">Figs 2G</figureCitation>
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,
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<figureCitation id="13512A0BFF9646748F482DE3401CFDAF" box="[264,294,511,537]" captionStart="FIGURE 4" captionStartId="7.[152,255,1524,1548]" captionTargetBox="[151,1436,932,1500]" captionTargetId="figure-75@7.[151,1436,932,1500]" captionTargetPageId="7" captionText="FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158." figureDoi="http://doi.org/10.5281/zenodo.7860368" httpUri="https://zenodo.org/record/7860368/files/figure.png" pageId="27" pageNumber="498">4F</figureCitation>
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,
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<figureCitation id="13512A0BFF9646748F712DE24075FDAF" box="[305,335,510,537]" captionStart="FIGURE 5" captionStartId="10.[152,255,1580,1604]" captionTargetBox="[179,1409,181,1556]" captionTargetId="figure-24@10.[179,1409,181,1556]" captionTargetPageId="10" captionText="FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154." figureDoi="http://doi.org/10.5281/zenodo.7860370" httpUri="https://zenodo.org/record/7860370/files/figure.png" pageId="27" pageNumber="498">5F</figureCitation>
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,
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<figureCitation id="13512A0BFF9646748F1B2DE34043FDAF" box="[347,377,511,537]" captionStart="FIGURE 6" captionStartId="11.[152,255,1885,1909]" captionTargetBox="[166,1421,309,1862]" captionTargetId="figure-87@11.[166,1421,309,1862]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154." figureDoi="http://doi.org/10.5281/zenodo.7860372" httpUri="https://zenodo.org/record/7860372/files/figure.png" pageId="27" pageNumber="498">6F</figureCitation>
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,
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<figureCitation id="13512A0BFF9646748FC42DE34098FDAF" box="[388,418,511,537]" captionStart="FIGURE 7" captionStartId="12.[151,250,1561,1585]" captionTargetBox="[151,1436,181,1537]" captionTargetId="figure-24@12.[151,1436,181,1537]" captionTargetPageId="12" captionText="FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154." figureDoi="http://doi.org/10.5281/zenodo.7860376" httpUri="https://zenodo.org/record/7860376/files/figure.png" pageId="27" pageNumber="498">7F</figureCitation>
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,
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<figureCitation id="13512A0BFF9646748FED2DE340F1FDAF" box="[429,459,511,537]" captionStart="FIGURE 8" captionStartId="13.[152,255,1891,1915]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-19@13.[151,1436,181,1868]" captionTargetPageId="13" captionText="FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154." figureDoi="http://doi.org/10.5281/zenodo.7860380" httpUri="https://zenodo.org/record/7860380/files/figure.png" pageId="27" pageNumber="498">8F</figureCitation>
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,
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<figureCitation id="13512A0BFF9646748F972DE3431CFDAF" box="[471,550,511,537]" captionStart="FIGURE 16" captionStartId="28.[152,255,768,792]" captionTargetBox="[151,1436,181,745]" captionTargetId="figure-24@28.[151,1436,181,745]" captionTargetPageId="28" captionText="FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (C–E). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). C–E. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158." figureDoi="http://doi.org/10.5281/zenodo.7860408" httpUri="https://zenodo.org/record/7860408/files/figure.png" pageId="27" pageNumber="498">16C–E</figureCitation>
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)
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</paragraph>
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<subSubSection id="C3706505FF9646748ED72E5B45FBF99B" pageId="27" pageNumber="498" type="materials_examined">
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<paragraph id="8BD5368EFF9646748ED72E5B4002FD1F" blockId="27.[151,1437,582,1941]" pageId="27" pageNumber="498">
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<emphasis id="B91EEA9CFF9646748ED72E5B407BFDD6" bold="true" box="[151,321,582,609]" pageId="27" pageNumber="498">Type material</emphasis>
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.
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<materialsCitation id="3B023CD3FF9646748F132E5B4002FD1F" ID-GBIF-Occurrence="4089552338" collectingDate="2018-09-08" collectionCode="MNHN" country="New Caledonia" latitude="-20.626667" location="Plateau Karembe" longLatPrecision="126" longitude="164.28833" municipality="Koumac" pageId="27" pageNumber="498" specimenCode="KM301, IM-2013-86159" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54D1882CFF9646748F132E5B4084FDD7" box="[339,446,583,609]" pageId="27" pageNumber="498" type="holotype">Holotype</typeStatus>
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,
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<location id="8EB56055FF9646748F8D2E5B43A9FDD7" LSID="urn:lsid:plazi:treatment:03C38798FF9646738ED72DC74231FC47:8EB56055FF9646748F8D2E5B43A9FDD7" box="[461,659,583,609]" country="New Caledonia" latitude="-20.626667" longLatPrecision="126" longitude="164.28833" municipality="Koumac" name="Plateau Karembe" pageId="27" pageNumber="498">Plateau Karembé</location>
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, north end,
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<collectingMunicipality id="6BB1ACF4FF9646748D612E5B42B9FDD7" box="[801,899,583,609]" pageId="27" pageNumber="498">Koumac</collectingMunicipality>
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,
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<collectingCountry id="F37D761EFF9646748DD22E5B4579FDD7" box="[914,1091,583,609]" name="New Caledonia" pageId="27" pageNumber="498">New Caledonia</collectingCountry>
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(
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<geoCoordinate id="EE5E5049FF9646748A152E5445FCFDD5" box="[1109,1222,584,611]" degrees="20" direction="south" minutes="37.6" orientation="latitude" pageId="27" pageNumber="498" precision="92" value="-20.626667">20°37.6′S</geoCoordinate>
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,
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<geoCoordinate id="EE5E5049FF9646748A962E54446DFDD5" box="[1238,1367,584,611]" degrees="164" direction="east" minutes="17.3" orientation="longitude" pageId="27" pageNumber="498" precision="92" value="164.28833">164°17.3′E</geoCoordinate>
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), 0 m depth [Koumac 2.1 stn.
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<specimenCode id="DBCC9EF5FF9646748FDF2E7740C3FD33" box="[415,505,619,645]" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="27" pageNumber="498">KM301</specimenCode>
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, coral blocks, sand, mud],
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<date id="FFD4104EFF9646748D5E2E7742A6FD33" box="[798,924,619,645]" pageId="27" pageNumber="498" value="2018-09-08">
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<collectingDate id="EF90E9A6FF9646748D5E2E7742A6FD33" box="[798,924,619,645]" pageId="27" pageNumber="498" value="2018-09-08">8 Sep 2018</collectingDate>
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</date>
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,
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<quantity id="4C929B6BFF9646748DE62E7742CEFD33" box="[934,1012,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="498" unit="mm" value="20.0">20 mm</quantity>
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long (shell) (
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<collectionCode id="ED7BAE4BFF9646748AC52E7745E6FD33" box="[1157,1244,619,645]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="498">MNHN</collectionCode>
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<specimenCode id="DBCC9EF5FF9646748AA22E7644A2FD33" box="[1250,1432,618,645]" collectionCode="IM-" pageId="27" pageNumber="498">IM-2013-86159</specimenCode>
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, isolate KF63).
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</materialsCitation>
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</paragraph>
|
||
<paragraph id="8BD5368EFF9646748E872EAE4397FCEB" blockId="27.[151,1437,582,1941]" pageId="27" pageNumber="498">
|
||
<emphasis id="B91EEA9CFF9646748E872EAE4092FD7A" bold="true" box="[199,424,690,716]" pageId="27" pageNumber="498">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="3B023CD3FF9646748FF52EAF40B4FCA3" ID-GBIF-Occurrence="4089552340" collectingDate="2018-09-08" collectingDateMax="2018-09-26" collectingDateMin="2018-09-08" collectionCode="MNHN" country="New Caledonia" latitude="-20.626667" location="Plateau Karembe" longLatPrecision="126" longitude="164.28833" municipality="Koumac" pageId="27" pageNumber="498" specimenCode="KM301, IM-2013- 86158, KM321, IM-2013-86154" specimenCount="2">
|
||
<location id="8EB56055FF9646748FF52EAF4342FD7B" LSID="urn:lsid:plazi:treatment:03C38798FF9646738ED72DC74231FC47:8EB56055FF9646748FF52EAF4342FD7B" box="[437,632,691,717]" country="New Caledonia" latitude="-20.626667" longLatPrecision="126" longitude="164.28833" municipality="Koumac" name="Plateau Karembe" pageId="27" pageNumber="498">Plateau Karembé</location>
|
||
, north end,
|
||
<collectingMunicipality id="6BB1ACF4FF9646748CB82EAF4260FD7B" box="[760,858,691,717]" pageId="27" pageNumber="498">Koumac</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F37D761EFF9646748D242EAF452BFD7B" box="[868,1041,691,717]" name="New Caledonia" pageId="27" pageNumber="498">New Caledonia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE5E5049FF9646748A5E2EA845AAFD79" box="[1054,1168,692,719]" degrees="20" direction="south" minutes="37.6" orientation="latitude" pageId="27" pageNumber="498" precision="92" value="-20.626667">20°37.6′S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E5049FF9646748ADA2EA84426FD79" box="[1178,1308,692,719]" degrees="164" direction="east" minutes="17.3" orientation="longitude" pageId="27" pageNumber="498" precision="92" value="164.28833">164°17.3′E</geoCoordinate>
|
||
), 0 m depth [Koumac 2.1 stn.
|
||
<specimenCode id="DBCC9EF5FF9646748F1D2ECB4082FD47" box="[349,440,727,753]" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="27" pageNumber="498">KM301</specimenCode>
|
||
, coral blocks, sand, mud],
|
||
<date id="FFD4104EFF9646748CA02ECB425AFD47" box="[736,864,727,753]" pageId="27" pageNumber="498" value="2018-09-08">
|
||
<collectingDate id="EF90E9A6FF9646748CA02ECB425AFD47" box="[736,864,727,753]" pageId="27" pageNumber="498" value="2018-09-08">8 Sep 2018</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D6CFD07FF9646748D2B2ECB42D3FD47" box="[875,1001,727,753]" pageId="27" pageNumber="498" type="generic">1 specimen</specimenCount>
|
||
<quantity id="4C929B6BFF9646748DAF2ECB4504FD47" box="[1007,1086,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="27" pageNumber="498" unit="mm" value="22.0">22 mm</quantity>
|
||
long (shell) (
|
||
<collectionCode id="ED7BAE4BFF9646748A912ECB4412FD47" box="[1233,1320,727,753]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="498">MNHN</collectionCode>
|
||
<specimenCode id="DBCC9EF5FF9646748B6F2ECB41DAFCA3" collectionCode="IM-" pageId="27" pageNumber="498">IM-2013- 86158</specimenCode>
|
||
, isolate KF62).
|
||
</materialsCitation>
|
||
<materialsCitation id="3B023CD3FF9646748FD52EE74393FCEB" ID-GBIF-Occurrence="4089552313" collectingDate="2018-09-08" collectingDateMax="2018-09-26" collectingDateMin="2018-09-08" collectionCode="MNHN" country="New Caledonia" latitude="-20.626667" location="Plateau Karembe" longLatPrecision="126" longitude="164.28833" municipality="Koumac" pageId="27" pageNumber="498" specimenCode="KM301, IM-2013- 86158, KM321, IM-2013-86154" specimenCount="2">
|
||
Plateau Karembé, north end barrier, Koumac,
|
||
<collectingCountry id="F37D761EFF9646748DDE2EE74577FCA3" box="[926,1101,763,789]" name="New Caledonia" pageId="27" pageNumber="498">New Caledonia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE5E5049FF9646748A1D2EE045F4FCA1" box="[1117,1230,764,791]" degrees="20" direction="south" minutes="37.6" orientation="latitude" pageId="27" pageNumber="498" precision="92" value="-20.626667">20°37.6′S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E5049FF9646748A9B2EE04466FCA1" box="[1243,1372,764,791]" degrees="164" direction="east" minutes="17.3" orientation="longitude" pageId="27" pageNumber="498" precision="92" value="164.28833">164°17.3′E</geoCoordinate>
|
||
), 0 m depth [Koumac 2.1 stn.
|
||
<specimenCode id="DBCC9EF5FF9646748FE92F034339FC8F" box="[425,515,799,825]" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="27" pageNumber="498">KM321</specimenCode>
|
||
, reef flat with living/dead coral, seagrass],
|
||
<date id="FFD4104EFF9646748DAD2F034544FC8F" box="[1005,1150,799,825]" pageId="27" pageNumber="498" value="2018-09-26">
|
||
<collectingDate id="EF90E9A6FF9646748DAD2F034544FC8F" box="[1005,1150,799,825]" pageId="27" pageNumber="498" value="2018-09-26">26 Sep 2018</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D6CFD07FF9646748ACB2F034431FC8F" box="[1163,1291,799,825]" pageId="27" pageNumber="498" type="generic">1 specimen</specimenCount>
|
||
<quantity id="4C929B6BFF9646748B532F034459FC8F" box="[1299,1379,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="498" unit="mm" value="18.0">18 mm</quantity>
|
||
long (shell) (
|
||
<collectionCode id="ED7BAE4BFF9646748EAD2F5F407EFCEB" box="[237,324,835,861]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="498">MNHN</collectionCode>
|
||
<specimenCode id="DBCC9EF5FF9646748F0C2F5E4338FCEB" box="[332,514,834,861]" collectionCode="IM-" pageId="27" pageNumber="498">IM-2013-86154</specimenCode>
|
||
, isolate KF69)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5368EFF9646748E872F7B43E4FAE3" blockId="27.[151,1437,582,1941]" pageId="27" pageNumber="498">
|
||
<emphasis id="B91EEA9CFF9646748E872F7B40F2FC37" bold="true" box="[199,456,871,897]" pageId="27" pageNumber="498">External morphology</emphasis>
|
||
. Body oval, about the same width throughout (
|
||
<figureCitation id="13512A0BFF9646748D9A2F7B450FFC37" box="[986,1077,871,897]" captionStart="FIGURE 2" captionStartId="6.[151,250,1786,1810]" captionTargetBox="[151,1436,181,1762]" captionTargetId="figure-24@6.[151,1436,181,1762]" captionTargetPageId="6" captionText="FIGURE 2. Photographs of live animals from Koumac, New Caledonia, dorsal view. A. Pupa solidula (Linnaeus, 1758), MHNH IM-2013-86155. B–C. Pupa sulcata (Gmelin, 1791), (B) MHNH IM-2013-86145, (C) MHNH IM-2013-86160. D. Pupa nitidula (Lamarck, 1822), MHNH IM-2013-86157. E. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. F. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. G. Pupa charlesi sp. nov., MHNH IM-2013-86159." figureDoi="http://doi.org/10.5281/zenodo.7860362" httpUri="https://zenodo.org/record/7860362/files/figure.png" pageId="27" pageNumber="498">Fig. 2G</figureCitation>
|
||
). Cephalic shield notched along entire length longitudinally, with two elongate posterior extensions covering anterior end of shell. Foot extending beyond cephalic shield laterally. Body color translucent white, with opaque white dots scattered all over dorsal surface of cephalic shield, foot. Shell delicate, narrow, elongate, widest at 2/3 from anterior end (
|
||
<figureCitation id="13512A0BFF9646748ABC2FCF44ABFC5B" box="[1276,1425,979,1005]" captionStart="FIGURE 16" captionStartId="28.[152,255,768,792]" captionTargetBox="[151,1436,181,745]" captionTargetId="figure-24@28.[151,1436,181,745]" captionTargetPageId="28" captionText="FIGURE 16. Photographs of shell specimens of Pupa niecaensis (Barnard, 1963) (A), Pupa pascuana Raines, 2003 (B) and Pupa charlesi sp. nov. (C–E). A. Holotype of Pupa niecaensis (Barnard, 1963) (SAM A6553), ©SAM, photo by R. Adams. B. Holotype of Pupa pascuana Raines, 2003 (LACM 2954). C–E. Specimens of Pupa charlesi sp. nov. from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158." figureDoi="http://doi.org/10.5281/zenodo.7860408" httpUri="https://zenodo.org/record/7860408/files/figure.png" pageId="27" pageNumber="498">Figs. 16C–E</figureCitation>
|
||
). Sides convex, rounded to elongate anterior end. Body whorl large, about 1/8 of total length. Spire short, conical, with 5 whorls. Suture channeled. Aperture elongate, wider anteriorly, narrowing gradually towards posterior end, ending at 1/7 of the posterior end of first whorl. Columellar margin thickened, slightly oblique, with single large, channeled anterior fold starting at anterior end of aperture. Apex of all specimens examined damaged, protoconch not observed. Sculpture composed of numerous punctuated spiral grooves. Punctuations conspicuous, irregular, oval, situated next to each other, fused together, within each groove. Grooves separated by gaps much wider than grooves. Shell color white to pale orange, checkered with dark grey to black, square to oval patches, situated on gaps between punctuated spiral grooves all over shell. Two bands on body whorl with smaller patches. Posterior whorls white with no dark patches. Punctuated spiral grooves often edged in orange. Operculum translucent to ochre, elongate, with faint longitudinal striations (
|
||
<figureCitation id="13512A0BFF9646748C3B292743EBFAE3" box="[635,721,1339,1365]" captionStart="FIGURE 4" captionStartId="7.[152,255,1524,1548]" captionTargetBox="[151,1436,932,1500]" captionTargetId="figure-75@7.[151,1436,932,1500]" captionTargetPageId="7" captionText="FIGURE 4. Photographs of the opercula of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86152. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM- 2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86158." figureDoi="http://doi.org/10.5281/zenodo.7860368" httpUri="https://zenodo.org/record/7860368/files/figure.png" pageId="27" pageNumber="498">Fig. 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD5368EFF9646748E87294345FBF99B" blockId="27.[151,1437,582,1941]" pageId="27" pageNumber="498">
|
||
<emphasis id="B91EEA9CFF9646748E87294340FAFACF" bold="true" box="[199,448,1375,1401]" pageId="27" pageNumber="498">Internal morphology</emphasis>
|
||
. Reproductive system with large, elongate penis (
|
||
<figureCitation id="13512A0BFF9646748DA42943450CFACF" box="[996,1078,1374,1401]" captionStart="FIGURE 5" captionStartId="10.[152,255,1580,1604]" captionTargetBox="[179,1409,181,1556]" captionTargetId="figure-24@10.[179,1409,181,1556]" captionTargetPageId="10" captionText="FIGURE 5. Photographs of the male reproductive organs of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154." figureDoi="http://doi.org/10.5281/zenodo.7860370" httpUri="https://zenodo.org/record/7860370/files/figure.png" pageId="27" pageNumber="498">Fig. 5F</figureCitation>
|
||
), otherwise indistinct. Digestive system with elongate buccal mass (
|
||
<figureCitation id="13512A0BFF9646748C70299F43B2FA2B" box="[560,648,1411,1437]" captionStart="FIGURE 6" captionStartId="11.[152,255,1885,1909]" captionTargetBox="[166,1421,309,1862]" captionTargetId="figure-87@11.[166,1421,309,1862]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of the anterior portion of the digestive system of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154." figureDoi="http://doi.org/10.5281/zenodo.7860372" httpUri="https://zenodo.org/record/7860372/files/figure.png" pageId="27" pageNumber="498">Fig. 6F</figureCitation>
|
||
). Oral tube conical, connected to long, curved buccal bulb. Salivary glands, esophagus, connected on proximal end of buccal bulb. Radular formula 121 × 6.0.
|
||
<quantity id="4C929B6BFF9646748AF129BB45DAFA77" box="[1201,1248,1447,1473]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="27" pageNumber="498" unit="in" value="6.0">6 in</quantity>
|
||
an
|
||
<quantity id="4C929B6BFF9646748B4E29BB445BFA77" box="[1294,1377,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="498" unit="mm" value="18.0">18 mm</quantity>
|
||
long specimen (
|
||
<collectingRegion id="49AEF86CFF9646748F4F29D7405CFA53" box="[271,358,1483,1509]" country="Ireland" name="Monaghan" pageId="27" pageNumber="498">MNHN</collectingRegion>
|
||
<collectionCode id="ED7BAE4BFF9646748F2E29D640A1FA53" box="[366,411,1482,1509]" pageId="27" pageNumber="498">IM-</collectionCode>
|
||
2013-86154). Lateral teeth similar in shape, increasing in size gradually towards outer edge (
|
||
<figureCitation id="13512A0BFF9646748EDF29F341CCF9BF" box="[159,246,1519,1545]" captionStart="FIGURE 7" captionStartId="12.[151,250,1561,1585]" captionTargetBox="[151,1436,181,1537]" captionTargetId="figure-24@12.[151,1436,181,1537]" captionTargetPageId="12" captionText="FIGURE 7. Scanning electron micrographs of the radulae of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154." figureDoi="http://doi.org/10.5281/zenodo.7860376" httpUri="https://zenodo.org/record/7860376/files/figure.png" pageId="27" pageNumber="498">Fig. 7F</figureCitation>
|
||
). Teeth triangular, with broad base; elongate, curved, sharp cusp; 4–7 denticles similar to cusp in shape, shorter. Jaws with short, wide elements, bearing 8–10 finger-like projections apically (
|
||
<figureCitation id="13512A0BFF9646748A1F2A0F458EF99B" box="[1119,1204,1555,1581]" captionStart="FIGURE 8" captionStartId="13.[152,255,1891,1915]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-19@13.[151,1436,181,1868]" captionTargetPageId="13" captionText="FIGURE 8. Scanning electron micrographs of the jaw elements of specimens from Koumac, New Caledonia. A. Pupa solidula (Linnaeus, 1758), MNHN IM-2013-86155. B. Pupa sulcata (Gmelin, 1791), MHNH IM-2013-86146. C. Pupa nitidula (Lamarck, 1822), MNHN IM-2013-86157. D. Pupa coccinata (Reeve, 1842), MHNH IM-2013-86143. E. Pupa affinis (A. Adams, 1855), MNHN IM-2013-86144. F. Pupa charlesi sp. nov., MNHN IM-2013-86154." figureDoi="http://doi.org/10.5281/zenodo.7860380" httpUri="https://zenodo.org/record/7860380/files/figure.png" pageId="27" pageNumber="498">Fig. 8F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3706505FF9646748E872A2B42C2F90B" pageId="27" pageNumber="498" type="etymology">
|
||
<paragraph id="8BD5368EFF9646748E872A2B41C7F92F" blockId="27.[151,1437,582,1941]" pageId="27" pageNumber="498">
|
||
<emphasis id="B91EEA9CFF9646748E872A2B4072F9E7" bold="true" box="[199,328,1591,1617]" pageId="27" pageNumber="498">Etymology</emphasis>
|
||
. This species is named in honor of Laurent Charles (MHNBx), a photographer during the Koumac 2.1 expedition who contributed excellent photographs of live specimens of
|
||
<taxonomicName id="4C6A4D0DFF9646748D8E2A474533F9C2" authorityName="Draparnaud" authorityYear="1801" box="[974,1033,1627,1652]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="27" pageNumber="498" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91EEA9CFF9646748D8E2A474533F9C2" box="[974,1033,1627,1652]" italics="true" pageId="27" pageNumber="498">Pupa</emphasis>
|
||
</taxonomicName>
|
||
as well as
|
||
<typeStatus id="54D1882CFF9646748A3C2A474596F9C3" box="[1148,1196,1627,1653]" pageId="27" pageNumber="498">type</typeStatus>
|
||
material photographs and data.
|
||
</paragraph>
|
||
<paragraph id="8BD5368EFF9646748E872ABE42C2F90B" blockId="27.[151,1437,582,1941]" box="[199,1016,1698,1725]" pageId="27" pageNumber="498">
|
||
<emphasis id="B91EEA9CFF9646748E872ABE40A4F90B" bold="true" box="[199,414,1698,1725]" pageId="27" pageNumber="498">Geographic range</emphasis>
|
||
. Possibly endemic to
|
||
<collectingCountry id="F37D761EFF9646748CD52ABF4279F90B" box="[661,835,1699,1725]" name="New Caledonia" pageId="27" pageNumber="498">New Caledonia</collectingCountry>
|
||
(present study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3706505FF9646738E872ADB4231FC47" lastPageId="28" lastPageNumber="499" pageId="27" pageNumber="498" type="discussion">
|
||
<paragraph id="8BD5368EFF9646748E872ADB45FBF823" blockId="27.[151,1437,582,1941]" pageId="27" pageNumber="498">
|
||
<emphasis id="B91EEA9CFF9646748E872ADB400EF957" bold="true" box="[199,308,1735,1761]" pageId="27" pageNumber="498">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C6A4D0DFF9646748F012ADB40E6F957" authority="Valdés & Feliciano & Malaquias, 2023" authorityName="Valdés & Feliciano & Malaquias" authorityYear="2023" box="[321,476,1735,1761]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="27" pageNumber="498" phylum="Mollusca" rank="species" species="charlesi" status="sp. nov.">
|
||
<emphasis id="B91EEA9CFF9646748F012ADB40E6F957" box="[321,476,1735,1761]" italics="true" pageId="27" pageNumber="498">Pupa charlesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91EEA9CFF9646748FA22ADB4300F957" bold="true" box="[482,570,1735,1761]" pageId="27" pageNumber="498">
|
||
<taxonomicNameLabel id="A22D57E7FF9646748FA22ADB4300F957" box="[482,570,1735,1761]" pageId="27" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was recovered as a distinct clade in the phylogenetic analyses and as a distinct species in the species delimitation analysis.
|
||
<taxonomicName id="4C6A4D0DFF9646748CC42AF7421AF8B3" authority="Valdés & Feliciano & Malaquias, 2023" authorityName="Valdés & Feliciano & Malaquias" authorityYear="2023" box="[644,800,1771,1797]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="27" pageNumber="498" phylum="Mollusca" rank="species" species="charlesi" status="sp. nov.">
|
||
<emphasis id="B91EEA9CFF9646748CC42AF7421AF8B3" box="[644,800,1771,1797]" italics="true" pageId="27" pageNumber="498">Pupa charlesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91EEA9CFF9646748D672AF742BAF8B3" bold="true" box="[807,896,1771,1797]" pageId="27" pageNumber="498">
|
||
<taxonomicNameLabel id="A22D57E7FF9646748D672AF742BAF8B3" box="[807,896,1771,1797]" pageId="27" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is externally very similar to
|
||
<taxonomicName id="4C6A4D0DFF9646748A822AF74448F8B2" baseAuthorityName="Reeve" baseAuthorityYear="1842" box="[1218,1394,1771,1796]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="27" pageNumber="498" phylum="Mollusca" rank="species" species="coccinata">
|
||
<emphasis id="B91EEA9CFF9646748A822AF74448F8B2" box="[1218,1394,1771,1796]" italics="true" pageId="27" pageNumber="498">Pupa coccinata</emphasis>
|
||
</taxonomicName>
|
||
but the shells are thinner and wider anteriorly, and the orange spots are typically further apart and less defined. These two species also differ in radular morphology, with
|
||
<taxonomicName id="4C6A4D0DFF9646748C9E2B2F426EF8FB" authority="Valdés & Feliciano & Malaquias, 2023" authorityName="Valdés & Feliciano & Malaquias" authorityYear="2023" box="[734,852,1843,1869]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="27" pageNumber="498" phylum="Mollusca" rank="species" species="charlesi" status="sp. nov.">
|
||
<emphasis id="B91EEA9CFF9646748C9E2B2F426EF8FB" box="[734,852,1843,1869]" italics="true" pageId="27" pageNumber="498">P. charlesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91EEA9CFF9646748D1C2B2F428CF8FB" bold="true" box="[860,950,1843,1869]" pageId="27" pageNumber="498">
|
||
<taxonomicNameLabel id="A22D57E7FF9646748D1C2B2F428CF8FB" box="[860,950,1843,1869]" pageId="27" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
having a 121 × 6.0.6 radular formula in an
|
||
<quantity id="4C929B6BFF9646748ED72B4B41DCF8C7" box="[151,230,1879,1905]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="498" unit="mm" value="18.0">18 mm</quantity>
|
||
long specimen and
|
||
<taxonomicName id="4C6A4D0DFF9646748F862B4B4375F8C6" baseAuthorityName="Reeve" baseAuthorityYear="1842" box="[454,591,1879,1904]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="27" pageNumber="498" phylum="Mollusca" rank="species" species="coccinata">
|
||
<emphasis id="B91EEA9CFF9646748F862B4B4375F8C6" box="[454,591,1879,1904]" italics="true" pageId="27" pageNumber="498">P. coccinata</emphasis>
|
||
</taxonomicName>
|
||
112 × 5.0.
|
||
<quantity id="4C929B6BFF9646748C872B4A43C8F8C7" box="[711,754,1878,1905]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="27" pageNumber="498" unit="in" value="5.0">5 in</quantity>
|
||
a
|
||
<quantity id="4C929B6BFF9646748D4C2B4B4261F8C7" box="[780,859,1879,1905]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="498" unit="mm" value="20.0">20 mm</quantity>
|
||
long specimen; the teeth of
|
||
<taxonomicName id="4C6A4D0DFF9646748AD92B4B4434F8C7" authority="Valdés & Feliciano & Malaquias, 2023" authorityName="Valdés & Feliciano & Malaquias" authorityYear="2023" box="[1177,1294,1879,1905]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="27" pageNumber="498" phylum="Mollusca" rank="species" species="charlesi" status="sp. nov.">
|
||
<emphasis id="B91EEA9CFF9646748AD92B4B4434F8C7" box="[1177,1294,1879,1905]" italics="true" pageId="27" pageNumber="498">P. charlesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22D57E7FF9646748B562B4B44A6F8C7" box="[1302,1436,1879,1905]" pageId="27" pageNumber="498" rank="species">new species</taxonomicNameLabel>
|
||
are much wider than those of
|
||
<taxonomicName id="4C6A4D0DFF9646748FA42B674356F822" baseAuthorityName="Reeve" baseAuthorityYear="1842" box="[484,620,1915,1940]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="27" pageNumber="498" phylum="Mollusca" rank="species" species="coccinata">
|
||
<emphasis id="B91EEA9CFF9646748FA42B674356F822" box="[484,620,1915,1940]" italics="true" pageId="27" pageNumber="498">P. coccinata</emphasis>
|
||
</taxonomicName>
|
||
and have more denticles, similar in size to the cusps.
|
||
</paragraph>
|
||
<caption id="DF156606FF9146738ED82F1C45BEFC32" ID-DOI="http://doi.org/10.5281/zenodo.7860408" ID-Zenodo-Dep="7860408" httpUri="https://zenodo.org/record/7860408/files/figure.png" pageId="28" pageNumber="499" startId="28.[152,255,768,792]" targetBox="[151,1436,181,745]" targetPageId="28" targetType="figure">
|
||
<paragraph id="8BD5368EFF9146738ED82F1C45BEFC32" blockId="28.[151,1437,768,901]" pageId="28" pageNumber="499">
|
||
<emphasis id="B91EEA9CFF9146738ED82F1C4025FCAE" bold="true" box="[152,287,768,792]" pageId="28" pageNumber="499">FIGURE 16</emphasis>
|
||
. Photographs of shell specimens of
|
||
<taxonomicName id="4C6A4D0DFF9146738CD42F1D42DEFCAE" authority="(Barnard, 1963)" baseAuthorityName="Barnard" baseAuthorityYear="1963" box="[660,996,768,792]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="28" pageNumber="499" phylum="Mollusca" rank="species" species="niecaensis">
|
||
<emphasis id="B91EEA9CFF9146738CD42F1D4201FCAE" box="[660,827,769,792]" italics="true" pageId="28" pageNumber="499">Pupa niecaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFB4B7FFF9146738D092F1C42E4FCAE" author="Barnard K. H." box="[841,990,768,792]" pageId="28" pageNumber="499" pagination="201 - 360" refId="ref21595" refString="Barnard K. H. (1963) Contributions to the knowledge of South African marine Mollusca. Part IV. Gastropoda: Prosobranchiata: Rhipidoglossa, Docoglossa. Tectibranchiata. Polyplacophora. Solenogastres. Scaphopoda. Annals of the South African Museum, 47, 201 - 360." type="journal article" year="1963">Barnard, 1963</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(A),
|
||
<taxonomicName id="4C6A4D0DFF9146738A5D2F1D4472FCAF" authority="Raines, 2003" authorityName="Raines" authorityYear="2003" box="[1053,1352,768,793]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="28" pageNumber="499" phylum="Mollusca" rank="species" species="pascuana">
|
||
<emphasis id="B91EEA9CFF9146738A5D2F1D4581FCAE" box="[1053,1211,769,792]" italics="true" pageId="28" pageNumber="499">Pupa pascuana</emphasis>
|
||
<bibRefCitation id="EFFB4B7FFF9146738A822F1C4472FCAF" author="Raines, B. K." box="[1218,1352,768,793]" pageId="28" pageNumber="499" pagination="51 - 53" refId="ref25687" refString="Raines, B. K. (2003) Pupa pascuana: A new species of Acteonidae from Easter Island. La Conchiglia, 34, 51 - 53." type="journal article" year="2003">Raines, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
(B) and
|
||
<taxonomicName id="4C6A4D0DFF9146738ED72F39401DFC8A" authority="Valdés & Feliciano & Malaquias, 2023" authorityName="Valdés & Feliciano & Malaquias" authorityYear="2023" box="[151,295,804,828]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="28" pageNumber="499" phylum="Mollusca" rank="species" species="charlesi" status="sp. nov.">
|
||
<emphasis id="B91EEA9CFF9146738ED72F39401DFC8A" box="[151,295,804,828]" italics="true" pageId="28" pageNumber="499">Pupa charlesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91EEA9CFF9146738F6E2F3940BAFC8B" bold="true" box="[302,384,805,829]" pageId="28" pageNumber="499">
|
||
<taxonomicNameLabel id="A22D57E7FF9146738F6E2F3940BAFC8B" box="[302,384,805,829]" pageId="28" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(C–E).
|
||
<emphasis id="B91EEA9CFF9146738F952F3840DDFC8A" bold="true" box="[469,487,804,828]" pageId="28" pageNumber="499">A</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C6A4D0DFF9146738C392F3942F1FC8A" authority="(Barnard, 1963)" baseAuthorityName="Barnard" baseAuthorityYear="1963" box="[633,971,804,828]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="28" pageNumber="499" phylum="Mollusca" rank="species" species="niecaensis">
|
||
<emphasis id="B91EEA9CFF9146738C392F39421AFC8A" box="[633,800,805,828]" italics="true" pageId="28" pageNumber="499">Pupa niecaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFB4B7FFF9146738D6F2F3842FEFC8A" author="Barnard K. H." box="[815,964,804,828]" pageId="28" pageNumber="499" pagination="201 - 360" refId="ref21595" refString="Barnard K. H. (1963) Contributions to the knowledge of South African marine Mollusca. Part IV. Gastropoda: Prosobranchiata: Rhipidoglossa, Docoglossa. Tectibranchiata. Polyplacophora. Solenogastres. Scaphopoda. Annals of the South African Museum, 47, 201 - 360." type="journal article" year="1963">Barnard, 1963</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(SAM A6553), ©SAM, photo by R. Adams.
|
||
<emphasis id="B91EEA9CFF9146738ED72F554192FCD7" bold="true" box="[151,168,841,865]" pageId="28" pageNumber="499">B</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C6A4D0DFF9146738F792F55435FFCD7" authority="Raines, 2003" authorityName="Raines" authorityYear="2003" box="[313,613,840,865]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="28" pageNumber="499" phylum="Mollusca" rank="species" species="pascuana">
|
||
<emphasis id="B91EEA9CFF9146738F792F5540EDFCD6" box="[313,471,841,864]" italics="true" pageId="28" pageNumber="499">Pupa pascuana</emphasis>
|
||
<bibRefCitation id="EFFB4B7FFF9146738F9F2F54435FFCD7" author="Raines, B. K." box="[479,613,840,865]" pageId="28" pageNumber="499" pagination="51 - 53" refId="ref25687" refString="Raines, B. K. (2003) Pupa pascuana: A new species of Acteonidae from Easter Island. La Conchiglia, 34, 51 - 53." type="journal article" year="2003">Raines, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
(LACM 2954).
|
||
<emphasis id="B91EEA9CFF9146738D4F2F544205FCD7" bold="true" box="[783,831,840,865]" pageId="28" pageNumber="499">C–E</emphasis>
|
||
. Specimens of
|
||
<taxonomicName id="4C6A4D0DFF9146738D9F2F554554FCD6" authority="Valdés & Feliciano & Malaquias, 2023" authorityName="Valdés & Feliciano & Malaquias" authorityYear="2023" box="[991,1134,840,864]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="28" pageNumber="499" phylum="Mollusca" rank="species" species="charlesi" status="sp. nov.">
|
||
<emphasis id="B91EEA9CFF9146738D9F2F554554FCD6" box="[991,1134,840,864]" italics="true" pageId="28" pageNumber="499">Pupa charlesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91EEA9CFF9146738A352F5545FDFCD7" bold="true" box="[1141,1223,841,865]" pageId="28" pageNumber="499">
|
||
<taxonomicNameLabel id="A22D57E7FF9146738A352F5545FDFCD7" box="[1141,1223,841,865]" pageId="28" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Koumac, New Caledonia, (C) MHNH IM-2013-86154, (D) MHNH IM-2013-86159, (E) MHNH IM-2013-86158.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD5368EFF9146738ED72FAF4231FC47" blockId="28.[151,1436,947,1009]" pageId="28" pageNumber="499">
|
||
A review of the literature has revealed no available names with a description matching that of
|
||
<taxonomicName id="4C6A4D0DFF9146738A942FA84448FC7B" authority="Valdés & Feliciano & Malaquias, 2023" authorityName="Valdés & Feliciano & Malaquias" authorityYear="2023" box="[1236,1394,947,973]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="28" pageNumber="499" phylum="Mollusca" rank="species" species="charlesi" status="sp. nov.">
|
||
<emphasis id="B91EEA9CFF9146738A942FA84448FC7B" box="[1236,1394,947,973]" italics="true" pageId="28" pageNumber="499">Pupa charlesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91EEA9CFF9146738B3B2FAF41F2FC47" bold="true" pageId="28" pageNumber="499">
|
||
<taxonomicNameLabel id="A22D57E7FF9146738B3B2FAF41F2FC47" pageId="28" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and therefore it is here described as a new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |