treatments-xml/data/C3/48/D6/C348D67BFFFF6348EECCC26EFEB106EB.xml

<document id="FD5FDD506F58C7431F323C6173A7C83E" ID-DOI="10.11646/zootaxa.4018.3.3" ID-GBIF-Dataset="a7f6a466-53f6-4df5-b235-a88978ac37bf" ID-ISSN="1175-5326" ID-Zenodo-Dep="232899" ID-ZooBank="07D5711E-F7E0-457A-8360-D4D3E7B95DD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461257399622" checkinUser="plazi" docAuthor="Maeda, Ken &amp; Palla, Herminie P." docDate="2015" docId="C348D67BFFFF6348EECCC26EFEB106EB" docLanguage="en" docName="zt04018p395.pdf" docOrigin="Zootaxa 4018 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stiphodon palawanensis Maeda &amp; Palla, 2015, new species" docType="treatment" docVersion="7" lastPageNumber="391" masterDocId="3F71AE03FFFE6342EE5BC650FFC90165" masterDocTitle="A new species of the genus Stiphodon from Palawan, Philippines (Gobiidae: Sicydiinae)" masterLastPageNumber="395" masterPageNumber="381" pageNumber="382" updateTime="1698608388314" updateUser="plazi">
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<mods:title id="BD3D77042AA1FCE68F7301EACDBE4F14">A new species of the genus Stiphodon from Palawan, Philippines (Gobiidae: Sicydiinae)</mods:title>
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<mods:namePart id="91E0AB55054F39E2457857102577E525">Maeda, Ken</mods:namePart>
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<mods:namePart id="663FF33544338F1C9AB2A314FED6110D">Palla, Herminie P.</mods:namePart>
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<paragraph id="4B5E676DFFFF6343EECCC26EFD93053D" blockId="1.[151,602,1086,1146]" box="[151,602,1086,1112]" pageId="1" pageNumber="382">
<heading id="1016D001FFFF6343EECCC26EFD93053D" bold="true" box="[151,602,1086,1112]" fontSize="11" level="1" pageId="1" pageNumber="382" reason="1">
<emphasis id="7995BB7FFFFF6343EECCC26EFD93053D" bold="true" box="[151,602,1086,1112]" pageId="1" pageNumber="382">
<taxonomicName id="8CE11CEEFFFF6343EECCC26EFE75053D" ID-CoL="52RQS" box="[151,444,1086,1112]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="382" phylum="Chordata" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="7995BB7FFFFF6343EECCC26EFE75053D" bold="true" box="[151,444,1086,1112]" italics="true" pageId="1" pageNumber="382">Stiphodon palawanensis</emphasis>
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, 
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(
<figureCitation id="D3DA7BE8FFFF6343EEC4C231FEC2051F" box="[159,267,1121,1146]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="3.[151,250,1935,1957]" captionStartId-1="4.[151,250,1696,1718]" captionStartId-2="5.[151,250,1194,1216]" captionStartId-3="6.[151,250,1952,1974]" captionStartId-4="7.[151,250,1660,1682]" captionTargetBox-0="[151,1436,481,1912]" captionTargetBox-1="[349,1239,204,1666]" captionTargetBox-2="[349,1236,184,1162]" captionTargetBox-3="[384,1184,211,1923]" captionTargetBox-4="[151,1436,193,1637]" captionTargetId-0="figure@3.[151,1436,481,1914]" captionTargetId-1="figure@4.[340,1247,193,1675]" captionTargetId-2="figure@5.[340,1247,173,1173]" captionTargetId-3="figure@6.[376,1206,193,1942]" captionTargetId-4="figure@7.[151,1436,193,1639]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Stiphodon palawanensis immediately after fixation. a, WPU-PPC-P 5, holotype (male, 62.2 mm SL); b, URM-P 48659, paratype (male, 51.1 mm SL); c, URM-P 48666, paratype (female, 58.5 mm SL); d, URM-P 48661, paratype (female, 41.1 mm SL)." captionText-1="FIGURE 2. Number of premaxillary and horizontal dentary teeth of Stiphodon palawanensis (red circles), S. alcedo (pink circles), S. atratus (black diamonds), S. imperiorientis (light blue triangles), S. maculidorsalis (light green triangles), S. multisquamus (deep blue circles), S. ornatus (orange triangles), S. pelewensis (gray diamonds), S. pulchellus (deep green triangles), and S. weberi (purple diamonds)." captionText-2="FIGURE 3. Dorsal scalation on head and nape in Stiphodon palawanensis. a, male (43.1 mm SL, CMK 11966); b, female (41.9 mm SL, CMK 11966)." captionText-3="FIGURE 4. Diagrammatic illustration of head showing arrangement of cephalic sensory pores and cutaneous sensory papillae in Stiphodon palawanensis (41.9 mm SL, CMK 11966). a, dorsal view; b, lateral view; c, ventral view. AN, anterior nostril; PN, posterior nostril." captionText-4="FIGURE 5. Males (a – d) and females (e, f) of live Stiphodon palawanensis observed in Balsahan Stream, Puerto Princesa City (b, e, f, 18 May 2015; a, c, 19 May 2015) and Barake Stream, Aborlan (d, 15 May 2015)." httpUri-0="https://zenodo.org/record/232900/files/figure.png" httpUri-1="https://zenodo.org/record/232901/files/figure.png" httpUri-2="https://zenodo.org/record/232902/files/figure.png" httpUri-3="https://zenodo.org/record/232903/files/figure.png" httpUri-4="https://zenodo.org/record/232904/files/figure.png" pageId="1" pageNumber="382">Figs. 1–5</figureCitation>
; Table 1)
</paragraph>
</subSubSection>
<subSubSection id="03FB34E6FFFF6343EECCC2F9FDF1062F" pageId="1" pageNumber="382" type="materials_examined">
<paragraph id="4B5E676DFFFF6343EECCC2F9FED40583" blockId="1.[151,1437,1193,2010]" pageId="1" pageNumber="382">
<emphasis id="7995BB7FFFFF6343EECCC2F9FE4E05A7" bold="true" box="[151,391,1193,1218]" pageId="1" pageNumber="382">Material examined.</emphasis>
<specimenCount id="5DE7ACE4FFFF6343EFCFC2F9FDFA05A7" box="[404,563,1193,1218]" pageId="1" pageNumber="382" type="generic">57 specimens</specimenCount>
(
<specimenCount id="5DE7ACE4FFFF6343EC12C2FAFD7C05A7" box="[585,693,1193,1218]" pageId="1" pageNumber="382" type="male">24 males</specimenCount>
and 
<specimenCount id="5DE7ACE4FFFF6343ECA1C2FAFCB705A7" box="[762,894,1193,1218]" pageId="1" pageNumber="382" type="female">33 females</specimenCount>
, 27.0– 
<quantity id="8C19CA88FFFF6343ED94C2F9FBF005A4" box="[975,1081,1193,1218]" metricMagnitude="-2" metricUnit="m" metricValue="6.39" pageId="1" pageNumber="382" unit="mm" value="63.9">63.9 mm</quantity>
SL) collected from Palawan, 
<collectingCountry id="33F627FDFFFF6343EECCC29DFED00583" box="[151,281,1229,1254]" name="Philippines" pageId="1" pageNumber="382">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="4B5E676DFFFF6343EE9CC2A1FC6B044B" blockId="1.[151,1437,1193,2010]" pageId="1" pageNumber="382">
<emphasis id="7995BB7FFFFF6343EE9CC2A1FEF2046F" bold="true" box="[199,315,1265,1290]" pageId="1" pageNumber="382">
<typeStatus id="945AD9CFFFFF6343EE9CC2A1FEFF046F" box="[199,310,1265,1290]" pageId="1" pageNumber="382" type="holotype">Holotype</typeStatus>
.
</emphasis>
WPU-PPC-P 5, male (
<quantity id="8C19CA88FFFF6343EC65C2A1FD6D046C" box="[574,676,1265,1290]" metricMagnitude="-2" metricUnit="m" metricValue="6.22" pageId="1" pageNumber="382" unit="mm" value="62.2">62.2 mm</quantity>
SL), Balsahan Stream in the Iwahig Prison and Penal Farm, Puerto Princesa City, Palawan, 
<date id="3F5F41ADFFFF6343EFFDC346FDF7044B" box="[422,574,1301,1326]" pageId="1" pageNumber="382" value="2015-05-18">18 May 2015</date>
, coll. K. Maeda and H. P. Palla.
</paragraph>
<paragraph id="4B5E676DFFFF6343EE9CC369FDF1062F" blockId="1.[151,1437,1193,2010]" pageId="1" pageNumber="382">
<emphasis id="7995BB7FFFFF6343EE9CC369FE8F0437" bold="true" box="[199,326,1337,1362]" pageId="1" pageNumber="382">
<typeStatus id="945AD9CFFFFF6343EE9CC369FE880437" box="[199,321,1337,1362]" pageId="1" pageNumber="382" type="paratype">Paratypes</typeStatus>
.
</emphasis>
CMK 11966, 
<specimenCount id="5DE7ACE4FFFF6343EFB3C36AFDF60437" box="[488,575,1337,1362]" pageId="1" pageNumber="382" type="male">2 males</specimenCount>
(37.3 and 
<quantity id="8C19CA88FFFF6343ECE3C36AFCD40434" box="[696,797,1338,1362]" metricMagnitude="-2" metricUnit="m" metricValue="4.31" pageId="1" pageNumber="382" unit="mm" value="43.1">43.1 mm</quantity>
SL) and 
<specimenCount id="5DE7ACE4FFFF6343EDDDC36AFC3B0437" box="[902,1010,1337,1362]" pageId="1" pageNumber="382" type="female">8 females</specimenCount>
(
<quantity id="8C19CA88FFFF6343EA59C369FB6E0434" box="[1026,1191,1337,1362]" metricMagnitude="-2" metricUnit="m" metricValue="3.785" metricValueMax="4.72" metricValueMin="2.85" pageId="1" pageNumber="382" unit="mm" value="37.85" valueMax="47.2" valueMin="28.5">28.5–47.2 mm</quantity>
SL), Malatgao River, Narra, Palawan, 
<date id="3F5F41ADFFFF6343EF0EC30EFDFC0413" box="[341,565,1373,1398]" pageId="1" pageNumber="382" value="1994-09-29">29 September 1994</date>
, coll. J. Margraf; CMK 11974, 
<specimenCount id="5DE7ACE4FFFF6343EDFBC30EFBC70413" box="[928,1038,1373,1398]" pageId="1" pageNumber="382" type="female">3 females</specimenCount>
(
<quantity id="8C19CA88FFFF6343EA45C30DFB0C0410" box="[1054,1221,1373,1398]" metricMagnitude="-2" metricUnit="m" metricValue="4.52" metricValueMax="4.85" metricValueMin="4.1899999999999995" pageId="1" pageNumber="382" unit="mm" value="45.2" valueMax="48.5" valueMin="41.9">41.9–48.5 mm</quantity>
SL), Estrella Falls (tributary of Malatgao River), Narra, Palawan, 
<date id="3F5F41ADFFFF6343ECEBC3D2FC5904FF" box="[688,912,1409,1434]" pageId="1" pageNumber="382" value="1994-09-29">29 September 1994</date>
, coll. J. Margraf; NSMT-P 45091, 45092, and 45094, 
<specimenCount id="5DE7ACE4FFFF6343EEB0C3F6FE8804DB" box="[235,321,1445,1470]" pageId="1" pageNumber="382" type="male">2 males</specimenCount>
(27.7 and 
<quantity id="8C19CA88FFFF6343EFE3C3F6FDD504D8" box="[440,540,1446,1470]" metricMagnitude="-2" metricUnit="m" metricValue="3.88" pageId="1" pageNumber="382" unit="mm" value="38.8">38.8 mm</quantity>
SL) and female (
<quantity id="8C19CA88FFFF6343EC85C3F5FC8A04D8" box="[734,835,1445,1470]" metricMagnitude="-2" metricUnit="m" metricValue="3.66" pageId="1" pageNumber="382" unit="mm" value="36.6">36.6 mm</quantity>
SL), Iwahig River, Puerto Princesa City, Palawan, 
<date id="3F5F41ADFFFF6343EBDBC3F6FE980487" pageId="1" pageNumber="382" value="1988-11-13">13 November 1988</date>
, coll. K. Matsuura; URM-P 31438, female (27.0 mm SL), Nagsagoiri River, Palawan, 
<date id="3F5F41ADFFFF6343EB6FC39AFF1B0763" pageId="1" pageNumber="382" value="1985-08-09">9 August 1985</date>
, coll. S. Shokita; URM-P 31439, 
<specimenCount id="5DE7ACE4FFFF6343EC3BC3BDFD730763" box="[608,698,1517,1542]" pageId="1" pageNumber="382" type="male">6 males</specimenCount>
(
<quantity id="8C19CA88FFFF6343EC97C3BDFCBD0760" box="[716,884,1517,1542]" metricMagnitude="-2" metricUnit="m" metricValue="3.545" metricValueMax="3.6799999999999997" metricValueMin="3.41" pageId="1" pageNumber="382" unit="mm" value="35.45" valueMax="36.8" valueMin="34.1">34.1–36.8 mm</quantity>
SL) and 
<specimenCount id="5DE7ACE4FFFF6343EDBDC3BEFB9F0763" box="[998,1110,1517,1542]" pageId="1" pageNumber="382" type="female">7 females</specimenCount>
(
<quantity id="8C19CA88FFFF6343EA33C3BDFAD90760" box="[1128,1296,1517,1542]" metricMagnitude="-2" metricUnit="m" metricValue="3.4299999999999997" metricValueMax="3.62" metricValueMin="3.2399999999999998" pageId="1" pageNumber="382" unit="mm" value="34.3" valueMax="36.2" valueMin="32.4">32.4–36.2 mm</quantity>
SL), Papait River, Palawan, 
<date id="3F5F41ADFFFF6343EF08C042FE37074F" box="[339,510,1553,1578]" pageId="1" pageNumber="382" value="1985-08-09">9 August 1985</date>
, coll. S. Shokita; URM-P 31440, 
<specimenCount id="5DE7ACE4FFFF6343EDD9C042FC39074F" box="[898,1008,1553,1578]" pageId="1" pageNumber="382" type="female">2 females</specimenCount>
(38.6 and 
<quantity id="8C19CA88FFFF6343EA36C042FB1A074C" box="[1133,1235,1554,1578]" metricMagnitude="-2" metricUnit="m" metricValue="4.07" pageId="1" pageNumber="382" unit="mm" value="40.7">40.7 mm</quantity>
SL), 
<collectingCountry id="33F627FDFFFF6343EB4DC042FA99074C" box="[1302,1360,1554,1577]" name="Iran" pageId="1" pageNumber="382">Iraan</collectingCountry>
River, Palawan, 
<date id="3F5F41ADFFFF6343EF5EC065FE79072B" box="[261,432,1589,1614]" pageId="1" pageNumber="382" value="1985-08-05">5 August 1985</date>
, coll. S. Shokita; URM-P 31441, 
<specimenCount id="5DE7ACE4FFFF6343ED69C065FC43072B" box="[818,906,1589,1614]" pageId="1" pageNumber="382" type="male">6 males</specimenCount>
(
<quantity id="8C19CA88FFFF6343EDC0C066FB880728" box="[923,1089,1590,1614]" metricMagnitude="-2" metricUnit="m" metricValue="3.635" metricValueMax="4.14" metricValueMin="3.13" pageId="1" pageNumber="382" unit="mm" value="36.35" valueMax="41.4" valueMin="31.3">31.3–41.4 mm</quantity>
SL) and 
<specimenCount id="5DE7ACE4FFFF6343EAF6C066FAD3072B" box="[1197,1306,1589,1614]" pageId="1" pageNumber="382" type="female">3 females</specimenCount>
(
<quantity id="8C19CA88FFFF6343EB71C066FF0A0714" metricMagnitude="-2" metricUnit="m" metricValue="3.4899999999999998" metricValueMax="3.91" metricValueMin="3.07" pageId="1" pageNumber="382" unit="mm" value="34.9" valueMax="39.1" valueMin="30.7">30.7–39.1 mm</quantity>
SL), Tagbariri, Palawan, 
<date id="3F5F41ADFFFF6343EFBCC00AFD580717" box="[487,657,1625,1650]" pageId="1" pageNumber="382" value="1985-08-09">9 August 1985</date>
, coll. S. Shokita; URM-P 48659–48662, 
<specimenCount id="5DE7ACE4FFFF6343EA3EC00AFB740717" box="[1125,1213,1625,1650]" pageId="1" pageNumber="382" type="male">2 males</specimenCount>
(46.2 and 
<quantity id="8C19CA88FFFF6343EB6CC009FA540714" box="[1335,1437,1625,1650]" metricMagnitude="-2" metricUnit="m" metricValue="5.11" pageId="1" pageNumber="382" unit="mm" value="51.1">51.1 mm</quantity>
SL) and 
<specimenCount id="5DE7ACE4FFFF6343EEA6C02EFEA207F3" box="[253,363,1661,1686]" pageId="1" pageNumber="382" type="female">2 females</specimenCount>
(41.1 and 
<quantity id="8C19CA88FFFF6343EFB7C02EFD9B07F0" box="[492,594,1662,1686]" metricMagnitude="-2" metricUnit="m" metricValue="4.72" pageId="1" pageNumber="382" unit="mm" value="47.2">47.2 mm</quantity>
SL), Barake Stream (tributary of Aborlan River), Barangay Magbabadil, Aborlan, Palawan, 
<date id="3F5F41ADFFFF6343EF2AC0F1FDC407DF" box="[369,525,1697,1722]" pageId="1" pageNumber="382" value="2015-05-15">15 May 2015</date>
, coll. K. Maeda and H. P. Palla; URM-P 48663–48666, 
<specimenCount id="5DE7ACE4FFFF6343EACEC0F2FB2407DF" box="[1173,1261,1697,1722]" pageId="1" pageNumber="382" type="male">2 males</specimenCount>
(59.7 and 
<quantity id="8C19CA88FFFF6343EB30C0F1FF0A07B8" metricMagnitude="-2" metricUnit="m" metricValue="6.39" pageId="1" pageNumber="382" unit="mm" value="63.9">63.9 mm</quantity>
SL) and 
<specimenCount id="5DE7ACE4FFFF6343EF76C096FE5307BB" box="[301,410,1733,1758]" pageId="1" pageNumber="382" type="female">2 females</specimenCount>
(58.5 and 
<quantity id="8C19CA88FFFF6343EC4DC095FDB307B8" box="[534,634,1733,1758]" metricMagnitude="-2" metricUnit="m" metricValue="6.36" pageId="1" pageNumber="382" unit="mm" value="63.6">63.6 mm</quantity>
SL), same data as 
<typeStatus id="945AD9CFFFFF6343ED08C095FC7107BB" box="[851,952,1733,1758]" pageId="1" pageNumber="382" type="holotype">holotype</typeStatus>
; WPU-PPC-P 2–4, 
<specimenCount id="5DE7ACE4FFFF6343EAC2C096FB3907BB" box="[1177,1264,1733,1758]" pageId="1" pageNumber="382" type="male">2 males</specimenCount>
(52.6 and 
<quantity id="8C19CA88FFFF6343EB30C095FF0A0664" metricMagnitude="-2" metricUnit="m" metricValue="5.92" pageId="1" pageNumber="382" unit="mm" value="59.2">59.2 mm</quantity>
SL) and female (
<quantity id="8C19CA88FFFF6343EFD0C0B9FE3B0664" box="[395,498,1769,1794]" metricMagnitude="-2" metricUnit="m" metricValue="4.63" pageId="1" pageNumber="382" unit="mm" value="46.3">46.3 mm</quantity>
SL), Barake Stream (tributary of Aborlan River), Barangay Magbabadil, Aborlan, Palawan, 
<date id="3F5F41ADFFFF6343EF58C15DFE530643" box="[259,410,1805,1830]" pageId="1" pageNumber="382" value="2015-05-15">15 May 2015</date>
, coll. K. Maeda and H. P. Palla; WPU-PPC-P 6–9, male (
<quantity id="8C19CA88FFFF6343EA4CC15DFBB50640" box="[1047,1148,1805,1830]" metricMagnitude="-2" metricUnit="m" metricValue="5.9399999999999995" pageId="1" pageNumber="382" unit="mm" value="59.4">59.4 mm</quantity>
SL) and 
<specimenCount id="5DE7ACE4FFFF6343EAB9C15EFA840643" box="[1250,1357,1805,1830]" pageId="1" pageNumber="382" type="female">3 females</specimenCount>
(
<quantity id="8C19CA88FFFF6343EB00C15DFF32062C" metricMagnitude="-2" metricUnit="m" metricValue="5.9799999999999995" metricValueMax="6.21" metricValueMin="5.75" pageId="1" pageNumber="382" unit="mm" value="59.8" valueMax="62.1" valueMin="57.5">57.5– 62.1 mm</quantity>
SL), same data as 
<typeStatus id="945AD9CFFFFF6343EF95C161FDFD062F" box="[462,564,1841,1866]" pageId="1" pageNumber="382" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="03FB34E6FFFF6340EE9CC105FD68019D" lastPageId="2" lastPageNumber="383" pageId="1" pageNumber="382" type="diagnosis">
<paragraph id="4B5E676DFFFF6340EE9CC105FD68019D" blockId="1.[151,1437,1193,2010]" lastBlockId="2.[151,1437,151,536]" lastPageId="2" lastPageNumber="383" pageId="1" pageNumber="382">
<emphasis id="7995BB7FFFFF6343EE9CC105FE88060B" bold="true" box="[199,321,1877,1902]" pageId="1" pageNumber="382">Diagnosis.</emphasis>
Second dorsal fin usually with one spine and nine segmented rays, pectoral fin usually with 15 segmented rays; first dorsal fin pointed in male; relatively large caudal fin (26–34% of SL) in male; premaxilla with 45–71 tricuspid teeth, dentary with 2–6 (male) or 1–4 (female) symphyseal teeth and 45–79 unicuspid horizontal teeth; nape and posterior part of occipital region covered by cycloid scales; 9–11 dusky transverse bars laterally on trunk and tail; first dorsal fin gray or dusky after preservation (orange or reddish brown in life) without any distinct markings, having a line of black blotches (male) or a black band (female) on distal part of second dorsal fin; pectoral fin without clear markings.
</paragraph>
</subSubSection>
<subSubSection id="03FB34E6FFFC634AEE9CC754FE5C0305" lastPageId="8" lastPageNumber="389" pageId="2" pageNumber="383" type="description">
<paragraph id="4B5E676DFFFC6340EE9CC754FBDC037D" blockId="2.[151,1437,151,536]" pageId="2" pageNumber="383">
<emphasis id="7995BB7FFFFC6340EE9CC754FE900078" bold="true" box="[199,345,260,285]" pageId="2" pageNumber="383">Description.</emphasis>
Morphometric measurements given in Table 1. Body elongate, cylindrical anteriorly and somewhat compressed posteriorly. Head somewhat depressed with a round snout protruding beyond upper lip. Anterior nostril tubular and short, posterior nostril round or oblong, not tubular. Mouth inferior with upper jaw projecting beyond lower jaw. Upper lip thick with small medial cleft. Premaxillary teeth 45–71, fine and tricuspid. Dentary with canine-like symphyseal teeth (number of teeth 
<quantity id="8C19CA88FFFC6340ED16C7C4FC5F00C8" box="[845,918,404,429]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.524" metricValueMin="0.508" pageId="2" pageNumber="383" unit="in" value="4.0" valueMax="6.0" valueMin="2.0">2–6 in</quantity>
male, 
<quantity id="8C19CA88FFFC6340EDBEC7C4FBE400C8" box="[997,1069,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="2" pageNumber="383" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1–4 in</quantity>
female) and a row of unicuspid horizontal teeth enclosed in fleshy sheath (number of teeth 45–79). Larger fish having more premaxillary and horizontal teeth (
<figureCitation id="D3DA7BE8FFFC6340EF39C78CFE670091" box="[354,430,476,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1696,1718]" captionTargetBox="[349,1239,204,1666]" captionTargetId="figure@4.[340,1247,193,1675]" captionTargetPageId="4" captionText="FIGURE 2. Number of premaxillary and horizontal dentary teeth of Stiphodon palawanensis (red circles), S. alcedo (pink circles), S. atratus (black diamonds), S. imperiorientis (light blue triangles), S. maculidorsalis (light green triangles), S. multisquamus (deep blue circles), S. ornatus (orange triangles), S. pelewensis (gray diamonds), S. pulchellus (deep green triangles), and S. weberi (purple diamonds)." httpUri="https://zenodo.org/record/232901/files/figure.png" pageId="2" pageNumber="383">Fig. 2</figureCitation>
). Urogenital papilla in male rounded with one cleft at posterior tip; that in female rectangular, bearing one small projection at each corner of the posterior margin.
</paragraph>
<paragraph id="4B5E676DFFFC6340EECCC418FC1C03E4" blockId="2.[151,1436,583,646]" pageId="2" pageNumber="383">
<emphasis id="7995BB7FFFFC6340EECCC418FEC3033A" bold="true" box="[151,266,584,607]" pageId="2" pageNumber="383">TABLE 1.</emphasis>
Morphometrics of 
<taxonomicName id="8CE11CEEFFFC6340EF81C417FD14033A" box="[474,733,583,607]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="383" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFFC6340EF81C417FD14033A" box="[474,733,583,607]" italics="true" pageId="2" pageNumber="383">Stiphodon palawanensis</emphasis>
</taxonomicName>
, expressed as a percentage of standard length. D1, first dorsal fin; D2, second dorsal fin; A, anal fin; C, caudal fin; P1, pectoral fin; P2, pelvic fin.
</paragraph>
<paragraph id="4B5E676DFFFC6340EEC4C499FACC064C" pageId="2" pageNumber="383">
<table id="39E195CDFFFC9CBDEECCC4F0FA55064C" box="[151,1436,672,1833]" gridcols="4" gridrows="8" pageId="2" pageNumber="383">
<tr id="F5D1652FFFFC9CBDEECCC4F0FA55026D" box="[151,1436,672,776]" gridrow="0" pageId="2" pageNumber="383">
<th id="B6000C53FFFC9CBDEECCC4F0FE39026D" box="[151,496,672,776]" gridcol="0" gridrow="0" pageId="2" pageNumber="383">Sex Number of specimens measured</th>
<th id="B6000C53FFFC9CBDECF1C4F0FCC1026D" box="[682,776,672,776]" gridcol="1" gridrow="0" pageId="2" pageNumber="383">Holotype Male 1</th>
<th id="B6000C53FFFC9CBDEDC3C4F0FC37026D" box="[920,1022,672,776]" gridcol="2" gridrow="0" pageId="2" pageNumber="383">Paratype Male 23</th>
<th id="B6000C53FFFC9CBDEAFBC4F0FA55026D" box="[1184,1436,672,776]" gridcol="3" gridrow="0" pageId="2" pageNumber="383">Female 33</th>
</tr>
<tr id="F5D1652FFFFC9CBDEECCC54CFA5502E1" box="[151,1436,796,900]" gridrow="1" pageId="2" pageNumber="383">
<th id="B6000C53FFFC9CBDEECCC54CFE3902E1" box="[151,496,796,900]" gridcol="0" gridrow="1" pageId="2" pageNumber="383">Standard length (mm) Head length Snout length</th>
<td id="B6000C53FFFC9CBDECF1C54CFCC102E1" box="[682,776,796,900]" gridcol="1" gridrow="1" pageId="2" pageNumber="383">62.2 22.8 9.2</td>
<td id="B6000C53FFFC9CBDEDC3C54CFC3702E1" box="[920,1022,796,900]" gridcol="2" gridrow="1" pageId="2" pageNumber="383">27.7–63.9 21.5–23.9 6.8–9.0</td>
<td id="B6000C53FFFC9CBDEAFBC54CFA5502E1" box="[1184,1436,796,900]" gridcol="3" gridrow="1" pageId="2" pageNumber="383">27.0–63.6 20.8–25.2 6.8–9.2</td>
</tr>
<tr id="F5D1652FFFFC9CBDEECCC5C8FA550565" box="[151,1436,920,1024]" gridrow="2" pageId="2" pageNumber="383">
<th id="B6000C53FFFC9CBDEECCC5C8FE390565" box="[151,496,920,1024]" gridcol="0" gridrow="2" pageId="2" pageNumber="383">Eye diameter Postorbital length of head Upper jaw length</th>
<td id="B6000C53FFFC9CBDECF1C5C8FCC10565" box="[682,776,920,1024]" gridcol="1" gridrow="2" pageId="2" pageNumber="383">4.3 10.9 10.3</td>
<td id="B6000C53FFFC9CBDEDC3C5C8FC370565" box="[920,1022,920,1024]" gridcol="2" gridrow="2" pageId="2" pageNumber="383">4.1–6.0 9.8–12.0 9.0–11.2</td>
<td id="B6000C53FFFC9CBDEAFBC5C8FA550565" box="[1184,1436,920,1024]" gridcol="3" gridrow="2" pageId="2" pageNumber="383">4.4–7.0 9.4–12.0 9.1–10.8</td>
</tr>
<tr id="F5D1652FFFFC9CBDEECCC244FA550530" box="[151,1436,1044,1109]" gridrow="3" pageId="2" pageNumber="383">
<th id="B6000C53FFFC9CBDEECCC244FE390530" box="[151,496,1044,1109]" gridcol="0" gridrow="3" pageId="2" pageNumber="383">Body depth at P2 origin Body depth at A origin</th>
<td id="B6000C53FFFC9CBDECF1C244FCC10530" box="[682,776,1044,1109]" gridcol="1" gridrow="3" pageId="2" pageNumber="383">15.4 16.2</td>
<td id="B6000C53FFFC9CBDEDC3C244FC370530" box="[920,1022,1044,1109]" gridcol="2" gridrow="3" pageId="2" pageNumber="383">12.9–15.2 14.9–17.2</td>
<td id="B6000C53FFFC9CBDEAFBC244FA550530" box="[1184,1436,1044,1109]" gridcol="3" gridrow="3" pageId="2" pageNumber="383">13.0–15.6 15.3–17.9</td>
</tr>
<tr id="F5D1652FFFFC9CBDEECCC239FA5505CD" box="[151,1436,1129,1192]" gridrow="4" pageId="2" pageNumber="383">
<th id="B6000C53FFFC9CBDEECCC239FE3905CD" box="[151,496,1129,1192]" gridcol="0" gridrow="4" pageId="2" pageNumber="383">Depth of caudal peduncle Length of caudal peduncle</th>
<td id="B6000C53FFFC9CBDECF1C239FCC105CD" box="[682,776,1129,1192]" gridcol="1" gridrow="4" pageId="2" pageNumber="383">13.0 19.3</td>
<td id="B6000C53FFFC9CBDEDC3C239FC3705CD" box="[920,1022,1129,1192]" gridcol="2" gridrow="4" pageId="2" pageNumber="383">11.3–12.6 18.2–21.1</td>
<td id="B6000C53FFFC9CBDEAFBC239FA5505CD" box="[1184,1436,1129,1192]" gridcol="3" gridrow="4" pageId="2" pageNumber="383">10.5–12.3 18.3–21.4</td>
</tr>
<tr id="F5D1652FFFFC9CBDEECCC2ECFA550732" box="[151,1436,1212,1623]" gridrow="5" pageId="2" pageNumber="383">
<th id="B6000C53FFFC9CBDEECCC2ECFE390732" box="[151,496,1212,1623]" gridcol="0" gridrow="5" pageId="2" pageNumber="383">Predorsal length Length of D1 base D1 length Length of longest spine of D1 Interval between D1 and D2 bases Length of D2 base D2 length Length of longest ray of D2 Preanal length Length of A base</th>
<td id="B6000C53FFFC9CBDECF1C2ECFCC10732" box="[682,776,1212,1623]" gridcol="1" gridrow="5" pageId="2" pageNumber="383">32.0 18.6 35.4 30.5 3.2 26.7 45.3 21.1 51.4 29.4</td>
<td id="B6000C53FFFC9CBDEDC3C2ECFC370732" box="[920,1022,1212,1623]" gridcol="2" gridrow="5" pageId="2" pageNumber="383">31.9–35.0 16.6–21.7 25.3–41.7 18.4–35.9 1.7–5.4 23.6–27.2 37.2–46.8 14.4–21.5 49.5–54.0 26.5–30.3</td>
<td id="B6000C53FFFC9CBDEAFBC2ECFA550732" box="[1184,1436,1212,1623]" gridcol="3" gridrow="5" pageId="2" pageNumber="383">33.0–37.0 17.0–23.0 18.1–23.0 13.4–17.8 2.5–7.2 21.4–25.7 28.5–36.7 12.7–16.7 49.4–56.0 18.6–27.5</td>
</tr>
<tr id="F5D1652FFFFC9CBDEECCC03BFA5507B6" box="[151,1436,1643,1747]" gridrow="6" pageId="2" pageNumber="383">
<th id="B6000C53FFFC9CBDEECCC03BFE3907B6" box="[151,496,1643,1747]" gridcol="0" gridrow="6" pageId="2" pageNumber="383">A length Length of longest ray of A Length from anus to A</th>
<td id="B6000C53FFFC9CBDECF1C03BFCC107B6" box="[682,776,1643,1747]" gridcol="1" gridrow="6" pageId="2" pageNumber="383">43.9 18.2 3.5</td>
<td id="B6000C53FFFC9CBDEDC3C03BFC3707B6" box="[920,1022,1643,1747]" gridcol="2" gridrow="6" pageId="2" pageNumber="383">38.6–47.0 13.7–18.3 2.9–4.2</td>
<td id="B6000C53FFFC9CBDEAFBC03BFA5507B6" box="[1184,1436,1643,1747]" gridcol="3" gridrow="6" pageId="2" pageNumber="383">31.6–37.7 11.1–14.4 3.0–5.6</td>
</tr>
<tr id="F5D1652FFFFC9CBDEECCC0B7FA55064C" box="[151,1436,1767,1833]" gridrow="7" pageId="2" pageNumber="383">
<th id="B6000C53FFFC9CBDEECCC0B7FE39064C" box="[151,496,1767,1833]" gridcol="0" gridrow="7" pageId="2" pageNumber="383">Length of longest ray of P1 C length</th>
<td id="B6000C53FFFC9CBDECF1C0B7FCC1064C" box="[682,776,1767,1833]" gridcol="1" gridrow="7" pageId="2" pageNumber="383">20.7 31.4</td>
<td id="B6000C53FFFC9CBDEDC3C0B7FC37064C" box="[920,1022,1767,1833]" gridcol="2" gridrow="7" pageId="2" pageNumber="383">18.2–23.7 26.4–34.2</td>
<td id="B6000C53FFFC9CBDEAFBC0B7FA55064C" box="[1184,1436,1767,1833]" gridcol="3" gridrow="7" pageId="2" pageNumber="383">17.9–21.9 21.3–26.1</td>
</tr>
</table>
</paragraph>
<paragraph id="4B5E676DFFFC6341EE9CC13DFE1300C8" blockId="2.[151,1436,1901,2034]" lastBlockId="3.[151,1436,151,429]" lastPageId="3" lastPageNumber="384" pageId="2" pageNumber="383">First dorsal fin with six spines, except two specimens with five spines; second dorsal fin usually with one spine and nine segmented rays (one specimen with one spine and eight segmented rays). In female, first dorsal fin rounded, almost semicircular, usually second and/or third spines longest, but its posteriormost tip never extending to origin of second dorsal fin. In male, first dorsal fin spines elongate (usually fourth spine longest) and posteriormost point of first dorsal fin (tip of fourth spine) extending to base of first to sixth segmented ray of second dorsal fin when depressed. Anal fin with one spine and ten segmented rays. In female, usually first or second and second or third segmented rays longest in second dorsal and anal fins, respectively; in male, posterior rays longer than anterior rays (last ray and/or penultimate ray usually longest). Caudal fin usually with 17 segmented rays, including 13 branched rays, posterior margin rounded; caudal fin relatively larger in male than in female (caudal-fin length 26–34% of SL in male, 21–26% of SL in female). Pectoral fin with 14 (n=7), 15 (n=47), or 16 (n=3) rays. Pelvic fin with one spine and five segmented rays; pelvic fins joined together to form strong, cuplike disk with fleshy frenum.</paragraph>
<caption id="1F9E37E5FFFD6341EECCC1DFFEED0687" httpUri="https://zenodo.org/record/232900/files/figure.png" pageId="3" pageNumber="384" targetBox="[151,1436,481,1912]" targetPageId="3">
<paragraph id="4B5E676DFFFD6341EECCC1DFFEED0687" blockId="3.[151,1436,1935,2018]" pageId="3" pageNumber="384">
<emphasis id="7995BB7FFFFD6341EECCC1DFFEDD06C1" bold="true" box="[151,276,1935,1957]" pageId="3" pageNumber="384">FIGURE 1.</emphasis>
<taxonomicName id="8CE11CEEFFFD6341EF40C1DFFDDA06C0" box="[283,531,1935,1957]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="384" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFFD6341EF40C1DFFDDA06C0" box="[283,531,1935,1957]" italics="true" pageId="3" pageNumber="384">Stiphodon palawanensis</emphasis>
</taxonomicName>
immediately after fixation. a, WPU-PPC-P 5, holotype (male, 62.2 mm SL); b, URM-P 48659, paratype (male, 51.1 mm SL); c, URM-P 48666, paratype (female, 58.5 mm SL); d, URM-P 48661, paratype (female, 41.1 mm SL).
</paragraph>
</caption>
<caption id="1F9E37E5FFFA6346EECCC0F0FD9D0677" httpUri="https://zenodo.org/record/232901/files/figure.png" pageId="4" pageNumber="385" targetBox="[349,1239,204,1666]" targetPageId="4">
<paragraph id="4B5E676DFFFA6346EECCC0F0FD9D0677" blockId="4.[151,1436,1696,1810]" pageId="4" pageNumber="385">
<emphasis id="7995BB7FFFFA6346EECCC0F0FEDC07D0" bold="true" box="[151,277,1696,1718]" pageId="4" pageNumber="385">FIGURE 2.</emphasis>
Number of premaxillary and horizontal dentary teeth of 
<taxonomicName id="8CE11CEEFFFA6346ED34C0F0FBA107D3" box="[879,1128,1696,1718]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFFA6346ED34C0F0FBA107D3" box="[879,1128,1696,1718]" italics="true" pageId="4" pageNumber="385">Stiphodon palawanensis</emphasis>
</taxonomicName>
(red circles), 
<taxonomicName id="8CE11CEEFFFA6346EAA6C0F1FA9707D3" box="[1277,1374,1696,1718]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="alcedo">
<emphasis id="7995BB7FFFFA6346EAA6C0F1FA9707D3" box="[1277,1374,1696,1718]" italics="true" pageId="4" pageNumber="385">S. alcedo</emphasis>
</taxonomicName>
(pink circles), 
<taxonomicName id="8CE11CEEFFFA6346EEA8C090FE9307B0" box="[243,346,1728,1749]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="atratus">
<emphasis id="7995BB7FFFFA6346EEA8C090FE9307B0" box="[243,346,1728,1749]" italics="true" pageId="4" pageNumber="385">S. atratus</emphasis>
</taxonomicName>
(black diamonds), 
<taxonomicName id="8CE11CEEFFFA6346EC77C090FD1207B0" box="[556,731,1728,1749]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="imperiorientis">
<emphasis id="7995BB7FFFFA6346EC77C090FD1207B0" box="[556,731,1728,1749]" italics="true" pageId="4" pageNumber="385">S. imperiorientis</emphasis>
</taxonomicName>
(light blue triangles), 
<taxonomicName id="8CE11CEEFFFA6346ED8AC090FB4C07B0" box="[977,1157,1727,1749]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="maculidorsalis">
<emphasis id="7995BB7FFFFA6346ED8AC090FB4C07B0" box="[977,1157,1727,1749]" italics="true" pageId="4" pageNumber="385">S. maculidorsalis</emphasis>
</taxonomicName>
(light green triangles), 
<taxonomicName id="8CE11CEEFFFA6346EBD2C090FEED0796" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="multisquamus">
<emphasis id="7995BB7FFFFA6346EBD2C090FEED0796" italics="true" pageId="4" pageNumber="385">S. multisquamus</emphasis>
</taxonomicName>
(deep blue circles), 
<taxonomicName id="8CE11CEEFFFA6346EC59C08EFDA40791" box="[514,621,1758,1780]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="7995BB7FFFFA6346EC59C08EFDA40791" box="[514,621,1758,1780]" italics="true" pageId="4" pageNumber="385">S. ornatus</emphasis>
</taxonomicName>
(orange triangles), 
<taxonomicName id="8CE11CEEFFFA6346ED1BC08EFC020796" box="[832,971,1757,1779]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="pelewensis">
<emphasis id="7995BB7FFFFA6346ED1BC08EFC020796" box="[832,971,1757,1779]" italics="true" pageId="4" pageNumber="385">S. pelewensis</emphasis>
</taxonomicName>
(gray diamonds), 
<taxonomicName id="8CE11CEEFFFA6346EACAC08EFADF0796" box="[1169,1302,1757,1779]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="7995BB7FFFFA6346EACAC08EFADF0796" box="[1169,1302,1757,1779]" italics="true" pageId="4" pageNumber="385">S. pulchellus</emphasis>
</taxonomicName>
(deep green triangles), and 
<taxonomicName id="8CE11CEEFFFA6346EF75C0ADFE450677" box="[302,396,1788,1810]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="weberi">
<emphasis id="7995BB7FFFFA6346EF75C0ADFE450677" box="[302,396,1788,1810]" italics="true" pageId="4" pageNumber="385">S. weberi</emphasis>
</taxonomicName>
(purple diamonds).
</paragraph>
</caption>
<caption id="1F9E37E5FFFB6347EECCC2FAFEB405BA" httpUri="https://zenodo.org/record/232902/files/figure.png" pageId="5" pageNumber="386" targetBox="[349,1236,184,1162]" targetPageId="5">
<paragraph id="4B5E676DFFFB6347EECCC2FAFEB405BA" blockId="5.[151,1436,1194,1247]" pageId="5" pageNumber="386">
<emphasis id="7995BB7FFFFB6347EECCC2FAFEDB05DA" bold="true" box="[151,274,1194,1216]" pageId="5" pageNumber="386">FIGURE 3.</emphasis>
Dorsal scalation on head and nape in 
<taxonomicName id="8CE11CEEFFFB6347ECD4C2FAFC4C05A5" box="[655,901,1194,1216]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFFB6347ECD4C2FAFC4C05A5" box="[655,901,1194,1216]" italics="true" pageId="5" pageNumber="386">Stiphodon palawanensis</emphasis>
</taxonomicName>
. a, male (43.1 mm SL, CMK 11966); b, female (41.9 mm SL, CMK 11966).
</paragraph>
</caption>
<paragraph id="4B5E676DFFFB6347EE9CC35BFD3B0745" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="386">
Scales in a longitudinal series 29 (n=4), 30 (n=13), 31 (n=19), 32 (n=15, including 
<typeStatus id="945AD9CFFFFB6347EAC9C35BFB300441" box="[1170,1273,1291,1316]" pageId="5" pageNumber="386" type="holotype">holotype</typeStatus>
), or 33 (n=6); scales in a transverse series 9 (n=1), 10 (n=3), 11 (n=52, including 
<typeStatus id="945AD9CFFFFB6347ED25C37FFC2D042D" box="[894,996,1327,1352]" pageId="5" pageNumber="386" type="holotype">holotype</typeStatus>
), 12 (n=1); circumpeduncular scales 15 (n=1) or 16 (n=55) (not including one broken specimen). Ctenoid scales covering tail, sides and dorsum of posterior trunk. Pectoral-fin base naked. Small naked area behind pectoral-fin base; some anteriormost scales on lateral sides of trunk cycloid. Belly covered with cycloid scales. Nape and posterior part of occipital region covered by cycloid scales (
<figureCitation id="D3DA7BE8FFFB6347EF37C3EFFE7804BD" box="[364,433,1471,1496]" captionStart="FIGURE 3" captionStartId="5.[151,250,1194,1216]" captionTargetBox="[349,1236,184,1162]" captionTargetId="figure@5.[340,1247,173,1173]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal scalation on head and nape in Stiphodon palawanensis. a, male (43.1 mm SL, CMK 11966); b, female (41.9 mm SL, CMK 11966)." httpUri="https://zenodo.org/record/232902/files/figure.png" pageId="5" pageNumber="386">Fig. 3</figureCitation>
); in female, scaled area slightly exceeding middle of occipital region; male similar, but sometimes not exceeding the middle. Cycloid scales also occurring on first and second dorsal-fin bases, anal-fin base, caudal-fin base, and proximal part of caudal fin.
</paragraph>
<paragraph id="4B5E676DFFFB6347EE9CC07BFE6D07E9" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="386">
Cephalic sensory pore system always with A´, B, C, D(S), F, H´, K´, L´, N´, and 
<collectingCountry id="33F627FDFFFB6347EA35C07CFB4D0721" box="[1134,1156,1580,1604]" name="Iceland" pageId="5" pageNumber="386">O</collectingCountry>
´(
<figureCitation id="D3DA7BE8FFFB6347EAC6C07BFB2D0721" box="[1181,1252,1579,1604]" captionStart="FIGURE 4" captionStartId="6.[151,250,1952,1974]" captionTargetBox="[384,1184,211,1923]" captionTargetId="figure@6.[376,1206,193,1942]" captionTargetPageId="6" captionText="FIGURE 4. Diagrammatic illustration of head showing arrangement of cephalic sensory pores and cutaneous sensory papillae in Stiphodon palawanensis (41.9 mm SL, CMK 11966). a, dorsal view; b, lateral view; c, ventral view. AN, anterior nostril; PN, posterior nostril." httpUri="https://zenodo.org/record/232903/files/figure.png" pageId="5" pageNumber="386">Fig. 4</figureCitation>
). Oculoscapular canal interrupted between pores H´and K´. Cutaneous sensory papillae developed over dorsal, lateral, and ventral surface of head (
<figureCitation id="D3DA7BE8FFFB6347EF0AC023FE5F07E9" box="[337,406,1651,1676]" captionStart="FIGURE 4" captionStartId="6.[151,250,1952,1974]" captionTargetBox="[384,1184,211,1923]" captionTargetId="figure@6.[376,1206,193,1942]" captionTargetPageId="6" captionText="FIGURE 4. Diagrammatic illustration of head showing arrangement of cephalic sensory pores and cutaneous sensory papillae in Stiphodon palawanensis (41.9 mm SL, CMK 11966). a, dorsal view; b, lateral view; c, ventral view. AN, anterior nostril; PN, posterior nostril." httpUri="https://zenodo.org/record/232903/files/figure.png" pageId="5" pageNumber="386">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="4B5E676DFFFB6347EE9CC0C7FEED07B1" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="386">
The largest male and female specimens were 63.9 and 
<quantity id="8C19CA88FFFB6347ED0CC0C7FC0907CA" box="[855,960,1687,1712]" metricMagnitude="-2" metricUnit="m" metricValue="6.36" pageId="5" pageNumber="386" unit="mm" value="63.6">63.6 mm</quantity>
SL (82.9 and 
<quantity id="8C19CA88FFFB6347EA29C0C8FB1207CA" box="[1138,1243,1688,1712]" metricMagnitude="-2" metricUnit="m" metricValue="7.79" pageId="5" pageNumber="386" unit="mm" value="77.9">77.9 mm</quantity>
in total length), respectively.
</paragraph>
<paragraph id="4B5E676DFFFB6347EE9CC08FFE9E0691" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="386">
<emphasis id="7995BB7FFFFB6347EE9CC08FFE0E0792" box="[199,455,1759,1783]" italics="true" pageId="5" pageNumber="386">Color in preservative.</emphasis>
In male, background brown; 9–11 (usually 10) dusky transverse bars laterally and dorsally on trunk and tail. First dorsal fin gray or dusky without distinct markings. Second dorsal fin gray or dusky; each ray having a black blotch surrounded by a translucent margin, these black blotches forming a line along second dorsal fin margin, but posterior part of the fin sometimes lacking this black blotch. Anal fin gray or dusky. Caudal fin with dusky longitudinal band on upper part; dorsal to this band, gray; ventrally along this band translucent; middle and lower parts of caudal fin gray. Pectoral fin pale gray without clear markings, but sometimes with one to four obscure, dusky spots on each of middle rays. Pelvic disk dusky, but margin of anterior and middle parts translucent.
</paragraph>
<caption id="1F9E37E5FFF86344EECCC1F0FEF60691" httpUri="https://zenodo.org/record/232903/files/figure.png" pageId="6" pageNumber="387" targetBox="[384,1184,211,1923]" targetPageId="6">
<paragraph id="4B5E676DFFF86344EECCC1F0FEF60691" blockId="6.[151,1436,1952,2036]" pageId="6" pageNumber="387">
<emphasis id="7995BB7FFFF86344EECCC1F0FEDA06D0" bold="true" box="[151,275,1952,1974]" pageId="6" pageNumber="387">FIGURE 4.</emphasis>
Diagrammatic illustration of head showing arrangement of cephalic sensory pores and cutaneous sensory papillae in 
<taxonomicName id="8CE11CEEFFF86344EEEBC1EFFE6F06B0" box="[176,422,1983,2005]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF86344EEEBC1EFFE6F06B0" box="[176,422,1983,2005]" italics="true" pageId="6" pageNumber="387">Stiphodon palawanensis</emphasis>
</taxonomicName>
(41.9 mm SL, CMK 11966). a, dorsal view; b, lateral view; c, ventral view. AN, anterior nostril; PN, posterior nostril.
</paragraph>
</caption>
<caption id="1F9E37E5FFF96345EECCC02CFBD707D4" httpUri="https://zenodo.org/record/232904/files/figure.png" pageId="7" pageNumber="388" targetBox="[151,1436,193,1637]" targetPageId="7">
<paragraph id="4B5E676DFFF96345EECCC02CFBD707D4" blockId="7.[151,1436,1660,1713]" pageId="7" pageNumber="388">
<emphasis id="7995BB7FFFF96345EECCC02CFEDA07F7" bold="true" box="[151,275,1660,1682]" pageId="7" pageNumber="388">FIGURE 5.</emphasis>
Males (a–d) and females (e, f) of live 
<taxonomicName id="8CE11CEEFFF96345ECC0C02CFC5807F7" box="[667,913,1660,1682]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF96345ECC0C02CFC5807F7" box="[667,913,1660,1682]" italics="true" pageId="7" pageNumber="388">Stiphodon palawanensis</emphasis>
</taxonomicName>
observed in Balsahan Stream, Puerto Princesa City (b, e, f, 18 May 2015; a, c, 19 May 2015) and Barake Stream, Aborlan (d, 15 May 2015).
</paragraph>
</caption>
<paragraph id="4B5E676DFFF96345EE9CC08EFA540697" blockId="7.[151,1437,1757,2034]" pageId="7" pageNumber="388">In female, background brown or yellowish brown; blackish longitudinal band extending from snout and upper lip to below eye and to middle of pectoral-fin base, band continuing from behind pectoral-fin base to posterior end of caudal peduncle through lateral midline; nine or ten (usually ten) dusky transverse bars laterally on trunk and tail intersecting with the mid-lateral longitudinal band. The longitudinal band often unclear if the transverse bars accentuated, and transverse bars often unclear if the longitudinal band accentuated. Additional blackish longitudinal band above the mid-lateral band from dorsoposterior edge of eye to base of upper procurrent caudalfin rays, but often obscure. First dorsal-fin gray without distinct markings. Second dorsal and anal fins gray with a black submarginal band and translucent margin; the black band on second dorsal fin thicker than that on anal fin.</paragraph>
<paragraph id="4B5E676DFFF6634AEECCC6C7FF1D0025" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="389">Black blotch on middle of proximal part of caudal fin; other part of caudal fin translucent with a black band (upside-down “L” shape) along dorsal and posterior margins and one to four black vertical stripes on middle part. Pectoral fin pale gray without clear markings, but sometimes with one or two obscure, dusky spots along middle rays. Pelvic fin translucent, but middle parts of rays, membranes, and frenum often dusky, forming a ring in ventral view.</paragraph>
<paragraph id="4B5E676DFFF6634AEE9CC71CFE5C0305" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="389">
<emphasis id="7995BB7FFFF6634AEE9CC71CFE900001" box="[199,345,332,356]" italics="true" pageId="8" pageNumber="389">Color in life.</emphasis>
Body and fin markings of male and female similar to those of preserved specimens, but in male (
<figureCitation id="D3DA7BE8FFF6634AEEC4C73FFF2D00ED" box="[159,228,367,392]" captionStart="FIGURE 5" captionStartId="7.[151,250,1660,1682]" captionTargetBox="[151,1436,193,1637]" captionTargetId="figure@7.[151,1436,193,1639]" captionTargetPageId="7" captionText="FIGURE 5. Males (a – d) and females (e, f) of live Stiphodon palawanensis observed in Balsahan Stream, Puerto Princesa City (b, e, f, 18 May 2015; a, c, 19 May 2015) and Barake Stream, Aborlan (d, 15 May 2015)." httpUri="https://zenodo.org/record/232904/files/figure.png" pageId="8" pageNumber="389">Fig. 5</figureCitation>
a–d), background of body grayish, purplish, or yellowish brown; first and second dorsal fins orange or reddish brown; black blotches on second dorsal fin surrounded by white or yellow margins; anal fin brown, gray, or orange; and dorsal part of caudal fin reddish or yellowish with bluish-white margin. In female (
<figureCitation id="D3DA7BE8FFF6634AEAB1C7E7FAE600B5" box="[1258,1327,439,464]" captionStart="FIGURE 5" captionStartId="7.[151,250,1660,1682]" captionTargetBox="[151,1436,193,1637]" captionTargetId="figure@7.[151,1436,193,1639]" captionTargetPageId="7" captionText="FIGURE 5. Males (a – d) and females (e, f) of live Stiphodon palawanensis observed in Balsahan Stream, Puerto Princesa City (b, e, f, 18 May 2015; a, c, 19 May 2015) and Barake Stream, Aborlan (d, 15 May 2015)." httpUri="https://zenodo.org/record/232904/files/figure.png" pageId="8" pageNumber="389">Fig. 5</figureCitation>
e, f), first dorsal fin reddish brown; second dorsal fin reddish brown with a black submarginal band and bluish white margin; white spots often lining proximal side of this band; anal fin brown with a black submarginal band and bluish white margin; upper and middle parts of caudal fin fringed by bluish white margin; area between the white margin and the black band orange.
</paragraph>
</subSubSection>
<subSubSection id="03FB34E6FFF6634BEE9CC43CFE5100B8" lastPageId="9" lastPageNumber="390" pageId="8" pageNumber="389" type="distribution">
<paragraph id="4B5E676DFFF6634AEE9CC43CFC140238" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="389">
<emphasis id="7995BB7FFFF6634AEE9CC43CFEA903E0" bold="true" box="[199,352,620,645]" pageId="8" pageNumber="389">Distribution.</emphasis>
The new species is currently known only from Palawan, in the western 
<collectingCountry id="33F627FDFFF6634AEACCC43CFAD003E0" box="[1175,1305,620,645]" name="Philippines" pageId="8" pageNumber="389">Philippines</collectingCountry>
. All known habitats are streams flowing into the Sulu Sea in the central part of the island (
<figureCitation id="D3DA7BE8FFF6634AEA6EC4DFFBB403CD" box="[1077,1149,655,680]" captionStart="FIGURE 6" captionStartId="8.[151,250,1866,1888]" captionTargetBox="[345,1240,917,1839]" captionTargetId="figure@8.[340,1248,913,1845]" captionTargetPageId="8" captionText="FIGURE 6. Location of Palawan in the Philippines (inset map) and sites where Stiphodon species have been found on Palawan. Red circles, S. palawanensis; green circles, S. pulchellus; blue circle, S. atropurpureus; and orange circle, S. percnopterygionus." httpUri="https://zenodo.org/record/232905/files/figure.png" pageId="8" pageNumber="389">Fig. 6</figureCitation>
). When we explored the Iwahig River where it flows into the South 
<collectingCountry id="33F627FDFFF6634AECEEC4E4FD3103A8" box="[693,760,692,717]" name="China" pageId="8" pageNumber="389">China</collectingCountry>
Sea at Quezon (
<figureCitation id="D3DA7BE8FFF6634AED97C4E4FBDF03A8" box="[972,1046,692,717]" captionStart="FIGURE 6" captionStartId="8.[151,250,1866,1888]" captionTargetBox="[345,1240,917,1839]" captionTargetId="figure@8.[340,1248,913,1845]" captionTargetPageId="8" captionText="FIGURE 6. Location of Palawan in the Philippines (inset map) and sites where Stiphodon species have been found on Palawan. Red circles, S. palawanensis; green circles, S. pulchellus; blue circle, S. atropurpureus; and orange circle, S. percnopterygionus." httpUri="https://zenodo.org/record/232905/files/figure.png" pageId="8" pageNumber="389">Fig. 6</figureCitation>
) for two days in 
<date id="3F5F41ADFFF6634AEAA3C4E5FABC03A8" box="[1272,1397,692,717]" pageId="8" pageNumber="389" value="2015-05">May 2015</date>
, 
<taxonomicName id="8CE11CEEFFF6634AEBDCC4E4FEF90395" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF6634AEBDCC4E4FEF90395" italics="true" pageId="8" pageNumber="389">S. palawanensis</emphasis>
</taxonomicName>
was not found, but other areas, such as the northern and southern parts of the island, have not yet been explored. Given its amphidromous life cycle with the high dispersal ability of the larvae suggested in this genus (
<bibRefCitation id="2F701A9CFFF6634AEEB7C54FFE3A025D" author="Yamasaki" box="[236,499,799,824]" pageId="8" pageNumber="389" refString="Yamasaki, N., Maeda, K. &amp; Tachihara, K. (2007) Pelagic larval duration and morphology at recruitment of Stiphodon percnopterygionus (Gobiidae: Sicydiinae). Raffles Bulletin of Zoology, 14 (Supplement), 209 - 214." type="journal article" year="2007">
Yamasaki 
<emphasis id="7995BB7FFFF6634AEF3CC571FE6C025D" box="[359,421,800,824]" italics="true" pageId="8" pageNumber="389">et al.</emphasis>
, 2007
</bibRefCitation>
; 
<bibRefCitation id="2F701A9CFFF6634AEC5FC54FFD31025D" author="Maeda" box="[516,760,799,824]" pageId="8" pageNumber="389" refString="Maeda, K., Mukai, T. &amp; Tachihara, K. (2012 b) A new species of amphidromous goby, Stiphodon alcedo, from the Ryukyu Archipelago (Gobiidae: Sicydiinae). Cybium, 35, 285 - 298." type="journal article" year="2012" yearSuffix="b">
Maeda 
<emphasis id="7995BB7FFFF6634AEC06C571FD53025D" box="[605,666,800,824]" italics="true" pageId="8" pageNumber="389">et al.</emphasis>
, 2012b
</bibRefCitation>
), and the geographically limited survey made to date, 
<taxonomicName id="8CE11CEEFFF6634AEBDCC570FEF90239" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF6634AEBDCC570FEF90239" italics="true" pageId="8" pageNumber="389">S. palawanensis</emphasis>
</taxonomicName>
is expected to be found elsewhere in the region in the future.
</paragraph>
<caption id="1F9E37E5FFF6634AEECCC11AFE9306F8" httpUri="https://zenodo.org/record/232905/files/figure.png" pageId="8" pageNumber="389" targetBox="[345,1240,917,1839]" targetPageId="8">
<paragraph id="4B5E676DFFF6634AEECCC11AFE9306F8" blockId="8.[151,1436,1866,1950]" pageId="8" pageNumber="389">
<emphasis id="7995BB7FFFF6634AEECCC11AFEDE063A" bold="true" box="[151,279,1866,1888]" pageId="8" pageNumber="389">FIGURE 6.</emphasis>
Location of Palawan in the Philippines (inset map) and sites where 
<taxonomicName id="8CE11CEEFFF6634AEA59C11AFBA10605" box="[1026,1128,1866,1888]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="389" phylum="Chordata" rank="genus">
<emphasis id="7995BB7FFFF6634AEA59C11AFBA10605" box="[1026,1128,1866,1888]" italics="true" pageId="8" pageNumber="389">Stiphodon</emphasis>
</taxonomicName>
species have been found on Palawan. Red circles, 
<taxonomicName id="8CE11CEEFFF6634AEFD3C13AFDF8061A" box="[392,561,1897,1919]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF6634AEFD3C13AFDF8061A" box="[392,561,1897,1919]" italics="true" pageId="8" pageNumber="389">S. palawanensis</emphasis>
</taxonomicName>
; green circles, 
<taxonomicName id="8CE11CEEFFF6634AEC86C13AFCAD061A" box="[733,868,1897,1919]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="7995BB7FFFF6634AEC86C13AFCAD061A" box="[733,868,1897,1919]" italics="true" pageId="8" pageNumber="389">S. pulchellus</emphasis>
</taxonomicName>
; blue circle, 
<taxonomicName id="8CE11CEEFFF6634AEDA1C13AFB60061A" box="[1018,1193,1898,1919]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="atropurpureus">
<emphasis id="7995BB7FFFF6634AEDA1C13AFB60061A" box="[1018,1193,1898,1919]" italics="true" pageId="8" pageNumber="389">S. atropurpureus</emphasis>
</taxonomicName>
; and orange circle, 
<taxonomicName id="8CE11CEEFFF6634AEBD2C13AFE9D06FB" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="percnopterygionus">
<emphasis id="7995BB7FFFF6634AEBD2C13AFE9D06FB" italics="true" pageId="8" pageNumber="389">S. percnopterygionus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="4B5E676DFFF7634BEE9CC6C7FE5100B8" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="390">
<emphasis id="7995BB7FFFF7634BEE9CC6C7FEE201D5" bold="true" box="[199,299,151,176]" pageId="9" pageNumber="390">Ecology.</emphasis>
The new species was one of the dominant fish species observed in the middle reaches of Balsahan Stream (Puerto Princesa City) and Barake Stream (Aborlan) in 
<date id="3F5F41ADFFF7634BED38C6EAFC1301B7" box="[867,986,185,210]" pageId="9" pageNumber="390" value="2015-05">May 2015</date>
. 
<taxonomicName id="8CE11CEEFFF7634BEDBEC6E9FB3F01B4" box="[997,1270,185,209]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BEDBEC6E9FB3F01B4" box="[997,1270,185,209]" italics="true" pageId="9" pageNumber="390">Stiphodon palawanensis</emphasis>
</taxonomicName>
inhabits pools with a substrate that is a mixture of boulders, gravel, and pebbles with exposed bedrock in some places. It also inhabits rapids. The water was clear. The fish were clinging to the rocks while feeding on algae. When disturbed, they swam to nearby rocks or hid in crevices under or between rocks. The maximum water depth of the sites was 
<quantity id="8C19CA88FFF7634BEECCC76EFF1E0033" box="[151,215,318,343]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="390" unit="m" value="1.5">1.5 m</quantity>
in May, but it could reach 3.0 m during the wet season. Normally, that occurs from June to December, while the dry season runs from January to May. In these two sites sampled, two cyprinid species 
<taxonomicName id="8CE11CEEFFF7634BEACCC730FED700FF" authority="Boulenger" authorityName="Boulenger" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palavanensis">
<emphasis id="7995BB7FFFF7634BEACCC730FA55001D" box="[1175,1436,352,376]" italics="true" pageId="9" pageNumber="390">Barbodes palavanensis</emphasis>
(Boulenger)
</taxonomicName>
and 
<taxonomicName id="8CE11CEEFFF7634BEF0CC7D1FD4400FF" authority="Boulenger" authorityName="Boulenger" box="[343,653,385,410]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="everetti">
<emphasis id="7995BB7FFFF7634BEF0CC7D1FDD800FC" box="[343,529,385,409]" italics="true" pageId="9" pageNumber="390">Rasbora everetti</emphasis>
Boulenger
</taxonomicName>
were very abundant with 
<taxonomicName id="8CE11CEEFFF7634BEDECC7D1FBA400FC" box="[951,1133,385,409]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BEDECC7D1FBA400FC" box="[951,1133,385,409]" italics="true" pageId="9" pageNumber="390">S. palawanensis</emphasis>
</taxonomicName>
. Other common species at these sites were two gobioids, 
<taxonomicName id="8CE11CEEFFF7634BEFB0C7F3FCB400DE" authority="Hoese &amp; Allen" authorityName="Hoese &amp; Allen" box="[491,893,418,443]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="illimis">
<emphasis id="7995BB7FFFF7634BEFB0C7F3FD1900DE" box="[491,720,419,443]" italics="true" pageId="9" pageNumber="390">Glossogobius illimis</emphasis>
Hoese &amp; Allen
</taxonomicName>
and 
<taxonomicName id="8CE11CEEFFF7634BEDE8C7F3FBF800DE" box="[947,1073,419,443]" class="Actinopterygii" family="Gobiidae" genus="Redigobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="genus">
<emphasis id="7995BB7FFFF7634BEDE8C7F3FBF800DE" box="[947,1073,419,443]" italics="true" pageId="9" pageNumber="390">Redigobius</emphasis>
</taxonomicName>
sp., and a halfbeak 
<taxonomicName id="8CE11CEEFFF7634BEB55C7F3FE5100B8" authority="Meisner." authorityName="Meisner." class="Actinopterygii" family="Zenarchopteridae" genus="Dermogenys" kingdom="Animalia" order="Beloniformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BEB55C7F3FEF900B9" italics="true" pageId="9" pageNumber="390">Dermogenys palawanensis</emphasis>
Meisner.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="03FB34E6FFF76348EE9CC7B5FEB106EB" lastPageId="10" lastPageNumber="391" pageId="9" pageNumber="390" type="etymology">
<paragraph id="4B5E676DFFF7634BEE9CC7B5FF16037B" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="390">
<emphasis id="7995BB7FFFF7634BEE9CC7B5FE87009B" bold="true" box="[199,334,485,510]" pageId="9" pageNumber="390">Etymology.</emphasis>
The name of the new species is derived from Palawan, the 
<typeStatus id="945AD9CFFFF7634BEA4DC7B6FB8F009B" box="[1046,1094,486,510]" pageId="9" pageNumber="390">type</typeStatus>
locality, and the Latin suffix 
<emphasis id="7995BB7FFFF7634BEECCC457FF11037B" box="[151,216,519,542]" italics="true" pageId="9" pageNumber="390">-ensis</emphasis>
.
</paragraph>
<paragraph id="4B5E676DFFF7634BEE9CC477FE0C02B5" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="390">
<emphasis id="7995BB7FFFF7634BEE9CC477FEA80325" bold="true" box="[199,353,551,576]" pageId="9" pageNumber="390">Comparison.</emphasis>
The new species was compared with congeners sharing the second dorsal- and pectoral-fin ray counts (one spine and nine segmented rays in the second dorsal fin, and usually 15 or 16 rays in the pectoral fin). 
<taxonomicName id="8CE11CEEFFF7634BEECCC43BFE6003E6" box="[151,425,619,643]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BEECCC43BFE6003E6" box="[151,425,619,643]" italics="true" pageId="9" pageNumber="390">Stiphodon palawanensis</emphasis>
</taxonomicName>
shares 9–11 dusky transverse bars laterally on the trunk and tail with 
<taxonomicName id="8CE11CEEFFF7634BEA8CC43BFEF803C1" authority="Maeda &amp; Tan" authorityName="Maeda &amp; Tan" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="maculidorsalis">
<emphasis id="7995BB7FFFF7634BEA8CC43BFA5203E6" box="[1239,1435,619,643]" italics="true" pageId="9" pageNumber="390">S. maculidorsalis</emphasis>
Maeda &amp; Tan
</taxonomicName>
and 
<taxonomicName id="8CE11CEEFFF7634BEF33C4DCFD5803C0" authority="Wu &amp; Ni" authorityName="Wu &amp; Ni" box="[360,657,652,677]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="multisquamus">
<emphasis id="7995BB7FFFF7634BEF33C4DCFDE903C1" box="[360,544,652,676]" italics="true" pageId="9" pageNumber="390">S. multisquamus</emphasis>
Wu &amp; Ni
</taxonomicName>
, but it differs from 
<taxonomicName id="8CE11CEEFFF7634BED36C4DCFBF903C1" box="[877,1072,652,676]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="maculidorsalis">
<emphasis id="7995BB7FFFF7634BED36C4DCFBF903C1" box="[877,1072,652,676]" italics="true" pageId="9" pageNumber="390">S. maculidorsalis</emphasis>
</taxonomicName>
in the lack of clear markings on the pectoral fin (vs. having fine black spots along rays), dorsal markings (black spots scattered dorsally on the head and tail of 
<taxonomicName id="8CE11CEEFFF7634BEF55C49FFE060383" box="[270,463,718,742]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="maculidorsalis">
<emphasis id="7995BB7FFFF7634BEF55C49FFE060383" box="[270,463,718,742]" italics="true" pageId="9" pageNumber="390">S. maculidorsalis</emphasis>
</taxonomicName>
, but not on 
<taxonomicName id="8CE11CEEFFF7634BEC0FC49FFCC10383" box="[596,776,718,742]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BEC0FC49FFCC10383" box="[596,776,718,742]" italics="true" pageId="9" pageNumber="390">S. palawanensis</emphasis>
</taxonomicName>
), and predorsal scalation (the posterior part of the occipital region is scaled in 
<taxonomicName id="8CE11CEEFFF7634BEF34C4A0FDED026D" box="[367,548,752,776]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BEF34C4A0FDED026D" box="[367,548,752,776]" italics="true" pageId="9" pageNumber="390">S. palawanensis</emphasis>
</taxonomicName>
, 
<figureCitation id="D3DA7BE8FFF7634BEC6FC4BFFDB0026D" box="[564,633,751,776]" captionStart="FIGURE 3" captionStartId="5.[151,250,1194,1216]" captionTargetBox="[349,1236,184,1162]" captionTargetId="figure@5.[340,1247,173,1173]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal scalation on head and nape in Stiphodon palawanensis. a, male (43.1 mm SL, CMK 11966); b, female (41.9 mm SL, CMK 11966)." httpUri="https://zenodo.org/record/232902/files/figure.png" pageId="9" pageNumber="390">Fig. 3</figureCitation>
, vs. an occipital region that is almost naked in 
<taxonomicName id="8CE11CEEFFF7634BEACDC4A0FA93026D" box="[1174,1370,752,776]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="maculidorsalis">
<emphasis id="7995BB7FFFF7634BEACDC4A0FA93026D" box="[1174,1370,752,776]" italics="true" pageId="9" pageNumber="390">S. maculidorsalis</emphasis>
</taxonomicName>
); and from 
<taxonomicName id="8CE11CEEFFF7634BEE8FC542FE42024C" box="[212,395,785,809]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="multisquamus">
<emphasis id="7995BB7FFFF7634BEE8FC542FE42024C" box="[212,395,785,809]" italics="true" pageId="9" pageNumber="390">S. multisquamus</emphasis>
</taxonomicName>
by the lack of distinct markings on the first dorsal and pectoral fins (vs. having a black blotch on the posterior part of the first dorsal fin in males, thick dusky lines along spines of the first dorsal fin in females, and fine black spots along the pectoral-fin rays in both sexes), having a line of black blotches (in males) or a black band (in females) on the distal part of the second dorsal fin (vs. no such black markings), and predorsal scalation of males (posterior part of the occipital region scaled in 
<taxonomicName id="8CE11CEEFFF7634BECAAC5C7FC6E02CB" box="[753,935,918,942]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BECAAC5C7FC6E02CB" box="[753,935,918,942]" italics="true" pageId="9" pageNumber="390">S. palawanensis</emphasis>
</taxonomicName>
, 
<figureCitation id="D3DA7BE8FFF7634BEDEEC5C6FC3002CA" box="[949,1017,918,943]" captionStart="FIGURE 3" captionStartId="5.[151,250,1194,1216]" captionTargetBox="[349,1236,184,1162]" captionTargetId="figure@5.[340,1247,173,1173]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal scalation on head and nape in Stiphodon palawanensis. a, male (43.1 mm SL, CMK 11966); b, female (41.9 mm SL, CMK 11966)." httpUri="https://zenodo.org/record/232902/files/figure.png" pageId="9" pageNumber="390">Fig. 3</figureCitation>
, vs. an occipital region that is almost naked in 
<taxonomicName id="8CE11CEEFFF7634BEEA6C5E8FE7C02B5" box="[253,437,952,976]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="multisquamus">
<emphasis id="7995BB7FFFF7634BEEA6C5E8FE7C02B5" box="[253,437,952,976]" italics="true" pageId="9" pageNumber="390">S. multisquamus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="4B5E676DFFF7634BEE9CC589FDD90512" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="390">
<taxonomicName id="8CE11CEEFFF7634BEE9CC589FD8B0297" authority="Maeda" authorityName="Maeda" box="[199,578,985,1010]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="niraikanaiensis">
<emphasis id="7995BB7FFFF7634BEE9CC589FE240294" box="[199,493,985,1009]" italics="true" pageId="9" pageNumber="390">Stiphodon niraikanaiensis</emphasis>
Maeda
</taxonomicName>
males have black longitudinal bands on the second dorsal and caudal fins, but females lack them. Although a line of black blotches on the second dorsal fin in 
<taxonomicName id="8CE11CEEFFF7634BEACEC5ABFA860576" box="[1173,1359,1019,1043]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BEACEC5ABFA860576" box="[1173,1359,1019,1043]" italics="true" pageId="9" pageNumber="390">S. palawanensis</emphasis>
</taxonomicName>
males resembles a black band when the fin is not fully open, 
<taxonomicName id="8CE11CEEFFF7634BED5FC24CFC700551" box="[772,953,1052,1076]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BED5FC24CFC700551" box="[772,953,1052,1076]" italics="true" pageId="9" pageNumber="390">S. palawanensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="8CE11CEEFFF7634BEA14C24CFAD00551" box="[1103,1305,1052,1076]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="niraikanaiensis">
<emphasis id="7995BB7FFFF7634BEA14C24CFAD00551" box="[1103,1305,1052,1076]" italics="true" pageId="9" pageNumber="390">S. niraikanaiensis</emphasis>
</taxonomicName>
also by the lack of conspicuous markings on the first dorsal and pectoral fins (vs. having black spots), and in the mode of the pectoral-fin ray count (15 vs. 16).
</paragraph>
<paragraph id="4B5E676DFFF7634BEE9CC2D0FB14074D" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="390">
<taxonomicName id="8CE11CEEFFF7634BEE9CC2D0FE1E05FD" box="[199,471,1152,1176]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BEE9CC2D0FE1E05FD" box="[199,471,1152,1176]" italics="true" pageId="9" pageNumber="390">Stiphodon palawanensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="8CE11CEEFFF7634BEC35C2D0FBDC05FD" authority="Maeda, Mukai &amp; Tachihara" authorityName="Maeda, Mukai &amp; Tachihara" box="[622,1045,1151,1176]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="alcedo">
<emphasis id="7995BB7FFFF7634BEC35C2D0FD1F05FD" box="[622,726,1152,1176]" italics="true" pageId="9" pageNumber="390">S. alcedo</emphasis>
Maeda, Mukai &amp; Tachihara
</taxonomicName>
by having a line of black blotches (in males) or a black band (in females) on the distal part of the second dorsal fin (vs. no such markings), having more premaxillary and horizontal teeth (
<figureCitation id="D3DA7BE8FFF7634BEC33C292FD6705BE" box="[616,686,1218,1243]" captionStart="FIGURE 2" captionStartId="4.[151,250,1696,1718]" captionTargetBox="[349,1239,204,1666]" captionTargetId="figure@4.[340,1247,193,1675]" captionTargetPageId="4" captionText="FIGURE 2. Number of premaxillary and horizontal dentary teeth of Stiphodon palawanensis (red circles), S. alcedo (pink circles), S. atratus (black diamonds), S. imperiorientis (light blue triangles), S. maculidorsalis (light green triangles), S. multisquamus (deep blue circles), S. ornatus (orange triangles), S. pelewensis (gray diamonds), S. pulchellus (deep green triangles), and S. weberi (purple diamonds)." httpUri="https://zenodo.org/record/232901/files/figure.png" pageId="9" pageNumber="390">Fig. 2</figureCitation>
), and the mode of the pectoral-fin ray count (15 vs. 16); from 
<taxonomicName id="8CE11CEEFFF7634BEBDCC293FEB10599" authority="Watson" authorityName="Watson" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="martenstyni">
<emphasis id="7995BB7FFFF7634BEBDCC293FED20599" italics="true" pageId="9" pageNumber="390">S. martenstyni</emphasis>
Watson
</taxonomicName>
(of which only the male 
<typeStatus id="945AD9CFFFF7634BECC1C2B4FD350598" box="[666,764,1252,1277]" pageId="9" pageNumber="390" type="holotype">holotype</typeStatus>
is known) by lack of clear markings on the first dorsal fin (vs. having two to five dusky spots along each spine) and having a line of black blotches on the second dorsal fin (vs. lack of it); from 
<taxonomicName id="8CE11CEEFFF7634BEF25C377FD8F045B" authority="Watson" authorityName="Watson" box="[382,582,1319,1342]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="atratus">
<emphasis id="7995BB7FFFF7634BEF25C377FE20045B" box="[382,489,1319,1342]" italics="true" pageId="9" pageNumber="390">S. atratus</emphasis>
Watson
</taxonomicName>
, 
<taxonomicName id="8CE11CEEFFF7634BEC0AC377FC0F045A" authority="Watson &amp; Chen" authorityName="Watson &amp; Chen" box="[593,966,1318,1343]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="imperiorientis">
<emphasis id="7995BB7FFFF7634BEC0AC377FCC2045B" box="[593,779,1319,1342]" italics="true" pageId="9" pageNumber="390">S. imperiorientis</emphasis>
Watson &amp; Chen
</taxonomicName>
, 
<taxonomicName id="8CE11CEEFFF7634BED8AC377FB66045A" authority="Meinken" authorityName="Meinken" box="[977,1199,1318,1343]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="7995BB7FFFF7634BED8AC377FB8B045B" box="[977,1090,1319,1342]" italics="true" pageId="9" pageNumber="390">S. ornatus</emphasis>
Meinken
</taxonomicName>
, 
<taxonomicName id="8CE11CEEFFF7634BEAE1C377FA51045B" authority="Herre" authorityName="Herre" box="[1210,1432,1318,1342]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="pelewensis">
<emphasis id="7995BB7FFFF7634BEAE1C377FA87045B" box="[1210,1358,1318,1342]" italics="true" pageId="9" pageNumber="390">S. pelewensis</emphasis>
Herre
</taxonomicName>
, 
<taxonomicName id="8CE11CEEFFF7634BEECCC318FEEE0405" box="[151,295,1352,1376]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="7995BB7FFFF7634BEECCC318FEEE0405" box="[151,295,1352,1376]" italics="true" pageId="9" pageNumber="390">S. pulchellus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="8CE11CEEFFF7634BEF3CC318FD300405" authority="Watson, Allen &amp; Kottelat" authorityName="Watson, Allen &amp; Kottelat" box="[359,761,1351,1376]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="weberi">
<emphasis id="7995BB7FFFF7634BEF3CC318FE190405" box="[359,464,1352,1376]" italics="true" pageId="9" pageNumber="390">S. weberi</emphasis>
Watson, Allen &amp; Kottelat
</taxonomicName>
by lack of distinct markings on the pectoral fin (vs. having many black spots along rays), having a line of black blotches (in males) or a black band (in females) on the second dorsal fin (vs. lack of it), and having more premaxillary and horizontal teeth (
<figureCitation id="D3DA7BE8FFF7634BEDACC3DAFBF504C6" box="[1015,1084,1418,1443]" captionStart="FIGURE 2" captionStartId="4.[151,250,1696,1718]" captionTargetBox="[349,1239,204,1666]" captionTargetId="figure@4.[340,1247,193,1675]" captionTargetPageId="4" captionText="FIGURE 2. Number of premaxillary and horizontal dentary teeth of Stiphodon palawanensis (red circles), S. alcedo (pink circles), S. atratus (black diamonds), S. imperiorientis (light blue triangles), S. maculidorsalis (light green triangles), S. multisquamus (deep blue circles), S. ornatus (orange triangles), S. pelewensis (gray diamonds), S. pulchellus (deep green triangles), and S. weberi (purple diamonds)." httpUri="https://zenodo.org/record/232901/files/figure.png" pageId="9" pageNumber="390">Fig. 2</figureCitation>
); and from 
<taxonomicName id="8CE11CEEFFF7634BEAE5C3DBFAB004C7" box="[1214,1401,1419,1442]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="atropurpureus">
<emphasis id="7995BB7FFFF7634BEAE5C3DBFAB004C7" box="[1214,1401,1419,1442]" italics="true" pageId="9" pageNumber="390">S. atropurpureus</emphasis>
</taxonomicName>
, 
<taxonomicName id="8CE11CEEFFF7634BEBDCC3DBFEF204A1" authority="Watson" authorityName="Watson" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="carisa">
<emphasis id="7995BB7FFFF7634BEBDCC3DBFF1404A1" italics="true" pageId="9" pageNumber="390">S. carisa</emphasis>
Watson
</taxonomicName>
, 
<taxonomicName id="8CE11CEEFFF7634BEF13C3FCFD3804A1" authority="Keith, Marquet &amp; Watson" authorityName="Keith, Marquet &amp; Watson" box="[328,753,1452,1477]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="kalfatak">
<emphasis id="7995BB7FFFF7634BEF13C3FCFE7604A1" box="[328,447,1452,1476]" italics="true" pageId="9" pageNumber="390">S. kalfatak</emphasis>
Keith, Marquet &amp; Watson
</taxonomicName>
, 
<taxonomicName id="8CE11CEEFFF7634BECA6C3FCFC0804A1" authority="Watson" authorityName="Watson" box="[765,961,1452,1476]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="larson">
<emphasis id="7995BB7FFFF7634BECA6C3FCFCAA04A1" box="[765,867,1452,1476]" italics="true" pageId="9" pageNumber="390">S. larson</emphasis>
Watson
</taxonomicName>
, and 
<taxonomicName id="8CE11CEEFFF7634BEDA5C3FCFB7504A0" authority="Weber" authorityName="Weber" box="[1022,1212,1452,1477]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="semoni">
<emphasis id="7995BB7FFFF7634BEDA5C3FCFBA204A1" box="[1022,1131,1452,1476]" italics="true" pageId="9" pageNumber="390">S. semoni</emphasis>
Weber
</taxonomicName>
by having a line of black blotches (in males) or a black band (in females) on the second dorsal fin (vs. lack of it), a pointed first dorsal fin in males (vs. rounded, except 
<taxonomicName id="8CE11CEEFFF7634BEC4CC3BFFDB30763" box="[535,634,1519,1542]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="carisa">
<emphasis id="7995BB7FFFF7634BEC4CC3BFFDB30763" box="[535,634,1519,1542]" italics="true" pageId="9" pageNumber="390">S. carisa</emphasis>
</taxonomicName>
males having a pointed fin), and the predorsal scalation of males (the posterior part of the occipital region is scaled, 
<figureCitation id="D3DA7BE8FFF7634BECC4C05FFD2B074D" box="[671,738,1551,1576]" captionStart="FIGURE 3" captionStartId="5.[151,250,1194,1216]" captionTargetBox="[349,1236,184,1162]" captionTargetId="figure@5.[340,1247,173,1173]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal scalation on head and nape in Stiphodon palawanensis. a, male (43.1 mm SL, CMK 11966); b, female (41.9 mm SL, CMK 11966)." httpUri="https://zenodo.org/record/232902/files/figure.png" pageId="9" pageNumber="390">Fig. 3</figureCitation>
, vs. an occipital region that is almost naked).
</paragraph>
<paragraph id="4B5E676DFFF7634BEE9CC061FB4107E9" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="390">
Briefly, the new species is most easily distinguished by its unique fin markings: any 
<taxonomicName id="8CE11CEEFFF7634BEAD0C061FB35072C" box="[1163,1276,1585,1609]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="genus">
<emphasis id="7995BB7FFFF7634BEAD0C061FB35072C" box="[1163,1276,1585,1609]" italics="true" pageId="9" pageNumber="390">Stiphodon</emphasis>
</taxonomicName>
with a line of black blotches (in males) or a black band (in females) on the distal part of the second dorsal fin, and without distinct markings on the first dorsal and pectoral fins can be identified as 
<taxonomicName id="8CE11CEEFFF7634BED97C024FB4907E9" box="[972,1152,1652,1676]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BED97C024FB4907E9" box="[972,1152,1652,1676]" italics="true" pageId="9" pageNumber="390">S. palawanensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4B5E676DFFF7634BEE9CC0C5FC5706F2" blockId="9.[151,1437,151,1943]" pageId="9" pageNumber="390">
<emphasis id="7995BB7FFFF7634BEE9CC0C5FD0207CB" bold="true" box="[199,715,1684,1710]" pageId="9" pageNumber="390">
Other 
<taxonomicName id="8CE11CEEFFF7634BEF4CC0C4FE4207C8" box="[279,395,1684,1709]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="genus">
<emphasis id="7995BB7FFFF7634BEF4CC0C4FE4207C8" bold="true" box="[279,395,1684,1709]" italics="true" pageId="9" pageNumber="390">Stiphodon</emphasis>
</taxonomicName>
species found in Palawan.
</emphasis>
During our two-day exploration in the 
<typeStatus id="945AD9CFFFF7634BEAD5C0C6FB7707CB" box="[1166,1214,1686,1710]" pageId="9" pageNumber="390">type</typeStatus>
locality of the new species (Balsahan Stream in the Iwahig Prison and Penal Farm, Sulu Sea side of Puerto Princesa City) in 
<date id="3F5F41ADFFF7634BEB32C0E7FF1B0795" pageId="9" pageNumber="390" value="2015-05">May 2015</date>
, two other 
<taxonomicName id="8CE11CEEFFF7634BEF38C088FE1D0795" box="[355,468,1752,1776]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="genus">
<emphasis id="7995BB7FFFF7634BEF38C088FE1D0795" box="[355,468,1752,1776]" italics="true" pageId="9" pageNumber="390">Stiphodon</emphasis>
</taxonomicName>
species were found; three female individuals of 
<taxonomicName id="8CE11CEEFFF7634BEA72C088FB250795" box="[1065,1260,1752,1776]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="atropurpureus">
<emphasis id="7995BB7FFFF7634BEA72C088FB250795" box="[1065,1260,1752,1776]" italics="true" pageId="9" pageNumber="390">S. atropurpureus</emphasis>
</taxonomicName>
and one male individual of 
<taxonomicName id="8CE11CEEFFF7634BEF69C0AAFD280677" authority="Watson &amp; Chen" authorityName="Watson &amp; Chen" box="[306,737,1785,1810]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="percnopterygionus">
<emphasis id="7995BB7FFFF7634BEF69C0AAFDE80674" box="[306,545,1786,1809]" italics="true" pageId="9" pageNumber="390">S. percnopterygionus</emphasis>
Watson &amp; Chen
</taxonomicName>
(
<figureCitation id="D3DA7BE8FFF7634BECA8C0A9FCF00677" box="[755,825,1785,1810]" captionStart="FIGURE 6" captionStartId="8.[151,250,1866,1888]" captionTargetBox="[345,1240,917,1839]" captionTargetId="figure@8.[340,1248,913,1845]" captionTargetPageId="8" captionText="FIGURE 6. Location of Palawan in the Philippines (inset map) and sites where Stiphodon species have been found on Palawan. Red circles, S. palawanensis; green circles, S. pulchellus; blue circle, S. atropurpureus; and orange circle, S. percnopterygionus." httpUri="https://zenodo.org/record/232905/files/figure.png" pageId="9" pageNumber="390">Fig. 6</figureCitation>
). Two of the three 
<taxonomicName id="8CE11CEEFFF7634BEA42C0AAFB1E0674" box="[1049,1239,1786,1809]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="atropurpureus">
<emphasis id="7995BB7FFFF7634BEA42C0AAFB1E0674" box="[1049,1239,1786,1809]" italics="true" pageId="9" pageNumber="390">S. atropurpureus</emphasis>
</taxonomicName>
individuals were collected (
<figureCitation id="D3DA7BE8FFF7634BEF50C14AFE840656" box="[267,333,1818,1843]" captionStart="FIGURE 7" captionStartId="10.[151,250,1241,1263]" captionTargetBox="[161,1420,201,1188]" captionTargetId="figure@10.[151,1436,193,1220]" captionTargetPageId="10" captionText="FIGURE 7. Stiphodon pulchellus (a, WPU-PPC-P 29, male, 45.9 mm SL; b, URM-P 48681, female, 39.6 mm SL) and Stiphodon atropurpureus (c, WPU-PPC-P 10, female, 43.7 mm SL) specimens collected from Palawan immediately after fixation." httpUri="https://zenodo.org/record/232906/files/figure.png" pageId="9" pageNumber="390">Fig. 7</figureCitation>
c) and listed as comparative material in this paper. 
<taxonomicName id="8CE11CEEFFF7634BEDDFC14BFB050657" box="[900,1228,1819,1843]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="percnopterygionus">
<emphasis id="7995BB7FFFF7634BEDDFC14BFB050657" box="[900,1228,1819,1843]" italics="true" pageId="9" pageNumber="390">Stiphodon percnopterygionus</emphasis>
</taxonomicName>
was not collected, however a wild individual was identified on site by snorkel based observation of characteristic markings of the body and fins (
<figureCitation id="D3DA7BE8FFF7634BEF1EC10DFE430613" box="[325,394,1885,1910]" captionStart="FIGURE 8" captionStartId="11.[151,250,1139,1161]" captionTargetBox="[151,1436,179,1117]" captionTargetId="figure@11.[151,1436,153,1117]" captionTargetPageId="11" captionText="FIGURE 8. Stiphodon percnopterygionus (a) with Stiphodon palawanensis observed in Balsahan Stream, Puerto Princesa City, Palawan, on 18 May 2015." httpUri="https://zenodo.org/record/232907/files/figure.png" pageId="9" pageNumber="390">Fig. 8</figureCitation>
) and its unique, high triangular first dorsal fin. While 
<taxonomicName id="8CE11CEEFFF7634BEDA4C10EFB7F0610" box="[1023,1206,1885,1909]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF7634BEDA4C10EFB7F0610" box="[1023,1206,1885,1909]" italics="true" pageId="9" pageNumber="390">S. palawanensis</emphasis>
</taxonomicName>
was very abundant, both 
<taxonomicName id="8CE11CEEFFF7634BEE94C12FFE4206F3" box="[207,395,1919,1942]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="atropurpureus">
<emphasis id="7995BB7FFFF7634BEE94C12FFE4206F3" box="[207,395,1919,1942]" italics="true" pageId="9" pageNumber="390">S. atropurpureus</emphasis>
</taxonomicName>
and 
<taxonomicName id="8CE11CEEFFF7634BEF99C12FFD6606F3" box="[450,687,1919,1942]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="percnopterygionus">
<emphasis id="7995BB7FFFF7634BEF99C12FFD6606F3" box="[450,687,1919,1942]" italics="true" pageId="9" pageNumber="390">S. percnopterygionus</emphasis>
</taxonomicName>
were rare at this site.
</paragraph>
<caption id="1F9E37E5FFF46348EECCC289FF240449" httpUri="https://zenodo.org/record/232906/files/figure.png" pageId="10" pageNumber="391" targetBox="[161,1420,201,1188]" targetPageId="10">
<paragraph id="4B5E676DFFF46348EECCC289FF240449" blockId="10.[151,1435,1241,1324]" pageId="10" pageNumber="391">
<emphasis id="7995BB7FFFF46348EECCC289FEDE058A" bold="true" box="[151,279,1241,1263]" pageId="10" pageNumber="391">FIGURE 7.</emphasis>
<taxonomicName id="8CE11CEEFFF46348EF79C289FE33058A" box="[290,506,1241,1263]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="7995BB7FFFF46348EF79C289FE33058A" box="[290,506,1241,1263]" italics="true" pageId="10" pageNumber="391">Stiphodon pulchellus</emphasis>
</taxonomicName>
(a, WPU-PPC-P 29, male, 45.9 mm SL; b, URM-P 48681, female, 39.6 mm SL) and 
<taxonomicName id="8CE11CEEFFF46348EECCC2A8FE51046B" box="[151,408,1272,1294]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="atropurpureus">
<emphasis id="7995BB7FFFF46348EECCC2A8FE51046B" box="[151,408,1272,1294]" italics="true" pageId="10" pageNumber="391">Stiphodon atropurpureus</emphasis>
</taxonomicName>
(c, WPU-PPC-P 10, female, 43.7 mm SL) specimens collected from Palawan immediately after fixation.
</paragraph>
</caption>
<paragraph id="4B5E676DFFF46348EE9CC30AFED80767" blockId="10.[151,1437,1369,1934]" pageId="10" pageNumber="391">
Four 
<taxonomicName id="8CE11CEEFFF46348EF59C309FEBA0414" box="[258,371,1369,1393]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="7995BB7FFFF46348EF59C309FEBA0414" box="[258,371,1369,1393]" italics="true" pageId="10" pageNumber="391">Stiphodon</emphasis>
</taxonomicName>
specimens collected in the Iwahig River at Puerto Princesa City in 1988 have been deposited in the National Museum of Nature and Science (Ibaraki, 
<collectingCountry id="33F627FDFFF46348ED4BC32EFC9C04F3" box="[784,853,1406,1430]" name="Japan" pageId="10" pageNumber="391">Japan</collectingCountry>
). The Iwahig is a relatively large river next to the Balsahan Stream. The specimens are composed of three 
<taxonomicName id="8CE11CEEFFF46348ED49C3F2FC0E04DC" box="[786,967,1441,1465]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF46348ED49C3F2FC0E04DC" box="[786,967,1441,1465]" italics="true" pageId="10" pageNumber="391">S. palawanensis</emphasis>
</taxonomicName>
(NSMT-P 45091, 45092, and 45094) and one 
<taxonomicName id="8CE11CEEFFF46348EE9CC396FE9104B8" box="[199,344,1477,1501]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="7995BB7FFFF46348EE9CC396FE9104B8" box="[199,344,1477,1501]" italics="true" pageId="10" pageNumber="391">S. pulchellus</emphasis>
</taxonomicName>
(NSMT-P 45093), but the latter species was not found in our exploration in Balsahan Stream in 
<date id="3F5F41ADFFF46348EECCC3BAFEC40767" box="[151,269,1513,1538]" pageId="10" pageNumber="391" value="2015-05">May 2015</date>
.
</paragraph>
<paragraph id="4B5E676DFFF46348EE9CC05EFCB0070B" blockId="10.[151,1437,1369,1934]" pageId="10" pageNumber="391">
In Barake Stream (Sulu Sea side of Aborlan), the only 
<taxonomicName id="8CE11CEEFFF46348ED72C05DFC530740" box="[809,922,1549,1573]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="7995BB7FFFF46348ED72C05DFC530740" box="[809,922,1549,1573]" italics="true" pageId="10" pageNumber="391">Stiphodon</emphasis>
</taxonomicName>
species found during our one-day exploration was 
<taxonomicName id="8CE11CEEFFF46348EE97C062FE4B072C" box="[204,386,1585,1609]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF46348EE97C062FE4B072C" box="[204,386,1585,1609]" italics="true" pageId="10" pageNumber="391">S. palawanensis</emphasis>
</taxonomicName>
. All collections from Aborlan and Narra, examined in the present study (CMK and URM), also comprised entirely of individuals of 
<taxonomicName id="8CE11CEEFFF46348EC38C006FCD10708" box="[611,792,1621,1645]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF46348EC38C006FCD10708" box="[611,792,1621,1645]" italics="true" pageId="10" pageNumber="391">S. palawanensis</emphasis>
</taxonomicName>
(
<figureCitation id="D3DA7BE8FFF46348ED7DC005FCA5070B" box="[806,876,1621,1646]" captionStart="FIGURE 6" captionStartId="8.[151,250,1866,1888]" captionTargetBox="[345,1240,917,1839]" captionTargetId="figure@8.[340,1248,913,1845]" captionTargetPageId="8" captionText="FIGURE 6. Location of Palawan in the Philippines (inset map) and sites where Stiphodon species have been found on Palawan. Red circles, S. palawanensis; green circles, S. pulchellus; blue circle, S. atropurpureus; and orange circle, S. percnopterygionus." httpUri="https://zenodo.org/record/232905/files/figure.png" pageId="10" pageNumber="391">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="4B5E676DFFF46348EE9CC02AFF1F07BF" blockId="10.[151,1437,1369,1934]" pageId="10" pageNumber="391">
We also explored another river bearing the name Iwahig at Quezon, on the South 
<collectingCountry id="33F627FDFFF46348EA3AC029FB6D07F7" box="[1121,1188,1657,1682]" name="China" pageId="10" pageNumber="391">China</collectingCountry>
Sea side of the island (
<figureCitation id="D3DA7BE8FFF46348EEC4C0CDFF2C07D3" box="[159,229,1693,1718]" captionStart="FIGURE 6" captionStartId="8.[151,250,1866,1888]" captionTargetBox="[345,1240,917,1839]" captionTargetId="figure@8.[340,1248,913,1845]" captionTargetPageId="8" captionText="FIGURE 6. Location of Palawan in the Philippines (inset map) and sites where Stiphodon species have been found on Palawan. Red circles, S. palawanensis; green circles, S. pulchellus; blue circle, S. atropurpureus; and orange circle, S. percnopterygionus." httpUri="https://zenodo.org/record/232905/files/figure.png" pageId="10" pageNumber="391">Fig. 6</figureCitation>
) in 
<date id="3F5F41ADFFF46348EF43C0CEFE5807D3" box="[280,401,1693,1718]" pageId="10" pageNumber="391" value="2015-05">May 2015</date>
. 
<taxonomicName id="8CE11CEEFFF46348EFC5C0CDFD4507D0" box="[414,652,1693,1717]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="7995BB7FFFF46348EFC5C0CDFD4507D0" box="[414,652,1693,1717]" italics="true" pageId="10" pageNumber="391">Stiphodon pulchellus</emphasis>
</taxonomicName>
(
<figureCitation id="D3DA7BE8FFF46348ECC4C0CDFD2C07D0" box="[671,741,1693,1718]" captionStart="FIGURE 7" captionStartId="10.[151,250,1241,1263]" captionTargetBox="[161,1420,201,1188]" captionTargetId="figure@10.[151,1436,193,1220]" captionTargetPageId="10" captionText="FIGURE 7. Stiphodon pulchellus (a, WPU-PPC-P 29, male, 45.9 mm SL; b, URM-P 48681, female, 39.6 mm SL) and Stiphodon atropurpureus (c, WPU-PPC-P 10, female, 43.7 mm SL) specimens collected from Palawan immediately after fixation." httpUri="https://zenodo.org/record/232906/files/figure.png" pageId="10" pageNumber="391">Fig. 7</figureCitation>
a, b) was abundant, but 
<taxonomicName id="8CE11CEEFFF46348EDA5C0CEFB7C07D0" box="[1022,1205,1693,1717]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF46348EDA5C0CEFB7C07D0" box="[1022,1205,1693,1717]" italics="true" pageId="10" pageNumber="391">S. palawanensis</emphasis>
</taxonomicName>
could not be found there.
</paragraph>
<paragraph id="4B5E676DFFF46348EE9CC0B5FEB106EB" blockId="10.[151,1437,1369,1934]" pageId="10" pageNumber="391">
Thus, the species composition of any 
<taxonomicName id="8CE11CEEFFF46348EC26C0B5FD270798" box="[637,750,1765,1789]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="7995BB7FFFF46348EC26C0B5FD270798" box="[637,750,1765,1789]" italics="true" pageId="10" pageNumber="391">Stiphodon</emphasis>
</taxonomicName>
assemblage seems to depend on the location on the island. 
<taxonomicName id="8CE11CEEFFF46348EECCC159FE630644" box="[151,426,1801,1825]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="palawanensis">
<emphasis id="7995BB7FFFF46348EECCC159FE630644" box="[151,426,1801,1825]" italics="true" pageId="10" pageNumber="391">Stiphodon palawanensis</emphasis>
</taxonomicName>
may be the dominant species in some areas, whereas 
<taxonomicName id="8CE11CEEFFF46348EA40C15AFB650644" box="[1051,1196,1801,1825]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="7995BB7FFFF46348EA40C15AFB650644" box="[1051,1196,1801,1825]" italics="true" pageId="10" pageNumber="391">S. pulchellus</emphasis>
</taxonomicName>
may be dominant in others. However, because only a small survey effort has been applied at each site and only a small part of the island has been explored so far, comprehensive surveys will be required to understand the distribution of 
<taxonomicName id="8CE11CEEFFF46348EB70C101FA55060C" box="[1323,1436,1873,1897]" class="Actinopterygii" family="Gobiidae" genus="Stiphodon" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="391" phylum="Chordata" rank="genus">
<emphasis id="7995BB7FFFF46348EB70C101FA55060C" box="[1323,1436,1873,1897]" italics="true" pageId="10" pageNumber="391">Stiphodon</emphasis>
</taxonomicName>
species on Palawan.
</paragraph>
</subSubSection>
</treatment>
</document>