treatments-xml/data/EA/68/F5/EA68F5315C664B7CE38264F17DE478EA.xml

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<mods:title id="367410549E5A90CC6132B8E28AC5BE36">Unesconia coibensis, gen. et sp. nov., a miniature sponge-associated shrimp from a biodiversity hotspot in the tropical eastern Pacific (Decapoda: Palaemonidae)</mods:title>
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<taxonomicName id="A5C13FA45C664B73E38264F17EE9792A" ID-CoL="761b249c-983b-4e16-96ed-26325eee9709" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[151,398,1510,1537]" class="Malacostraca" family="Palaemonidae" genus="Unesconia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="coibensis" status="sp. nov.">
<emphasis id="50B598355C664B73E38264F17EE9792A" bold="true" box="[151,398,1510,1537]" italics="true" pageId="4" pageNumber="119">Unesconia coibensis</emphasis>
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<paragraph id="627E44275C664B73E382671D7E6A790F" blockId="4.[151,499,1510,1572]" box="[151,269,1546,1572]" pageId="4" pageNumber="119">
<figureCitation id="FAFA58A25C664B73E382671D7E6A790F" box="[151,269,1546,1572]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="5.[151,250,1133,1158]" captionStartId-1="6.[151,250,1756,1781]" captionStartId-2="7.[151,250,1641,1666]" captionStartId-3="8.[151,250,1177,1202]" captionStartId-4="9.[151,250,1592,1617]" captionStartId-5="10.[151,250,1647,1672]" captionStartId-6="11.[151,250,1384,1409]" captionStartId-7="12.[151,250,1791,1816]" captionStartId-8="13.[151,250,1044,1069]" captionStartId-9="14.[151,250,1887,1912]" captionTargetBox-0="[177,1368,335,1076]" captionTargetBox-1="[227,1363,425,1711]" captionTargetBox-2="[165,1423,185,1615]" captionTargetBox-3="[173,1387,192,1106]" captionTargetBox-4="[163,1425,417,1536]" captionTargetBox-5="[183,1412,211,1601]" captionTargetBox-6="[151,1436,206,1331]" captionTargetBox-7="[196,1382,196,1757]" captionTargetBox-8="[151,1436,174,1024]" captionTargetBox-9="[151,1436,517,1867]" captionTargetId-0="figure@5.[151,1436,301,1114]" captionTargetId-1="figure@6.[214,1372,409,1736]" captionTargetId-2="figure@7.[151,1435,174,1622]" captionTargetId-3="figure@8.[151,1435,174,1158]" captionTargetId-4="figure@9.[151,1435,393,1548]" captionTargetId-5="figure@10.[157,1429,174,1628]" captionTargetId-6="figure@11.[151,1436,206,1331]" captionTargetId-7="figure@12.[184,1403,174,1772]" captionTargetId-8="figure@13.[151,1436,174,1024]" captionTargetId-9="figure@14.[151,1436,517,1867]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionTargetPageId-9="14" captionText-0="FIGURE 1. Unesconia coibensis gen. et sp. nov.: holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51926): habitus, lateral view." captionText-1="FIGURE 2. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–E]; holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from the same locality (FLMNH UF 51926) [F]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [G–I]: A—frontal region, dorsal view [left eye somewhat abnormal, i.e. smaller and partly shrunk, in the illustrated specimen (see Fig. 10B), shown as normal based on the right eye]; B—same, lateral view; C—sixth pleonite and tail fan, dorsal view; D—telson, posterior region, dorsal view; E—uropodal exopod, detail of lateral angle of diaeresis, dorsal view; F—frontal region of carapace and left eyestalk, lateral view; G—frontal region, dorsal view; H—same, lateral view; I—uropodal exopod, detail of lateral angle of diaeresis, dorsal view." captionText-2="FIGURE 3. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right mandible, mesial view; B—left mandible, lateral view; C—same, mesial view; D—maxillule, lateral view; E—maxilla, lateral view; F—first maxilliped, lateral view; G—second maxilliped, lateral view; H—third maxilliped, lateral view; I—epistome, ventral view." captionText-3="FIGURE 4. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right first pereiopod, lateral view; B—same, mesial view; C—same, chela, mesial view; D—same, chela with fingers opened, setae omitted for clarity, dorsolateral view; E—same, chela with fingers opened, setae omitted for clarity, ventromesial view." captionText-4="FIGURE 5. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—major (left) second pereiopod, mesial (dorsal) view; B—same, carpus and chela with fingers closed, lateral (ventral) view; C—same, carpus and chela with fingers opened, lateral (ventral) view; D—same, chela fingers opened, setae omitted for clarity, lateral (ventral) view; E—minor (right) second pereiopod, lateral (ventral) view; F—same, carpus and chela, lateral (ventral) view." captionText-5="FIGURE 6. Unesconia coibensis gen. et sp. nov., allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927): A—major (left) second pereiopod, dorsomesial view; B—same, carpus and chela with fingers closed, lateral (ventral) view; C–H—same, chela fingers opened viewed from various angles mesially (C–F) or laterally (G, H), setae omitted for clarity." captionText-6="FIGURE 7. Unesconia coibensis gen. et sp. nov., allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927): A—major (left) second pereiopod, chela fingers opened viewed from various angles mesially (A–C) or laterally (D). See also Fig. 6." captionText-7="FIGURE 8. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–H]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [I, J]: A—third pereiopod, lateral view; B—same, propodus and dactylus, mesial view; C—same, distal portion of propodus and dactylus, lateral view; D—fourth pereiopod, lateral view; E—same, propodus and dactylus, mesial view; F—same, distal portion of propodus and dactylus, lateral view; G—fifth pereiopod, lateral view; H—same, propodus and dactylus, mesial view; I—male second pleopod, posterior (“lateral”) view; J—same, detail of appendix masculina and appendix interna, drawn without scale." captionText-8="FIGURE 9. Unesconia coibensis gen. et sp. nov.: holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51926), shrimp alive, dorsal view. Photograph by the author." captionText-9="FIGURE 10. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928), shrimp alive, dorsal [A] and lateral [B] views; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927), shrimp alive, dorsal view, right (minor) cheliped regenerating [C]. Photographs by the author." figureDoi-0="http://doi.org/10.5281/zenodo.3635670" figureDoi-1="http://doi.org/10.5281/zenodo.3635672" figureDoi-2="http://doi.org/10.5281/zenodo.3635674" figureDoi-3="http://doi.org/10.5281/zenodo.3635676" figureDoi-4="http://doi.org/10.5281/zenodo.3635678" figureDoi-5="http://doi.org/10.5281/zenodo.3635680" figureDoi-6="http://doi.org/10.5281/zenodo.3635682" figureDoi-7="http://doi.org/10.5281/zenodo.3635684" figureDoi-8="http://doi.org/10.5281/zenodo.3635686" figureDoi-9="http://doi.org/10.5281/zenodo.3635688" httpUri-0="https://zenodo.org/record/3635670/files/figure.png" httpUri-1="https://zenodo.org/record/3635672/files/figure.png" httpUri-2="https://zenodo.org/record/3635674/files/figure.png" httpUri-3="https://zenodo.org/record/3635676/files/figure.png" httpUri-4="https://zenodo.org/record/3635678/files/figure.png" httpUri-5="https://zenodo.org/record/3635680/files/figure.png" httpUri-6="https://zenodo.org/record/3635682/files/figure.png" httpUri-7="https://zenodo.org/record/3635684/files/figure.png" httpUri-8="https://zenodo.org/record/3635686/files/figure.png" httpUri-9="https://zenodo.org/record/3635688/files/figure.png" pageId="4" pageNumber="119">Figs. 1–10</figureCitation>
</paragraph>
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<emphasis id="50B598355C664B73E38267457E597947" bold="true" box="[151,318,1617,1644]" pageId="4" pageNumber="119">Type material</emphasis>
. 
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: ovigerous female (pocl 2.0 mm, cl 
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), 
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, 
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, 
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coast, 
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, 
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Canales 
<collectorName id="CF3421F15C664B73E2C267617D2379BB" box="[471,580,1654,1680]" pageId="4" pageNumber="119">de Afuera</collectorName>
</location>
(channel between Isla 
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and 
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<collectingMunicipality id="821ADE5D5C664B73E0E167617B3179BA" box="[1012,1110,1654,1681]" pageId="4" pageNumber="119">Afuerita</collectingMunicipality>
), channel between small islet and northern side of Isla 
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, 
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, shallow subtidal sand flat with abundance of fragmented coral rubble and living colonies of 
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<emphasis id="50B598355C664B73E10967A97DC579F3" box="[540,674,1726,1752]" italics="true" pageId="4" pageNumber="119">Pocillopora</emphasis>
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, depth: 
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at low tide, in large piece of coral rubble, presumably associated with yellowish incrusting sponge, leg. 
<collectorName id="CF3421F15C664B73E1AF67F57C7879D7" box="[698,799,1762,1788]" pageId="4" pageNumber="119">A. Anker</collectorName>
&amp; 
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, 
<date id="167F62E75C664B73E0E167F57BDE79D7" box="[1012,1209,1762,1788]" pageId="4" pageNumber="119" value="2019-02-19">
<collectingDate id="063B9B0F5C664B73E0E167F57BDE79D7" box="[1012,1209,1762,1788]" pageId="4" pageNumber="119" value="2019-02-19">19 February 2019</collectingDate>
</date>
[fcn PAN-99]
</materialsCitation>
. 
<materialsCitation id="D2A94E7A5C664B73E67467F57D79786F" ID-GBIF-Occurrence="2562577887" collectingDate="2019-02-19" collectionCode="FLMNH" collectorName="de Afuera &amp; A. Anker &amp; P. G. P. Pachelle" country="Panama" county="Afuera" elevation="2" latitude="7.695249" location="Pacific" longLatPrecision="1" longitude="-81.63571" municipality="Afuerita" pageId="4" pageNumber="119" specimenCode="UF 51927" specimenCount="1" typeStatus="allotype">
<typeStatus id="BD7AFA855C664B73E67467F57FAC780A" pageId="4" pageNumber="119" type="allotype">Allotype</typeStatus>
: male (pocl 
<quantity id="A539E9C25C664B73E24766117ECE780A" box="[338,425,1798,1825]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="119" unit="mm" value="1.7">1.7 mm</quantity>
, cl 
<quantity id="A539E9C25C664B73E2DB66117D42780A" box="[462,549,1798,1825]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="119" unit="mm" value="2.8">2.8 mm</quantity>
, minor second cheliped regenerating), 
<collectionCode id="04D0DCE25C664B73E0C166107B51780A" box="[980,1078,1799,1825]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="4" pageNumber="119" type="Museum">FLMNH</collectionCode>
<specimenCode id="3267EC5C5C664B73E72966107BC8780B" box="[1084,1199,1797,1825]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="4" pageNumber="119" type="Museum">UF 51927</specimenCode>
, same collection data as for the holotype [fcn PAN-100]
</materialsCitation>
. 
<materialsCitation id="D2A94E7A5C664B73E13F663C7C2D78A7" ID-GBIF-Occurrence="2562577885" collectingDate="2019-02-19" collectionCode="FLMNH" collectorName="de Afuera &amp; A. Anker &amp; P. G. P. Pachelle" country="Panama" county="Afuera" elevation="2" latitude="7.695249" location="Pacific" longLatPrecision="1" longitude="-81.63571" municipality="Afuerita" pageId="4" pageNumber="119" specimenCode="UF 51928, UF 51929" specimenCount="2" typeStatus="paratype">
<typeStatus id="BD7AFA855C664B73E13F663C7DFC786E" box="[554,667,1835,1861]" pageId="4" pageNumber="119" type="paratype">Paratypes</typeStatus>
: 1 ovigerous female (pocl 
<quantity id="A539E9C25C664B73E0DC663D7B45786E" box="[969,1058,1834,1861]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="119" unit="mm" value="2.1">2.1 mm</quantity>
, cl 3.0 mm), 
<collectionCode id="04D0DCE25C664B73E7AC663C7A7C786E" box="[1209,1307,1835,1861]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="4" pageNumber="119" type="Museum">FLMNH</collectionCode>
<specimenCode id="3267EC5C5C664B73E636663C7AFF786F" box="[1315,1432,1833,1861]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="4" pageNumber="119" type="Museum">UF 51928</specimenCode>
, same collection data as for the holotype [fcn PAN-98]; 1 ovigerous female (pocl 
<quantity id="A539E9C25C664B73E73666597B1C7842" box="[1059,1147,1870,1897]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="119" unit="mm" value="1.7">1.7 mm</quantity>
, cl 
<quantity id="A539E9C25C664B73E7B766597B9B7842" box="[1186,1276,1870,1897]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="119" unit="mm" value="2.8">2.8 mm</quantity>
), 
<collectionCode id="04D0DCE25C664B73E61A66587A167842" box="[1295,1393,1871,1897]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="4" pageNumber="119" type="Museum">FLMNH</collectionCode>
<specimenCode id="3267EC5C5C664B73E66D66587F8778A7" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="4" pageNumber="119" type="Museum">UF 51929</specimenCode>
, same collection data as for the 
<typeStatus id="BD7AFA855C664B73E15366657DCF78A7" box="[582,680,1906,1932]" pageId="4" pageNumber="119" type="holotype">holotype</typeStatus>
[fcn PAN-97]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2ADB17AC5C664B7DE3D266827CA97882" lastPageId="10" lastPageNumber="125" pageId="4" pageNumber="119" type="description">
<paragraph id="627E44275C664B72E3D266827DD27E34" blockId="4.[151,1437,1617,2041]" lastBlockId="5.[151,1436,153,287]" lastPageId="5" lastPageNumber="120" pageId="4" pageNumber="119">
<emphasis id="50B598355C664B73E3D266827E36789B" bold="true" box="[199,337,1941,1968]" pageId="4" pageNumber="119">Description</emphasis>
. Small-sized pontoniine shrimp with dorsoventrally somewhat flattened body (
<figureCitation id="FAFA58A25C664B73E7C266817A7D789B" box="[1239,1306,1942,1968]" captionStart="FIGURE 1" captionStartId="5.[151,250,1133,1158]" captionTargetBox="[177,1368,335,1076]" captionTargetId="figure@5.[151,1436,301,1114]" captionTargetPageId="5" captionText="FIGURE 1. Unesconia coibensis gen. et sp. nov.: holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51926): habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635670" httpUri="https://zenodo.org/record/3635670/files/figure.png" pageId="4" pageNumber="119">Fig. 1</figureCitation>
). Carapace smooth, glabrous, neither setose nor pubescent (
<figureCitation id="FAFA58A25C664B73E1B866AD7D8978FF" box="[685,750,1978,2004]" captionStart="FIGURE 1" captionStartId="5.[151,250,1133,1158]" captionTargetBox="[177,1368,335,1076]" captionTargetId="figure@5.[151,1436,301,1114]" captionTargetPageId="5" captionText="FIGURE 1. Unesconia coibensis gen. et sp. nov.: holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51926): habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635670" httpUri="https://zenodo.org/record/3635670/files/figure.png" pageId="4" pageNumber="119">Fig. 1</figureCitation>
). Rostrum well developed; dorsal carina strong, dentate, with typically four teeth, sometimes only two teeth; ventral carina unarmed; tip reaching to at least distal margin of first article of antennular peduncle, usually to end of antennular peduncle; lateral carinae proximally expanded, each side with one strong, anteriorly pointing, simple supraorbital tooth (
<figureCitation id="FAFA58A25C674B72E04C61AA7CB57FFC" box="[857,978,189,215]" captionStart="FIGURE 2" captionStartId="6.[151,250,1756,1781]" captionTargetBox="[227,1363,425,1711]" captionTargetId="figure@6.[214,1372,409,1736]" captionTargetPageId="6" captionText="FIGURE 2. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–E]; holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from the same locality (FLMNH UF 51926) [F]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [G–I]: A—frontal region, dorsal view [left eye somewhat abnormal, i.e. smaller and partly shrunk, in the illustrated specimen (see Fig. 10B), shown as normal based on the right eye]; B—same, lateral view; C—sixth pleonite and tail fan, dorsal view; D—telson, posterior region, dorsal view; E—uropodal exopod, detail of lateral angle of diaeresis, dorsal view; F—frontal region of carapace and left eyestalk, lateral view; G—frontal region, dorsal view; H—same, lateral view; I—uropodal exopod, detail of lateral angle of diaeresis, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3635672" httpUri="https://zenodo.org/record/3635672/files/figure.png" pageId="5" pageNumber="120">Fig. 2A, B</figureCitation>
, F–H). Orbit well developed; inferior orbital angle distinct; epigastric and hepatic teeth absent; antennal tooth strong, acute; pterygostomial angle distinctly protruding, acute or subacute (
<figureCitation id="FAFA58A25C674B72E2F860127D007E34" box="[493,615,261,287]" captionStart="FIGURE 2" captionStartId="6.[151,250,1756,1781]" captionTargetBox="[227,1363,425,1711]" captionTargetId="figure@6.[214,1372,409,1736]" captionTargetPageId="6" captionText="FIGURE 2. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–E]; holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from the same locality (FLMNH UF 51926) [F]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [G–I]: A—frontal region, dorsal view [left eye somewhat abnormal, i.e. smaller and partly shrunk, in the illustrated specimen (see Fig. 10B), shown as normal based on the right eye]; B—same, lateral view; C—sixth pleonite and tail fan, dorsal view; D—telson, posterior region, dorsal view; E—uropodal exopod, detail of lateral angle of diaeresis, dorsal view; F—frontal region of carapace and left eyestalk, lateral view; G—frontal region, dorsal view; H—same, lateral view; I—uropodal exopod, detail of lateral angle of diaeresis, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3635672" httpUri="https://zenodo.org/record/3635672/files/figure.png" pageId="5" pageNumber="120">Fig. 2A, B</figureCitation>
, F–H).
</paragraph>
<caption id="36BE14AF5C674B72E382657A7CFD7B81" ID-DOI="http://doi.org/10.5281/zenodo.3635670" ID-Zenodo-Dep="3635670" httpUri="https://zenodo.org/record/3635670/files/figure.png" pageId="5" pageNumber="120" startId="5.[151,250,1133,1158]" targetBox="[177,1368,335,1076]" targetPageId="5">
<paragraph id="627E44275C674B72E382657A7CFD7B81" blockId="5.[151,1436,1133,1194]" pageId="5" pageNumber="120">
<emphasis id="50B598355C674B72E382657A7E6A7BAD" bold="true" box="[151,269,1133,1158]" pageId="5" pageNumber="120">FIGURE 1</emphasis>
. 
<taxonomicName id="A5C13FA45C674B72E20F65787E8D7BAD" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[282,490,1134,1158]" class="Malacostraca" family="Palaemonidae" genus="Unesconia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="coibensis" status="gen. et sp. nov.">
<emphasis id="50B598355C674B72E20F65787E8D7BAD" box="[282,490,1134,1158]" italics="true" pageId="5" pageNumber="120">Unesconia coibensis</emphasis>
</taxonomicName>
<emphasis id="50B598355C674B72E2E465797DE97BAD" bold="true" box="[497,654,1134,1158]" pageId="5" pageNumber="120">
<taxonomicNameLabel id="4B86254E5C674B72E2E465797DE97BAD" box="[497,654,1134,1158]" pageId="5" pageNumber="120" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
: holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51926): habitus, lateral view.
</paragraph>
</caption>
<paragraph id="627E44275C674B72E3D265CF7B607AC4" blockId="5.[151,1437,1240,2023]" pageId="5" pageNumber="120">
All thoracic sternites unarmed, fourth without median process. Pleon smooth, glabrous; posteroventral margin of first four pleura rounded, that of fifth pleuron angular, that of sixth pleonite subacutely produced (
<figureCitation id="FAFA58A25C674B72E7EC65EB7A5D7A3D" box="[1273,1338,1276,1302]" captionStart="FIGURE 1" captionStartId="5.[151,250,1133,1158]" captionTargetBox="[177,1368,335,1076]" captionTargetId="figure@5.[151,1436,301,1114]" captionTargetPageId="5" captionText="FIGURE 1. Unesconia coibensis gen. et sp. nov.: holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51926): habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635670" httpUri="https://zenodo.org/record/3635670/files/figure.png" pageId="5" pageNumber="120">Fig. 1</figureCitation>
). Telson relatively short and broad, about 1.5 times as long as maximal (proximal) width; dorsal surface with two pairs of very stout, long cuspidate setae, inserted close to lateral margin and reaching to or well beyond it, anterior pair inserted at about 0.3 length of telson, posterior pair at about 0.7 length of telson; posterior margin rounded, with three pairs of robust, elongate spiniform setae; most lateral spiniform setae about as long as dorsal cuspidate setae; intermediate spiniform setae longest, more than twice as long as lateral, slightly stouter; median spiniform setae slenderest, longer than lateral but much shorter than intermediate (
<figureCitation id="FAFA58A25C674B72E06A64C37C9D7AC4" box="[895,1018,1492,1519]" captionStart="FIGURE 2" captionStartId="6.[151,250,1756,1781]" captionTargetBox="[227,1363,425,1711]" captionTargetId="figure@6.[214,1372,409,1736]" captionTargetPageId="6" captionText="FIGURE 2. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–E]; holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from the same locality (FLMNH UF 51926) [F]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [G–I]: A—frontal region, dorsal view [left eye somewhat abnormal, i.e. smaller and partly shrunk, in the illustrated specimen (see Fig. 10B), shown as normal based on the right eye]; B—same, lateral view; C—sixth pleonite and tail fan, dorsal view; D—telson, posterior region, dorsal view; E—uropodal exopod, detail of lateral angle of diaeresis, dorsal view; F—frontal region of carapace and left eyestalk, lateral view; G—frontal region, dorsal view; H—same, lateral view; I—uropodal exopod, detail of lateral angle of diaeresis, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3635672" httpUri="https://zenodo.org/record/3635672/files/figure.png" pageId="5" pageNumber="120">Fig. 2C, D</figureCitation>
).
</paragraph>
<paragraph id="627E44275C674B72E3D264EE7A0C7939" blockId="5.[151,1437,1240,2023]" box="[199,1387,1528,1555]" pageId="5" pageNumber="120">
Eye peduncles robust; cornea well developed, globular; accessory eye (Nebenauge) small (
<figureCitation id="FAFA58A25C674B72E7D464EF7A397939" box="[1217,1374,1528,1555]" captionStart="FIGURE 2" captionStartId="6.[151,250,1756,1781]" captionTargetBox="[227,1363,425,1711]" captionTargetId="figure@6.[214,1372,409,1736]" captionTargetPageId="6" captionText="FIGURE 2. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–E]; holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from the same locality (FLMNH UF 51926) [F]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [G–I]: A—frontal region, dorsal view [left eye somewhat abnormal, i.e. smaller and partly shrunk, in the illustrated specimen (see Fig. 10B), shown as normal based on the right eye]; B—same, lateral view; C—sixth pleonite and tail fan, dorsal view; D—telson, posterior region, dorsal view; E—uropodal exopod, detail of lateral angle of diaeresis, dorsal view; F—frontal region of carapace and left eyestalk, lateral view; G—frontal region, dorsal view; H—same, lateral view; I—uropodal exopod, detail of lateral angle of diaeresis, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3635672" httpUri="https://zenodo.org/record/3635672/files/figure.png" pageId="5" pageNumber="120">Fig. 2A, B, G</figureCitation>
).
</paragraph>
<paragraph id="627E44275C674B72E3D2670B7DE079C0" blockId="5.[151,1437,1240,2023]" pageId="5" pageNumber="120">
Antennule robust, stout, short, with first article longest and widest, ending in stout, simple, sharp distolateral tooth; stylocerite small, well developed, acute distally; second and third articles very short; lateral flagellum with short accessory ramus and several groups of aesthetascs (
<figureCitation id="FAFA58A25C674B72E00667737CA87954" box="[787,975,1636,1663]" captionStart="FIGURE 2" captionStartId="6.[151,250,1756,1781]" captionTargetBox="[227,1363,425,1711]" captionTargetId="figure@6.[214,1372,409,1736]" captionTargetPageId="6" captionText="FIGURE 2. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–E]; holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from the same locality (FLMNH UF 51926) [F]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [G–I]: A—frontal region, dorsal view [left eye somewhat abnormal, i.e. smaller and partly shrunk, in the illustrated specimen (see Fig. 10B), shown as normal based on the right eye]; B—same, lateral view; C—sixth pleonite and tail fan, dorsal view; D—telson, posterior region, dorsal view; E—uropodal exopod, detail of lateral angle of diaeresis, dorsal view; F—frontal region of carapace and left eyestalk, lateral view; G—frontal region, dorsal view; H—same, lateral view; I—uropodal exopod, detail of lateral angle of diaeresis, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3635672" httpUri="https://zenodo.org/record/3635672/files/figure.png" pageId="5" pageNumber="120">Fig. 2A, B, G, H</figureCitation>
). Antenna with relatively small basicerite armed with small subacute tooth on distoventral margin; carpocerite short, not reaching distal margin of first article of antennular peduncle; scaphocerite well developed, with broad blade and strong distolateral tooth, reaching far beyond end of antennule (
<figureCitation id="FAFA58A25C674B72E2AE67C77D1D79C0" box="[443,634,1744,1771]" captionStart="FIGURE 2" captionStartId="6.[151,250,1756,1781]" captionTargetBox="[227,1363,425,1711]" captionTargetId="figure@6.[214,1372,409,1736]" captionTargetPageId="6" captionText="FIGURE 2. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–E]; holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from the same locality (FLMNH UF 51926) [F]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [G–I]: A—frontal region, dorsal view [left eye somewhat abnormal, i.e. smaller and partly shrunk, in the illustrated specimen (see Fig. 10B), shown as normal based on the right eye]; B—same, lateral view; C—sixth pleonite and tail fan, dorsal view; D—telson, posterior region, dorsal view; E—uropodal exopod, detail of lateral angle of diaeresis, dorsal view; F—frontal region of carapace and left eyestalk, lateral view; G—frontal region, dorsal view; H—same, lateral view; I—uropodal exopod, detail of lateral angle of diaeresis, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3635672" httpUri="https://zenodo.org/record/3635672/files/figure.png" pageId="5" pageNumber="120">Fig. 2A, B, G, H</figureCitation>
).
</paragraph>
<paragraph id="627E44275C674B71E3D267E37A417EA0" blockId="5.[151,1437,1240,2023]" lastBlockId="6.[151,1436,153,396]" lastPageId="6" lastPageNumber="121" pageId="5" pageNumber="120">
Mouthparts of non-filtering 
<typeStatus id="BD7AFA855C674B72E11267E27D5C7824" box="[519,571,1781,1807]" pageId="5" pageNumber="120">type</typeStatus>
. Mandible without palp; incisor process with at least two distal teeth; molar process with distal micro-serration and fields of short setae (
<figureCitation id="FAFA58A25C674B72E028660F7C197819" box="[829,894,1816,1842]" captionStart="FIGURE 3" captionStartId="7.[151,250,1641,1666]" captionTargetBox="[165,1423,185,1615]" captionTargetId="figure@7.[151,1435,174,1622]" captionTargetPageId="7" captionText="FIGURE 3. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right mandible, mesial view; B—left mandible, lateral view; C—same, mesial view; D—maxillule, lateral view; E—maxilla, lateral view; F—first maxilliped, lateral view; G—second maxilliped, lateral view; H—third maxilliped, lateral view; I—epistome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3635674" httpUri="https://zenodo.org/record/3635674/files/figure.png" pageId="5" pageNumber="120">Fig. 3</figureCitation>
A–C). Maxillule with both laciniae fairly broad; palp bilobed distally, ventral lobe more developed, with short hook-shaped seta, dorsal lobe short, without seta (
<figureCitation id="FAFA58A25C674B72E67A662B7FDA7851" captionStart="FIGURE 3" captionStartId="7.[151,250,1641,1666]" captionTargetBox="[165,1423,185,1615]" captionTargetId="figure@7.[151,1435,174,1622]" captionTargetPageId="7" captionText="FIGURE 3. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right mandible, mesial view; B—left mandible, lateral view; C—same, mesial view; D—maxillule, lateral view; E—maxilla, lateral view; F—first maxilliped, lateral view; G—second maxilliped, lateral view; H—third maxilliped, lateral view; I—epistome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3635674" httpUri="https://zenodo.org/record/3635674/files/figure.png" pageId="5" pageNumber="120">Fig. 3D</figureCitation>
). Maxilla with palp (endopod) moderately broad, not subdivided; endite with deep cleft; scaphognathite fairly broad (
<figureCitation id="FAFA58A25C674B72E3F366937E5A78B5" box="[230,317,1924,1950]" captionStart="FIGURE 3" captionStartId="7.[151,250,1641,1666]" captionTargetBox="[165,1423,185,1615]" captionTargetId="figure@7.[151,1435,174,1622]" captionTargetPageId="7" captionText="FIGURE 3. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right mandible, mesial view; B—left mandible, lateral view; C—same, mesial view; D—maxillule, lateral view; E—maxilla, lateral view; F—first maxilliped, lateral view; G—second maxilliped, lateral view; H—third maxilliped, lateral view; I—epistome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3635674" httpUri="https://zenodo.org/record/3635674/files/figure.png" pageId="5" pageNumber="120">Fig. 3E</figureCitation>
). First maxilliped with palp (endopod) narrow, short, tapering distally, not subdivided; endite broad, with small notch indicating subdivision; exopod short, caridean lobe well-developed, broad; epipod moderately developed, bilobed (
<figureCitation id="FAFA58A25C674B72E29666DB7EBC78CD" box="[387,475,1996,2022]" captionStart="FIGURE 3" captionStartId="7.[151,250,1641,1666]" captionTargetBox="[165,1423,185,1615]" captionTargetId="figure@7.[151,1435,174,1622]" captionTargetPageId="7" captionText="FIGURE 3. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right mandible, mesial view; B—left mandible, lateral view; C—same, mesial view; D—maxillule, lateral view; E—maxilla, lateral view; F—first maxilliped, lateral view; G—second maxilliped, lateral view; H—third maxilliped, lateral view; I—epistome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3635674" httpUri="https://zenodo.org/record/3635674/files/figure.png" pageId="5" pageNumber="120">Fig. 3F</figureCitation>
). Second maxilliped not especially modified; shape of endopod typical for family; exopod moderately long; epipod moderately developed, subquadrate, without podobranch (
<figureCitation id="FAFA58A25C644B71E78B618E7B9E7F98" box="[1182,1273,153,179]" captionStart="FIGURE 3" captionStartId="7.[151,250,1641,1666]" captionTargetBox="[165,1423,185,1615]" captionTargetId="figure@7.[151,1435,174,1622]" captionTargetPageId="7" captionText="FIGURE 3. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right mandible, mesial view; B—left mandible, lateral view; C—same, mesial view; D—maxillule, lateral view; E—maxilla, lateral view; F—first maxilliped, lateral view; G—second maxilliped, lateral view; H—third maxilliped, lateral view; I—epistome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3635674" httpUri="https://zenodo.org/record/3635674/files/figure.png" pageId="6" pageNumber="121">Fig. 3G</figureCitation>
). Third maxilliped pediform, moderately stout; lateral plate of coxa broadly rounded, not protruding; antepenultimate article (ischiomerus) and basis distinct, separated from each other by narrow complete suture; antepenultimate article with rounded lateral ridge, about 4.5 times as long as wide; penultimate article somewhat shorter than antepenultimate article; ultimate article shortest, about 0.6 times length of penultimate article, tapering distally; exopod falling short of distal margin of antepenultimate article; arthrobranch apparently absent (or greatly reduced and detached during dissection) (
<figureCitation id="FAFA58A25C644B71E23560667E1D7EA0" box="[288,378,369,395]" captionStart="FIGURE 3" captionStartId="7.[151,250,1641,1666]" captionTargetBox="[165,1423,185,1615]" captionTargetId="figure@7.[151,1435,174,1622]" captionTargetPageId="7" captionText="FIGURE 3. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right mandible, mesial view; B—left mandible, lateral view; C—same, mesial view; D—maxillule, lateral view; E—maxilla, lateral view; F—first maxilliped, lateral view; G—second maxilliped, lateral view; H—third maxilliped, lateral view; I—epistome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3635674" httpUri="https://zenodo.org/record/3635674/files/figure.png" pageId="6" pageNumber="121">Fig. 3H</figureCitation>
). Epistome (median lip) unarmed (
<figureCitation id="FAFA58A25C644B71E01560667C297EA0" box="[768,846,369,395]" captionStart="FIGURE 3" captionStartId="7.[151,250,1641,1666]" captionTargetBox="[165,1423,185,1615]" captionTargetId="figure@7.[151,1435,174,1622]" captionTargetPageId="7" captionText="FIGURE 3. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right mandible, mesial view; B—left mandible, lateral view; C—same, mesial view; D—maxillule, lateral view; E—maxilla, lateral view; F—first maxilliped, lateral view; G—second maxilliped, lateral view; H—third maxilliped, lateral view; I—epistome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3635674" httpUri="https://zenodo.org/record/3635674/files/figure.png" pageId="6" pageNumber="121">Fig. 3I</figureCitation>
); paragnaths without remarkable features.
</paragraph>
<caption id="36BE14AF5C644B71E38267CB7EB778DA" ID-DOI="http://doi.org/10.5281/zenodo.3635672" ID-Zenodo-Dep="3635672" httpUri="https://zenodo.org/record/3635672/files/figure.png" pageId="6" pageNumber="121" startId="6.[151,250,1756,1781]" targetBox="[227,1363,425,1711]" targetPageId="6">
<paragraph id="627E44275C644B71E38267CB7EB778DA" blockId="6.[151,1437,1756,2033]" pageId="6" pageNumber="121">
<emphasis id="50B598355C644B71E38267CB7E6B79DE" bold="true" box="[151,268,1756,1781]" pageId="6" pageNumber="121">FIGURE 2</emphasis>
. 
<taxonomicName id="A5C13FA45C644B71E20267CA7E8179DF" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[279,486,1756,1780]" class="Malacostraca" family="Palaemonidae" genus="Unesconia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="coibensis" status="gen. et sp. nov.">
<emphasis id="50B598355C644B71E20267CA7E8179DF" box="[279,486,1756,1780]" italics="true" pageId="6" pageNumber="121">Unesconia coibensis</emphasis>
</taxonomicName>
<emphasis id="50B598355C644B71E2F967CB7DE279DF" bold="true" box="[492,645,1756,1780]" pageId="6" pageNumber="121">
<taxonomicNameLabel id="4B86254E5C644B71E2F967CB7DE279DF" box="[492,645,1756,1780]" pageId="6" pageNumber="121" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–E]; holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from the same locality (FLMNH UF 51926) [F]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [G–I]: A—frontal region, dorsal view [left eye somewhat abnormal, i.e. smaller and partly shrunk, in the illustrated specimen (see Fig. 10B), shown as normal based on the right eye]; B—same, lateral view; C—sixth pleonite and tail fan, dorsal view; D—telson, posterior region, dorsal view; E—uropodal exopod, detail of lateral angle of diaeresis, dorsal view; F—frontal region of carapace and left eyestalk, lateral view; G—frontal region, dorsal view; H—same, lateral view; I—uropodal exopod, detail of lateral angle of diaeresis, dorsal view.
</paragraph>
</caption>
<caption id="36BE14AF5C654B70E382677E7CEF79C5" ID-DOI="http://doi.org/10.5281/zenodo.3635674" ID-Zenodo-Dep="3635674" httpUri="https://zenodo.org/record/3635674/files/figure.png" pageId="7" pageNumber="122" startId="7.[151,250,1641,1666]" targetBox="[165,1423,185,1615]" targetPageId="7">
<paragraph id="627E44275C654B70E382677E7CEF79C5" blockId="7.[151,1437,1641,1774]" pageId="7" pageNumber="122">
<emphasis id="50B598355C654B70E382677E7E6B79A9" bold="true" box="[151,268,1641,1666]" pageId="7" pageNumber="122">FIGURE 3</emphasis>
. 
<taxonomicName id="A5C13FA45C654B70E202677D7E8179AA" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[279,486,1641,1665]" class="Malacostraca" family="Palaemonidae" genus="Unesconia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="coibensis" status="gen. et sp. nov.">
<emphasis id="50B598355C654B70E202677D7E8179AA" box="[279,486,1641,1665]" italics="true" pageId="7" pageNumber="122">Unesconia coibensis</emphasis>
</taxonomicName>
<emphasis id="50B598355C654B70E2F9677D7DE279A9" bold="true" box="[492,645,1642,1666]" pageId="7" pageNumber="122">
<taxonomicNameLabel id="4B86254E5C654B70E2F9677D7DE279A9" box="[492,645,1642,1666]" pageId="7" pageNumber="122" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right mandible, mesial view; B—left mandible, lateral view; C—same, mesial view; D—maxillule, lateral view; E—maxilla, lateral view; F—first maxilliped, lateral view; G—second maxilliped, lateral view; H—third maxilliped, lateral view; I—epistome, ventral view.
</paragraph>
</caption>
<paragraph id="627E44275C654B70E3D2660B7B1678ED" blockId="7.[151,1436,1820,1990]" pageId="7" pageNumber="122">
First pereiopods symmetrical in shape and equal in size, relatively short and stout; coxa with stout, anteriorly protruding process on anteroventral surface; ischium stout, widening distally; merus about four times as long as maximal width; carpus about 0.8 times as long as merus, depressed ventrally; chela as long as carpus; fingers noticeably shorter than palm, broadened, subspatulate (deeply excavated on cutting surfaces), fingertips strongly tridentate, with teeth sharp, fitting into corresponding hiatuses on opposing tips (
<figureCitation id="FAFA58A25C654B70E70A66BB7B0378ED" box="[1055,1124,1964,1990]" captionStart="FIGURE 4" captionStartId="8.[151,250,1177,1202]" captionTargetBox="[173,1387,192,1106]" captionTargetId="figure@8.[151,1435,174,1158]" captionTargetPageId="8" captionText="FIGURE 4. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right first pereiopod, lateral view; B—same, mesial view; C—same, chela, mesial view; D—same, chela with fingers opened, setae omitted for clarity, dorsolateral view; E—same, chela with fingers opened, setae omitted for clarity, ventromesial view." figureDoi="http://doi.org/10.5281/zenodo.3635676" httpUri="https://zenodo.org/record/3635676/files/figure.png" pageId="7" pageNumber="122">Fig. 4</figureCitation>
).
</paragraph>
<caption id="36BE14AF5C6A4B7FE382658E7DBD7A35" ID-DOI="http://doi.org/10.5281/zenodo.3635676" ID-Zenodo-Dep="3635676" httpUri="https://zenodo.org/record/3635676/files/figure.png" pageId="8" pageNumber="123" startId="8.[151,250,1177,1202]" targetBox="[173,1387,192,1106]" targetPageId="8">
<paragraph id="627E44275C6A4B7FE382658E7DBD7A35" blockId="8.[151,1437,1177,1310]" pageId="8" pageNumber="123">
<emphasis id="50B598355C6A4B7FE382658E7E6A7B99" bold="true" box="[151,269,1177,1202]" pageId="8" pageNumber="123">FIGURE 4</emphasis>
. 
<taxonomicName id="A5C13FA45C6A4B7FE20F658D7E8C7B9A" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[282,491,1177,1201]" class="Malacostraca" family="Palaemonidae" genus="Unesconia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="123" phylum="Arthropoda" rank="species" species="coibensis" status="gen. et sp. nov.">
<emphasis id="50B598355C6A4B7FE20F658D7E8C7B9A" box="[282,491,1177,1201]" italics="true" pageId="8" pageNumber="123">Unesconia coibensis</emphasis>
</taxonomicName>
<emphasis id="50B598355C6A4B7FE2E4658D7DE87B99" bold="true" box="[497,655,1178,1202]" pageId="8" pageNumber="123">
<taxonomicNameLabel id="4B86254E5C6A4B7FE2E4658D7DE87B99" box="[497,655,1178,1202]" pageId="8" pageNumber="123" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—right first pereiopod, lateral view; B—same, mesial view; C—same, chela, mesial view; D—same, chela with fingers opened, setae omitted for clarity, dorsolateral view; E—same, chela with fingers opened, setae omitted for clarity, ventromesial view.
</paragraph>
</caption>
<paragraph id="627E44275C6A4B7FE3D2645B7BA279E4" blockId="8.[151,1437,1356,2031]" pageId="8" pageNumber="123">
Both second pereiopods (chelipeds) enlarged, robust; chelae more or less swollen, unequal in size, asymmetrical in shape, moderately setose; major chela more swollen than minor chela, with very different finger armature; sexual dimorphism of major cheliped not significant. Major cheliped with ischium very short, widening distally; merus stout, less than twice as long as maximal width, ventromesial margin with small rugosities; carpus short, cupshaped; chela moderately swollen (slightly more swollen in male), surface smooth except for minute tubercles along ventral and dorsal surfaces of palm and ventral margin of pollex; fingers about half as long as palm, cutting edges with two opposing deep fossae, one on pollex and one on dactylus (
<figureCitation id="FAFA58A25C6A4B7FE09B67337CBD7915" box="[910,986,1571,1598]" captionStart="FIGURE 5" captionStartId="9.[151,250,1592,1617]" captionTargetBox="[163,1425,417,1536]" captionTargetId="figure@9.[151,1435,393,1548]" captionTargetPageId="9" captionText="FIGURE 5. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—major (left) second pereiopod, mesial (dorsal) view; B—same, carpus and chela with fingers closed, lateral (ventral) view; C—same, carpus and chela with fingers opened, lateral (ventral) view; D—same, chela fingers opened, setae omitted for clarity, lateral (ventral) view; E—minor (right) second pereiopod, lateral (ventral) view; F—same, carpus and chela, lateral (ventral) view." figureDoi="http://doi.org/10.5281/zenodo.3635678" httpUri="https://zenodo.org/record/3635678/files/figure.png" pageId="8" pageNumber="123">Figs. 5</figureCitation>
A–D, 6, 7). Female minor cheliped with ischium, merus and carpus very similar to those of major cheliped; carpus slightly longer; chela much less swollen, slenderer, surface smooth, except for minute tubercles along ventral and dorsal surfaces of palm and near base of pollex; fingers subequal in length to palm, cutting edges unarmed; pollex shallowly excavated on each side of cutting edge (
<figureCitation id="FAFA58A25C6A4B7FE21B67A37EE579E5" box="[270,386,1715,1742]" captionStart="FIGURE 5" captionStartId="9.[151,250,1592,1617]" captionTargetBox="[163,1425,417,1536]" captionTargetId="figure@9.[151,1435,393,1548]" captionTargetPageId="9" captionText="FIGURE 5. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—major (left) second pereiopod, mesial (dorsal) view; B—same, carpus and chela with fingers closed, lateral (ventral) view; C—same, carpus and chela with fingers opened, lateral (ventral) view; D—same, chela fingers opened, setae omitted for clarity, lateral (ventral) view; E—minor (right) second pereiopod, lateral (ventral) view; F—same, carpus and chela, lateral (ventral) view." figureDoi="http://doi.org/10.5281/zenodo.3635678" httpUri="https://zenodo.org/record/3635678/files/figure.png" pageId="8" pageNumber="123">Fig. 5E, F</figureCitation>
). 
<collectingRegion id="A0058AC55C6A4B7FE28367A37EB779E5" box="[406,464,1716,1742]" country="Maldives" name="Maale" pageId="8" pageNumber="123">Male</collectingRegion>
minor cheliped presently unknown (regenerating in male 
<typeStatus id="BD7AFA855C6A4B7FE74F67A37BDE79E4" box="[1114,1209,1716,1743]" pageId="8" pageNumber="123" type="allotype">allotype</typeStatus>
).
</paragraph>
<paragraph id="627E44275C6A4B7EE3D267CF7B967E4C" blockId="8.[151,1437,1356,2031]" lastBlockId="9.[151,1437,153,359]" lastPageId="9" lastPageNumber="124" pageId="8" pageNumber="123">
Third to fifth (ambulatory) pereiopods relatively slender, diminishing in robustness from third to fifth, similar in shape. Third pereiopod with ischium about three times as long as wide, unarmed; merus slightly swollen, with feebly convex margins, 4.5 times as long as maximal width, unarmed; carpus noticeably slenderer than merus, slightly less than half-length of merus, unarmed; propodus twice as long as carpus, slender, ventral margin with four small but stout spiniform setae, including terminal one at propodo-dactylar articulation; dactylus moderately stout, compressed, slightly more than 0.2 length of propodus; unguis slender, acute; corpus with one large distal tooth, stouter than unguis, almost parallel to unguis; ventral margin of corpus with two or three smaller articulated spinules (sometimes minute) (
<figureCitation id="FAFA58A25C6A4B7FE29D66C37EAD78C5" box="[392,458,2004,2030]" captionStart="FIGURE 8" captionStartId="12.[151,250,1791,1816]" captionTargetBox="[196,1382,196,1757]" captionTargetId="figure@12.[184,1403,174,1772]" captionTargetPageId="12" captionText="FIGURE 8. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–H]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [I, J]: A—third pereiopod, lateral view; B—same, propodus and dactylus, mesial view; C—same, distal portion of propodus and dactylus, lateral view; D—fourth pereiopod, lateral view; E—same, propodus and dactylus, mesial view; F—same, distal portion of propodus and dactylus, lateral view; G—fifth pereiopod, lateral view; H—same, propodus and dactylus, mesial view; I—male second pleopod, posterior (“lateral”) view; J—same, detail of appendix masculina and appendix interna, drawn without scale." figureDoi="http://doi.org/10.5281/zenodo.3635684" httpUri="https://zenodo.org/record/3635684/files/figure.png" pageId="8" pageNumber="123">Fig. 8</figureCitation>
A–C). Fourth pereiopod generally similar to third pereiopod, but noticeably slenderer; merus not swollen, with straight margins; propodus and dactylus similar to those of third pereiopod (
<figureCitation id="FAFA58A25C6B4B7EE603618E7A3F7F98" box="[1302,1368,153,179]" captionStart="FIGURE 8" captionStartId="12.[151,250,1791,1816]" captionTargetBox="[196,1382,196,1757]" captionTargetId="figure@12.[184,1403,174,1772]" captionTargetPageId="12" captionText="FIGURE 8. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–H]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [I, J]: A—third pereiopod, lateral view; B—same, propodus and dactylus, mesial view; C—same, distal portion of propodus and dactylus, lateral view; D—fourth pereiopod, lateral view; E—same, propodus and dactylus, mesial view; F—same, distal portion of propodus and dactylus, lateral view; G—fifth pereiopod, lateral view; H—same, propodus and dactylus, mesial view; I—male second pleopod, posterior (“lateral”) view; J—same, detail of appendix masculina and appendix interna, drawn without scale." figureDoi="http://doi.org/10.5281/zenodo.3635684" httpUri="https://zenodo.org/record/3635684/files/figure.png" pageId="9" pageNumber="124">Fig. 8</figureCitation>
D–F). Fifth pereiopod slenderest; ischium almost four times as long as wide; merus not swollen, with straight margins, more than six times as long as maximal width; propodus much more slender and proportionally longer than that of third or fourth pereiopod, with two slender spiniform setae (sometimes not visible in lateral view), including terminal one at propodo-dactylar articulation; distal ventrolateral surface with four rows of serrulate setae forming propodal cleaning brush; dactylus similar to that of third pereiopod, somewhat stouter (
<figureCitation id="FAFA58A25C6B4B7EE77D605A7B837E4C" box="[1128,1252,333,359]" captionStart="FIGURE 8" captionStartId="12.[151,250,1791,1816]" captionTargetBox="[196,1382,196,1757]" captionTargetId="figure@12.[184,1403,174,1772]" captionTargetPageId="12" captionText="FIGURE 8. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–H]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [I, J]: A—third pereiopod, lateral view; B—same, propodus and dactylus, mesial view; C—same, distal portion of propodus and dactylus, lateral view; D—fourth pereiopod, lateral view; E—same, propodus and dactylus, mesial view; F—same, distal portion of propodus and dactylus, lateral view; G—fifth pereiopod, lateral view; H—same, propodus and dactylus, mesial view; I—male second pleopod, posterior (“lateral”) view; J—same, detail of appendix masculina and appendix interna, drawn without scale." figureDoi="http://doi.org/10.5281/zenodo.3635684" httpUri="https://zenodo.org/record/3635684/files/figure.png" pageId="9" pageNumber="124">Fig. 8G, H</figureCitation>
).
</paragraph>
<caption id="36BE14AF5C6B4B7EE382672F7C4C79CA" ID-DOI="http://doi.org/10.5281/zenodo.3635678" ID-Zenodo-Dep="3635678" httpUri="https://zenodo.org/record/3635678/files/figure.png" pageId="9" pageNumber="124" startId="9.[151,250,1592,1617]" targetBox="[163,1425,417,1536]" targetPageId="9">
<paragraph id="627E44275C6B4B7EE382672F7C4C79CA" blockId="9.[151,1437,1592,1761]" pageId="9" pageNumber="124">
<emphasis id="50B598355C6B4B7EE382672F7E6B797A" bold="true" box="[151,268,1592,1617]" pageId="9" pageNumber="124">FIGURE 5</emphasis>
. 
<taxonomicName id="A5C13FA45C6B4B7EE202672E7E81797B" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[279,486,1592,1616]" class="Malacostraca" family="Palaemonidae" genus="Unesconia" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="124" phylum="Arthropoda" rank="species" species="coibensis" status="gen. et sp. nov.">
<emphasis id="50B598355C6B4B7EE202672E7E81797B" box="[279,486,1592,1616]" italics="true" pageId="9" pageNumber="124">Unesconia coibensis</emphasis>
</taxonomicName>
<emphasis id="50B598355C6B4B7EE2F9672E7DE2797A" bold="true" box="[492,645,1593,1617]" pageId="9" pageNumber="124">
<taxonomicNameLabel id="4B86254E5C6B4B7EE2F9672E7DE2797A" box="[492,645,1593,1617]" pageId="9" pageNumber="124" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928): A—major (left) second pereiopod, mesial (dorsal) view; B—same, carpus and chela with fingers closed, lateral (ventral) view; C—same, carpus and chela with fingers opened, lateral (ventral) view; D—same, chela fingers opened, setae omitted for clarity, lateral (ventral) view; E—minor (right) second pereiopod, lateral (ventral) view; F—same, carpus and chela, lateral (ventral) view.
</paragraph>
</caption>
<paragraph id="627E44275C6B4B7EE3D266187B4578F6" blockId="9.[151,1436,1807,2013]" pageId="9" pageNumber="124">
<collectingRegion id="A0058AC55C6B4B7EE3D266187E667802" box="[199,257,1807,1833]" country="Maldives" name="Maale" pageId="9" pageNumber="124">Male</collectingRegion>
second pleopod with well-developed appendix masculina furnished with at least six stiff setae on apex and margin opposed to endopod; appendix interna longer than appendix masculine masculina (
<figureCitation id="FAFA58A25C6B4B7EE78466247B9F7866" box="[1169,1272,1843,1869]" captionStart="FIGURE 8" captionStartId="12.[151,250,1791,1816]" captionTargetBox="[196,1382,196,1757]" captionTargetId="figure@12.[184,1403,174,1772]" captionTargetPageId="12" captionText="FIGURE 8. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928) [A–H]; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927) [I, J]: A—third pereiopod, lateral view; B—same, propodus and dactylus, mesial view; C—same, distal portion of propodus and dactylus, lateral view; D—fourth pereiopod, lateral view; E—same, propodus and dactylus, mesial view; F—same, distal portion of propodus and dactylus, lateral view; G—fifth pereiopod, lateral view; H—same, propodus and dactylus, mesial view; I—male second pleopod, posterior (“lateral”) view; J—same, detail of appendix masculina and appendix interna, drawn without scale." figureDoi="http://doi.org/10.5281/zenodo.3635684" httpUri="https://zenodo.org/record/3635684/files/figure.png" pageId="9" pageNumber="124">Fig. 8I, J</figureCitation>
). Uropod with lateral lobe of protopod acutely produced; exopod broadly ovoid, with small distolateral tooth; diaeresis (transverse suture) abruptly curved near its lateral end, typically with five strong spiniform setae, third (most central) spiniform seta strongest and longest, sometimes with four spiniform setae; none of spiniform setae extending to exopod’s lateral margin; endopod narrowly ovoid, without specific features (Fig; 2C, E, I).
</paragraph>
<caption id="36BE14AF5C684B7DE38267787FDF79DF" ID-DOI="http://doi.org/10.5281/zenodo.3635680" ID-Zenodo-Dep="3635680" httpUri="https://zenodo.org/record/3635680/files/figure.png" pageId="10" pageNumber="125" startId="10.[151,250,1647,1672]" targetBox="[183,1412,211,1601]" targetPageId="10">
<paragraph id="627E44275C684B7DE38267787FDF79DF" blockId="10.[151,1436,1647,1780]" pageId="10" pageNumber="125">
<emphasis id="50B598355C684B7DE38267787E6B79A3" bold="true" box="[151,268,1647,1672]" pageId="10" pageNumber="125">FIGURE 6</emphasis>
. 
<taxonomicName id="A5C13FA45C684B7DE20D67667E8F79A3" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[280,488,1648,1672]" class="Malacostraca" family="Palaemonidae" genus="Unesconia" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="125" phylum="Arthropoda" rank="species" species="coibensis" status="gen. et sp. nov.">
<emphasis id="50B598355C684B7DE20D67667E8F79A3" box="[280,488,1648,1672]" italics="true" pageId="10" pageNumber="125">Unesconia coibensis</emphasis>
</taxonomicName>
<emphasis id="50B598355C684B7DE2FB67677DEE79A3" bold="true" box="[494,649,1648,1672]" pageId="10" pageNumber="125">
<taxonomicNameLabel id="4B86254E5C684B7DE2FB67677DEE79A3" box="[494,649,1648,1672]" pageId="10" pageNumber="125" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927): A—major (left) second pereiopod, dorsomesial view; B—same, carpus and chela with fingers closed, lateral (ventral) view; C–H—same, chela fingers opened viewed from various angles mesially (C–F) or laterally (G, H), setae omitted for clarity.
</paragraph>
</caption>
<paragraph id="627E44275C684B7DE3D266357CA97882" blockId="10.[151,1437,1826,2033]" pageId="10" pageNumber="125">
<emphasis id="50B598355C684B7DE3D266357E1B7816" bold="true" box="[199,380,1826,1853]" pageId="10" pageNumber="125">Colour pattern</emphasis>
. Females: body semi-translucent, pale to more intense yolk-yellow, second pereiopod chelae lemon-yellow, walking legs and tail fan transparent with yellow tinge (
<figureCitation id="FAFA58A25C684B7DE0BD66517C8B784A" box="[936,1004,1862,1889]" captionStart="FIGURE 9" captionStartId="13.[151,250,1044,1069]" captionTargetBox="[151,1436,174,1024]" captionTargetId="figure@13.[151,1436,174,1024]" captionTargetPageId="13" captionText="FIGURE 9. Unesconia coibensis gen. et sp. nov.: holotype, ovigerous female (pocl 2.0 mm, cl 3.1 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51926), shrimp alive, dorsal view. Photograph by the author." figureDoi="http://doi.org/10.5281/zenodo.3635686" httpUri="https://zenodo.org/record/3635686/files/figure.png" pageId="10" pageNumber="125">Fig. 9</figureCitation>
, 
<figureCitation id="FAFA58A25C684B7DE0E266507B2D784A" box="[1015,1098,1863,1889]" captionStart="FIGURE 10" captionStartId="14.[151,250,1887,1912]" captionTargetBox="[151,1436,517,1867]" captionTargetId="figure@14.[151,1436,517,1867]" captionTargetPageId="14" captionText="FIGURE 10. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928), shrimp alive, dorsal [A] and lateral [B] views; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927), shrimp alive, dorsal view, right (minor) cheliped regenerating [C]. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3635688" httpUri="https://zenodo.org/record/3635688/files/figure.png" pageId="10" pageNumber="125">10A, B</figureCitation>
). 
<collectingRegion id="A0058AC55C684B7DE74B66517BFB784B" box="[1118,1180,1862,1888]" country="Maldives" name="Maale" pageId="10" pageNumber="125">Male</collectingRegion>
: body and appendages, including second pereiopod chelae semi-translucent lemon-yellow; waking legs, antennae and proximal half of palm of second pereiopod chela with conspicuous white spots (
<figureCitation id="FAFA58A25C684B7DE04E66997CA57882" box="[859,962,1934,1961]" captionStart="FIGURE 10" captionStartId="14.[151,250,1887,1912]" captionTargetBox="[151,1436,517,1867]" captionTargetId="figure@14.[151,1436,517,1867]" captionTargetPageId="14" captionText="FIGURE 10. Unesconia coibensis gen. et sp. nov., paratype, ovigerous female (pocl 2.1 mm, cl 3.0 mm) from Canales de Afuera, Coiba Archipelago, Panama (FLMNH UF 51928), shrimp alive, dorsal [A] and lateral [B] views; allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927), shrimp alive, dorsal view, right (minor) cheliped regenerating [C]. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3635688" httpUri="https://zenodo.org/record/3635688/files/figure.png" pageId="10" pageNumber="125">Fig. 10C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2ADB17AC5C684B7DE3D266A47C6D78DB" pageId="10" pageNumber="125" type="etymology">
<paragraph id="627E44275C684B7DE3D266A47C6D78DB" blockId="10.[151,1437,1826,2033]" pageId="10" pageNumber="125">
<emphasis id="50B598355C684B7DE3D266A47E2F78E6" bold="true" box="[199,328,1971,1997]" pageId="10" pageNumber="125">Etymology</emphasis>
. The specific epithet refers to the Coiba Archipelago / Coiba National Park, which includes Canales de Afuera islands, its 
<typeStatus id="BD7AFA855C684B7DE29C66C07EDE78DA" box="[393,441,2007,2033]" pageId="10" pageNumber="125">type</typeStatus>
locality; used as an adjective.
</paragraph>
</subSubSection>
<caption id="36BE14AF5C694B7CE382647F7E547AE2" ID-DOI="http://doi.org/10.5281/zenodo.3635682" ID-Zenodo-Dep="3635682" httpUri="https://zenodo.org/record/3635682/files/figure.png" pageId="11" pageNumber="126" startId="11.[151,250,1384,1409]" targetBox="[151,1436,206,1331]" targetPageId="11">
<paragraph id="627E44275C694B7CE382647F7E547AE2" blockId="11.[151,1437,1384,1481]" pageId="11" pageNumber="126">
<emphasis id="50B598355C694B7CE382647F7E6B7AAA" bold="true" box="[151,268,1384,1409]" pageId="11" pageNumber="126">FIGURE 7</emphasis>
. 
<taxonomicName id="A5C13FA45C694B7CE20D647E7E8F7AAB" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[280,488,1384,1408]" class="Malacostraca" family="Palaemonidae" genus="Unesconia" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="126" phylum="Arthropoda" rank="species" species="coibensis" status="gen. et sp. nov.">
<emphasis id="50B598355C694B7CE20D647E7E8F7AAB" box="[280,488,1384,1408]" italics="true" pageId="11" pageNumber="126">Unesconia coibensis</emphasis>
</taxonomicName>
<emphasis id="50B598355C694B7CE2FB647E7DEE7AAA" bold="true" box="[494,649,1385,1409]" pageId="11" pageNumber="126">
<taxonomicNameLabel id="4B86254E5C694B7CE2FB647E7DEE7AAA" box="[494,649,1385,1409]" pageId="11" pageNumber="126" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, allotype, male (pocl 1.7 mm, cl 2.8 mm) from the same locality (FLMNH UF 51927): A—major (left) second pereiopod, chela fingers opened viewed from various angles mesially (A–C) or laterally (D). See also Fig. 6.
</paragraph>
</caption>
<subSubSection id="2ADB17AC5C694B7CE3D264E07B59793A" box="[199,1086,1526,1553]" pageId="11" pageNumber="126" type="materials_examined">
<paragraph id="627E44275C694B7CE3D264E07B59793A" blockId="11.[151,1437,1526,1985]" box="[199,1086,1526,1553]" pageId="11" pageNumber="126">
<materialsCitation id="D2A94E7A5C694B7CE3D264E07B5E793A" ID-GBIF-Occurrence="2562577888" box="[199,1081,1526,1553]" country="Panama" county="Pacific" location="Canales de Afuera" municipality="Coiba Archipelago" pageId="11" pageNumber="126" specimenCount="1" typeStatus="holotype">
<emphasis id="50B598355C694B7CE3D264E07E07793A" bold="true" box="[199,352,1526,1553]" pageId="11" pageNumber="126">
<typeStatus id="BD7AFA855C694B7CE3D264E07E65793A" box="[199,258,1527,1553]" pageId="11" pageNumber="126">Type</typeStatus>
locality
</emphasis>
. 
<location id="671E12FC5C694B7CE27B64E07D59793A" LSID="urn:lsid:plazi:treatment:EA68F5315C664B7CE38264F17DE478EA:671E12FC5C694B7CE27B64E07D59793A" box="[366,574,1527,1553]" country="Panama" county="Pacific" municipality="Coiba Archipelago" name="Canales de Afuera" pageId="11" pageNumber="126">Canales de Afuera</location>
, 
<collectingMunicipality id="821ADE5D5C694B7CE15C64E07C79793A" box="[585,798,1527,1553]" pageId="11" pageNumber="126">Coiba Archipelago</collectingMunicipality>
, 
<collectingCounty id="8B1F3CAB5C694B7CE03F64E07C1F793A" box="[810,888,1527,1553]" pageId="11" pageNumber="126">Pacific</collectingCounty>
coast of 
<collectingCountry id="1AD604B75C694B7CE0C864E07B5E793A" box="[989,1081,1527,1553]" name="Panama" pageId="11" pageNumber="126">Panama</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2ADB17AC5C694B7CE3D2670D7BDB791E" box="[199,1212,1562,1589]" pageId="11" pageNumber="126" type="distribution">
<paragraph id="627E44275C694B7CE3D2670D7BDB791E" blockId="11.[151,1437,1526,1985]" box="[199,1212,1562,1589]" pageId="11" pageNumber="126">
<emphasis id="50B598355C694B7CE3D2670D7E3E791E" bold="true" box="[199,345,1562,1589]" pageId="11" pageNumber="126">Distribution</emphasis>
. Presently known only from the 
<typeStatus id="BD7AFA855C694B7CE1DC670C7D9E791E" box="[713,761,1563,1589]" pageId="11" pageNumber="126">type</typeStatus>
locality on the Pacific coast of 
<collectingCountry id="1AD604B75C694B7CE74E670C7BD0791E" box="[1115,1207,1563,1589]" name="Panama" pageId="11" pageNumber="126">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="2ADB17AC5C694B7CE3D267287DE478EA" pageId="11" pageNumber="126" type="biology_ecology">
<paragraph id="627E44275C694B7CE3D267287C8A78B6" blockId="11.[151,1437,1526,1985]" pageId="11" pageNumber="126">
<emphasis id="50B598355C694B7CE3D267287E417972" bold="true" box="[199,294,1599,1625]" pageId="11" pageNumber="126">Ecology</emphasis>
. All specimens of 
<taxonomicName id="A5C13FA45C694B7CE2E367577DBA7972" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[502,733,1599,1625]" class="Malacostraca" family="Palaemonidae" genus="Unesconia" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="126" phylum="Arthropoda" rank="species" species="coibensis" status="gen. et sp. nov.">
<emphasis id="50B598355C694B7CE2E367577DBA7972" box="[502,733,1599,1625]" italics="true" pageId="11" pageNumber="126">Unesconia coibensis</emphasis>
</taxonomicName>
<emphasis id="50B598355C694B7CE1F167287CF67972" bold="true" box="[740,913,1599,1625]" pageId="11" pageNumber="126">
<taxonomicNameLabel id="4B86254E5C694B7CE1F167287CF67972" box="[740,913,1599,1625]" pageId="11" pageNumber="126" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
were extracted from a large (about 
<quantity id="A539E9C25C694B7CE60A67287A037972" box="[1311,1380,1599,1625]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="126" unit="cm" value="70.0">70 cm</quantity>
long and 
<quantity id="A539E9C25C694B7CE3DD67757E687956" box="[200,271,1634,1661]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="126" unit="cm" value="50.0">50 cm</quantity>
wide) dead coral head (
<taxonomicName id="A5C13FA45C694B7CE13367747C557956" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[550,818,1635,1661]" class="Anthozoa" family="Pocilloporidae" genus="Pocillopora" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="126" phylum="Cnidaria" rank="species" species="damicornis">
<emphasis id="50B598355C694B7CE13367747C557956" box="[550,818,1635,1661]" italics="true" pageId="11" pageNumber="126">Pocillopora damicornis</emphasis>
</taxonomicName>
), which was partly immerged into surrounding coarse sand mixed with smaller fragments of coral rubble, in very shallow water (
<quantity id="A539E9C25C694B7CE0FF67907B54798A" box="[1002,1075,1671,1697]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="11" pageNumber="126" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
at low tide). The collection site was also characterised by a noticeably stronger and colder current due to the constant water flow between a small islet and the main island of 
<collectingCountry id="1AD604B75C694B7CE2DA67D87E9079C2" box="[463,503,1743,1769]" name="Iceland" pageId="11" pageNumber="126">Isla</collectingCountry>
Afuera. The shrimps were likely associated with a yellow incrusting sponge growing deep inside the dead coral crevices; a small fragment of this sponge preserved in ethanol was deposited in the FLMNH sponge collection for further studies (FLMNH Porifera 4384). However, it must be noted that the specimens were falling out of the broken-up dead coral head, which possibly contained other incrusting sponges. The association of 
<taxonomicName id="A5C13FA45C694B7CE22E66777EA17852" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[315,454,1887,1913]" class="Malacostraca" family="Palaemonidae" genus="Unesconia" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="126" phylum="Arthropoda" rank="species" species="coibensis" status="sp. nov.">
<emphasis id="50B598355C694B7CE22E66777EA17852" box="[315,454,1887,1913]" italics="true" pageId="11" pageNumber="126">U. coibensis</emphasis>
</taxonomicName>
<emphasis id="50B598355C694B7CE2DB66487D4E7852" bold="true" box="[462,553,1887,1913]" pageId="11" pageNumber="126">
<taxonomicNameLabel id="4B86254E5C694B7CE2DB66487D4E7852" box="[462,553,1887,1913]" pageId="11" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with that particular yellow sponge was presumed based on its large extension inside the coral head and on the similarly yellow colouration of the shrimps.
</paragraph>
<paragraph id="627E44275C694B7CE3D266B17DE478EA" blockId="11.[151,1437,1526,1985]" box="[199,643,1958,1985]" pageId="11" pageNumber="126">
<emphasis id="50B598355C694B7CE3D266B17ED178EA" bold="true" box="[199,438,1958,1985]" pageId="11" pageNumber="126">Taxonomic remarks</emphasis>
. See under genus.
</paragraph>
</subSubSection>
</treatment>
</document>