629 lines
87 KiB
XML
629 lines
87 KiB
XML
<document id="F05A3E7C708FC373E710F8FA9CFA5278" ID-DOI="10.11646/zootaxa.5383.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350767" ID-ZooBank="7E434B40-6346-4E6D-BC55-38EBAE24BD52" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1702283478140" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2023" docId="386ABE42FFACFFB1FF06ADBB2C8FFD68" docLanguage="en" docName="zootaxa.5383.2.5.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudathanas Bruce 1983" docType="treatment" docVersion="3" lastPageNumber="209" masterDocId="C453C63AFFB0FFAFFF91AE4D2E20FFA5" masterDocTitle="Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="215" masterPageNumber="179" pageNumber="207" updateTime="1702658426235" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="3B82679A2700598B7202C7FB76793004">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
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<mods:namePart id="45242230E8B7C7484652FA9AFBFC3A45">Anker, Arthur</mods:namePart>
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<mods:affiliation id="2350D93BF4A7D06EB9C6961D63D276EA">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil & Red Sea Research Center, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia</mods:affiliation>
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<mods:title id="1D255618B8660D119749E18709E707AD">Zootaxa</mods:title>
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<mods:date id="217E4DEB294DC3C6E047478AF11AF45D">2023</mods:date>
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<mods:number id="5F2DD47EDFFC60A6CA88D229F1DBF899">2023-12-11</mods:number>
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<mods:classification id="29EB21009ED7C0845EE0E2B8EE5F84A5">journal article</mods:classification>
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<subSubSection id="F8D95CDFFFACFFB3FF06ADBB2C14FBB5" box="[151,564,1014,1041]" pageId="28" pageNumber="207" type="nomenclature">
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<paragraph id="B07C0F54FFACFFB3FF06ADBB2C14FBB5" blockId="28.[151,564,1014,1041]" box="[151,564,1014,1041]" pageId="28" pageNumber="207">
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<heading id="EB34B838FFACFFB3FF06ADBB2C14FBB5" bold="true" box="[151,564,1014,1041]" fontSize="11" level="1" pageId="28" pageNumber="207" reason="1">
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<emphasis id="82B7D346FFACFFB3FF06ADBB2C14FBB5" bold="true" box="[151,564,1014,1041]" pageId="28" pageNumber="207">
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Genus
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<taxonomicName id="77C374D7FFACFFB3FF7FADBB2C14FBB5" ID-CoL="85HN4" authority="Bruce, 1983" authorityName="Bruce" authorityYear="1983" box="[238,564,1014,1041]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="genus">
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<emphasis id="82B7D346FFACFFB3FF7FADBB2FB8FBB4" bold="true" box="[238,408,1014,1041]" italics="true" pageId="28" pageNumber="207">Pseudathanas</emphasis>
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<bibRefCitation id="D45272A5FFACFFB3FE0EADBA2C14FBB5" author="Bruce, A. J." box="[415,564,1014,1041]" pageId="28" pageNumber="207" pagination="463 - 471" refId="ref21222" refString="Bruce, A. J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463 - 471. https: // doi. org / 10.2307 / 1548146" type="journal article" year="1983">Bruce, 1983</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F8D95CDFFFACFFB3FF06AA762C24FBF6" box="[151,516,1083,1107]" pageId="28" pageNumber="207" type="reference_group">
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<paragraph id="B07C0F54FFACFFB3FF06AA762C24FBF6" blockId="28.[151,516,1083,1107]" box="[151,516,1083,1107]" pageId="28" pageNumber="207">
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<treatmentCitationGroup id="90D3287AFFACFFB3FF06AA762C24FBF6" box="[151,516,1083,1107]" pageId="28" pageNumber="207">
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<treatmentCitation id="31622945FFACFFB3FF06AA762C20FBF6" author="Kazmi, Q. B. & Kazmi, M. A." box="[151,512,1083,1107]" page="233" pageId="28" pageNumber="207" year="2010">
|
||
<taxonomicName id="77C374D7FFACFFB3FF06AA762FEEFBF6" ID-CoL="85HN4" authority="Kazmi & Kazmi, 2010" authorityName="Kazmi & Kazmi" authorityYear="2010" box="[151,462,1083,1107]" class="Malacostraca" family="Alpheidae" genus="Alberta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFACFFB3FF06AA762EC3FBF6" box="[151,227,1083,1107]" italics="true" pageId="28" pageNumber="207">Alberta</emphasis>
|
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<bibRefCitation id="D45272A5FFACFFB3FF7BAA762C20FBF6" author="Kazmi, Q. B. & Kazmi, M. A." box="[234,512,1083,1107]" pageId="28" pageNumber="207" refId="ref22537" refString="Kazmi, Q. B. & Kazmi, M. A. (2010) Biodiversity and biogeography of caridean shrimps of Pakistan. Higher Education Commission, Islamabad, xvii + 516 pp." type="book" year="2010">Kazmi & Kazmi 2010: 233</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="F8D95CDFFFACFFB2FF06AA362BACFD3B" lastPageId="29" lastPageNumber="208" pageId="28" pageNumber="207" type="diagnosis">
|
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<paragraph id="B07C0F54FFACFFB3FF06AA362D86F850" blockId="28.[151,1437,1146,2037]" pageId="28" pageNumber="207">
|
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<emphasis id="82B7D346FFACFFB3FF06AA362FA2FB31" bold="true" box="[151,386,1146,1173]" pageId="28" pageNumber="207">Emended diagnosis</emphasis>
|
||
. Body not compressed, not particularly slender. Carapace smooth, unarmed dorsally and laterally. Rostrum broadly triangular, unarmed dorsally and ventrally. Orbital teeth absent or reduced. Pterygostomial angle rounded, not protruding; posteroventral margin of branchiostegite fringed with long plumose setae; cardiac notch well developed. All pleonites with distoventrally rounded pleura; sixth pleuron with articulated plate. Telson moderately broad, with two pairs of stout spiniform setae on dorsal surface; posterior margin rounded, with two pairs of spiniform setae; anal tubercles absent. Eyes well developed, partly visible or concealed in dorsal view; cornea large, well pigmented. Antennular peduncle stout, relatively short; stylocerite well developed, distally acute, overreaching mid-length of second article of peduncle; ventromesial carina armed with tooth; second article not particularly elongate; lateral flagellum with short fused portion and well-developed accessory ramus with aesthetascs. Antenna with basicerite stout, unarmed or armed with small sharp tooth; scaphocerite with well-developed blade and small distolateral tooth. Mandible with two-articulated palp; molar and incisor processes well developed. Third maxilliped with well-developed exopod; coxa with acutely produced lateral plate; ultimate article tapering into corneous tip, unarmed. First pereiopods (= chelipeds) enlarged in both sexes, unequal or subequal in size, asymmetrical or subsymmetrical in shape, carried flexed beneath body when not in use; ischium robust, with dorsal and ventral margins rugose and armed with short spiniform setae; merus with ventrolateral margin rugose and armed with prominent tooth near its mid-length, ventral surface deeply excavated; carpus vase- or cup-shaped, without setal rows on mesial surface; chelae more or less swollen; palm with tubercles and row of setae on ventral surface; fingers of major chela armed with large teeth, without snapping mechanism. Second pereiopod with ischium unarmed; carpus with four or five subarticles; chela simple, without modifications. Third and fourth pereiopods moderately slender; ischium armed with one or two spiniform seta(e); merus unarmed; carpus with one spiniform seta on distoventral margin; propodus with several spiniform setae; dactylus simple, conical. Fifth pereiopod with ischium armed with one spiniform seta; merus unarmed; carpus with one spiniform seta on distoventral margin; propodus with at least two spiniform setae and short cleaning brush; dactylus simple. Second male pleopod with appendix masculina subequal in length to appendix interna; second female pleopod with appendix interna only. Uropodal exopod with diaeresis armed with row of stout spiniform setae. Gill-exopod formula provided in
|
||
<tableCitation id="FD413AEFFFACFFB3FCDCA9962D81F850" box="[845,929,2011,2037]" captionStart="TABLE 3" captionStartId="29.[152,244,151,175]" captionTargetPageId="29" captionText="TABLE 3. Gill-exopod formula of Pseudathanas Bruce, 1983." pageId="28" pageNumber="207">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="E4BC5FDCFFADFFB2FF09AEDA2D1BFF0A" box="[152,827,151,175]" pageId="29" pageNumber="208" startId="29.[152,244,151,175]" targetBox="[163,1384,191,537]" targetIsTable="true" targetPageId="29" targetType="table">
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<paragraph id="B07C0F54FFADFFB2FF09AEDA2D1BFF0A" blockId="29.[152,827,151,175]" box="[152,827,151,175]" pageId="29" pageNumber="208">
|
||
<emphasis id="82B7D346FFADFFB2FF09AEDA2F28FF0A" bold="true" box="[152,264,151,175]" pageId="29" pageNumber="208">TABLE 3</emphasis>
|
||
. Gill-exopod formula of
|
||
<taxonomicName id="77C374D7FFADFFB2FD84AEDA2D17FF0A" authority="Bruce, 1983" authorityName="Bruce" authorityYear="1983" box="[533,823,151,175]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FD84AEDA2C8BFF0A" box="[533,683,151,175]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
<bibRefCitation id="D45272A5FFADFFB2FD20AED52D17FF0A" author="Bruce, A. J." box="[689,823,151,175]" pageId="29" pageNumber="208" pagination="463 - 471" refId="ref21222" refString="Bruce, A. J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463 - 471. https: // doi. org / 10.2307 / 1548146" type="journal article" year="1983">Bruce, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B07C0F54FFADFFB2FF32AEF22B7EFDBC" pageId="29" pageNumber="208">
|
||
<table id="C2C3FDF4FFAD0050FF32AEF22B48FDBC" box="[163,1384,191,537]" gridcols="9" gridrows="9" pageId="29" pageNumber="208">
|
||
<tr id="0EF30D16FFAD0050FF32AEF22B48FF72" box="[163,1384,191,215]" gridrow="0" pageId="29" pageNumber="208" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="4D22646AFFAD0050FF32AEF22F6BFF72" box="[163,331,191,215]" gridcol="0" gridrow="0" pageId="29" pageNumber="208">Gills/exopods</th>
|
||
<th id="4D22646AFFAD0050FD9BAEF22CA3FF72" box="[522,643,191,215]" gridcol="2" gridrow="0" pageId="29" pageNumber="208">Maxillipeds</th>
|
||
<th id="4D22646AFFAD0050FB8EAEF22AB0FF72" box="[1055,1168,191,215]" gridcol="6" gridrow="0" pageId="29" pageNumber="208">Pereiopods</th>
|
||
</tr>
|
||
<tr id="0EF30D16FFAD0050FF32AEAA2B48FEA5" box="[163,1384,231,256]" gridrow="1" pageId="29" pageNumber="208" rowspan-0="1">
|
||
<td id="4D22646AFFAD0050FE32AEAA2FFFFEA5" box="[419,479,231,256]" gridcol="1" gridrow="1" pageId="29" pageNumber="208">Mxp1</td>
|
||
<td id="4D22646AFFAD0050FD9BAEAA2CA3FEA5" box="[522,643,231,256]" gridcol="2" gridrow="1" pageId="29" pageNumber="208">Mxp2</td>
|
||
<td id="4D22646AFFAD0050FD23AEAA2CCFFEA5" box="[690,751,231,256]" gridcol="3" gridrow="1" pageId="29" pageNumber="208">Mxp3</td>
|
||
<td id="4D22646AFFAD0050FCD8AEAA2D43FEA5" box="[841,867,231,256]" gridcol="4" gridrow="1" pageId="29" pageNumber="208">P1</td>
|
||
<td id="4D22646AFFAD0050FC5DAEAA2DC7FEA5" box="[972,999,231,256]" gridcol="5" gridrow="1" pageId="29" pageNumber="208">P2</td>
|
||
<td id="4D22646AFFAD0050FB8EAEAA2AB0FEA5" box="[1055,1168,231,256]" gridcol="6" gridrow="1" pageId="29" pageNumber="208">P3</td>
|
||
<td id="4D22646AFFAD0050FB5AAEAA2AC6FEA5" box="[1227,1254,231,256]" gridcol="7" gridrow="1" pageId="29" pageNumber="208">P4</td>
|
||
<td id="4D22646AFFAD0050FADCAEAA2B48FEA5" box="[1357,1384,231,256]" gridcol="8" gridrow="1" pageId="29" pageNumber="208">P5</td>
|
||
</tr>
|
||
<tr id="0EF30D16FFAD0050FF32AF5D2B48FE8D" box="[163,1384,272,296]" gridrow="2" pageId="29" pageNumber="208">
|
||
<th id="4D22646AFFAD0050FF32AF5D2F6BFE8D" box="[163,331,272,296]" gridcol="0" gridrow="2" pageId="29" pageNumber="208">Pleurobranchs</th>
|
||
<td id="4D22646AFFAD0050FE32AF5D2FFFFE8D" box="[419,479,272,296]" gridcol="1" gridrow="2" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FD9BAF5D2CA3FE8D" box="[522,643,272,296]" gridcol="2" gridrow="2" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FD23AF5D2CCFFE8D" box="[690,751,272,296]" gridcol="3" gridrow="2" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FCD8AF5D2D43FE8D" box="[841,867,272,296]" gridcol="4" gridrow="2" pageId="29" pageNumber="208">1</td>
|
||
<td id="4D22646AFFAD0050FC5DAF5D2DC7FE8D" box="[972,999,272,296]" gridcol="5" gridrow="2" pageId="29" pageNumber="208">1</td>
|
||
<td id="4D22646AFFAD0050FB8EAF5D2AB0FE8D" box="[1055,1168,272,296]" gridcol="6" gridrow="2" pageId="29" pageNumber="208">1</td>
|
||
<td id="4D22646AFFAD0050FB5AAF5D2AC6FE8D" box="[1227,1254,272,296]" gridcol="7" gridrow="2" pageId="29" pageNumber="208">1</td>
|
||
<td id="4D22646AFFAD0050FADCAF5D2B48FE8D" box="[1357,1384,272,296]" gridcol="8" gridrow="2" pageId="29" pageNumber="208">1</td>
|
||
</tr>
|
||
<tr id="0EF30D16FFAD0050FF32AF752B48FEF5" box="[163,1384,312,336]" gridrow="3" pageId="29" pageNumber="208">
|
||
<th id="4D22646AFFAD0050FF32AF752F6BFEF5" box="[163,331,312,336]" gridcol="0" gridrow="3" pageId="29" pageNumber="208">Arthrobranchs</th>
|
||
<td id="4D22646AFFAD0050FE32AF752FFFFEF5" box="[419,479,312,336]" gridcol="1" gridrow="3" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FD9BAF752CA3FEF5" box="[522,643,312,336]" gridcol="2" gridrow="3" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FD23AF752CCFFEF5" box="[690,751,312,336]" gridcol="3" gridrow="3" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FCD8AF752D43FEF5" box="[841,867,312,336]" gridcol="4" gridrow="3" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FC5DAF752DC7FEF5" box="[972,999,312,336]" gridcol="5" gridrow="3" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FB8EAF752AB0FEF5" box="[1055,1168,312,336]" gridcol="6" gridrow="3" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FB5AAF752AC6FEF5" box="[1227,1254,312,336]" gridcol="7" gridrow="3" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FADCAF752B48FEF5" box="[1357,1384,312,336]" gridcol="8" gridrow="3" pageId="29" pageNumber="208">-</td>
|
||
</tr>
|
||
<tr id="0EF30D16FFAD0050FF32AF2D2B48FEDD" box="[163,1384,352,376]" gridrow="4" pageId="29" pageNumber="208">
|
||
<th id="4D22646AFFAD0050FF32AF2D2F6BFEDD" box="[163,331,352,376]" gridcol="0" gridrow="4" pageId="29" pageNumber="208">Podobranchs</th>
|
||
<td id="4D22646AFFAD0050FE32AF2D2FFFFEDD" box="[419,479,352,376]" gridcol="1" gridrow="4" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FD9BAF2D2CA3FEDD" box="[522,643,352,376]" gridcol="2" gridrow="4" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FD23AF2D2CCFFEDD" box="[690,751,352,376]" gridcol="3" gridrow="4" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FCD8AF2D2D43FEDD" box="[841,867,352,376]" gridcol="4" gridrow="4" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FC5DAF2D2DC7FEDD" box="[972,999,352,376]" gridcol="5" gridrow="4" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FB8EAF2D2AB0FEDD" box="[1055,1168,352,376]" gridcol="6" gridrow="4" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FB5AAF2D2AC6FEDD" box="[1227,1254,352,376]" gridcol="7" gridrow="4" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FADCAF2D2B48FEDD" box="[1357,1384,352,376]" gridcol="8" gridrow="4" pageId="29" pageNumber="208">-</td>
|
||
</tr>
|
||
<tr id="0EF30D16FFAD0050FF32AFCA2B48FE05" box="[163,1384,391,416]" gridrow="5" pageId="29" pageNumber="208">
|
||
<th id="4D22646AFFAD0050FF32AFCA2F6BFE05" box="[163,331,391,416]" gridcol="0" gridrow="5" pageId="29" pageNumber="208">Epipods</th>
|
||
<td id="4D22646AFFAD0050FE32AFCA2FFFFE05" box="[419,479,391,416]" gridcol="1" gridrow="5" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FD9BAFCA2CA3FE05" box="[522,643,391,416]" gridcol="2" gridrow="5" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FD23AFCA2CCFFE05" box="[690,751,391,416]" gridcol="3" gridrow="5" pageId="29" pageNumber="208">+1</td>
|
||
<td id="4D22646AFFAD0050FCD8AFCA2D43FE05" box="[841,867,391,416]" gridcol="4" gridrow="5" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FC5DAFCA2DC7FE05" box="[972,999,391,416]" gridcol="5" gridrow="5" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FB8EAFCA2AB0FE05" box="[1055,1168,391,416]" gridcol="6" gridrow="5" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FB5AAFCA2AC6FE05" box="[1227,1254,391,416]" gridcol="7" gridrow="5" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FADCAFCA2B48FE05" box="[1357,1384,391,416]" gridcol="8" gridrow="5" pageId="29" pageNumber="208">-</td>
|
||
</tr>
|
||
<tr id="0EF30D16FFAD0050FF32AFFD2B48FE6C" box="[163,1384,432,457]" gridrow="6" pageId="29" pageNumber="208">
|
||
<th id="4D22646AFFAD0050FF32AFFD2F6BFE6C" box="[163,331,432,457]" gridcol="0" gridrow="6" pageId="29" pageNumber="208">Mastigobranchs2</th>
|
||
<td id="4D22646AFFAD0050FE32AFFD2FFFFE6C" box="[419,479,432,457]" gridcol="1" gridrow="6" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FD9BAFFD2CA3FE6C" box="[522,643,432,457]" gridcol="2" gridrow="6" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FD23AFFD2CCFFE6C" box="[690,751,432,457]" gridcol="3" gridrow="6" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FCD8AFFD2D43FE6C" box="[841,867,432,457]" gridcol="4" gridrow="6" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FC5DAFFD2DC7FE6C" box="[972,999,432,457]" gridcol="5" gridrow="6" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FB8EAFFD2AB0FE6C" box="[1055,1168,432,457]" gridcol="6" gridrow="6" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FB5AAFFD2AC6FE6C" box="[1227,1254,432,457]" gridcol="7" gridrow="6" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FADCAFFD2B48FE6C" box="[1357,1384,432,457]" gridcol="8" gridrow="6" pageId="29" pageNumber="208">-</td>
|
||
</tr>
|
||
<tr id="0EF30D16FFAD0050FF32AF942B48FE54" box="[163,1384,473,497]" gridrow="7" pageId="29" pageNumber="208">
|
||
<th id="4D22646AFFAD0050FF32AF942F6BFE54" box="[163,331,473,497]" gridcol="0" gridrow="7" pageId="29" pageNumber="208">Setobranchs</th>
|
||
<td id="4D22646AFFAD0050FE32AF942FFFFE54" box="[419,479,473,497]" gridcol="1" gridrow="7" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FD9BAF942CA3FE54" box="[522,643,473,497]" gridcol="2" gridrow="7" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FD23AF942CCFFE54" box="[690,751,473,497]" gridcol="3" gridrow="7" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FCD8AF942D43FE54" box="[841,867,473,497]" gridcol="4" gridrow="7" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FC5DAF942DC7FE54" box="[972,999,473,497]" gridcol="5" gridrow="7" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FB8EAF942AB0FE54" box="[1055,1168,473,497]" gridcol="6" gridrow="7" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FB5AAF942AC6FE54" box="[1227,1254,473,497]" gridcol="7" gridrow="7" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FADCAF942B48FE54" box="[1357,1384,473,497]" gridcol="8" gridrow="7" pageId="29" pageNumber="208">-</td>
|
||
</tr>
|
||
<tr id="0EF30D16FFAD0050FF32AC4D2B48FDBC" box="[163,1384,512,537]" gridrow="8" pageId="29" pageNumber="208">
|
||
<th id="4D22646AFFAD0050FF32AC4D2F6BFDBC" box="[163,331,512,537]" gridcol="0" gridrow="8" pageId="29" pageNumber="208">Exopods</th>
|
||
<td id="4D22646AFFAD0050FE32AC4D2FFFFDBC" box="[419,479,512,537]" gridcol="1" gridrow="8" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FD9BAC4D2CA3FDBC" box="[522,643,512,537]" gridcol="2" gridrow="8" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FD23AC4D2CCFFDBC" box="[690,751,512,537]" gridcol="3" gridrow="8" pageId="29" pageNumber="208">+</td>
|
||
<td id="4D22646AFFAD0050FCD8AC4D2D43FDBC" box="[841,867,512,537]" gridcol="4" gridrow="8" pageId="29" pageNumber="208">-3</td>
|
||
<td id="4D22646AFFAD0050FC5DAC4D2DC7FDBC" box="[972,999,512,537]" gridcol="5" gridrow="8" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FB8EAC4D2AB0FDBC" box="[1055,1168,512,537]" gridcol="6" gridrow="8" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FB5AAC4D2AC6FDBC" box="[1227,1254,512,537]" gridcol="7" gridrow="8" pageId="29" pageNumber="208">-</td>
|
||
<td id="4D22646AFFAD0050FADCAC4D2B48FDBC" box="[1357,1384,512,537]" gridcol="8" gridrow="8" pageId="29" pageNumber="208">-</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFADFFB2FF06AC682E8FFD96" blockId="29.[151,721,549,575]" box="[151,175,549,563]" pageId="29" pageNumber="208">
|
||
<superScript id="47B6A21CFFADFFB2FF06AC682E8FFD96" attach="left" box="[151,175,549,563]" fontSize="6" pageId="29" pageNumber="208">1, 2</superScript>
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFADFFB2FF24AC6A2CF1FD9A" blockId="29.[151,721,549,575]" box="[181,721,549,575]" pageId="29" pageNumber="208">
|
||
same remarks as for Table 1.
|
||
<superScript id="47B6A21CFFADFFB2FE7FAC682FD6FD96" attach="right" box="[494,502,549,563]" fontSize="6" pageId="29" pageNumber="208">3</superScript>
|
||
needs confirmation.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFADFFB2FF56ACCE2BACFD3B" blockId="29.[151,1437,643,2038]" box="[199,1420,643,670]" pageId="29" pageNumber="208">
|
||
<emphasis id="82B7D346FFADFFB2FF56ACCE2FACFD38" bold="true" box="[199,396,643,669]" pageId="29" pageNumber="208">Species included</emphasis>
|
||
.
|
||
<typeStatus id="6F78B1F6FFADFFB2FE08ACC92FF1FD3B" box="[409,465,644,670]" pageId="29" pageNumber="208">Type</typeStatus>
|
||
species
|
||
<taxonomicName id="77C374D7FFADFFB2FDA1ACC92D4DFD3B" authority="Bruce, 1983" authorityName="Bruce" authorityYear="1983" box="[560,877,643,670]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="darwiniensis">
|
||
<emphasis id="82B7D346FFADFFB2FDA1ACC92CFAFD38" box="[560,730,643,669]" italics="true" pageId="29" pageNumber="208">P. darwiniensis</emphasis>
|
||
<bibRefCitation id="D45272A5FFADFFB2FD73ACC92D4DFD3B" author="Bruce, A. J." box="[738,877,644,670]" pageId="29" pageNumber="208" pagination="463 - 471" refId="ref21222" refString="Bruce, A. J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463 - 471. https: // doi. org / 10.2307 / 1548146" type="journal article" year="1983">Bruce, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="77C374D7FFADFFB2FCE8ACC92B25FD3B" authority="(Kazmi & Kazmi, 2010)" authorityName="Anker" authorityYear="2023" baseAuthorityName="Kazmi & Kazmi" baseAuthorityYear="2010" box="[889,1285,643,670]" class="Magnoliopsida" family="Rubiaceae" genus="Alberta" kingdom="Plantae" order="Gentianales" pageId="29" pageNumber="208" phylum="Tracheophyta" rank="species" species="banneri" status="comb. nov.">
|
||
<emphasis id="82B7D346FFADFFB2FCE8ACC92DCCFD38" box="[889,1004,643,669]" italics="true" pageId="29" pageNumber="208">P. banneri</emphasis>
|
||
(
|
||
<bibRefCitation id="D45272A5FFADFFB2FC6DACCE2ADDFD3B" author="Kazmi, Q. B. & Kazmi, M. A." box="[1020,1277,643,670]" pageId="29" pageNumber="208" refId="ref22537" refString="Kazmi, Q. B. & Kazmi, M. A. (2010) Biodiversity and biogeography of caridean shrimps of Pakistan. Higher Education Commission, Islamabad, xvii + 516 pp." type="book" year="2010">Kazmi & Kazmi, 2010</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFADFFB2FA9AACC92BACFD3B" bold="true" box="[1291,1420,644,670]" pageId="29" pageNumber="208">
|
||
<taxonomicNameLabel id="99846E3DFFADFFB2FA9AACC92BACFD3B" box="[1291,1420,644,670]" pageId="29" pageNumber="208" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8D95CDFFFADFFB1FF56ACE52C8FFD68" lastPageId="30" lastPageNumber="209" pageId="29" pageNumber="208" type="discussion">
|
||
<paragraph id="B07C0F54FFADFFB2FF56ACE52DF5FC1B" blockId="29.[151,1437,643,2038]" pageId="29" pageNumber="208">
|
||
<emphasis id="82B7D346FFADFFB2FF56ACE52F14FD67" bold="true" box="[199,308,680,706]" pageId="29" pageNumber="208">Remarks</emphasis>
|
||
. The alpheid genus
|
||
<taxonomicName id="77C374D7FFADFFB2FD8BACEA2C97FD64" authorityName="Bruce" authorityYear="1983" box="[538,695,679,705]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FD8BACEA2C97FD64" box="[538,695,679,705]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
was originally established to accommodate only the
|
||
<typeStatus id="6F78B1F6FFADFFB2FA82ACE52B63FD67" box="[1299,1347,680,706]" pageId="29" pageNumber="208">type</typeStatus>
|
||
species
|
||
<taxonomicName id="77C374D7FFADFFB2FF06AC862C7BFD43" authority="Bruce, 1983" authorityName="Bruce" authorityYear="1983" box="[151,603,715,742]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="darwiniensis">
|
||
<emphasis id="82B7D346FFADFFB2FF06AC862FEAFD40" box="[151,458,715,741]" italics="true" pageId="29" pageNumber="208">Pseudathanas darwiniensis</emphasis>
|
||
<bibRefCitation id="D45272A5FFADFFB2FE41AC812C7BFD43" author="Bruce, A. J." box="[464,603,716,742]" pageId="29" pageNumber="208" pagination="463 - 471" refId="ref21222" refString="Bruce, A. J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463 - 471. https: // doi. org / 10.2307 / 1548146" type="journal article" year="1983">Bruce, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
, a small, peculiar, intertidal shrimp found in silted rock pools near Darwin,
|
||
<collectingRegion id="7207C1B6FFADFFB2FF06ACA22F49FCAC" box="[151,361,751,777]" country="Australia" name="Northern Territory" pageId="29" pageNumber="208">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingCountry id="C8D44FC4FFADFFB2FEE4ACA22FFCFCAC" box="[373,476,751,777]" name="Australia" pageId="29" pageNumber="208">Australia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="D45272A5FFADFFB2FE7CACBD2C57FCAF" author="Bruce, A. J." box="[493,631,752,778]" pageId="29" pageNumber="208" pagination="463 - 471" refId="ref21222" refString="Bruce, A. J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463 - 471. https: // doi. org / 10.2307 / 1548146" type="journal article" year="1983">Bruce 1983</bibRefCitation>
|
||
).
|
||
<taxonomicName id="77C374D7FFADFFB2FD1CACA22D0AFCAC" authorityName="Bruce" authorityYear="1983" box="[653,810,751,777]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FD1CACA22D0AFCAC" box="[653,810,751,777]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
was separated from the closely related genus
|
||
<taxonomicName id="77C374D7FFADFFB2FAD1ACA22BBCFCAC" authorityName="Leach" authorityYear="1814" box="[1344,1436,751,777]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FAD1ACA22BBCFCAC" box="[1344,1436,751,777]" italics="true" pageId="29" pageNumber="208">Athanas</emphasis>
|
||
</taxonomicName>
|
||
by the much shorter rostrum, the greatly reduced orbital teeth, the proximally thickened antennal flagella, the unequal and asymmetrical chelipeds, and the presence of stout spiniform setae on the transverse suture (diaeresis) of the uropodal exopod (
|
||
<bibRefCitation id="D45272A5FFADFFB2FE29AD112C62FCD3" author="Bruce, A. J." box="[440,578,860,886]" pageId="29" pageNumber="208" pagination="463 - 471" refId="ref21222" refString="Bruce, A. J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463 - 471. https: // doi. org / 10.2307 / 1548146" type="journal article" year="1983">Bruce 1983</bibRefCitation>
|
||
), this latter feature being unique within the
|
||
<taxonomicName id="77C374D7FFADFFB2FBD7AD162A99FCD0" authorityName="Rafinesque" authorityYear="1815" box="[1094,1209,859,885]" class="Malacostraca" family="Alpheidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="family">Alpheidae</taxonomicName>
|
||
and therefore most diagnostic of the genus. No other specimens of
|
||
<taxonomicName id="77C374D7FFADFFB2FD3EAD322D6CFC3C" authorityName="Bruce" authorityYear="1983" box="[687,844,895,921]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FD3EAD322D6CFC3C" box="[687,844,895,921]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
have been reported since
|
||
<bibRefCitation id="D45272A5FFADFFB2FBE0ADCD2B38FC3F" author="Bruce, A. J." box="[1137,1304,896,922]" pageId="29" pageNumber="208" pagination="463 - 471" refId="ref21222" refString="Bruce, A. J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463 - 471. https: // doi. org / 10.2307 / 1548146" type="journal article" year="1983">Bruce’s (1983)</bibRefCitation>
|
||
description of
|
||
<taxonomicName id="77C374D7FFADFFB2FF27ADE92F40FC18" authorityName="Bruce" authorityYear="1983" box="[182,352,931,957]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="darwiniensis">
|
||
<emphasis id="82B7D346FFADFFB2FF27ADE92F40FC18" box="[182,352,931,957]" italics="true" pageId="29" pageNumber="208">P. darwiniensis</emphasis>
|
||
</taxonomicName>
|
||
and the genus remained monotypic for almost 40 years.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFADFFB2FF56AD852A42FA7C" blockId="29.[151,1437,643,2038]" pageId="29" pageNumber="208">
|
||
In the poorly known, not peer-reviewed publication (book) on the caridean shrimps of
|
||
<collectingCountry id="C8D44FC4FFADFFB2FB50AD8A2B03FC44" box="[1217,1315,967,993]" name="Pakistan" pageId="29" pageNumber="208">Pakistan</collectingCountry>
|
||
,
|
||
<bibRefCitation id="D45272A5FFADFFB2FAA0AD8A2F15FBA3" author="Kazmi, Q. B. & Kazmi, M. A." pageId="29" pageNumber="208" refId="ref22537" refString="Kazmi, Q. B. & Kazmi, M. A. (2010) Biodiversity and biogeography of caridean shrimps of Pakistan. Higher Education Commission, Islamabad, xvii + 516 pp." type="book" year="2010">Kazmi & Kazmi (2010)</bibRefCitation>
|
||
established the alpheid genus
|
||
<taxonomicName id="77C374D7FFADFFB2FD1EADA62DCEFBA3" authority="Kazmi & Kazmi, 2010" authorityName="Kazmi & Kazmi" authorityYear="2010" box="[655,1006,1003,1030]" class="Magnoliopsida" family="Rubiaceae" genus="Alberta" kingdom="Plantae" order="Gentianales" pageId="29" pageNumber="208" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FD1EADA62CC3FBA0" box="[655,739,1003,1029]" italics="true" pageId="29" pageNumber="208">Alberta</emphasis>
|
||
<bibRefCitation id="D45272A5FFADFFB2FD7DADA62DCEFBA3" author="Kazmi, Q. B. & Kazmi, M. A." box="[748,1006,1003,1030]" pageId="29" pageNumber="208" refId="ref22537" refString="Kazmi, Q. B. & Kazmi, M. A. (2010) Biodiversity and biogeography of caridean shrimps of Pakistan. Higher Education Commission, Islamabad, xvii + 516 pp." type="book" year="2010">Kazmi & Kazmi, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
for
|
||
<taxonomicName id="77C374D7FFADFFB2FB8EADA62EF2FB8F" authority="Kazmi & Kazmi, 2010" authorityName="Kazmi & Kazmi" authorityYear="2010" class="Magnoliopsida" family="Rubiaceae" genus="Alberta" kingdom="Plantae" order="Gentianales" pageId="29" pageNumber="208" phylum="Tracheophyta" rank="species" species="banneri">
|
||
<emphasis id="82B7D346FFADFFB2FB8EADA62AF2FBA0" box="[1055,1234,1003,1029]" italics="true" pageId="29" pageNumber="208">Alberta banneri</emphasis>
|
||
<bibRefCitation id="D45272A5FFADFFB2FB4BADA62EF2FB8F" author="Kazmi, Q. B. & Kazmi, M. A." pageId="29" pageNumber="208" refId="ref22537" refString="Kazmi, Q. B. & Kazmi, M. A. (2010) Biodiversity and biogeography of caridean shrimps of Pakistan. Higher Education Commission, Islamabad, xvii + 516 pp." type="book" year="2010">Kazmi & Kazmi, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
, based on a single ovigerous female from Bulleji, near Karachi. All material listed in this study, including the
|
||
<typeStatus id="6F78B1F6FFADFFB2FF52AA7E2F05FBE8" box="[195,293,1075,1101]" pageId="29" pageNumber="208" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="77C374D7FFADFFB2FEDEAA792FE7FBE8" authorityName="Kazmi & Kazmi" authorityYear="2010" box="[335,455,1075,1102]" class="Magnoliopsida" family="Rubiaceae" genus="Alberta" kingdom="Plantae" order="Gentianales" pageId="29" pageNumber="208" phylum="Tracheophyta" rank="species" species="banneri">
|
||
<emphasis id="82B7D346FFADFFB2FEDEAA792FE7FBE8" box="[335,455,1075,1102]" italics="true" pageId="29" pageNumber="208">A. banneri</emphasis>
|
||
</taxonomicName>
|
||
, was supposedly deposited in the Marine Reference Collection and Resource Centre, University of Karachi,
|
||
<collectingCountry id="C8D44FC4FFADFFB2FE0DAA1A2FDFFBD4" box="[412,511,1111,1137]" name="Pakistan" pageId="29" pageNumber="208">Pakistan</collectingCountry>
|
||
. The generic diagnosis of
|
||
<taxonomicName id="77C374D7FFADFFB2FCB8AA1A2D5DFBD4" authorityName="Kazmi & Kazmi" authorityYear="2010" box="[809,893,1111,1137]" class="Magnoliopsida" family="Rubiaceae" genus="Alberta" kingdom="Plantae" order="Gentianales" pageId="29" pageNumber="208" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FCB8AA1A2D5DFBD4" box="[809,893,1111,1137]" italics="true" pageId="29" pageNumber="208">Alberta</emphasis>
|
||
</taxonomicName>
|
||
, as well as the description of its type species
|
||
<taxonomicName id="77C374D7FFADFFB2FA15AA152ECEFB30" authorityName="Kazmi & Kazmi" authorityYear="2010" class="Magnoliopsida" family="Rubiaceae" genus="Alberta" kingdom="Plantae" order="Gentianales" pageId="29" pageNumber="208" phylum="Tracheophyta" rank="species" species="banneri">
|
||
<emphasis id="82B7D346FFADFFB2FA15AA152ECEFB30" italics="true" pageId="29" pageNumber="208">A. banneri</emphasis>
|
||
</taxonomicName>
|
||
, are very poor compared to modern standards in caridean taxonomy. Nevertheless, the semi-diagrammatic illustrations of
|
||
<taxonomicName id="77C374D7FFADFFB2FED2AAED2F9AFB1C" authorityName="Kazmi & Kazmi" authorityYear="2010" box="[323,442,1183,1210]" class="Magnoliopsida" family="Rubiaceae" genus="Alberta" kingdom="Plantae" order="Gentianales" pageId="29" pageNumber="208" phylum="Tracheophyta" rank="species" species="banneri">
|
||
<emphasis id="82B7D346FFADFFB2FED2AAED2F9AFB1C" box="[323,442,1183,1210]" italics="true" pageId="29" pageNumber="208">A. banneri</emphasis>
|
||
</taxonomicName>
|
||
provided by
|
||
<bibRefCitation id="D45272A5FFADFFB2FDC3AAD22D41FB1F" author="Kazmi, Q. B. & Kazmi, M. A." box="[594,865,1183,1210]" pageId="29" pageNumber="208" refId="ref22537" refString="Kazmi, Q. B. & Kazmi, M. A. (2010) Biodiversity and biogeography of caridean shrimps of Pakistan. Higher Education Commission, Islamabad, xvii + 516 pp." type="book" year="2010">Kazmi & Kazmi (2010)</bibRefCitation>
|
||
are sufficient to suggest that this taxon belongs to the genus
|
||
<taxonomicName id="77C374D7FFADFFB2FE9BAA8E2F87FB78" authorityName="Bruce" authorityYear="1983" box="[266,423,1219,1245]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FE9BAA8E2F87FB78" box="[266,423,1219,1245]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
. Most importantly, the diaeresis of the uropodal exopod of
|
||
<taxonomicName id="77C374D7FFADFFB2FBD7AA892A9DFB78" authorityName="Kazmi & Kazmi" authorityYear="2010" box="[1094,1213,1219,1246]" class="Magnoliopsida" family="Rubiaceae" genus="Alberta" kingdom="Plantae" order="Gentianales" pageId="29" pageNumber="208" phylum="Tracheophyta" rank="species" species="banneri">
|
||
<emphasis id="82B7D346FFADFFB2FBD7AA892A9DFB78" box="[1094,1213,1219,1246]" italics="true" pageId="29" pageNumber="208">A. banneri</emphasis>
|
||
</taxonomicName>
|
||
is adorned with the same row of strong spiniform setae as in
|
||
<taxonomicName id="77C374D7FFADFFB2FDFCAAA52D38FAA4" authorityName="Bruce" authorityYear="1983" box="[621,792,1255,1281]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="darwiniensis">
|
||
<emphasis id="82B7D346FFADFFB2FDFCAAA52D38FAA4" box="[621,792,1255,1281]" italics="true" pageId="29" pageNumber="208">P. darwiniensis</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="D45272A5FFADFFB2FCDEAAA52DF6FAA7" author="Bruce, A. J." box="[847,982,1256,1282]" pageId="29" pageNumber="208" pagination="463 - 471" refId="ref21222" refString="Bruce, A. J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463 - 471. https: // doi. org / 10.2307 / 1548146" type="journal article" year="1983">Bruce 1983</bibRefCitation>
|
||
: fig. 2F, 5C;
|
||
<bibRefCitation id="D45272A5FFADFFB2FBFAAAAA2B4AFAA7" author="Kazmi, Q. B. & Kazmi, M. A." box="[1131,1386,1255,1282]" pageId="29" pageNumber="208" refId="ref22537" refString="Kazmi, Q. B. & Kazmi, M. A. (2010) Biodiversity and biogeography of caridean shrimps of Pakistan. Higher Education Commission, Islamabad, xvii + 516 pp." type="book" year="2010">Kazmi & Kazmi 2010</bibRefCitation>
|
||
: fig. 108B). The collection of
|
||
<specimenCount id="A6C5C4DDFFADFFB2FE3DAB462C7EFA80" box="[428,606,1291,1317]" count="3" pageId="29" pageNumber="208" type="generic">three specimens</specimenCount>
|
||
(one subsequently lost) of a species with all characteristics of
|
||
<taxonomicName id="77C374D7FFADFFB2FA9DAB412BA1FA80" authorityName="Kazmi & Kazmi" authorityYear="2010" box="[1292,1409,1291,1318]" class="Magnoliopsida" family="Rubiaceae" genus="Alberta" kingdom="Plantae" order="Gentianales" pageId="29" pageNumber="208" phylum="Tracheophyta" rank="species" species="banneri">
|
||
<emphasis id="82B7D346FFADFFB2FA9DAB412BA1FA80" box="[1292,1409,1291,1318]" italics="true" pageId="29" pageNumber="208">A. banneri</emphasis>
|
||
</taxonomicName>
|
||
in northern
|
||
<collectingCountry id="C8D44FC4FFADFFB2FF6DAB7D2F60FAEF" box="[252,320,1328,1354]" name="Oman" pageId="29" pageNumber="208">Oman</collectingCountry>
|
||
(see below) eliminated any lingering doubts about its identity. Therefore,
|
||
<taxonomicName id="77C374D7FFADFFB2FBE9AB622AECFAEC" authorityName="Kazmi & Kazmi" authorityYear="2010" box="[1144,1228,1327,1353]" class="Magnoliopsida" family="Rubiaceae" genus="Alberta" kingdom="Plantae" order="Gentianales" pageId="29" pageNumber="208" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FBE9AB622AECFAEC" box="[1144,1228,1327,1353]" italics="true" pageId="29" pageNumber="208">Alberta</emphasis>
|
||
</taxonomicName>
|
||
is herein placed in the synonymy of
|
||
<taxonomicName id="77C374D7FFADFFB2FEC5AB1E2FD1FAC8" authorityName="Bruce" authorityYear="1983" box="[340,497,1363,1389]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FEC5AB1E2FD1FAC8" box="[340,497,1363,1389]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
and the Omani material is reported as
|
||
<taxonomicName id="77C374D7FFADFFB2FC02AB1E2BBDFACB" authority="(Kazmi & Kazmi, 2010)" authorityName="Anker" authorityYear="2023" baseAuthorityName="Kazmi & Kazmi" baseAuthorityYear="2010" box="[915,1437,1363,1390]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="banneri" status="comb. nov.">
|
||
<emphasis id="82B7D346FFADFFB2FC02AB1E2AACFAC8" box="[915,1164,1363,1389]" italics="true" pageId="29" pageNumber="208">Pseudathanas banneri</emphasis>
|
||
(
|
||
<bibRefCitation id="D45272A5FFADFFB2FB0BAB1E2BB5FACB" author="Kazmi, Q. B. & Kazmi, M. A." box="[1178,1429,1363,1390]" pageId="29" pageNumber="208" refId="ref22537" refString="Kazmi, Q. B. & Kazmi, M. A. (2010) Biodiversity and biogeography of caridean shrimps of Pakistan. Higher Education Commission, Islamabad, xvii + 516 pp." type="book" year="2010">Kazmi & Kazmi, 2010</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFADFFB2FF06AB352F37FA37" bold="true" box="[151,279,1400,1426]" pageId="29" pageNumber="208">
|
||
<taxonomicNameLabel id="99846E3DFFADFFB2FF06AB352F37FA37" box="[151,279,1400,1426]" pageId="29" pageNumber="208" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The generic diagnosis of
|
||
<taxonomicName id="77C374D7FFADFFB2FDA9AB3A2CF5FA34" authorityName="Bruce" authorityYear="1983" box="[568,725,1399,1425]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FDA9AB3A2CF5FA34" box="[568,725,1399,1425]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
provided above was emended to accommodate several features of
|
||
<taxonomicName id="77C374D7FFADFFB2FF26ABD12F0BFA10" authorityName="Anker" authorityYear="2023" baseAuthorityName="Kazmi & Kazmi" baseAuthorityYear="2010" box="[183,299,1435,1461]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="banneri">
|
||
<emphasis id="82B7D346FFADFFB2FF26ABD12F0BFA10" box="[183,299,1435,1461]" italics="true" pageId="29" pageNumber="208">P. banneri</emphasis>
|
||
</taxonomicName>
|
||
that are novel for the genus, especially the dorsally covered eyes, the subequal and subsymmetrical chelipeds, and the second pereiopod carpus with four subarticles (carpal subdivisions).
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFADFFB1FF56ABA92C82FF70" blockId="29.[151,1437,643,2038]" lastBlockId="30.[151,1437,151,717]" lastPageId="30" lastPageNumber="209" pageId="29" pageNumber="208">
|
||
As suggested by
|
||
<bibRefCitation id="D45272A5FFADFFB2FE14ABA92C3CFA5B" author="Bruce, A. J." box="[389,540,1508,1534]" pageId="29" pageNumber="208" pagination="463 - 471" refId="ref21222" refString="Bruce, A. J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463 - 471. https: // doi. org / 10.2307 / 1548146" type="journal article" year="1983">Bruce (1983)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="77C374D7FFADFFB2FDB9ABAE2CE5FA58" authorityName="Bruce" authorityYear="1983" box="[552,709,1507,1533]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FDB9ABAE2CE5FA58" box="[552,709,1507,1533]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
is closely related to
|
||
<taxonomicName id="77C374D7FFADFFB2FC3AABAE2A27FA58" authorityName="Leach" authorityYear="1814" box="[939,1031,1507,1533]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FC3AABAE2A27FA58" box="[939,1031,1507,1533]" italics="true" pageId="29" pageNumber="208">Athanas</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs essentially by the armature of the diaeresis.All other morphological characters listed by
|
||
<bibRefCitation id="D45272A5FFADFFB2FC58A8452A7DF987" author="Bruce, A. J." box="[969,1117,1544,1570]" pageId="29" pageNumber="208" pagination="463 - 471" refId="ref21222" refString="Bruce, A. J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463 - 471. https: // doi. org / 10.2307 / 1548146" type="journal article" year="1983">Bruce (1983)</bibRefCitation>
|
||
to distinguish
|
||
<taxonomicName id="77C374D7FFADFFB2FB6EA84A2BBCF984" authorityName="Bruce" authorityYear="1983" box="[1279,1436,1543,1569]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FB6EA84A2BBCF984" box="[1279,1436,1543,1569]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="77C374D7FFADFFB2FF48A8662F15F9E0" authorityName="Leach" authorityYear="1814" box="[217,309,1579,1605]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FF48A8662F15F9E0" box="[217,309,1579,1605]" italics="true" pageId="29" pageNumber="208">Athanas</emphasis>
|
||
</taxonomicName>
|
||
are invalid as they are present in some more recently described species of the latter genus (see also above). These characters are the marked reduction of the rostrum (also observed in
|
||
<taxonomicName id="77C374D7FFADFFB2FB15A81D2B21F9CC" authorityName="Anker, Naderloo & Marin" authorityYear="2010" box="[1156,1281,1616,1642]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis id="82B7D346FFADFFB2FB15A81D2B21F9CC" box="[1156,1281,1616,1642]" italics="true" pageId="29" pageNumber="208">A. iranicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFADFFB2FA81A81D2B54F9CC" authorityName="Anker" authorityYear="2011" box="[1296,1396,1615,1642]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="daviei">
|
||
<emphasis id="82B7D346FFADFFB2FA81A81D2B54F9CC" box="[1296,1396,1615,1642]" italics="true" pageId="29" pageNumber="208">A. daviei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFADFFB2FA15A81D2F32F928" authorityName="Duris & Anker" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="82B7D346FFADFFB2FA15A81D2F32F928" italics="true" pageId="29" pageNumber="208">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFADFFB2FE8EA8392FEBF928" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[287,459,1651,1678]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
|
||
<emphasis id="82B7D346FFADFFB2FE8EA8392FEBF928" box="[287,459,1651,1678]" italics="true" pageId="29" pageNumber="208">A. claereboudti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFADFFB2FE40A8392C0AF92B" bold="true" box="[465,554,1652,1678]" pageId="29" pageNumber="208">
|
||
<taxonomicNameLabel id="99846E3DFFADFFB2FE40A8392C0AF92B" box="[465,554,1652,1678]" pageId="29" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
); the absence of supra-, extra-, or infra-corneal teeth (absent or greatly reduced in
|
||
<taxonomicName id="77C374D7FFADFFB2FF26A8D52F15F914" authorityName="Anker, Naderloo & Marin" authorityYear="2010" box="[183,309,1688,1714]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis id="82B7D346FFADFFB2FF26A8D52F15F914" box="[183,309,1688,1714]" italics="true" pageId="29" pageNumber="208">A. iranicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFADFFB2FED7A8D52F8DF914" authorityName="Anker" authorityYear="2011" box="[326,429,1687,1714]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="daviei">
|
||
<emphasis id="82B7D346FFADFFB2FED7A8D52F8DF914" box="[326,429,1687,1714]" italics="true" pageId="29" pageNumber="208">A. daviei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFADFFB2FE2FA8D52C7BF914" authorityName="Duris & Anker" authorityYear="2014" box="[446,603,1687,1714]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="82B7D346FFADFFB2FE2FA8D52C7BF914" box="[446,603,1687,1714]" italics="true" pageId="29" pageNumber="208">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFADFFB2FD09A8D52D67F914" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[664,839,1687,1714]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
|
||
<emphasis id="82B7D346FFADFFB2FD09A8D52D67F914" box="[664,839,1687,1714]" italics="true" pageId="29" pageNumber="208">A. claereboudti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFADFFB2FCC0A8D52D8EF917" bold="true" box="[849,942,1688,1714]" pageId="29" pageNumber="208">
|
||
<taxonomicNameLabel id="99846E3DFFADFFB2FCC0A8D52D8EF917" box="[849,942,1688,1714]" pageId="29" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
); the well-developed, dissimilar chelipeds, apparently in both males and females (female minor cheliped unknown in
|
||
<taxonomicName id="77C374D7FFADFFB2FC64A8F12ABAF970" authorityName="Bruce" authorityYear="1983" box="[1013,1178,1723,1749]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="darwinensis">
|
||
<emphasis id="82B7D346FFADFFB2FC64A8F12ABAF970" box="[1013,1178,1723,1749]" italics="true" pageId="29" pageNumber="208">P. darwinensis</emphasis>
|
||
</taxonomicName>
|
||
) (variable in
|
||
<taxonomicName id="77C374D7FFADFFB2FAA8A8F62BB5F970" authorityName="Leach" authorityYear="1814" box="[1337,1429,1723,1749]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FAA8A8F62BB5F970" box="[1337,1429,1723,1749]" italics="true" pageId="29" pageNumber="208">Athanas</emphasis>
|
||
</taxonomicName>
|
||
, with similar condition found, for instance, in
|
||
<taxonomicName id="77C374D7FFADFFB2FD37A8AD2D74F95C" authorityName="Anker" authorityYear="2011" box="[678,852,1759,1786]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="shawnsmithi">
|
||
<emphasis id="82B7D346FFADFFB2FD37A8AD2D74F95C" box="[678,852,1759,1786]" italics="true" pageId="29" pageNumber="208">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
); and the proximally thickened antennal flagellum (also observed in
|
||
<taxonomicName id="77C374D7FFADFFB2FEC8A9492FF3F8B8" authorityName="Anker, Naderloo & Marin" authorityYear="2010" box="[345,467,1796,1822]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="iranicus">
|
||
<emphasis id="82B7D346FFADFFB2FEC8A9492FF3F8B8" box="[345,467,1796,1822]" italics="true" pageId="29" pageNumber="208">A. iranicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFADFFB2FE71A9492CA9F8B8" authorityName="Anker" authorityYear="2011" box="[480,649,1795,1822]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="shawnsmithi">
|
||
<emphasis id="82B7D346FFADFFB2FE71A9492CA9F8B8" box="[480,649,1795,1822]" italics="true" pageId="29" pageNumber="208">A. shawnsmithi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFADFFB2FD07A9492D0FF8B8" authorityName="Duris & Anker" authorityYear="2014" box="[662,815,1795,1822]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="manticolus">
|
||
<emphasis id="82B7D346FFADFFB2FD07A9492D0FF8B8" box="[662,815,1795,1822]" italics="true" pageId="29" pageNumber="208">A. manticolus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFADFFB2FCF2A9492A2DF8B8" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[867,1037,1795,1822]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
|
||
<emphasis id="82B7D346FFADFFB2FCF2A9492A2DF8B8" box="[867,1037,1795,1822]" italics="true" pageId="29" pageNumber="208">A. claereboudti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFADFFB2FB82A9492A4BF8BB" bold="true" box="[1043,1131,1796,1822]" pageId="29" pageNumber="208">
|
||
<taxonomicNameLabel id="99846E3DFFADFFB2FB82A9492A4BF8BB" box="[1043,1131,1796,1822]" pageId="29" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
<taxonomicName id="77C374D7FFADFFB2FB10A94E2B3EF8B8" authorityName="Bruce" authorityYear="1983" box="[1153,1310,1795,1821]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FB10A94E2B3EF8B8" box="[1153,1310,1795,1821]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
also shares many features with
|
||
<taxonomicName id="77C374D7FFADFFB2FEE8A96A2C86F8E7" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" box="[377,678,1831,1858]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FEE8A96A2FD5F8E4" box="[377,501,1831,1857]" italics="true" pageId="29" pageNumber="208">Athanopsis</emphasis>
|
||
<bibRefCitation id="D45272A5FFADFFB2FE6FA96A2C86F8E7" author="Coutiere, H." box="[510,678,1831,1858]" pageId="29" pageNumber="208" pagination="301 - 303" refId="ref21484" refString="Coutiere, H. (1897) Note sur un nouveau genre d'Alpheides. Bulletin du Museum d'Histoire Naturelle, 1 (3), 301 - 303. https: // doi. org / 10.5962 / bhl. part. 19260" type="journal article" year="1897">Coutière, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFADFFB2FD71A96A2AB5F8E7" authority="De Grave & Anker, 2008" authorityName="De Grave & Anker" authorityYear="2008" box="[736,1173,1831,1858]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FD71A96A2D4AF8E4" box="[736,874,1831,1857]" italics="true" pageId="29" pageNumber="208">Leptathanas</emphasis>
|
||
<bibRefCitation id="D45272A5FFADFFB2FCE2A9652AB5F8E7" author="De Grave, S. & Anker, A." box="[883,1173,1831,1858]" pageId="29" pageNumber="208" pagination="43 - 52" refId="ref21714" refString="De Grave, S. & Anker, A. (2008) Leptathanas powelli gen. nov., sp. nov., a new infaunal alpheid shrimp associated with upogebiid mudshrimps in Nigeria (Crustacea, Decapoda). Zootaxa, 1750 (1), 43 - 52. https: // doi. org / 10.11646 / zootaxa. 1750.1.4" type="journal article" year="2008">De Grave & Anker, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
, differing from both of these genera by the unique armature of the uropodal diaeresis; from
|
||
<taxonomicName id="77C374D7FFADFFB2FC02A9062A2FF8C0" authorityName="Coutiere" authorityYear="1897" box="[915,1039,1867,1893]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FC02A9062A2FF8C0" box="[915,1039,1867,1893]" italics="true" pageId="29" pageNumber="208">Athanopsis</emphasis>
|
||
</taxonomicName>
|
||
by the distally pointed rostrum (
|
||
<emphasis id="82B7D346FFADFFB2FAEFA9012BB5F8C0" box="[1406,1429,1868,1893]" italics="true" pageId="29" pageNumber="208">vs</emphasis>
|
||
. rounded in
|
||
<taxonomicName id="77C374D7FFADFFB2FE88A9222FB5F82C" authorityName="Coutiere" authorityYear="1897" box="[281,405,1903,1929]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FE88A9222FB5F82C" box="[281,405,1903,1929]" italics="true" pageId="29" pageNumber="208">Athanopsis</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName id="77C374D7FFADFFB2FD8CA9222C87F82C" authorityName="De Grave & Anker" authorityYear="2008" box="[541,679,1903,1929]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FD8CA9222C87F82C" box="[541,679,1903,1929]" italics="true" pageId="29" pageNumber="208">Leptathanas</emphasis>
|
||
</taxonomicName>
|
||
by the relatively well developed rostrum (reduced in
|
||
<taxonomicName id="77C374D7FFADFFB2FA99A9222BB2F82C" authorityName="De Grave & Anker" authorityYear="2008" box="[1288,1426,1903,1929]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FA99A9222BB2F82C" box="[1288,1426,1903,1929]" italics="true" pageId="29" pageNumber="208">Leptathanas</emphasis>
|
||
</taxonomicName>
|
||
) and the much slenderer walking legs and antennular peduncles (very stout in
|
||
<taxonomicName id="77C374D7FFADFFB2FC7AA9DE2A55F808" authorityName="De Grave & Anker" authorityYear="2008" box="[1003,1141,1939,1965]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FC7AA9DE2A55F808" box="[1003,1141,1939,1965]" italics="true" pageId="29" pageNumber="208">Leptathanas</emphasis>
|
||
</taxonomicName>
|
||
) (cf.
|
||
<bibRefCitation id="D45272A5FFADFFB2FB20A9DE2B72F80B" author="Coutiere, H." box="[1201,1362,1939,1966]" pageId="29" pageNumber="208" pagination="301 - 303" refId="ref21484" refString="Coutiere, H. (1897) Note sur un nouveau genre d'Alpheides. Bulletin du Museum d'Histoire Naturelle, 1 (3), 301 - 303. https: // doi. org / 10.5962 / bhl. part. 19260" type="journal article" year="1897">Coutière 1897</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="D45272A5FFADFFB2FACDA9D92BB7F80B" author="Coutiere, H." box="[1372,1431,1940,1966]" pageId="29" pageNumber="208" pagination="1 - 6" refId="ref21527" refString="Coutiere, H. (1899) Les " Alpheidae ", morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie, Serie 8, Vol. 9. Masson, Paris, 559 pp., pls. 1 - 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="book chapter" year="1899">1899</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D45272A5FFADFFB2FF06A9FA2FBAF877" author="Anker, A. & Ahyong, S. T." box="[151,410,1975,2002]" pageId="29" pageNumber="208" pagination="685 - 697" refId="ref19649" refString="Anker, A. & Ahyong, S. T. (2007) A rediagnosis of Athanopsis australis Banner & Banner, 1982, a rare alpheid shrimp from southern Australia, with a phylogeny of Athanopsis Coutiere, 1897 and remarks on antitropical distributions in the Alpheidae (Decapoda, Caridea). Crustaceana, 80, 685 - 697. https: // doi. org / 10.1163 / 156854007781360612" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D45272A5FFADFFB2FE34A9F52C99F877" author="De Grave, S. & Anker, A." box="[421,697,1975,2002]" pageId="29" pageNumber="208" pagination="43 - 52" refId="ref21714" refString="De Grave, S. & Anker, A. (2008) Leptathanas powelli gen. nov., sp. nov., a new infaunal alpheid shrimp associated with upogebiid mudshrimps in Nigeria (Crustacea, Decapoda). Zootaxa, 1750 (1), 43 - 52. https: // doi. org / 10.11646 / zootaxa. 1750.1.4" type="journal article" year="2008">De Grave & Anker 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D45272A5FFADFFB2FD52A9FA2D76F874" author="Anker, A." box="[707,854,1975,2001]" pageId="29" pageNumber="208" pagination="1 - 22" refId="ref19338" refString="Anker, A. (2011 b) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2734.1.1" type="journal article" year="2011">Anker 2011b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="D45272A5FFADFFB2FCF0A9F52DBDF877" author="Anker, A." box="[865,925,1976,2002]" pageId="29" pageNumber="208" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D45272A5FFADFFB2FC39A9FA2A4BF877" author="Marin, I. & Sheibani, R. & Sari, A." box="[936,1131,1975,2002]" pageId="29" pageNumber="208" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. & Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
|
||
Marin
|
||
<emphasis id="82B7D346FFADFFB2FC62A9F52A0AF874" box="[1011,1066,1975,2001]" italics="true" pageId="29" pageNumber="208">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
, see also
|
||
<figureCitation id="28F813D1FFADFFB2FB42A9FA2B05F877" box="[1235,1317,1975,2002]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="29" pageNumber="208">Fig. 10</figureCitation>
|
||
). The long plumose setae on the posteroventral margin of the branchiostegite are present in
|
||
<taxonomicName id="77C374D7FFADFFB2FBB4A9962AE2F850" authorityName="Bruce" authorityYear="1983" box="[1061,1218,2011,2037]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="29" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFADFFB2FBB4A9962AE2F850" box="[1061,1218,2011,2037]" italics="true" pageId="29" pageNumber="208">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
, but also in the not closely related
|
||
<taxonomicName id="77C374D7FFAEFFB1FEAEAEDA2D20FF14" authority="De Grave & Anker, 2000" authorityName="De Grave & Anker" authorityYear="2000" box="[319,768,151,177]" class="Malacostraca" family="Alpheidae" genus="Orygmalpheus" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FEAEAEDA2FC4FF14" box="[319,484,151,177]" italics="true" pageId="30" pageNumber="209">Orygmalpheus</emphasis>
|
||
<bibRefCitation id="D45272A5FFAEFFB1FE7BAEDA2D20FF14" author="De Grave, S. & Anker, A." box="[490,768,151,177]" pageId="30" pageNumber="209" pagination="249 - 256" refId="ref21671" refString="De Grave, S. & Anker, A. (2000) A new genus and new species of alpheid shrimp (Crustacea: Decapoda) from Papua New Guinea. Raffles Bulletin of Zoology, 48, 249 - 256." type="journal article" year="2000">De Grave & Anker, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
and (with lesser development) in some other alpheid genera (
|
||
<bibRefCitation id="D45272A5FFAEFFB1FF0EAEF62F98FF70" author="De Grave, S. & Anker, A." box="[159,440,187,213]" pageId="30" pageNumber="209" pagination="249 - 256" refId="ref21671" refString="De Grave, S. & Anker, A. (2000) A new genus and new species of alpheid shrimp (Crustacea: Decapoda) from Papua New Guinea. Raffles Bulletin of Zoology, 48, 249 - 256." type="journal article" year="2000">De Grave & Anker 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D45272A5FFAEFFB1FE50AEF62CB7FF70" author="Anker, A. & Ahyong, S. T. & Palmer, A. R. & Noel, P. Y." box="[449,663,187,213]" pageId="30" pageNumber="209" pagination="2507 - 2528" refId="ref19722" refString="Anker, A., Ahyong, S. T., Palmer, A. R. & Noel, P. Y. (2006 a) Morphological phylogeny of alpheid shrimps: parallel preadaptation and the origin of a key morphological innovation, the snapping claw. Evolution, 60, 2507 - 2528. https: // doi. org / 10.1111 / j. 0014 - 3820.2006. tb 01886. x" type="journal article" year="2006">
|
||
Anker
|
||
<emphasis id="82B7D346FFAEFFB1FD9FAEF62C66FF70" box="[526,582,187,213]" italics="true" pageId="30" pageNumber="209">et al.</emphasis>
|
||
2006a
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFAEFFB1FF56AE922C8FFD68" blockId="30.[151,1437,151,717]" pageId="30" pageNumber="209">
|
||
<bibRefCitation id="D45272A5FFAEFFB1FF56AE922F8FFF5C" author="Anker, A. & Ahyong, S. T. & Palmer, A. R. & Noel, P. Y." box="[199,431,223,249]" pageId="30" pageNumber="209" pagination="2507 - 2528" refId="ref19722" refString="Anker, A., Ahyong, S. T., Palmer, A. R. & Noel, P. Y. (2006 a) Morphological phylogeny of alpheid shrimps: parallel preadaptation and the origin of a key morphological innovation, the snapping claw. Evolution, 60, 2507 - 2528. https: // doi. org / 10.1111 / j. 0014 - 3820.2006. tb 01886. x" type="journal article" year="2006">
|
||
Anker
|
||
<emphasis id="82B7D346FFAEFFB1FE87AE922F6FFF5C" box="[278,335,223,249]" italics="true" pageId="30" pageNumber="209">et al.</emphasis>
|
||
(2006a)
|
||
</bibRefCitation>
|
||
performed a cladistic analysis of morphological characters and recovered
|
||
<taxonomicName id="77C374D7FFAEFFB1FB6EAE922BBCFF5C" authorityName="Bruce" authorityYear="1983" box="[1279,1436,223,249]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FB6EAE922BBCFF5C" box="[1279,1436,223,249]" italics="true" pageId="30" pageNumber="209">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFAEFFB1FF58AF4E2F05FEB8" authorityName="Leach" authorityYear="1814" box="[201,293,259,285]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FF58AF4E2F05FEB8" box="[201,293,259,285]" italics="true" pageId="30" pageNumber="209">Athanas</emphasis>
|
||
</taxonomicName>
|
||
in sister position within a larger “athanoid” clade (clade AP), together with
|
||
<taxonomicName id="77C374D7FFAEFFB1FB02AF4E2ACFFEB8" authorityName="Leach" authorityYear="1814" box="[1171,1263,259,285]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FB02AF4E2ACFFEB8" box="[1171,1263,259,285]" italics="true" pageId="30" pageNumber="209">Athanas</emphasis>
|
||
</taxonomicName>
|
||
(paraphyletic),
|
||
<taxonomicName id="77C374D7FFAEFFB1FF06AF6A2FAAFEE4" authority="Stimpson, 1860" authorityName="Stimpson" authorityYear="1860" box="[151,394,295,321]" class="Malacostraca" family="Alpheidae" genus="Arete" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FF06AF6A2EF3FEE4" box="[151,211,295,321]" italics="true" pageId="30" pageNumber="209">Arete</emphasis>
|
||
<bibRefCitation id="D45272A5FFAEFFB1FF4BAF6A2FAAFEE4" author="Stimpson, W." box="[218,394,295,321]" pageId="30" pageNumber="209" pagination="22 - 47" refId="ref23493" refString="Stimpson, W. (1860) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VIII. Crustacea Macrura. Proceedings of the Academy of Natural Sciences of Philadelphia, 1860, 22 - 47." type="journal article" year="1860">Stimpson, 1860</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFAEFFB1FE50AF6A2CF5FEE4" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" box="[449,725,295,321]" class="Malacostraca" family="Alpheidae" genus="Aretopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FE50AF6A2C0AFEE4" box="[449,554,295,321]" italics="true" pageId="30" pageNumber="209">Aretopsis</emphasis>
|
||
<bibRefCitation id="D45272A5FFAEFFB1FDA0AF6A2CF5FEE4" author="De Man, J. G." box="[561,725,295,321]" pageId="30" pageNumber="209" pagination="287 - 319" refId="ref22004" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the " Siboga-Expedition ", V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
. In the molecular analysis of the
|
||
<taxonomicName id="77C374D7FFAEFFB1FBD6AF6A2BBCFEE4" authority=", Chow et al. (2021)" authorityName="Chow" authorityYear="2021" box="[1095,1436,295,321]" class="Malacostraca" family="Alpheidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="family">
|
||
Alpheidae,
|
||
<bibRefCitation id="D45272A5FFAEFFB1FB59AF6A2BBCFEE4" author="Chow, L. H. & De Grave, S. & Anker, A. & Poon, K. K. Y. & Ma, K. & Chu, K. & Chan, T. Y. & Tsang, L. M." box="[1224,1436,295,321]" pageId="30" pageNumber="209" pagination="2898 - 2910" refId="ref21336" refString="Chow, L. H., De Grave, S., Anker, A., Poon, K. K. Y., Ma, K. Y, Chu, K. H, Chan, T. Y. & Tsang, L. M. (2021) Distinct suites of pre- and post-adaptations indicate independent evolutionary pathways of snapping claws in the shrimp family Alpheidae (Decapoda: Caridea). Evolution, 75 (11), 2898 - 2910. https: // doi. org / 10.1111 / evo. 14351" type="journal article" year="2021">
|
||
Chow
|
||
<emphasis id="82B7D346FFAEFFB1FA83AF6A2B6BFEE4" box="[1298,1355,295,321]" italics="true" pageId="30" pageNumber="209">et al.</emphasis>
|
||
(2021)
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
also recovered an athanoid clade (clade S-V), which further included
|
||
<taxonomicName id="77C374D7FFAEFFB1FC35AF062EEFFE2C" authority="Anker, Poddoubtchenko & Jeng, 2006" authorityName="Anker, Poddoubtchenko & Jeng" authorityYear="2006" class="Malacostraca" family="Alpheidae" genus="Acanthanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FC35AF062A08FEC0" box="[932,1064,331,357]" italics="true" pageId="30" pageNumber="209">Acanthanas</emphasis>
|
||
<bibRefCitation id="D45272A5FFAEFFB1FBBFAF062EEFFE2C" author="Anker, A. & Poddoubtchenko, D. & Jeng, M. S." pageId="30" pageNumber="209" pagination="341 - 348" refId="ref20458" refString="Anker, A., Poddoubtchenko, D. & Jeng, M. S. (2006 b) Acanthanas pusillus, new genus, new species, a miniature alpheid shrimp with spiny eyes from the Philippines (Crustacea: Decapoda). Raffles Bulletin of Zoology, 54, 341 - 348." type="journal article" year="2006">Anker, Poddoubtchenko & Jeng, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="D45272A5FFAEFFB1FF4FAF222F99FE2C" author="Anker, A. & Poddoubtchenko, D. & Jeng, M. S." box="[222,441,367,393]" pageId="30" pageNumber="209" pagination="341 - 348" refId="ref20458" refString="Anker, A., Poddoubtchenko, D. & Jeng, M. S. (2006 b) Acanthanas pusillus, new genus, new species, a miniature alpheid shrimp with spiny eyes from the Philippines (Crustacea: Decapoda). Raffles Bulletin of Zoology, 54, 341 - 348." type="journal article" year="2006">
|
||
Anker
|
||
<emphasis id="82B7D346FFAEFFB1FEBFAF222F47FE2C" box="[302,359,367,393]" italics="true" pageId="30" pageNumber="209">et al.</emphasis>
|
||
2006b
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="77C374D7FFAEFFB1FE5CAF222C4AFE2C" authorityName="Bruce" authorityYear="1983" box="[461,618,367,393]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FE5CAF222C4AFE2C" box="[461,618,367,393]" italics="true" pageId="30" pageNumber="209">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFAEFFB1FD33AF222D0CFE2C" authorityName="De Grave & Anker" authorityYear="2008" box="[674,812,367,393]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FD33AF222D0CFE2C" box="[674,812,367,393]" italics="true" pageId="30" pageNumber="209">Leptathanas</emphasis>
|
||
</taxonomicName>
|
||
were not included in their analysis because no suitable material of these genera was available at that time. Interestingly,
|
||
<bibRefCitation id="D45272A5FFAEFFB1FCE4AFDE2A6BFE08" author="Chow, L. H. & De Grave, S. & Anker, A. & Poon, K. K. Y. & Ma, K. & Chu, K. & Chan, T. Y. & Tsang, L. M." box="[885,1099,403,429]" pageId="30" pageNumber="209" pagination="2898 - 2910" refId="ref21336" refString="Chow, L. H., De Grave, S., Anker, A., Poon, K. K. Y., Ma, K. Y, Chu, K. H, Chan, T. Y. & Tsang, L. M. (2021) Distinct suites of pre- and post-adaptations indicate independent evolutionary pathways of snapping claws in the shrimp family Alpheidae (Decapoda: Caridea). Evolution, 75 (11), 2898 - 2910. https: // doi. org / 10.1111 / evo. 14351" type="journal article" year="2021">
|
||
Chow
|
||
<emphasis id="82B7D346FFAEFFB1FC51AFDE2DD9FE08" box="[960,1017,403,429]" italics="true" pageId="30" pageNumber="209">et al.</emphasis>
|
||
(2021)
|
||
</bibRefCitation>
|
||
showed
|
||
<taxonomicName id="77C374D7FFAEFFB1FB21AFDE2B2CFE08" authorityName="Leach" authorityYear="1814" box="[1200,1292,403,429]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FB21AFDE2B2CFE08" box="[1200,1292,403,429]" italics="true" pageId="30" pageNumber="209">Athanas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFAEFFB1FAD2AFDE2B5FFE08" authorityName="Stimpson" authorityYear="1860" box="[1347,1407,403,429]" class="Malacostraca" family="Alpheidae" genus="Arete" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FAD2AFDE2B5FFE08" box="[1347,1407,403,429]" italics="true" pageId="30" pageNumber="209">Arete</emphasis>
|
||
</taxonomicName>
|
||
to be non-monophyletic due to the positions of two species of
|
||
<taxonomicName id="77C374D7FFAEFFB1FCD5AFFA2DE0FE74" authorityName="Coutiere" authorityYear="1897" box="[836,960,439,465]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FCD5AFFA2DE0FE74" box="[836,960,439,465]" italics="true" pageId="30" pageNumber="209">Athanopsis</emphasis>
|
||
</taxonomicName>
|
||
and one species of
|
||
<taxonomicName id="77C374D7FFAEFFB1FB35AFFA2B20FE74" authorityName="Leach" authorityYear="1814" box="[1188,1280,439,465]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FB35AFFA2B20FE74" box="[1188,1280,439,465]" italics="true" pageId="30" pageNumber="209">Athanas</emphasis>
|
||
</taxonomicName>
|
||
, respectively. Surprisingly, and difficult to explain from the morphological point of view,
|
||
<taxonomicName id="77C374D7FFAEFFB1FC6BAF962B48FE50" authority="Anker & Jeng 2007" authorityName="Anker & Jeng" authorityYear="2007" box="[1018,1384,475,501]" class="Malacostraca" family="Alpheidae" genus="Rugathanas" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FC6BAF962AA0FE50" box="[1018,1152,475,501]" italics="true" pageId="30" pageNumber="209">Rugathanas</emphasis>
|
||
<bibRefCitation id="D45272A5FFAEFFB1FB16AF962B48FE50" author="Anker, A. & Jeng, M. S." box="[1159,1384,475,501]" pageId="30" pageNumber="209" pagination="454 - 472" refId="ref20209" refString="Anker, A. & Jeng, M. S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinitions of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker & Jeng 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
was excluded from clade S-V despite the presence of several synapomorphic features linking it to other athanoid genera (
|
||
<bibRefCitation id="D45272A5FFAEFFB1FF0EAC6E2FA6FD98" author="Anker, A. & Jeng, M. S." box="[159,390,547,573]" pageId="30" pageNumber="209" pagination="454 - 472" refId="ref20209" refString="Anker, A. & Jeng, M. S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinitions of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker & Jeng 2007</bibRefCitation>
|
||
). Based on its overall morphology,
|
||
<taxonomicName id="77C374D7FFAEFFB1FC8AAC6E2D98FD98" authorityName="Bruce" authorityYear="1983" box="[795,952,547,573]" class="Malacostraca" family="Alpheidae" genus="Pseudathanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FC8AAC6E2D98FD98" box="[795,952,547,573]" italics="true" pageId="30" pageNumber="209">Pseudathanas</emphasis>
|
||
</taxonomicName>
|
||
appears to be closest to
|
||
<taxonomicName id="77C374D7FFAEFFB1FB45AC6E2B70FD98" authorityName="Coutiere" authorityYear="1897" box="[1236,1360,547,573]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FB45AC6E2B70FD98" box="[1236,1360,547,573]" italics="true" pageId="30" pageNumber="209">Athanopsis</emphasis>
|
||
</taxonomicName>
|
||
and is most likely nested within the
|
||
<taxonomicName id="77C374D7FFAEFFB1FE70AC0A2C1DFDC4" authorityName="Leach" authorityYear="1814" box="[481,573,583,609]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FE70AC0A2C1DFDC4" box="[481,573,583,609]" italics="true" pageId="30" pageNumber="209">Athanas</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="77C374D7FFAEFFB1FDCAAC0A2CF7FDC4" authorityName="Coutiere" authorityYear="1897" box="[603,727,583,609]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FDCAAC0A2CF7FDC4" box="[603,727,583,609]" italics="true" pageId="30" pageNumber="209">Athanopsis</emphasis>
|
||
</taxonomicName>
|
||
lineage of
|
||
<bibRefCitation id="D45272A5FFAEFFB1FCC5AC0A2A0AFDC4" author="Chow, L. H. & De Grave, S. & Anker, A. & Poon, K. K. Y. & Ma, K. & Chu, K. & Chan, T. Y. & Tsang, L. M." box="[852,1066,583,609]" pageId="30" pageNumber="209" pagination="2898 - 2910" refId="ref21336" refString="Chow, L. H., De Grave, S., Anker, A., Poon, K. K. Y., Ma, K. Y, Chu, K. H, Chan, T. Y. & Tsang, L. M. (2021) Distinct suites of pre- and post-adaptations indicate independent evolutionary pathways of snapping claws in the shrimp family Alpheidae (Decapoda: Caridea). Evolution, 75 (11), 2898 - 2910. https: // doi. org / 10.1111 / evo. 14351" type="journal article" year="2021">
|
||
Chow
|
||
<emphasis id="82B7D346FFAEFFB1FC0FAC0A2DF7FDC4" box="[926,983,583,609]" italics="true" pageId="30" pageNumber="209">et al.</emphasis>
|
||
(2021)
|
||
</bibRefCitation>
|
||
, whilst the phylogenetic position of the highly derived
|
||
<taxonomicName id="77C374D7FFAEFFB1FE1CAC262C37FD20" authorityName="De Grave & Anker" authorityYear="2008" box="[397,535,619,645]" class="Malacostraca" family="Alpheidae" genus="Leptathanas" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFAEFFB1FE1CAC262C37FD20" box="[397,535,619,645]" italics="true" pageId="30" pageNumber="209">Leptathanas</emphasis>
|
||
</taxonomicName>
|
||
remains more enigmatic. The entire athanoid clade will definitively need more phylogenetic efforts devoted to it, in view of interesting evolutionary convergences due to symbiotic lifestyles (both infaunal and epibiotic, see
|
||
<bibRefCitation id="D45272A5FFAEFFB1FE52ACFE2C84FD68" author="Anker, A. & Jeng, M. S." box="[451,676,691,717]" pageId="30" pageNumber="209" pagination="454 - 472" refId="ref20209" refString="Anker, A. & Jeng, M. S. (2007) Establishment of a new genus for Arete borradailei Coutiere, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinitions of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454 - 472." type="journal article" year="2007">Anker & Jeng 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |