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<document id="50C6F9961ECD6BCC05D24F714AA5711A" ID-DOI="10.11646/zootaxa.5383.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350767" ID-ZooBank="7E434B40-6346-4E6D-BC55-38EBAE24BD52" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702283478140" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2023" docId="386ABE42FFB1FFACFF06AB872EC2FE08" docLanguage="en" docName="zootaxa.5383.2.5.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athanas Leach 1814" docType="treatment" docVersion="2" lastPageNumber="182" masterDocId="C453C63AFFB0FFAFFF91AE4D2E20FFA5" masterDocTitle="Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="215" masterPageNumber="179" pageNumber="180" updateTime="1702284319693" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BA14778406AAF311A0F3B60A2D05CECE">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
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<mods:namePart id="5E8914605D6C21FA55FC410566E3099A">Anker, Arthur</mods:namePart>
<mods:affiliation id="49B3C702919AB84678362C30C31CD14C">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil &amp; Red Sea Research Center, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia</mods:affiliation>
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<subSubSection id="F8D95CDFFFB1FFAEFF06AB872FD1FA41" box="[151,497,1482,1509]" pageId="1" pageNumber="180" type="nomenclature">
<paragraph id="B07C0F54FFB1FFAEFF06AB872FD1FA41" blockId="1.[151,497,1482,1509]" box="[151,497,1482,1509]" pageId="1" pageNumber="180">
<heading id="EB34B838FFB1FFAEFF06AB872FD1FA41" bold="true" box="[151,497,1482,1509]" fontSize="11" level="1" pageId="1" pageNumber="180" reason="1">
<emphasis id="82B7D346FFB1FFAEFF06AB872FD1FA41" bold="true" box="[151,497,1482,1509]" pageId="1" pageNumber="180">
Genus
<taxonomicName id="77C374D7FFB1FFAEFF7FAB872FD1FA41" ID-CoL="84LMQ" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[238,497,1482,1509]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB1FFAEFF7FAB872F73FA40" bold="true" box="[238,339,1482,1509]" italics="true" pageId="1" pageNumber="180">Athanas</emphasis>
<bibRefCitation id="D45272A5FFB1FFAEFECAAB862FD1FA41" author="Leach, W. E." box="[347,497,1482,1509]" pageId="1" pageNumber="180" pagination="383 - 437" refId="ref22744" refString="Leach, W. E. (1814) Crustaceology. In: Brewster, D. (Ed.), The Edinburgh Encyclopaedia, 7, pp. 383 - 437." type="journal article" year="1814">Leach, 1814</bibRefCitation>
</taxonomicName>
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</paragraph>
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<paragraph id="B07C0F54FFB1FFAEFF06A8422F82F982" blockId="1.[151,418,1551,1575]" box="[151,418,1551,1575]" pageId="1" pageNumber="180">
<treatmentCitationGroup id="90D3287AFFB1FFAEFF06A8422F82F982" box="[151,418,1551,1575]" pageId="1" pageNumber="180">
<treatmentCitation id="31622945FFB1FFAEFF06A8422FBEF982" author="Leach, W. E." box="[151,414,1551,1575]" page="401" pageId="1" pageNumber="180" year="1814">
<taxonomicName id="77C374D7FFB1FFAEFF06A8422FBEF982" ID-CoL="84LMQ" authority="Leach 1814: 401" authorityName="Leach" authorityPageNumber="401" authorityYear="1814" box="[151,414,1551,1575]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB1FFAEFF06A8422ECAF982" box="[151,234,1551,1575]" italics="true" pageId="1" pageNumber="180">Athanas</emphasis>
<bibRefCitation id="D45272A5FFB1FFAEFF60A8422FBEF982" author="Leach, W. E." box="[241,414,1551,1575]" pageId="1" pageNumber="180" pagination="383 - 437" refId="ref22744" refString="Leach, W. E. (1814) Crustaceology. In: Brewster, D. (Ed.), The Edinburgh Encyclopaedia, 7, pp. 383 - 437." type="journal article" year="1814">Leach 1814: 401</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB1FFADFF06A8182AB6F9DD" lastPageId="2" lastPageNumber="181" pageId="1" pageNumber="180" type="diagnosis">
<paragraph id="B07C0F54FFB1FFADFF06A8182C75FC48" blockId="1.[151,1437,1621,2008]" lastBlockId="2.[151,1437,151,1005]" lastPageId="2" lastPageNumber="181" pageId="1" pageNumber="180">
<emphasis id="82B7D346FFB1FFAEFF06A8182F5CF9CA" bold="true" box="[151,380,1621,1647]" pageId="1" pageNumber="180">Emended diagnosis</emphasis>
(simplified and updated from
<bibRefCitation id="D45272A5FFB1FFAEFD56A8182D82F9CA" author="Anker, A. &amp; Jeng, M. S." box="[711,930,1621,1648]" pageId="1" pageNumber="180" pagination="454 - 472" refId="ref20209" refString="Anker, A. &amp; Jeng, M. S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinitions of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker &amp; Jeng 2007</bibRefCitation>
). Body not compressed, slender to moderately stout. Carapace smooth, glabrous or moderately setose, typically unarmed dorsally, sometimes with small postrostral tubercle. Rostrum typically elongate, sometimes lanceolate in dorsal view, rarely short, subtriangular; dorsal margin unarmed or with minute teeth; ventral margin unarmed. Orbital teeth usually present as one pair of extra-corneal tooth, often well developed and sometimes protruding beyond anterior margin of eye, and one pair of infra-corneal teeth; occasionally infra-corneal teeth reduced or absent; extra-corneal teeth occasionally reduced to small bumps; supra-corneal teeth absent (often) or present (rarely). Pterygostomial angle rounded or angular, sometimes produced as sharp tooth; branchiostegite with scarce setae; cardiac notch well developed. First to fourth pleonites with distoventrally rounded or bluntly angular pleura; fifth pleonite with pleuron angular or subacute distoventrally; sixth pleuron with articulated plate. Telson moderately slender to broad, tapering distally, with two pairs of stout spiniform setae on dorsal surface; posterior margin rounded or straight, with two pairs of spiniform setae; anal tubercles absent. Eyes well developed, at least partly visible in dorsal view; cornea large, well pigmented, sometimes slightly reduced. Antennular peduncle stout, relatively short; stylocerite well developed, distally acute or subacute, usually overreaching mid-length of second article, sometimes its distal margin; ventromesial carina armed with tooth; second article not or variously elongate; lateral flagellum with short fused portion and well-developed accessory ramus with aesthetascs.Antenna with basicerite moderately to very stout, armed with sharp tooth; scaphocerite with well-developed blade and variously developed distolateral tooth. Mandible with two-articulated palp; molar and incisor processes well developed. Third maxilliped with well-developed exopod; coxa with moderately produced lateral plate; ultimate article tapering into corneous tip, often with small spiniform seta(e). First pereiopods (= chelipeds) highly variable in development, often sexually dimorphic, typically greatly enlarged in males, sometimes in both sexes, subequal or very unequal in size, subsymmetrical or highly asymmetrical in shape, carried extended or flexed beneath body when not in use; basis typically with rudimentary exopod; ischium robust, with dorsal margin or both dorsal and ventral margins often armed with spiniform setae; merus with ventrolateral margin smooth or dentate, ventral surface flattened or deeply excavated; carpus variable, cylindrical, vase- or cup-shaped, without setal rows on mesial surface; chelae more or less enlarged and swollen, palm with or without tubercles and row of setae on ventral surface; fingers of chelae unarmed or armed with teeth, in males or in both sexes, without snapping mechanism. Second pereiopod with ischium unarmed; carpus typically with five, rarely four or six, subarticles; chela simple, without modifications. Third and fourth pereiopods varying from very slender to moderately stout; ischium usually armed with one spiniform seta, rarely with two spiniform setae or unarmed; merus unarmed; propodus with several spiniform setae or short stiff setae on ventral margin; dactylus more or less slender, simple or biunguiculate. Fifth pereiopod with ischium unarmed or armed with one spiniform seta; merus unarmed; propodus with or without spiniform setae and well-developed cleaning brush; dactylus simple or biunguiculate. Second male pleopod with appendix masculina equal in length or exceeding appendix interna, sometimes reaching beyond endopod; second female pleopod with appendix interna only. Uropodal exopod with unarmed, sinuous or straight diaeresis. Gill-exopod formula summarised in
<tableCitation id="FD413AEFFFB2FFADFE6AAD9E2C6FFC48" box="[507,591,979,1005]" captionStart="TABLE 1" captionStartId="2.[152,244,1052,1076]" captionText="TABLE 1. Gill-exopod formula of Athanas Leach, 1814." pageId="2" pageNumber="181">Table 1</tableCitation>
.
</paragraph>
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<paragraph id="B07C0F54FFB2FFADFF09AA512CDEFB90" blockId="2.[152,766,1052,1077]" box="[152,766,1052,1077]" pageId="2" pageNumber="181">
<emphasis id="82B7D346FFB2FFADFF09AA512F28FB91" bold="true" box="[152,264,1052,1076]" pageId="2" pageNumber="181">TABLE 1</emphasis>
. Gill-exopod formula of
<taxonomicName id="77C374D7FFB2FFADFD84AA512CDBFB90" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[533,763,1052,1077]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB2FFADFD84AA512C4CFB91" box="[533,620,1052,1076]" italics="true" pageId="2" pageNumber="181">Athanas</emphasis>
<bibRefCitation id="D45272A5FFB2FFADFDE2AA512CDBFB90" author="Leach, W. E." box="[627,763,1052,1077]" pageId="2" pageNumber="181" pagination="383 - 437" refId="ref22744" refString="Leach, W. E. (1814) Crustaceology. In: Brewster, D. (Ed.), The Edinburgh Encyclopaedia, 7, pp. 383 - 437." type="journal article" year="1814">Leach, 1814</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="B07C0F54FFB2FFADFF32AA082B7FFA3B" pageId="2" pageNumber="181">
<table id="C2C3FDF4FFB20050FF32AA082B4AFA3A" box="[163,1386,1093,1439]" gridcols="9" gridrows="9" pageId="2" pageNumber="181">
<tr id="0EF30D16FFB20050FF32AA082B4AFBF8" box="[163,1386,1093,1117]" gridrow="0" pageId="2" pageNumber="181" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
<th id="4D22646AFFB20050FF32AA082F6BFBF8" box="[163,331,1093,1117]" gridcol="0" gridrow="0" pageId="2" pageNumber="181">Gills/exopods</th>
<th id="4D22646AFFB20050FD9DAA082CA5FBF8" box="[524,645,1093,1117]" gridcol="2" gridrow="0" pageId="2" pageNumber="181">Maxillipeds</th>
<th id="4D22646AFFB20050FBB3AA082AB3FBF8" box="[1058,1171,1093,1117]" gridcol="6" gridrow="0" pageId="2" pageNumber="181">Pereiopods</th>
</tr>
<tr id="0EF30D16FFB20050FF32AA202B4AFB20" box="[163,1386,1133,1157]" gridrow="1" pageId="2" pageNumber="181" rowspan-0="1">
<td id="4D22646AFFB20050FE31AA202FFDFB20" box="[416,477,1133,1157]" gridcol="1" gridrow="1" pageId="2" pageNumber="181">Mxp1</td>
<td id="4D22646AFFB20050FD9DAA202CA5FB20" box="[524,645,1133,1157]" gridcol="2" gridrow="1" pageId="2" pageNumber="181">Mxp2</td>
<td id="4D22646AFFB20050FD25AA202CD1FB20" box="[692,753,1133,1157]" gridcol="3" gridrow="1" pageId="2" pageNumber="181">Mxp3</td>
<td id="4D22646AFFB20050FCD8AA202D4EFB20" box="[841,878,1133,1157]" gridcol="4" gridrow="1" pageId="2" pageNumber="181">P1</td>
<td id="4D22646AFFB20050FC5EAA202DCAFB20" box="[975,1002,1133,1157]" gridcol="5" gridrow="1" pageId="2" pageNumber="181">P2</td>
<td id="4D22646AFFB20050FBB3AA202AB3FB20" box="[1058,1171,1133,1157]" gridcol="6" gridrow="1" pageId="2" pageNumber="181">P3</td>
<td id="4D22646AFFB20050FB5CAA202ACAFB20" box="[1229,1258,1133,1157]" gridcol="7" gridrow="1" pageId="2" pageNumber="181">P4</td>
<td id="4D22646AFFB20050FADCAA202B4AFB20" box="[1357,1386,1133,1157]" gridcol="8" gridrow="1" pageId="2" pageNumber="181">P5</td>
</tr>
<tr id="0EF30D16FFB20050FF32AAD82B4AFB08" box="[163,1386,1173,1197]" gridrow="2" pageId="2" pageNumber="181">
<th id="4D22646AFFB20050FF32AAD82F6BFB08" box="[163,331,1173,1197]" gridcol="0" gridrow="2" pageId="2" pageNumber="181">Pleurobranchs</th>
<td id="4D22646AFFB20050FE31AAD82FFDFB08" box="[416,477,1173,1197]" gridcol="1" gridrow="2" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FD9DAAD82CA5FB08" box="[524,645,1173,1197]" gridcol="2" gridrow="2" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FD25AAD82CD1FB08" box="[692,753,1173,1197]" gridcol="3" gridrow="2" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FCD8AAD82D4EFB08" box="[841,878,1173,1197]" gridcol="4" gridrow="2" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FC5EAAD82DCAFB08" box="[975,1002,1173,1197]" gridcol="5" gridrow="2" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FBB3AAD82AB3FB08" box="[1058,1171,1173,1197]" gridcol="6" gridrow="2" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FB5CAAD82ACAFB08" box="[1229,1258,1173,1197]" gridcol="7" gridrow="2" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FADCAAD82B4AFB08" box="[1357,1386,1173,1197]" gridcol="8" gridrow="2" pageId="2" pageNumber="181">+</td>
</tr>
<tr id="0EF30D16FFB20050FF32AAF02B4AFB70" box="[163,1386,1213,1237]" gridrow="3" pageId="2" pageNumber="181">
<th id="4D22646AFFB20050FF32AAF02F6BFB70" box="[163,331,1213,1237]" gridcol="0" gridrow="3" pageId="2" pageNumber="181">Arthrobranchs</th>
<td id="4D22646AFFB20050FE31AAF02FFDFB70" box="[416,477,1213,1237]" gridcol="1" gridrow="3" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FD9DAAF02CA5FB70" box="[524,645,1213,1237]" gridcol="2" gridrow="3" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FD25AAF02CD1FB70" box="[692,753,1213,1237]" gridcol="3" gridrow="3" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FCD8AAF02D4EFB70" box="[841,878,1213,1237]" gridcol="4" gridrow="3" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FC5EAAF02DCAFB70" box="[975,1002,1213,1237]" gridcol="5" gridrow="3" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FBB3AAF02AB3FB70" box="[1058,1171,1213,1237]" gridcol="6" gridrow="3" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FB5CAAF02ACAFB70" box="[1229,1258,1213,1237]" gridcol="7" gridrow="3" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FADCAAF02B4AFB70" box="[1357,1386,1213,1237]" gridcol="8" gridrow="3" pageId="2" pageNumber="181">-</td>
</tr>
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<th id="4D22646AFFB20050FF32AAAB2F6BFB5B" box="[163,331,1254,1278]" gridcol="0" gridrow="4" pageId="2" pageNumber="181">Podobranchs</th>
<td id="4D22646AFFB20050FE31AAAB2FFDFB5B" box="[416,477,1254,1278]" gridcol="1" gridrow="4" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FD9DAAAB2CA5FB5B" box="[524,645,1254,1278]" gridcol="2" gridrow="4" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FD25AAAB2CD1FB5B" box="[692,753,1254,1278]" gridcol="3" gridrow="4" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FCD8AAAB2D4EFB5B" box="[841,878,1254,1278]" gridcol="4" gridrow="4" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FC5EAAAB2DCAFB5B" box="[975,1002,1254,1278]" gridcol="5" gridrow="4" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FBB3AAAB2AB3FB5B" box="[1058,1171,1254,1278]" gridcol="6" gridrow="4" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FB5CAAAB2ACAFB5B" box="[1229,1258,1254,1278]" gridcol="7" gridrow="4" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FADCAAAB2B4AFB5B" box="[1357,1386,1254,1278]" gridcol="8" gridrow="4" pageId="2" pageNumber="181">-</td>
</tr>
<tr id="0EF30D16FFB20050FF32AB402B4AFA83" box="[163,1386,1293,1318]" gridrow="5" pageId="2" pageNumber="181">
<th id="4D22646AFFB20050FF32AB402F6BFA83" box="[163,331,1293,1318]" gridcol="0" gridrow="5" pageId="2" pageNumber="181">Epipods</th>
<td id="4D22646AFFB20050FE31AB402FFDFA83" box="[416,477,1293,1318]" gridcol="1" gridrow="5" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FD9DAB402CA5FA83" box="[524,645,1293,1318]" gridcol="2" gridrow="5" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FD25AB402CD1FA83" box="[692,753,1293,1318]" gridcol="3" gridrow="5" pageId="2" pageNumber="181">+1</td>
<td id="4D22646AFFB20050FCD8AB402D4EFA83" box="[841,878,1293,1318]" gridcol="4" gridrow="5" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FC5EAB402DCAFA83" box="[975,1002,1293,1318]" gridcol="5" gridrow="5" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FBB3AB402AB3FA83" box="[1058,1171,1293,1318]" gridcol="6" gridrow="5" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FB5CAB402ACAFA83" box="[1229,1258,1293,1318]" gridcol="7" gridrow="5" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FADCAB402B4AFA83" box="[1357,1386,1293,1318]" gridcol="8" gridrow="5" pageId="2" pageNumber="181">-</td>
</tr>
<tr id="0EF30D16FFB20050FF32AB782B4AFAEB" box="[163,1386,1333,1358]" gridrow="6" pageId="2" pageNumber="181">
<th id="4D22646AFFB20050FF32AB782F6BFAEB" box="[163,331,1333,1358]" gridcol="0" gridrow="6" pageId="2" pageNumber="181">Mastigobranchs2</th>
<td id="4D22646AFFB20050FE31AB782FFDFAEB" box="[416,477,1333,1358]" gridcol="1" gridrow="6" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FD9DAB782CA5FAEB" box="[524,645,1333,1358]" gridcol="2" gridrow="6" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FD25AB782CD1FAEB" box="[692,753,1333,1358]" gridcol="3" gridrow="6" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FCD8AB782D4EFAEB" box="[841,878,1333,1358]" gridcol="4" gridrow="6" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FC5EAB782DCAFAEB" box="[975,1002,1333,1358]" gridcol="5" gridrow="6" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FBB3AB782AB3FAEB" box="[1058,1171,1333,1358]" gridcol="6" gridrow="6" pageId="2" pageNumber="181">+/-</td>
<td id="4D22646AFFB20050FB5CAB782ACAFAEB" box="[1229,1258,1333,1358]" gridcol="7" gridrow="6" pageId="2" pageNumber="181">+/-</td>
<td id="4D22646AFFB20050FADCAB782B4AFAEB" box="[1357,1386,1333,1358]" gridcol="8" gridrow="6" pageId="2" pageNumber="181">-</td>
</tr>
<tr id="0EF30D16FFB20050FF32AB132B4AFAD3" box="[163,1386,1374,1398]" gridrow="7" pageId="2" pageNumber="181">
<th id="4D22646AFFB20050FF32AB132F6BFAD3" box="[163,331,1374,1398]" gridcol="0" gridrow="7" pageId="2" pageNumber="181">Setobranchs</th>
<td id="4D22646AFFB20050FE31AB132FFDFAD3" box="[416,477,1374,1398]" gridcol="1" gridrow="7" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FD9DAB132CA5FAD3" box="[524,645,1374,1398]" gridcol="2" gridrow="7" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FD25AB132CD1FAD3" box="[692,753,1374,1398]" gridcol="3" gridrow="7" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FCD8AB132D4EFAD3" box="[841,878,1374,1398]" gridcol="4" gridrow="7" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FC5EAB132DCAFAD3" box="[975,1002,1374,1398]" gridcol="5" gridrow="7" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FBB3AB132AB3FAD3" box="[1058,1171,1374,1398]" gridcol="6" gridrow="7" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FB5CAB132ACAFAD3" box="[1229,1258,1374,1398]" gridcol="7" gridrow="7" pageId="2" pageNumber="181">+/-</td>
<td id="4D22646AFFB20050FADCAB132B4AFAD3" box="[1357,1386,1374,1398]" gridcol="8" gridrow="7" pageId="2" pageNumber="181">+/-</td>
</tr>
<tr id="0EF30D16FFB20050FF32ABCB2B4AFA3A" box="[163,1386,1414,1439]" gridrow="8" pageId="2" pageNumber="181">
<th id="4D22646AFFB20050FF32ABCB2F6BFA3A" box="[163,331,1414,1439]" gridcol="0" gridrow="8" pageId="2" pageNumber="181">Exopods</th>
<td id="4D22646AFFB20050FE31ABCB2FFDFA3A" box="[416,477,1414,1439]" gridcol="1" gridrow="8" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FD9DABCB2CA5FA3A" box="[524,645,1414,1439]" gridcol="2" gridrow="8" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FD25ABCB2CD1FA3A" box="[692,753,1414,1439]" gridcol="3" gridrow="8" pageId="2" pageNumber="181">+</td>
<td id="4D22646AFFB20050FCD8ABCB2D4EFA3A" box="[841,878,1414,1439]" gridcol="4" gridrow="8" pageId="2" pageNumber="181">+3/-</td>
<td id="4D22646AFFB20050FC5EABCB2DCAFA3A" box="[975,1002,1414,1439]" gridcol="5" gridrow="8" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FBB3ABCB2AB3FA3A" box="[1058,1171,1414,1439]" gridcol="6" gridrow="8" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FB5CABCB2ACAFA3A" box="[1229,1258,1414,1439]" gridcol="7" gridrow="8" pageId="2" pageNumber="181">-</td>
<td id="4D22646AFFB20050FADCABCB2B4AFA3A" box="[1357,1386,1414,1439]" gridcol="8" gridrow="8" pageId="2" pageNumber="181">-</td>
</tr>
</table>
</paragraph>
<paragraph id="B07C0F54FFB2FFADFF06ABE72A02FA61" blockId="2.[151,1058,1450,1476]" box="[151,1058,1450,1476]" pageId="2" pageNumber="181">
<tableNote id="4D250EDAFFB2FFADFF06ABE72A02FA61" box="[151,1058,1450,1476]" pageId="2" pageNumber="181" targetBox="[163,1386,1093,1439]" targetPageId="2">
<superScript id="47B6A21CFFB2FFADFF06ABE72EBFFA1D" attach="left" box="[151,159,1450,1464]" fontSize="6" pageId="2" pageNumber="181">1</superScript>
lateral plate on coxa;
<superScript id="47B6A21CFFB2FFADFE1AABE72FB3FA1D" attach="right" box="[395,403,1450,1464]" fontSize="6" pageId="2" pageNumber="181">2</superScript>
strap-like epipods of
<bibRefCitation id="D45272A5FFB2FFADFDEAABE12D2FFA61" author="Chace, F. A. Jr." box="[635,783,1452,1476]" pageId="2" pageNumber="181" pagination="1 - 99" refId="ref21274" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, i - v + 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace (1988)</bibRefCitation>
;
<superScript id="47B6A21CFFB2FFADFC8BABE72D02FA1D" attach="right" box="[794,802,1450,1464]" fontSize="6" pageId="2" pageNumber="181">3</superScript>
if present, rudimentary.
</tableNote>
</paragraph>
<paragraph id="B07C0F54FFB2FFADFF56A85B2AB6F9DD" blockId="2.[151,1437,1558,2017]" pageId="2" pageNumber="181">
<emphasis id="82B7D346FFB2FFADFF56A85B2FADF995" bold="true" box="[199,397,1558,1584]" pageId="2" pageNumber="181">Species included</emphasis>
.
<typeStatus id="6F78B1F6FFB2FFADFE0BA85A2FF2F994" box="[410,466,1559,1585]" pageId="2" pageNumber="181">Type</typeStatus>
species
<taxonomicName id="77C374D7FFB2FFADFDA3A85A2D40F995" authority="(Leach, 1814)" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[562,864,1558,1585]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="nitescens">
<emphasis id="82B7D346FFB2FFADFDA3A85A2C99F995" box="[562,697,1559,1585]" italics="true" pageId="2" pageNumber="181">A. nitescens</emphasis>
(
<bibRefCitation id="D45272A5FFB2FFADFD56A85B2D78F995" author="Leach, W. E." box="[711,856,1558,1584]" pageId="2" pageNumber="181" pagination="383 - 437" refId="ref22744" refString="Leach, W. E. (1814) Crustaceology. In: Brewster, D. (Ed.), The Edinburgh Encyclopaedia, 7, pp. 383 - 437." type="journal article" year="1814">Leach, 1814</bibRefCitation>
)
</taxonomicName>
; all species listed in
<bibRefCitation id="D45272A5FFB2FFADFBDBA85A2BA5F995" author="De Grave, S. &amp; Fransen, C. H. J. M." box="[1098,1413,1558,1585]" pageId="2" pageNumber="181" pagination="195 - 589" refId="ref21881" refString="De Grave, S. &amp; Fransen, C. H. J. M. (2011) Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). Zoologische Mededelingen, Leiden, 85 (9), 195 - 589." type="journal article" year="2011">De Grave &amp; Fransen (2011)</bibRefCitation>
+
<taxonomicName id="77C374D7FFB2FFADFF06A8762C0AF9F1" authority="Duris &amp; Anker, 2014" authorityName="Duris &amp; Anker" authorityYear="2014" box="[151,554,1594,1623]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="manticolus">
<emphasis id="82B7D346FFB2FFADFF06A8762F12F9F1" box="[151,306,1594,1621]" italics="true" pageId="2" pageNumber="181">A. manticolus</emphasis>
<bibRefCitation id="D45272A5FFB2FFADFEA8A8712C0AF9F1" author="Duris, Z. &amp; Anker, A." box="[313,554,1594,1623]" pageId="2" pageNumber="181" pagination="550 - 558" refId="ref22106" refString="Duris, Z. &amp; Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš &amp; Anker, 2014</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="77C374D7FFB2FFADFDA4A8762D76F9F0" authority="Ahyong, 2015" authorityName="Ahyong" authorityYear="2015" box="[565,854,1594,1621]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="82B7D346FFB2FFADFDA4A8762C8CF9F1" box="[565,684,1594,1621]" italics="true" pageId="2" pageNumber="181">A. mendax</emphasis>
<bibRefCitation id="D45272A5FFB2FFADFD23A8772D76F9F0" author="Ahyong, S. T." box="[690,854,1594,1621]" pageId="2" pageNumber="181" pagination="405 - 442" refId="ref19152" refString="Ahyong, S. T. (2015) Decapod Crustacea of the Kermadec Biodiversity Expedition 2011. Bulletin of the Auckland Museum, 20, 405 - 442." type="journal article" year="2015">Ahyong, 2015</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="77C374D7FFB2FFADFCF3A8762AE1F9F1" authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[866,1217,1594,1621]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="alpheusophilus">
<emphasis id="82B7D346FFB2FFADFCF3A8762A0CF9F1" box="[866,1068,1594,1621]" italics="true" pageId="2" pageNumber="181">A. alpheusophilus</emphasis>
<bibRefCitation id="D45272A5FFB2FFADFBA2A8772AE1F9F1" author="Marin, I." box="[1075,1217,1594,1620]" pageId="2" pageNumber="181" pagination="50 - 62" refId="ref22879" refString="Marin, I. (2017) Athanas alpheusophilus sp. nov. (Decapoda: Alpheidae) - a new Alpheus - associated shrimp from the Russian coast of the Sea of Japan. Zootaxa, 4324 (1), 50 - 62. https: // doi. org / 10.11646 / zootaxa. 4324.1.3" type="journal article" year="2017">Marin, 2017</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="77C374D7FFB2FFADFB5CA8762F15F9DD" authority="Anker &amp; Duris 2022" authorityName="Anker &amp; Duris" authorityYear="2022" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="philippei">
<emphasis id="82B7D346FFB2FFADFB5CA8762B71F9F1" box="[1229,1361,1594,1621]" italics="true" pageId="2" pageNumber="181">A. philippei</emphasis>
<bibRefCitation id="D45272A5FFB2FFADFAC7A8772F15F9DD" author="Anker, A. &amp; Duris, Z." pageId="2" pageNumber="181" pagination="288 - 300" refId="ref20021" refString="Anker, A. &amp; Duris, Z. (2022) Athanas philippei n. sp., a new alpheid shrimp associated with stomatopods in New Caledonia (Decapoda: Caridea). Zootaxa, 5094 (2), 288 - 300. https: // doi. org / 10.11646 / zootaxa. 5094.2.4" type="journal article" year="2022">Anker &amp; Ďuriš 2022</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="77C374D7FFB2FFADFED1A8122C86F9DD" authority="Komai &amp; Henmi, 2023" authorityName="Komai &amp; Henmi" authorityYear="2023" box="[320,678,1630,1657]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="82B7D346FFB2FFADFED1A8122FBAF9DD" box="[320,410,1630,1657]" italics="true" pageId="2" pageNumber="181">A. exilis</emphasis>
<bibRefCitation id="D45272A5FFB2FFADFE30A8132C86F9DD" author="Komai, T. &amp; Henmi, Y." box="[417,678,1630,1656]" pageId="2" pageNumber="181" pagination="363 - 373" refId="ref22686" refString="Komai, T. &amp; Henmi, Y. (2023) A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from the Sea of Japan. Zootaxa, 5277 (2), 363 - 373. https: // doi. org / 10.11646 / zootaxa. 5277.2.6" type="journal article" year="2023">Komai &amp; Henmi, 2023</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="77C374D7FFB2FFADFD23A8122D7DF9DD" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[690,861,1630,1657]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB2FFADFD23A8122D7DF9DD" box="[690,861,1630,1657]" italics="true" pageId="2" pageNumber="181">A. claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFB2FFADFCF4A8132D9EF9DD" bold="true" box="[869,958,1630,1656]" pageId="2" pageNumber="181">
<taxonomicNameLabel id="99846E3DFFB2FFADFCF4A8132D9EF9DD" box="[869,958,1630,1656]" pageId="2" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(described below).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB2FFACFF56A8CF2EC2FE08" lastPageId="3" lastPageNumber="182" pageId="2" pageNumber="181" type="discussion">
<paragraph id="B07C0F54FFB2FFADFF56A8CF2CE3F845" blockId="2.[151,1437,1558,2017]" pageId="2" pageNumber="181">
<emphasis id="82B7D346FFB2FFADFF56A8CF2F14F939" bold="true" box="[199,308,1666,1692]" pageId="2" pageNumber="181">Remarks</emphasis>
.
<taxonomicName id="77C374D7FFB2FFADFED4A8CF2F81F939" authorityName="Leach" authorityYear="1814" box="[325,417,1666,1692]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB2FFADFED4A8CF2F81F939" box="[325,417,1666,1692]" italics="true" pageId="2" pageNumber="181">Athanas</emphasis>
</taxonomicName>
and allied genera were reviewed and taxonomically redefined by
<bibRefCitation id="D45272A5FFB2FFADFB0FA8CF2BB8F939" author="Anker, A. &amp; Jeng, M. S." box="[1182,1432,1666,1693]" pageId="2" pageNumber="181" pagination="454 - 472" refId="ref20209" refString="Anker, A. &amp; Jeng, M. S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinitions of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker &amp; Jeng (2007)</bibRefCitation>
. However, the rapidly increasing number of species, especially among the infaunal “commensals”, resulted in the much greater morphological heterogeneity of
<taxonomicName id="77C374D7FFB2FFADFD30A8872CDDF941" authorityName="Leach" authorityYear="1814" box="[673,765,1738,1764]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB2FFADFD30A8872CDDF941" box="[673,765,1738,1764]" italics="true" pageId="2" pageNumber="181">Athanas</emphasis>
</taxonomicName>
compared to the definition of the genus given by
<bibRefCitation id="D45272A5FFB2FFADFAA9A8872F3FF8AD" author="Anker, A. &amp; Jeng, M. S." pageId="2" pageNumber="181" pagination="454 - 472" refId="ref20209" refString="Anker, A. &amp; Jeng, M. S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinitions of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker &amp; Jeng (2007)</bibRefCitation>
. In the herein emended generic diagnosis of
<taxonomicName id="77C374D7FFB2FFADFC9DA8A32D48F8AD" authorityName="Leach" authorityYear="1814" box="[780,872,1774,1800]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB2FFADFC9DA8A32D48F8AD" box="[780,872,1774,1800]" italics="true" pageId="2" pageNumber="181">Athanas</emphasis>
</taxonomicName>
, the author attempted to take into consideration all important novel features described in the genus (including the below-described new species) since its redefinition by
<bibRefCitation id="D45272A5FFB2FFADFF29A97B2F89F8F5" author="Anker, A. &amp; Jeng, M. S." box="[184,425,1846,1873]" pageId="2" pageNumber="181" pagination="454 - 472" refId="ref20209" refString="Anker, A. &amp; Jeng, M. S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinitions of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker &amp; Jeng (2007)</bibRefCitation>
. It is interesting to note that the infaunal lifestyle that evolved in some lineages of
<taxonomicName id="77C374D7FFB2FFADFAD1A97B2BBCF8F5" authorityName="Leach" authorityYear="1814" box="[1344,1436,1846,1872]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB2FFADFAD1A97B2BBCF8F5" box="[1344,1436,1846,1872]" italics="true" pageId="2" pageNumber="181">Athanas</emphasis>
</taxonomicName>
and
<taxonomicName id="77C374D7FFB2FFADFF58A9172F62F8D1" authorityName="Holthuis" authorityYear="1955" box="[201,322,1882,1908]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB2FFADFF58A9172F62F8D1" box="[201,322,1882,1908]" italics="true" pageId="2" pageNumber="181">Salmoneus</emphasis>
</taxonomicName>
resulted in remarkable convergences between these two genera, for instance, in the shape of the frontal margin of the carapace (decrease in length of the rostrum and reduction of orbital teeth), reduction of the cornea size, strengthening of the antennal flagella, and appearance of either setal brushes (
<taxonomicName id="77C374D7FFB2FFADFB0CA9EF2AD9F819" authorityName="Leach" authorityYear="1814" box="[1181,1273,1954,1980]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB2FFADFB0CA9EF2AD9F819" box="[1181,1273,1954,1980]" italics="true" pageId="2" pageNumber="181">Athanas</emphasis>
</taxonomicName>
) or setal rows (
<taxonomicName id="77C374D7FFB2FFADFF30A98B2F3AF845" authorityName="Holthuis" authorityYear="1955" box="[161,282,1990,2016]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB2FFADFF30A98B2F3AF845" box="[161,282,1990,2016]" italics="true" pageId="2" pageNumber="181">Salmoneus</emphasis>
</taxonomicName>
) on the chelae of the first pereiopods.
</paragraph>
<paragraph id="B07C0F54FFB3FFACFF56AEDA2EC2FE08" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="182">
The recent phylogenetic hypothesis for the
<taxonomicName id="77C374D7FFB3FFACFD27AEDA2D09FF14" authorityName="Rafinesque" authorityYear="1815" box="[694,809,151,177]" class="Malacostraca" family="Alpheidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="182" phylum="Arthropoda" rank="family">Alpheidae</taxonomicName>
based on molecular data (
<bibRefCitation id="D45272A5FFB3FFACFBC6AEDA2B00FF14" author="Chow, L. H. &amp; De Grave, S. &amp; Anker, A. &amp; Poon, K. K. Y. &amp; Ma, K. &amp; Chu, K. &amp; Chan, T. Y. &amp; Tsang, L. M." box="[1111,1312,151,177]" pageId="3" pageNumber="182" pagination="2898 - 2910" refId="ref21336" refString="Chow, L. H., De Grave, S., Anker, A., Poon, K. K. Y., Ma, K. Y, Chu, K. H, Chan, T. Y. &amp; Tsang, L. M. (2021) Distinct suites of pre- and post-adaptations indicate independent evolutionary pathways of snapping claws in the shrimp family Alpheidae (Decapoda: Caridea). Evolution, 75 (11), 2898 - 2910. https: // doi. org / 10.1111 / evo. 14351" type="journal article" year="2021">
Chow
<emphasis id="82B7D346FFB3FFACFB34AEDA2AFFFF14" box="[1189,1247,151,177]" italics="true" pageId="3" pageNumber="182">et al.</emphasis>
2021
</bibRefCitation>
) showed a non-monophyly of
<taxonomicName id="77C374D7FFB3FFACFEFFAEF62FEAFF70" authorityName="Leach" authorityYear="1814" box="[366,458,187,213]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB3FFACFEFFAEF62FEAFF70" box="[366,458,187,213]" italics="true" pageId="3" pageNumber="182">Athanas</emphasis>
</taxonomicName>
in its present composition (see also discussion under
<taxonomicName id="77C374D7FFB3FFACFB8AAEF62A98FF70" authorityName="Bruce" authorityYear="1983" box="[1051,1208,187,213]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB3FFACFB8AAEF62A98FF70" box="[1051,1208,187,213]" italics="true" pageId="3" pageNumber="182">Pseudathanas</emphasis>
</taxonomicName>
). Another important result of this study was the lack of support for the two currently recognised, informal species groups within the genus, which were in use since
<bibRefCitation id="D45272A5FFB3FFACFE6EAF4E2C94FEB8" author="Coutiere, H." box="[511,692,259,285]" pageId="3" pageNumber="182" pagination="1 - 6" refId="ref21527" refString="Coutiere, H. (1899) Les &quot; Alpheidae &quot;, morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie, Serie 8, Vol. 9. Masson, Paris, 559 pp., pls. 1 - 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="book chapter" year="1899">Coutière (1899)</bibRefCitation>
, namely the
<taxonomicName id="77C374D7FFB3FFACFCD7AF4E2A53FEB8" authority="(Leach, 1814)" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[838,1139,259,285]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="nitescens">
<emphasis id="82B7D346FFB3FFACFCD7AF4E2DEDFEB9" box="[838,973,259,285]" italics="true" pageId="3" pageNumber="182">A. nitescens</emphasis>
(
<bibRefCitation id="D45272A5FFB3FFACFC4DAF4E2A4AFEB8" author="Leach, W. E." box="[988,1130,259,285]" pageId="3" pageNumber="182" pagination="383 - 437" refId="ref22744" refString="Leach, W. E. (1814) Crustaceology. In: Brewster, D. (Ed.), The Edinburgh Encyclopaedia, 7, pp. 383 - 437." type="journal article" year="1814">Leach, 1814</bibRefCitation>
)
</taxonomicName>
group (containing species with extended chelipeds) and the
<taxonomicName id="77C374D7FFB3FFACFD8EAF6A2D50FEE4" authority="Ortmann, 1890" authorityName="Ortmann" authorityYear="1890" box="[543,880,295,321]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis id="82B7D346FFB3FFACFD8EAF6A2C99FEE4" box="[543,697,295,321]" italics="true" pageId="3" pageNumber="182">A. dimorphus</emphasis>
<bibRefCitation id="D45272A5FFB3FFACFD52AF6A2D50FEE4" author="Ortmann, A." box="[707,880,295,321]" pageId="3" pageNumber="182" pagination="437 - 542" refId="ref23182" refString="Ortmann, A. (1890) Die Unterordnung Natantia Boas. Theil I. Die Decapoden Krebse des Strassburger Museums, mit besonderer Beru ¨ cksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. Zoologische Jahrbu ¨ cher, Systematik, 5, 437 - 542, pls. 36, 37." type="book chapter" year="1890">Ortmann, 1890</bibRefCitation>
</taxonomicName>
group (containing species with ventrally folded chelipeds). Therefore, the herein provided diagnosis of
<taxonomicName id="77C374D7FFB3FFACFC82AF062D4FFEC0" authorityName="Leach" authorityYear="1814" box="[787,879,331,357]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB3FFACFC82AF062D4FFEC0" box="[787,879,331,357]" italics="true" pageId="3" pageNumber="182">Athanas</emphasis>
</taxonomicName>
should be used with some reservation as further taxonomic rearrangements, with redefinitions of genera and their diagnostic features, are expected in the near future.
</paragraph>
</subSubSection>
</treatment>
</document>
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