treatments-xml/data/C3/41/8A/C3418A72FF9F7E46FF73FF4FFA3DFD5F.xml

<document id="104C5F274BC8B6E5A807E20B5F94C341" ID-DOI="10.11646/phytotaxa.616.2.6" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695981262274" checkinUser="plazi" docAuthor="Jara-Muñoz, O. A., Moonlight, P. W. &amp; Zabala-Rivera, J. C." docDate="2023" docId="C3418A72FF9F7E46FF73FF4FFA3DFD5F" docLanguage="en" docName="phytotaxa.616.2.6.pdf" docOrigin="Phytotaxa 616 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.616.2.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Begonia betancurii Jara &amp; Moonlight. A. Habit 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="173" masterDocId="3F78F20AFF9E7E42FFFBFFBBFF9DFFEF" masterDocTitle="A New Species of Begonia (Begoniaceae) from the Eastern Cordillera of Colombia" masterLastPageNumber="175" masterPageNumber="169" pageNumber="170" updateTime="1695981330486" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="19E7B29E5327486B4A8170E0846A0129">A New Species of Begonia (Begoniaceae) from the Eastern Cordillera of Colombia</mods:title>
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<mods:namePart id="F994F632DC2F8069981F3A32AC8D240B">Jara-Muñoz, O. A.</mods:namePart>
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<mods:affiliation id="81D7CEDEFA5F395E910FD664402BF174">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Carrera 45 N ° 26 - 85, Bogotá, D. C., Colombia Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland &amp; oajaram @ unal. edu. co; https: // orcid. org / 0000 - 0002 - 7123 - 124 X</mods:affiliation>
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<mods:namePart id="B714CECDC173BF0EEDD0718BC567E513">Moonlight, P. W.</mods:namePart>
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<mods:affiliation id="A4CC69BDCB1CAF45FB9E972F6408CF82">moonligp @ tcd. ie; https: // orcid. org / 0000 - 0003 - 4342 - 2089</mods:affiliation>
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<mods:namePart id="F70FB10592A2F3F173FA3F388964DE5E">Zabala-Rivera, J. C.</mods:namePart>
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<mods:affiliation id="E738DDFB22A29CC5F810135488ED56D8">Grupo Interdisciplinario en Ciencias GICI-DIN, Universidad Pedagógica y Tecnológica de Colombia &amp; juan. zabala @ uptc. edu. com; http: // orcid. org / 0000 - 0003 - 2395 - 4205</mods:affiliation>
<mods:nameIdentifier id="FD2388D5D95A400F2417AE0D2FE4B5C0" type="email">juan.zabala@uptc.edu.com</mods:nameIdentifier>
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<mods:date id="362593E4F9D5A5C4F1CDF73FD894AD52">2023</mods:date>
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<taxonomicName id="8CE840E7FF9F7E43FF73FF4FFDDAFEE1" authority="Jara &amp; Moonlight" authorityName="Jara &amp; Moonlight. A. Habit" authorityYear="2023" box="[136,583,244,271]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="betancurii" status="sp. nov.">
<emphasis id="799CE776FF9F7E43FF73FF4FFEFBFEE0" bold="true" box="[136,358,244,271]" italics="true" pageId="1" pageNumber="170">Begonia betancurii</emphasis>
Jara &amp; Moonlight
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,
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<taxonomicNameLabel id="62AF5A0DFF9F7E43FDA8FF4EFD35FEE1" box="[595,680,245,270]" pageId="1" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D3D327E1FF9F7E43FD4DFF4FFCA0FEE1" box="[694,829,244,270]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[136,229,1992,2014]" captionStartId-1="4.[136,229,1648,1670]" captionTargetBox-0="[205,1383,187,1965]" captionTargetBox-1="[151,1436,713,1623]" captionTargetId-0="figure-24@2.[205,1383,187,1965]" captionTargetId-1="figure-290@4.[151,1436,713,1623]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1. Begonia betancurii Jara &amp; Moonlight. A. Habit; B. Staminate flower (front view); C. Pistillate flowers (front view). D. Pistillate flowers (side view), showing ovary and bract. E. Mature fruit (side view). F. Immature Fruit in cross-section showing bifid placentae. (photos A to E, by J.C. Zabala; F by M. Fernandez)." captionText-1="FIGURE 2.Distribution of Begonia betancurii Jara &amp;Moonlight inColombia,showing borders of Boyacá and Cundinamarca departments, and red dots representing records of presence of the species." pageId="1" pageNumber="170">Figures 1–2</figureCitation>
)
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</paragraph>
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<emphasis id="799CE776FF9F7E43FF73FEFBFF27FEBA" bold="true" box="[136,186,320,341]" pageId="1" pageNumber="170">Type</emphasis>
:— 
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<collectingCountry id="33FF7BF4FF9F7E43FF2CFEFBFEC0FEBA" box="[215,349,320,341]" name="Colombia" pageId="1" pageNumber="170">COLOMBIA</collectingCountry>
. 
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, 
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, via 
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, entrada a mano izquierda antes del tunel 
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, vereda 
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, finca 
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, cuenca de la quebrada 
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<location id="4E376DBFFF9F7E43FCD5FEDFFC28FE95" LSID="urn:lsid:plazi:treatment:C3418A72FF9F7E46FF73FF4FFA3DFD5F:4E376DBFFF9F7E43FCD5FEDFFC28FE95" box="[814,949,356,378]" country="Colombia" county="Municipio de Santa Maria" latitude="4.9066668" longLatPrecision="130" longitude="-73.295" municipality="Santa Maria-Las Juntas" name="La Esmeralda" pageId="1" pageNumber="170" stateProvince="Boyaca">La Esmeralda</location>
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, 
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, 
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, 
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, 
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<collectingDate id="2F12E44CFF9F7E43FADCFEDFFF72FE72" pageId="1" pageNumber="170" value="2009-09-25">25 September 2009</collectingDate>
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, 
<collectionCode id="2DF9A3A1FF9F7E43FF07FE33FE97FE72" box="[252,266,392,413]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="1" pageNumber="170" type="Herbarium">
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<emphasis id="799CE776FF9F7E43FEF1FE33FE3AFE72" box="[266,423,392,413]" italics="true" pageId="1" pageNumber="170">
. Betancur, 
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. 
<collectionCode id="2DF9A3A1FF9F7E43FE68FE33FE3EFE72" box="[403,419,392,413]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="1" pageNumber="170" type="Herbarium">F</collectionCode>
.
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<materialsCitation id="FB803139FF9F7E43FE54FE33FE44FE2D" collectingDate="2009-02" collectionCode="COL, HUA" collectorName="Gonzalez &amp; de Angiospermas" country="Colombia" location="Taxonomia de Angiospermas" pageId="1" pageNumber="170" specimenCode="COL000452537, HUA216209" specimenCount="1" stateProvince="Boyaca" typeStatus="holotype">
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<collectorName id="E61D5EB2FF9F7E43FE54FE33FD95FE72" box="[431,520,392,413]" pageId="1" pageNumber="170">Gonzalez</collectorName>
&amp; estudiantes 
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Taxonomía 
<collectorName id="E61D5EB2FF9F7E43FCEFFE33FC20FE72" box="[788,957,392,413]" pageId="1" pageNumber="170">de Angiospermas</collectorName>
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</date>
14304
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(
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[
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]; isotype 
<collectionCode id="2DF9A3A1FF9F7E43FEF2FE17FEA1FE2E" box="[265,316,428,449]" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15677" lsid="urn:lsid:biocol.org:col:15677" name="Universidad de Antioquia" pageId="1" pageNumber="170" type="Herbarium">HUA</collectionCode>
[
<specimenCode id="1B4E931FFF9F7E43FEBCFE17FE51FE2D" box="[327,460,428,450]" collectionCode="HUA" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15677" lsid="urn:lsid:biocol.org:col:15677" name="Universidad de Antioquia" pageId="1" pageNumber="170" type="Herbarium">HUA216209</specimenCode>
])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="03F268EFFF9F7E43FF73FE4BFEF9F93D" pageId="1" pageNumber="170" type="diagnosis">
<paragraph id="4B573B64FF9F7E43FF73FE4BFC41FD75" blockId="1.[136,1452,496,1999]" pageId="1" pageNumber="170">
<emphasis id="799CE776FF9F7E43FF73FE4BFF66FDE5" bold="true" box="[136,251,496,522]" pageId="1" pageNumber="170">Diagnosis</emphasis>
:— 
<taxonomicName id="8CE840E7FF9F7E43FEE4FE4AFD42FDE5" authority="Jara &amp; Moonlight" authorityName="Jara &amp; Moonlight. A. Habit" authorityYear="2023" box="[287,735,496,523]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="betancurii">
<emphasis id="799CE776FF9F7E43FEE4FE4AFE60FDE5" box="[287,509,496,522]" italics="true" pageId="1" pageNumber="170">Begonia betancurii</emphasis>
Jara &amp; Moonlight
</taxonomicName>
can be differentiated from the other species in the section 
<taxonomicName id="8CE840E7FF9F7E43FF73FDAEFEBFFDC1" box="[136,290,533,558]" pageId="1" pageNumber="170" rank="section" section="Ruizopavonia">
<emphasis id="799CE776FF9F7E43FF73FDAEFEBFFDC1" box="[136,290,533,558]" italics="true" pageId="1" pageNumber="170">Ruizopavonia</emphasis>
</taxonomicName>
bearing staminate flowers with 4 and pistillate flowers with 3 tepals, namely: 
<taxonomicName id="8CE840E7FF9F7E43FB45FDAEFE0CFDBD" authority="L. B. Sm. &amp; B. G. Schub." authorityName="L. B. Sm. &amp; B. G. Schub." class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="cymbalifera">
<emphasis id="799CE776FF9F7E43FB45FDAEFA31FDC1" box="[1214,1452,532,558]" italics="true" pageId="1" pageNumber="170">Begonia cymbalifera</emphasis>
L.B.Sm. &amp; B.G.Schub.
</taxonomicName>
, and 
<taxonomicName id="8CE840E7FF9F7E43FE35FD82FC92FDBD" authority="Irmsch." authorityName="Irmsch." box="[462,783,568,594]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="consobrina">
<emphasis id="799CE776FF9F7E43FE35FD82FD2CFDBD" box="[462,689,568,594]" italics="true" pageId="1" pageNumber="170">Begonia consobrina</emphasis>
Irmsch.
</taxonomicName>
, by its larger outer staminate (9.5–17.4 × 
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vs. ca. 5 × 
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) and pistillate (13.9–18.7 × 13.8–20.1 vs. 
<quantity id="8C109681FF9F7E43FCF7FDE7FCF1FD99" box="[780,876,604,631]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="1" pageNumber="170" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
) tepals. In addition, from 
<taxonomicName id="8CE840E7FF9F7E43FB62FDE6FADDFD99" box="[1177,1344,604,630]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="cymbalifera">
<emphasis id="799CE776FF9F7E43FB62FDE6FADDFD99" box="[1177,1344,604,630]" italics="true" pageId="1" pageNumber="170">B. cymbalifera</emphasis>
</taxonomicName>
it can be distinguished by its palmate-pinnate venation pattern (vs. pinnate venation).
</paragraph>
<paragraph id="4B573B64FF9F7E43FF46FD1FFDF6F965" blockId="1.[136,1452,496,1999]" pageId="1" pageNumber="170">
Caulescent terrestrial herb. Stem erect, to 
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tall, frequently branching, internodes 
<quantity id="8C109681FF9F7E43FB78FD1FFB45FD50" box="[1155,1240,676,703]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="1" pageNumber="170" unit="cm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 cm</quantity>
long, 
<quantity id="8C109681FF9F7E43FAD8FD1FFA31FD50" box="[1315,1452,676,703]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" metricValueMax="3.5" metricValueMin="2.7" pageId="1" pageNumber="170" unit="mm" value="3.1" valueMax="3.5" valueMin="2.7">2.7–3.5 mm</quantity>
in diameter, glabrous, smooth, greenish to reddish-green, browning with age. Stipules early deciduous, lanceolate, ca. 10 × 
<quantity id="8C109681FF9F7E43FF15FD57FEDAFCE8" box="[238,327,748,775]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="170" unit="mm" value="3.5">3.5 mm</quantity>
, apex acute, margin entire, glabrous. Leaves alternate, basifixed; petioles joining blade at an angle of ca. 90°–120°, 
<quantity id="8C109681FF9F7E43FEB1FCABFE59FCC4" box="[330,452,784,811]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.7" metricValueMin="1.2" pageId="1" pageNumber="170" unit="mm" value="24.5" valueMax="37.0" valueMin="12.0">12–37 mm</quantity>
long, glabrous; blade asymmetrical, elliptic, oblong to obovate, 9.4–18.7 × 
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, apex acuminate, base lobate on the broad side of the lamina, obtuse to rounded on the narrow side, margin serrate, upper surface green, glabrous, lower surface glabrous, green with reddish-green veins, palmate-pinnate-nerved, 11- to 14-veined on the broad side of the lamina, 9- to 11-veined on the narrow side of the lamina. Inflorescences 1–2 per branch, axillary but the youngest arising close to the apex of the stem and appearing terminal, 2–4 times dichotomous ending in cymes, with central staminate flowers developing first fallowed by lateral pistilate ones; peduncle 
<quantity id="8C109681FF9F7E43FACFFC7FFA31FC30" box="[1332,1452,964,991]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" metricValueMax="7.7" metricValueMin="3.4" pageId="1" pageNumber="170" unit="mm" value="55.5" valueMax="77.0" valueMin="34.0">34–77 mm</quantity>
long, internodes elongate, 
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long, glabrous, pale green–flushed red. Staminate flowers: pedicels 
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long, glabrous, white to pale green; tepals 4, outer two mostly white and pinkish-white on veins and margins, ovate, 9.5–17.4 × 
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, base cordate, apex rounded, margin entire, glabrous; inner two white, narrowly oblanceolate, 3.1–4.2 × 
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, apex rounded, margin entire, glabrous; stamens ca. 55, free, filaments 1.4–2.0 mm long, anthers symmetrically basifixed, base rounded or sagittate, oblong, 
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long, dehiscing via lateral slits, connectives extended to 
<quantity id="8C109681FF9F7E43FEE3FB27FEE9FB58" box="[280,372,1180,1207]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="170" unit="mm" value="0.9">0.9 mm</quantity>
. Pistillate flowers: bracteoles 2, deciduous, reddish-green, broadly ovate, margin entire, apex denticulate, 13–18.2 × ca. 
<quantity id="8C109681FF9F7E43FE40FB7BFD9CFB34" box="[443,513,1216,1243]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="170" unit="mm" value="6.0">6 mm</quantity>
; pedicel 
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long, glabrous, pale green to reddish-green; locular region of the ovary slightly obovoid to ellipsoid, 8.1–16.2 × 
<quantity id="8C109681FF9F7E43FD31FB5FFCD0FB10" box="[714,845,1252,1279]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="10.0" metricValueMin="3.7" pageId="1" pageNumber="170" unit="mm" value="6.85" valueMax="10.0" valueMin="3.7">3.7–10 mm</quantity>
, glabrous, 3-winged (rarely 4), wings white or green, unequal, largest wing asymmetric, proximal side concave, describing an arc, distal side nearly straight, forming an angle 40–60º to the perpendicular of the body apex, margin entire, 
<quantity id="8C109681FF9F7E43FC69FA97FB93FAA8" box="[914,1038,1324,1351]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="1" pageNumber="170" unit="mm" value="26.5" valueMax="27.0" valueMin="26.0">26–27 mm</quantity>
long, 
<quantity id="8C109681FF9F7E43FBA2FA97FB48FAA8" box="[1113,1237,1324,1351]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="1" pageNumber="170" unit="mm" value="14.5" valueMax="16.0" valueMin="13.0">13–16 mm</quantity>
wide, smaller two wings semicircular, margins entire, apex rounded, 
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tall, 
<quantity id="8C109681FF9F7E43FC9EFAEBFC5CFA84" box="[869,961,1360,1387]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="1" pageNumber="170" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
wide; 3-locular (rarely 4-locular), placentae axilary, bifid, bearing ovules on both surfaces; tepals 3, outer two spreading to reflexed, pinkish- white, glabrous, suborbicular, 13.9–18.7 × 
<quantity id="8C109681FF9F7E43FE7FFA23FDBAFA5C" box="[388,551,1432,1459]" metricMagnitude="-2" metricUnit="m" metricValue="1.6950000000000003" metricValueMax="2.0100000000000002" metricValueMin="1.3800000000000001" pageId="1" pageNumber="170" unit="mm" value="16.95" valueMax="20.1" valueMin="13.8">13.8–20.1 mm</quantity>
, base slightly cordate, apex rounded, margin entire; inner one white, oblanceolate or obovate, glabrous, ca. 5.5 × 
<quantity id="8C109681FF9F7E43FE17FA07FDDBFA38" box="[492,582,1468,1495]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="170" unit="mm" value="1.5">1.5 mm</quantity>
; styles 3, multifid, divided near to the base into two or three main branches, each branch ca. 
<quantity id="8C109681FF9F7E43FEC4FA5BFE08FA14" box="[319,405,1504,1531]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="170" unit="mm" value="2.9">2.9 mm</quantity>
long, then divided into two or three twisted papillose bands (stigmatic surface), 
<quantity id="8C109681FF9F7E43FADFFA5BFA31FA14" box="[1316,1452,1504,1531]" metricMagnitude="-3" metricUnit="m" metricValue="2.9499999999999997" metricValueMax="4.1" metricValueMin="1.8" pageId="1" pageNumber="170" unit="mm" value="2.9499999999999997" valueMax="4.1" valueMin="1.8">1.8–4.1 mm</quantity>
long. Fruiting pedicel to 
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long. Fruit pendant, body as in the ovary, dehiscent near the back of the locules, expanding to 16.3 × 
<quantity id="8C109681FF9F7E43FE8EF993FE52F9AC" box="[373,463,1576,1603]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="1" pageNumber="170" unit="mm" value="8.3">8.3 mm</quantity>
, light brown, glabrous, the largest wing expanding to 32 × 
<quantity id="8C109681FF9F7E43FB8EF993FB5AF9AC" box="[1141,1223,1576,1603]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="170" unit="mm" value="11.0">11 mm</quantity>
, similar to the wing in the ovary, green, pink or brown, longer wing expanding to 
<quantity id="8C109681FF9F7E43FCB0F9F7FC2CF988" box="[843,945,1612,1639]" metricMagnitude="-2" metricUnit="m" metricValue="2.51" pageId="1" pageNumber="170" unit="mm" value="25.1">25.1 mm</quantity>
long, and 
<quantity id="8C109681FF9F7E43FBD0F9F7FB0CF988" box="[1067,1169,1612,1639]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="1" pageNumber="170" unit="mm" value="14.3">14.3 mm</quantity>
wide, shorter two wings expanding to 
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long, and 
<quantity id="8C109681FF9F7E43FE1EF9CBFDBBF964" box="[485,550,1648,1675]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="170" unit="mm" value="8.0">8 mm</quantity>
wide.
</paragraph>
<paragraph id="4B573B64FF9F7E43FF46F92FFEF9F93D" blockId="1.[136,1452,496,1999]" pageId="1" pageNumber="170">
<emphasis id="799CE776FF9F7E43FF46F92FFEA4F941" bold="true" box="[189,313,1684,1710]" pageId="1" pageNumber="170">Phenology</emphasis>
:— 
<taxonomicName id="8CE840E7FF9F7E43FEA5F92EFDAAF941" authorityName="Jara &amp; Moonlight. A. Habit" authorityYear="2023" box="[350,567,1684,1710]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="betancurii">
<emphasis id="799CE776FF9F7E43FEA5F92EFDAAF941" box="[350,567,1684,1710]" italics="true" pageId="1" pageNumber="170">Begonia betancurii</emphasis>
</taxonomicName>
has been collected flowering and fruiting from March to October, and likely does so year-round.
</paragraph>
</subSubSection>
<subSubSection id="03F268EFFF9F7E46FF46F967FA3DFD5F" lastPageId="4" lastPageNumber="173" pageId="1" pageNumber="170" type="etymology">
<paragraph id="4B573B64FF9F7E43FF46F967FDE4F8D0" blockId="1.[136,1452,496,1999]" pageId="1" pageNumber="170">
<emphasis id="799CE776FF9F7E43FF46F967FEA3F919" bold="true" box="[189,318,1756,1782]" pageId="1" pageNumber="170">Etymology</emphasis>
:—The epithet “ 
<taxonomicName id="8CE840E7FF9F7E43FE06F967FDEDF919" authorityName="Jara &amp; Moonlight. A. Habit" authorityYear="2023" box="[509,624,1756,1782]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="betancurii">
<emphasis id="799CE776FF9F7E43FE06F967FDEDF919" box="[509,624,1756,1782]" italics="true" pageId="1" pageNumber="170">betancurii</emphasis>
</taxonomicName>
” honors the Colombian botanist Julio C. Betancur, a prolific collector of plants and for long time curator of the Colombian National Herbarium (COL), he also collected several of the specimens of this species, included the 
<typeStatus id="945385C6FF9F7E43FDBAF89EFDE8F8D0" box="[577,629,1829,1855]" pageId="1" pageNumber="170">type</typeStatus>
.
</paragraph>
<paragraph id="4B573B64FF9F7E43FF46F8F3FB8FF821" blockId="1.[136,1452,496,1999]" pageId="1" pageNumber="170">
<emphasis id="799CE776FF9F7E43FF46F8F3FE6FF88D" bold="true" box="[189,498,1864,1890]" pageId="1" pageNumber="170">Conservation Assessment</emphasis>
:— 
<taxonomicName id="8CE840E7FF9F7E43FDEDF8F2FD69F88D" authorityName="Jara &amp; Moonlight. A. Habit" authorityYear="2023" box="[534,756,1864,1890]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="betancurii">
<emphasis id="799CE776FF9F7E43FDEDF8F2FD69F88D" box="[534,756,1864,1890]" italics="true" pageId="1" pageNumber="170">Begonia betancurii</emphasis>
</taxonomicName>
is known from populations in 
<collectingRegion id="892CF586FF9F7E43FB8AF8F3FA89F88D" box="[1137,1300,1864,1890]" country="Colombia" name="Cundinamarca" pageId="1" pageNumber="170">Cundinamarca</collectingRegion>
and 
<collectingRegion id="892CF586FF9F7E43FAACF8F3FA36F88D" box="[1367,1451,1864,1890]" country="Colombia" name="Boyaca" pageId="1" pageNumber="170">Boyacá</collectingRegion>
departments, which together have an Extent of Occurrence (EOO) of ca. 
<quantity id="8C109681FF9F7E43FC39F8D7FBBBF869" box="[962,1062,1900,1927]" metricMagnitude="6" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="170" unit="km" value="1050.0">1050 km</quantity>
<superScript id="BC9D962CFF9F7E43FBDEF8D7FBB0F895" attach="left" box="[1061,1069,1900,1914]" fontSize="6" pageId="1" pageNumber="170">2</superScript>
. It is known only from fragments of montane forest and is not known from any areas&gt; 
<quantity id="8C109681FF9F7E43FD23F82BFC8CF845" box="[728,785,1936,1962]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="170" unit="km" value="1.0">1 km</quantity>
from roads or urban areas. As such, we provisionally assess 
<taxonomicName id="8CE840E7FF9F7E43FF73F80EFE87F821" authorityName="Jara &amp; Moonlight. A. Habit" authorityYear="2023" box="[136,282,1972,1998]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="170" phylum="Tracheophyta" rank="species" species="betancurii">
<emphasis id="799CE776FF9F7E43FF73F80EFE87F821" box="[136,282,1972,1998]" italics="true" pageId="1" pageNumber="170">B. betancurii</emphasis>
</taxonomicName>
as Endangered (EN B1a,b(iii)) under IUCN Criteria (
<bibRefCitation id="2F794695FF9F7E43FC8EF80FFB9BF821" author="IUCN" box="[885,1030,1972,1998]" pageId="1" pageNumber="170" refId="ref3663" refString="IUCN (2022) The IUCN Red List of Threatened Species, version 2022 - 2. [http: // www. iucnredlist. org /]" type="url" year="2022">IUCN, 2022</bibRefCitation>
).
</paragraph>
<caption id="1F976BECFF9C7E40FF73F873FD7FF7C9" pageId="2" pageNumber="171" startId="2.[136,229,1992,2014]" targetBox="[205,1383,187,1965]" targetPageId="2" targetType="figure">
<paragraph id="4B573B64FF9C7E40FF73F873FD7FF7C9" blockId="2.[136,1452,1992,2086]" pageId="2" pageNumber="171">
<emphasis id="799CE776FF9C7E40FF73F873FF62F832" bold="true" box="[136,255,1992,2014]" pageId="2" pageNumber="171">FIGURE 1.</emphasis>
<taxonomicName id="8CE840E7FF9C7E40FEFCF873FD70F831" authority="Jara &amp; Moonlight. A. Habit" authorityName="Jara &amp; Moonlight. A. Habit" authorityYear="2023" box="[263,749,1992,2014]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="2" pageNumber="171" phylum="Tracheophyta" rank="species" species="betancurii">
<emphasis id="799CE776FF9C7E40FEFCF873FD33F832" bold="true" box="[263,686,1992,2014]" pageId="2" pageNumber="171">
<emphasis id="799CE776FF9C7E40FEFCF873FE5BF831" bold="true" box="[263,454,1992,2014]" italics="true" pageId="2" pageNumber="171">Begonia betancurii</emphasis>
Jara &amp; Moonlight. A.
</emphasis>
Habit
</taxonomicName>
; 
<emphasis id="799CE776FF9C7E40FD02F873FC92F832" bold="true" box="[761,783,1992,2013]" pageId="2" pageNumber="171">B.</emphasis>
Staminate flower (front view); 
<emphasis id="799CE776FF9C7E40FBBCF873FBC2F831" bold="true" box="[1095,1119,1992,2014]" pageId="2" pageNumber="171">C.</emphasis>
Pistillate flowers (front view). 
<emphasis id="799CE776FF9C7E40FA6FF873FA31F832" bold="true" box="[1428,1452,1992,2013]" pageId="2" pageNumber="171">D.</emphasis>
Pistillate flowers (side view), showing ovary and bract. 
<emphasis id="799CE776FF9C7E40FD48F857FD54F7EE" bold="true" box="[691,713,2028,2049]" pageId="2" pageNumber="171">E.</emphasis>
Mature fruit (side view). 
<emphasis id="799CE776FF9C7E40FC37F857FC43F7EE" bold="true" box="[972,990,2028,2049]" pageId="2" pageNumber="171">F.</emphasis>
Immature Fruit in cross-section showing bifid placentae. (photos A to E, by J.C. Zabala; F by M. Fernandez).
</paragraph>
</caption>
<caption id="1F976BECFF9D7E41FF73FDACF864FDDE" box="[136,2041,535,561]" pageId="3" pageNumber="172" startId="3.[136,224,535,561]" targetType="table">
<paragraph id="4B573B64FF9D7E41FF73FDACF864FDDE" blockId="3.[136,2041,535,561]" box="[136,2041,535,561]" pageId="3" pageNumber="172">
<emphasis id="799CE776FF9D7E41FF73FDACFF66FDDE" bold="true" box="[136,251,535,561]" pageId="3" pageNumber="172">TABLE 1.</emphasis>
Comparison of diagnostic characters used to distinguish 
<taxonomicName id="8CE840E7FF9D7E41FCA6FDACFBB6FDDE" authorityName="Jara &amp; Moonlight. A. Habit" authorityYear="2023" box="[861,1067,535,561]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="betancurii">
<emphasis id="799CE776FF9D7E41FCA6FDACFBB6FDDE" box="[861,1067,535,561]" italics="true" pageId="3" pageNumber="172">Begonia betancurii</emphasis>
</taxonomicName>
and the most similar species in sections 
<taxonomicName id="8CE840E7FF9D7E41FA27FDACF9F2FDDE" box="[1500,1647,535,561]" pageId="3" pageNumber="173" rank="section" section="Ruizopavonia">
<emphasis id="799CE776FF9D7E41FA27FDACF9F2FDDE" box="[1500,1647,535,561]" italics="true" pageId="3" pageNumber="172">Ruizopavonia</emphasis>
</taxonomicName>
and 
<taxonomicName id="8CE840E7FF9D7E41F958FDACF8A7FDDE" authorityName="Klotzsch (1855: 220) De Candolle" authorityYear="1864" box="[1699,1850,535,561]" class="Magnoliopsida" family="Begoniaceae" genus="Cyathocnemis" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="genus">
<emphasis id="799CE776FF9D7E41F958FDACF8A7FDDE" box="[1699,1850,535,561]" italics="true" pageId="3" pageNumber="172">Cyathocnemis</emphasis>
</taxonomicName>
(
<taxonomicName id="8CE840E7FF9D7E41F8B1FDACF874FDDE" box="[1866,2025,535,561]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="magdalenae">
<emphasis id="799CE776FF9D7E41F8B1FDACF874FDDE" box="[1866,2025,535,561]" italics="true" pageId="3" pageNumber="172">B. magdalenae</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="4B573B64FF9D7E41FE69FDE9F8EFF92A" pageId="3" pageNumber="172">
<table id="39E8C9C4FF9D81BDFF5DFDE9F878F92A" box="[166,2021,594,1733]" gridcols="9" gridrows="9" pageId="3" pageNumber="172">
<tr id="F5D83926FF9D81BDFF5DFDE9F878FD69" box="[166,2021,594,646]" gridrow="0" pageId="3" pageNumber="172" rowspan-0="1">
<th id="B609505AFF9D81BDFE69FDE9FDB4FD69" box="[402,553,594,646]" gridcol="1" gridrow="0" pageId="3" pageNumber="172">
<emphasis id="799CE776FF9D7E41FE69FDE9FE41FD69" bold="true" italics="true" pageId="3" pageNumber="172">
<taxonomicName id="8CE840E7FF9D7E41FE69FDE9FD8EFD86" authority="Jara-Muñoz &amp; Moonlight &amp; Zabala-Rivera, 2023" authorityName="Jara-Muñoz &amp; Moonlight &amp; Zabala-Rivera" authorityYear="2023" box="[402,531,594,617]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="betancurii" status="sp. nov.">B. betancurii</taxonomicName>
<taxonomicNameLabel id="62AF5A0DFF9D7E41FE69FDD4FE41FD69" box="[402,476,623,646]" pageId="3" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="B609505AFF9D81BDFD8BFDE9FD79FD69" box="[624,740,594,646]" gridcol="2" gridrow="0" pageId="3" pageNumber="172">
<taxonomicName id="8CE840E7FF9D7E41FD8BFDE9FD7DFD86" box="[624,736,594,617]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="alnifolia">
<emphasis id="799CE776FF9D7E41FD8BFDE9FD7DFD86" bold="true" box="[624,736,594,617]" italics="true" pageId="3" pageNumber="172">B. alnifolia</emphasis>
</taxonomicName>
</th>
<th id="B609505AFF9D81BDFCD2FDE9FC22FD69" box="[809,959,594,646]" gridcol="3" gridrow="0" pageId="3" pageNumber="172">
<taxonomicName id="8CE840E7FF9D7E41FCD2FDE9FC29FD86" box="[809,948,594,617]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="consobrina">
<emphasis id="799CE776FF9D7E41FCD2FDE9FC29FD86" bold="true" box="[809,948,594,617]" italics="true" pageId="3" pageNumber="172">B. consobrina</emphasis>
</taxonomicName>
</th>
<th id="B609505AFF9D81BDFC06FDE9FB13FD69" box="[1021,1166,594,646]" gridcol="4" gridrow="0" pageId="3" pageNumber="172">
<taxonomicName id="8CE840E7FF9D7E41FC06FDE9FB13FD86" box="[1021,1166,594,617]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="cymbalifera">
<emphasis id="799CE776FF9D7E41FC06FDE9FB13FD86" bold="true" box="[1021,1166,594,617]" italics="true" pageId="3" pageNumber="172">B. cymbalifera</emphasis>
</taxonomicName>
</th>
<th id="B609505AFF9D81BDFB37FDE9FAC2FD69" box="[1228,1375,594,646]" gridcol="5" gridrow="0" pageId="3" pageNumber="172">
<taxonomicName id="8CE840E7FF9D7E41FB37FDE9FAC2FD86" box="[1228,1375,594,617]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="dugandiana">
<emphasis id="799CE776FF9D7E41FB37FDE9FAC2FD86" bold="true" box="[1228,1375,594,617]" italics="true" pageId="3" pageNumber="172">B. dugandiana</emphasis>
</taxonomicName>
</th>
<th id="B609505AFF9D81BDFA65FDE9F99BFD69" box="[1438,1542,594,646]" gridcol="6" gridrow="0" pageId="3" pageNumber="172">
<taxonomicName id="8CE840E7FF9D7E41FA65FDE9F998FD86" box="[1438,1541,594,617]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="extensa">
<emphasis id="799CE776FF9D7E41FA65FDE9F998FD86" bold="true" box="[1438,1541,594,617]" italics="true" pageId="3" pageNumber="172">B. extensa</emphasis>
</taxonomicName>
</th>
<th id="B609505AFF9D81BDF9B6FDE9F898FD69" box="[1613,1797,594,646]" gridcol="7" gridrow="0" pageId="3" pageNumber="172">
<taxonomicName id="8CE840E7FF9D7E41F9B6FDE9F89CFD86" box="[1613,1793,594,617]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="pseudodendron">
<emphasis id="799CE776FF9D7E41F9B6FDE9F89CFD86" bold="true" box="[1613,1793,594,617]" italics="true" pageId="3" pageNumber="172">B. pseudodendron</emphasis>
</taxonomicName>
</th>
<th id="B609505AFF9D81BDF8BAFDE9F878FD69" box="[1857,2021,594,646]" gridcol="8" gridrow="0" pageId="3" pageNumber="172">
<emphasis id="799CE776FF9D7E41F8BAFDE9F848FD86" bold="true" box="[1857,2005,594,617]" italics="true" pageId="3" pageNumber="172">B. magdalenae</emphasis>
</th>
</tr>
<tr id="F5D83926FF9D81BDFF5DFD06F878FCE1" box="[166,2021,701,782]" gridrow="1" pageId="3" pageNumber="172">
<th id="B609505AFF9D81BDFF5DFD06FEC8FCE1" box="[166,341,701,782]" gridcol="0" gridrow="1" pageId="3" pageNumber="172">Leaf position relative to the petiole</th>
<td id="B609505AFF9D81BDFE69FD06FDB4FCE1" box="[402,553,701,782]" gridcol="1" gridrow="1" pageId="3" pageNumber="172">Transverse</td>
<td id="B609505AFF9D81BDFD8BFD06FD79FCE1" box="[624,740,701,782]" gridcol="2" gridrow="1" pageId="3" pageNumber="172">Straight</td>
<td id="B609505AFF9D81BDFCD2FD06FC22FCE1" box="[809,959,701,782]" gridcol="3" gridrow="1" pageId="3" pageNumber="172">Transverse</td>
<td id="B609505AFF9D81BDFC06FD06FB13FCE1" box="[1021,1166,701,782]" gridcol="4" gridrow="1" pageId="3" pageNumber="172">Straight or transverse</td>
<td id="B609505AFF9D81BDFB37FD06FAC2FCE1" box="[1228,1375,701,782]" gridcol="5" gridrow="1" pageId="3" pageNumber="172">Straight to slightly transverse</td>
<td id="B609505AFF9D81BDFA65FD06F99BFCE1" box="[1438,1542,701,782]" gridcol="6" gridrow="1" pageId="3" pageNumber="172">Transverse</td>
<td id="B609505AFF9D81BDF9B6FD06F898FCE1" box="[1613,1797,701,782]" gridcol="7" gridrow="1" pageId="3" pageNumber="172">Straight</td>
<td id="B609505AFF9D81BDF8BAFD06F878FCE1" box="[1857,2021,701,782]" gridcol="8" gridrow="1" pageId="3" pageNumber="172">Transverse</td>
</tr>
<tr id="F5D83926FF9D81BDFF5DFCFFF878FC3E" box="[166,2021,836,977]" gridrow="2" pageId="3" pageNumber="172">
<th id="B609505AFF9D81BDFF5DFCFFFEC8FC3E" box="[166,341,836,977]" gridcol="0" gridrow="2" pageId="3" pageNumber="172">Leaf venation and number of veins on the broad /narrow sides of the lamina</th>
<td id="B609505AFF9D81BDFE69FCFFFDB4FC3E" box="[402,553,836,977]" gridcol="1" gridrow="2" pageId="3" pageNumber="172">Palmate- pinnate-nerved; 9–10 / 5–6</td>
<td id="B609505AFF9D81BDFD8BFCFFFD79FC3E" box="[624,740,836,977]" gridcol="2" gridrow="2" pageId="3" pageNumber="172">Pinnate- nerved; 13–17 / 13–15</td>
<td id="B609505AFF9D81BDFCD2FCFFFC22FC3E" box="[809,959,836,977]" gridcol="3" gridrow="2" pageId="3" pageNumber="172">Palmate- pinnate-nerved, 9–12 / 7–9</td>
<td id="B609505AFF9D81BDFC06FCFFFB13FC3E" box="[1021,1166,836,977]" gridcol="4" gridrow="2" pageId="3" pageNumber="172">Pinnate- nerved; 8–12 / 6–8</td>
<td id="B609505AFF9D81BDFB37FCFFFAC2FC3E" box="[1228,1375,836,977]" gridcol="5" gridrow="2" pageId="3" pageNumber="172">Pinnate- nerved; 6–8 / 4–6</td>
<td id="B609505AFF9D81BDFA65FCFFF99BFC3E" box="[1438,1542,836,977]" gridcol="6" gridrow="2" pageId="3" pageNumber="172">Palmate- pinnate- nerved; 9 / 6–8</td>
<td id="B609505AFF9D81BDF9B6FCFFF898FC3E" box="[1613,1797,836,977]" gridcol="7" gridrow="2" pageId="3" pageNumber="172">Pinnate-nerved; 7–9 / 5–7</td>
<td id="B609505AFF9D81BDF8BAFCFFF878FC3E" box="[1857,2021,836,977]" gridcol="8" gridrow="2" pageId="3" pageNumber="172">Palmate-pinnatenerved; 10–12 / 6–7</td>
</tr>
<tr id="F5D83926FF9D81BDFF5DFBBCF878FBB7" box="[166,2021,1031,1112]" gridrow="3" pageId="3" pageNumber="172">
<th id="B609505AFF9D81BDFF5DFBBCFEC8FBB7" box="[166,341,1031,1112]" gridcol="0" gridrow="3" pageId="3" pageNumber="172">Flowering peduncle, long (mm)</th>
<td id="B609505AFF9D81BDFE69FBBCFDB4FBB7" box="[402,553,1031,1112]" gridcol="1" gridrow="3" pageId="3" pageNumber="172">34–77</td>
<td id="B609505AFF9D81BDFD8BFBBCFD79FBB7" box="[624,740,1031,1112]" gridcol="2" gridrow="3" pageId="3" pageNumber="172">25–66</td>
<td id="B609505AFF9D81BDFCD2FBBCFC22FBB7" box="[809,959,1031,1112]" gridcol="3" gridrow="3" pageId="3" pageNumber="172">20–45</td>
<td id="B609505AFF9D81BDFC06FBBCFB13FBB7" box="[1021,1166,1031,1112]" gridcol="4" gridrow="3" pageId="3" pageNumber="172">ca. 50</td>
<td id="B609505AFF9D81BDFB37FBBCFAC2FBB7" box="[1228,1375,1031,1112]" gridcol="5" gridrow="3" pageId="3" pageNumber="172">14–20</td>
<td id="B609505AFF9D81BDFA65FBBCF99BFBB7" box="[1438,1542,1031,1112]" gridcol="6" gridrow="3" pageId="3" pageNumber="172">37</td>
<td id="B609505AFF9D81BDF9B6FBBCF898FBB7" box="[1613,1797,1031,1112]" gridcol="7" gridrow="3" pageId="3" pageNumber="172">15–45</td>
<td id="B609505AFF9D81BDF8BAFBBCF878FBB7" box="[1857,2021,1031,1112]" gridcol="8" gridrow="3" pageId="3" pageNumber="172">50–60</td>
</tr>
<tr id="F5D83926FF9D81BDFF5DFB34F878FB0F" box="[166,2021,1167,1248]" gridrow="4" pageId="3" pageNumber="172">
<th id="B609505AFF9D81BDFF5DFB34FEC8FB0F" box="[166,341,1167,1248]" gridcol="0" gridrow="4" pageId="3" pageNumber="172">Number of bifurcations of the inflorescence</th>
<td id="B609505AFF9D81BDFE69FB34FDB4FB0F" box="[402,553,1167,1248]" gridcol="1" gridrow="4" pageId="3" pageNumber="172">2–4</td>
<td id="B609505AFF9D81BDFD8BFB34FD79FB0F" box="[624,740,1167,1248]" gridcol="2" gridrow="4" pageId="3" pageNumber="172">3–5</td>
<td id="B609505AFF9D81BDFCD2FB34FC22FB0F" box="[809,959,1167,1248]" gridcol="3" gridrow="4" pageId="3" pageNumber="172">2–3</td>
<td id="B609505AFF9D81BDFC06FB34FB13FB0F" box="[1021,1166,1167,1248]" gridcol="4" gridrow="4" pageId="3" pageNumber="172">4</td>
<td id="B609505AFF9D81BDFB37FB34FAC2FB0F" box="[1228,1375,1167,1248]" gridcol="5" gridrow="4" pageId="3" pageNumber="172">2–3</td>
<td id="B609505AFF9D81BDFA65FB34F99BFB0F" box="[1438,1542,1167,1248]" gridcol="6" gridrow="4" pageId="3" pageNumber="172">2</td>
<td id="B609505AFF9D81BDF9B6FB34F898FB0F" box="[1613,1797,1167,1248]" gridcol="7" gridrow="4" pageId="3" pageNumber="172">6–8</td>
<td id="B609505AFF9D81BDF8BAFB34F878FB0F" box="[1857,2021,1167,1248]" gridcol="8" gridrow="4" pageId="3" pageNumber="172">3–4</td>
</tr>
<tr id="F5D83926FF9D81BDFF5DFAADF878FA6A" box="[166,2021,1302,1413]" gridrow="5" pageId="3" pageNumber="172">
<th id="B609505AFF9D81BDFF5DFAADFEC8FA6A" box="[166,341,1302,1413]" gridcol="0" gridrow="5" pageId="3" pageNumber="172">Number of tepals in staminate and pistillate flowers</th>
<td id="B609505AFF9D81BDFE69FAADFDB4FA6A" box="[402,553,1302,1413]" gridcol="1" gridrow="5" pageId="3" pageNumber="172">4 in staminate, 3 in pistillate</td>
<td id="B609505AFF9D81BDFD8BFAADFD79FA6A" box="[624,740,1302,1413]" gridcol="2" gridrow="5" pageId="3" pageNumber="172">2 in staminate, 2 in pistillate</td>
<td id="B609505AFF9D81BDFCD2FAADFC22FA6A" box="[809,959,1302,1413]" gridcol="3" gridrow="5" pageId="3" pageNumber="172">4 in staminate, 3 in pistillate</td>
<td id="B609505AFF9D81BDFC06FAADFB13FA6A" box="[1021,1166,1302,1413]" gridcol="4" gridrow="5" pageId="3" pageNumber="172">4 in staminate, 3 in pistillate</td>
<td id="B609505AFF9D81BDFB37FAADFAC2FA6A" box="[1228,1375,1302,1413]" gridcol="5" gridrow="5" pageId="3" pageNumber="172">4 in staminate, 2 in pistillate</td>
<td id="B609505AFF9D81BDFA65FAADF99BFA6A" box="[1438,1542,1302,1413]" gridcol="6" gridrow="5" pageId="3" pageNumber="172">2 in staminate, 3 in pistillate</td>
<td id="B609505AFF9D81BDF9B6FAADF898FA6A" box="[1613,1797,1302,1413]" gridcol="7" gridrow="5" pageId="3" pageNumber="172">4 in staminate, 2 in pistillate</td>
<td id="B609505AFF9D81BDF8BAFAADF878FA6A" box="[1857,2021,1302,1413]" gridcol="8" gridrow="5" pageId="3" pageNumber="172">2 in staminate, 2–3 in pistillate</td>
</tr>
<tr id="F5D83926FF9D81BDFF5DFA00F878F9E2" box="[166,2021,1467,1549]" gridrow="6" pageId="3" pageNumber="172">
<th id="B609505AFF9D81BDFF5DFA00FEC8F9E2" box="[166,341,1467,1549]" gridcol="0" gridrow="6" pageId="3" pageNumber="172">Length of staminate outer tepals (mm)</th>
<td id="B609505AFF9D81BDFE69FA00FDB4F9E2" box="[402,553,1467,1549]" gridcol="1" gridrow="6" pageId="3" pageNumber="172">ca. 15</td>
<td id="B609505AFF9D81BDFD8BFA00FD79F9E2" box="[624,740,1467,1549]" gridcol="2" gridrow="6" pageId="3" pageNumber="172">4–5</td>
<td id="B609505AFF9D81BDFCD2FA00FC22F9E2" box="[809,959,1467,1549]" gridcol="3" gridrow="6" pageId="3" pageNumber="172">4–5</td>
<td id="B609505AFF9D81BDFC06FA00FB13F9E2" box="[1021,1166,1467,1549]" gridcol="4" gridrow="6" pageId="3" pageNumber="172">ca. 5</td>
<td id="B609505AFF9D81BDFB37FA00FAC2F9E2" box="[1228,1375,1467,1549]" gridcol="5" gridrow="6" pageId="3" pageNumber="172">ca. 5.5</td>
<td id="B609505AFF9D81BDFA65FA00F99BF9E2" box="[1438,1542,1467,1549]" gridcol="6" gridrow="6" pageId="3" pageNumber="172">ca. 16</td>
<td id="B609505AFF9D81BDF9B6FA00F898F9E2" box="[1613,1797,1467,1549]" gridcol="7" gridrow="6" pageId="3" pageNumber="172">3–6</td>
<td id="B609505AFF9D81BDF8BAFA00F878F9E2" box="[1857,2021,1467,1549]" gridcol="8" gridrow="6" pageId="3" pageNumber="172">7</td>
</tr>
<tr id="F5D83926FF9D81BDFF5DF9F8F878F997" box="[166,2021,1603,1656]" gridrow="7" pageId="3" pageNumber="172">
<th id="B609505AFF9D81BDFF5DF9F8FEC8F997" box="[166,341,1603,1656]" gridcol="0" gridrow="7" pageId="3" pageNumber="172">Number of stamens</th>
<td id="B609505AFF9D81BDFE69F9F8FDB4F997" box="[402,553,1603,1656]" gridcol="1" gridrow="7" pageId="3" pageNumber="172">ca. 55</td>
<td id="B609505AFF9D81BDFD8BF9F8FD79F997" box="[624,740,1603,1656]" gridcol="2" gridrow="7" pageId="3" pageNumber="172">ca. 30</td>
<td id="B609505AFF9D81BDFCD2F9F8FC22F997" box="[809,959,1603,1656]" gridcol="3" gridrow="7" pageId="3" pageNumber="172">&gt;25</td>
<td id="B609505AFF9D81BDFC06F9F8FB13F997" box="[1021,1166,1603,1656]" gridcol="4" gridrow="7" pageId="3" pageNumber="172">&gt;25</td>
<td id="B609505AFF9D81BDFB37F9F8FAC2F997" box="[1228,1375,1603,1656]" gridcol="5" gridrow="7" pageId="3" pageNumber="172">ca. 20</td>
<td id="B609505AFF9D81BDFA65F9F8F99BF997" box="[1438,1542,1603,1656]" gridcol="6" gridrow="7" pageId="3" pageNumber="172">ca. 50</td>
<td id="B609505AFF9D81BDF9B6F9F8F898F997" box="[1613,1797,1603,1656]" gridcol="7" gridrow="7" pageId="3" pageNumber="172">ca. 20</td>
<td id="B609505AFF9D81BDF8BAF9F8F878F997" box="[1857,2021,1603,1656]" gridcol="8" gridrow="7" pageId="3" pageNumber="172">ca. 13</td>
</tr>
<tr id="F5D83926FF9D81BDFF5DF915F878F92A" box="[166,2021,1710,1733]" gridrow="8" pageId="3" pageNumber="172">
<th id="B609505AFF9D81BDFF5DF915FEC8F92A" box="[166,341,1710,1733]" gridcol="0" gridrow="8" pageId="3" pageNumber="172">Form of the styles</th>
<td id="B609505AFF9D81BDFE69F915FDB4F92A" box="[402,553,1710,1733]" gridcol="1" gridrow="8" pageId="3" pageNumber="172">Multifid</td>
<td id="B609505AFF9D81BDFD8BF915FD79F92A" box="[624,740,1710,1733]" gridcol="2" gridrow="8" pageId="3" pageNumber="172">Multifid</td>
<td id="B609505AFF9D81BDFCD2F915FC22F92A" box="[809,959,1710,1733]" gridcol="3" gridrow="8" pageId="3" pageNumber="172">Multifid</td>
<td id="B609505AFF9D81BDFC06F915FB13F92A" box="[1021,1166,1710,1733]" gridcol="4" gridrow="8" pageId="3" pageNumber="172">Multifid</td>
<td id="B609505AFF9D81BDFB37F915FAC2F92A" box="[1228,1375,1710,1733]" gridcol="5" gridrow="8" pageId="3" pageNumber="172">Multifid</td>
<td id="B609505AFF9D81BDFA65F915F99BF92A" box="[1438,1542,1710,1733]" gridcol="6" gridrow="8" pageId="3" pageNumber="172">Bifid</td>
<td id="B609505AFF9D81BDF9B6F915F898F92A" box="[1613,1797,1710,1733]" gridcol="7" gridrow="8" pageId="3" pageNumber="172">Bifid</td>
<td id="B609505AFF9D81BDF8BAF915F878F92A" box="[1857,2021,1710,1733]" gridcol="8" gridrow="8" pageId="3" pageNumber="172">Bifid</td>
</tr>
</table>
</paragraph>
<paragraph id="4B573B64FF9A7E46FF46FF25FBD5FE5A" blockId="4.[136,1452,158,689]" pageId="4" pageNumber="173">
<emphasis id="799CE776FF9A7E46FF46FF25FE18FF57" bold="true" box="[189,389,158,184]" pageId="4" pageNumber="173">Taxonomic notes</emphasis>
:—Section 
<taxonomicName id="8CE840E7FF9A7E46FDF8FF24FD00FF57" box="[515,669,159,184]" pageId="4" pageNumber="173" rank="section" section="Ruizopavonia">
<emphasis id="799CE776FF9A7E46FDF8FF24FD00FF57" box="[515,669,159,184]" italics="true" pageId="4" pageNumber="173">Ruizopavonia</emphasis>
</taxonomicName>
was split by 
<bibRefCitation id="2F794695FF9A7E46FCD6FF25FBA9FF57" author="Moonlight, P. W. &amp; Ardi, W. H. &amp; Arroyo Padilla, L. &amp; Chung, K. - F. &amp; Fuller, D. &amp; Girmansyah, D. &amp; Hollands, R. &amp; Jara-Munoz, O. A. &amp; Kiew, R. &amp; Leong, W. - C. &amp; Liu, Y. &amp; Mahardika, A. &amp; Marasinghe, L. D. K. &amp; O'Connor, M. &amp; Peng, C. &amp; Perez, A. J. &amp; Phutthai, T. &amp; Pullan, M. &amp; Rajbhandary, S. &amp; Reynel, C. &amp; Rubite, R. R. &amp; Sang, J. &amp; Scherberich, D. &amp; Shui, Y. - M. &amp; Tebbitt, M. C. &amp; Thomas, D. C. &amp; Wilson, H. P. &amp; Zaini, N. H. &amp; Hughes, M." box="[813,1076,158,184]" pageId="4" pageNumber="173" pagination="267 - 323" refId="ref3808" refString="Moonlight, P. W., Ardi, W. H., Arroyo Padilla, L., Chung, K. - F., Fuller, D., Girmansyah, D., Hollands, R., Jara-Munoz, O. A., Kiew, R., Leong, W. - C., Liu, Y., Mahardika, A., Marasinghe, L. D. K., O'Connor, M., Peng, C. - I, Perez, A. J., Phutthai, T., Pullan, M., Rajbhandary, S., Reynel, C., Rubite, R. R., Sang, J., Scherberich, D., Shui, Y. - M., Tebbitt, M. C., Thomas, D. C., Wilson, H. P., Zaini, N. H. &amp; Hughes, M. (2018) Dividing and conquering the fastest-growing genus: towards a natural sectional classification of the mega-diverse genus Begonia (Begoniaceae). Taxon 67: 267 - 323. https: // doi. org / 10.12705 / 672.3" type="journal article" year="2018">
Moonlight 
<emphasis id="799CE776FF9A7E46FC52FF24FC7CFF57" box="[937,993,158,184]" italics="true" pageId="4" pageNumber="173">et al.</emphasis>
(2018)
</bibRefCitation>
, so its size was reduced in respect from the earlier concept of the section by Doorembos 
<emphasis id="799CE776FF9A7E46FD1FFF78FC81FF33" box="[740,796,194,220]" italics="true" pageId="4" pageNumber="173">et al.</emphasis>
(1998). After the rearrangement, 26 species were included in 
<taxonomicName id="8CE840E7FF9A7E46FF5FFF5CFEA3FEEF" box="[164,318,231,256]" pageId="4" pageNumber="173" rank="section" section="Ruizopavonia">
<emphasis id="799CE776FF9A7E46FF5FFF5CFEA3FEEF" box="[164,318,231,256]" italics="true" pageId="4" pageNumber="173">Ruizopavonia</emphasis>
</taxonomicName>
, while the remainder were placed in sections 
<taxonomicName id="8CE840E7FF9A7E46FCC3FF5DFF23FECB" authority="Klotzsch (1855: 220) De Candolle (1864: 332)" authorityName="Klotzsch (1855: 220) De Candolle" authorityYear="1864" class="Magnoliopsida" family="Begoniaceae" genus="Cyathocnemis" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="genus">
<emphasis id="799CE776FF9A7E46FCC3FF5DFC4AFEEF" box="[824,983,230,256]" italics="true" pageId="4" pageNumber="173">Cyathocnemis</emphasis>
<bibRefCitation id="2F794695FF9A7E46FC26FF5DFB57FEEF" author="Klotzsch, J. F." box="[989,1226,230,257]" pageId="4" pageNumber="173" pagination="121 - 255" refId="ref3755" refString="Klotzsch, J. F. (1855) Begoniaceeen - Gattungen und Arden. Abhandlungen der Koniglichen Akademie der Wissenschaften zu Berlin: aus dem Jahre 1854: 121 - 255." type="journal article" year="1855">Klotzsch (1855: 220)</bibRefCitation>
<bibRefCitation id="2F794695FF9A7E46FB2BFF5CFF23FECB" author="De Candolle, A. L. P. P." pageId="4" pageNumber="173" pagination="265 - 408" refId="ref3522" refString="De Candolle, A. L. P. P. (1864) Begoniaceae. In: De Candolle, A. L. P. P. (Ed.) Prodromus systematis naturalis regni vegetabilis sive enumeratio contracta ordinum, generum specierumque plantarum huc usque cognitarum, juxta methodi naturalis normas digesta. Sumptibus sociorum Treuttel et Wurtz, Paris, pp. 265 - 408." type="book chapter" year="1864">De Candolle (1864: 332)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="8CE840E7FF9A7E46FF36FEB1FCF8FECB" authority="Klotzsch (1854: 127) De Candolle (1859: 127)" authorityName="Klotzsch (1854: 127) De Candolle" authorityYear="1859" box="[205,869,266,293]" class="Magnoliopsida" family="Begoniaceae" genus="Donaldia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="genus">
<emphasis id="799CE776FF9A7E46FF36FEB1FEAAFECB" box="[205,311,266,292]" italics="true" pageId="4" pageNumber="173">Donaldia</emphasis>
Klotzsch (1854: 127) 
<bibRefCitation id="2F794695FF9A7E46FDBBFEB0FCF8FECB" author="De Candolle, A. L. P. P." box="[576,869,266,293]" pageId="4" pageNumber="173" pagination="115 - 145" refId="ref3489" refString="De Candolle, A. L. P. P. (1859) Memoire Sur La Famille Des Begoniacees. Annales Des Sciences Naturelles. Botanique 11: 115 - 145." type="journal article" year="1859">De Candolle (1859: 127)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="8CE840E7FF9A7E46FC88FEB0FF23FEA7" authority="Klotzsch (1854: 123) De Candolle (1859: 139)" authorityName="Klotzsch (1854: 123) De Candolle" authorityYear="1859" class="Magnoliopsida" family="Begoniaceae" genus="Lepsia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="genus">
<emphasis id="799CE776FF9A7E46FC88FEB0FC23FECB" box="[883,958,267,292]" italics="true" pageId="4" pageNumber="173">Lepsia</emphasis>
Klotzsch (1854: 123) 
<bibRefCitation id="2F794695FF9A7E46FB33FEB0FF23FEA7" author="De Candolle, A. L. P. P." pageId="4" pageNumber="173" pagination="115 - 145" refId="ref3489" refString="De Candolle, A. L. P. P. (1859) Memoire Sur La Famille Des Begoniacees. Annales Des Sciences Naturelles. Botanique 11: 115 - 145." type="journal article" year="1859">De Candolle (1859: 139)</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="8CE840E7FF9A7E46FF01FE95FCEBFEA7" authority="Klotzsch (1854: 126) De Candolle (1859: 137)" authorityName="Klotzsch (1854: 126) De Candolle" authorityYear="1859" box="[250,886,302,329]" class="Magnoliopsida" family="Begoniaceae" genus="Pritzelia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="genus">
<emphasis id="799CE776FF9A7E46FF01FE95FEC7FEA7" box="[250,346,302,328]" italics="true" pageId="4" pageNumber="173">Pritzelia</emphasis>
Klotzsch (1854: 126) 
<bibRefCitation id="2F794695FF9A7E46FDA2FE94FCEBFEA7" author="De Candolle, A. L. P. P." box="[601,886,302,329]" pageId="4" pageNumber="173" pagination="115 - 145" refId="ref3489" refString="De Candolle, A. L. P. P. (1859) Memoire Sur La Famille Des Begoniacees. Annales Des Sciences Naturelles. Botanique 11: 115 - 145." type="journal article" year="1859">De Candolle (1859: 137)</bibRefCitation>
</taxonomicName>
. However, uncertainty persisted in the placements of Bolivian members of 
<taxonomicName id="8CE840E7FF9A7E46FE60FEE8FDA8FE83" box="[411,565,339,364]" pageId="4" pageNumber="173" rank="section" section="Ruizopavonia">
<emphasis id="799CE776FF9A7E46FE60FEE8FDA8FE83" box="[411,565,339,364]" italics="true" pageId="4" pageNumber="173">Ruizopavonia</emphasis>
</taxonomicName>
, which were not sampled in the phylogeny. The placement of northern Andean species of section 
<taxonomicName id="8CE840E7FF9A7E46FE9BFECDFE62FE7F" authorityName="Klotzsch (1855: 220) De Candolle" authorityYear="1864" box="[352,511,374,400]" class="Magnoliopsida" family="Begoniaceae" genus="Cyathocnemis" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="genus">
<emphasis id="799CE776FF9A7E46FE9BFECDFE62FE7F" box="[352,511,374,400]" italics="true" pageId="4" pageNumber="173">Cyathocnemis</emphasis>
</taxonomicName>
(e.g. 
<taxonomicName id="8CE840E7FF9A7E46FDBDFECCFBC6FE7F" authority="Smith &amp; Schubert (1946: 90))" authorityName="Smith &amp; Schubert" authorityPageNumber="90" authorityYear="1946" box="[582,1115,374,400]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="species" species="magdalenae">
<emphasis id="799CE776FF9A7E46FDBDFECCFD6FFE7F" box="[582,754,374,400]" italics="true" pageId="4" pageNumber="173">B. magdalenae</emphasis>
<bibRefCitation id="2F794695FF9A7E46FD07FECDFBC6FE7F" author="Smith, L. B. &amp; Schubert, B. G." box="[764,1115,374,400]" pageId="4" pageNumber="173" pagination="18 - 20" refId="ref4078" refString="Smith, L. B. &amp; Schubert, B. G. (1946) The Begoniaceae of Colombia. Caldasia 4: 18 - 20." type="journal article" year="1946">Smith &amp; Schubert (1946: 90))</bibRefCitation>
</taxonomicName>
is also uncertain, and future phylogenetic evidence may result in these species being placed in sect. 
<taxonomicName id="8CE840E7FF9A7E46FC5CFE20FBDCFE5B" box="[935,1089,411,436]" pageId="4" pageNumber="173" rank="section" section="Ruizopavonia">
<emphasis id="799CE776FF9A7E46FC5CFE20FBDCFE5B" box="[935,1089,411,436]" italics="true" pageId="4" pageNumber="173">Ruizopavonia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4B573B64FF9A7E46FF46FE04FA3DFD5F" blockId="4.[136,1452,158,689]" pageId="4" pageNumber="173">
We provide a comparative table (
<tableCitation id="066A0EDFFF9A7E46FDCFFE05FD15FE37" box="[564,648,446,472]" captionStart="TABLE 1" captionStartId="3.[136,224,535,561]" captionText="TABLE 1. Comparison of diagnostic characters used to distinguish Begonia betancurii and the most similar species in sections Ruizopavonia and Cyathocnemis (B. magdalenae)." pageId="4" pageNumber="173">Table 1</tableCitation>
) with selected characters of 
<taxonomicName id="8CE840E7FF9A7E46FC2BFE04FBFEFE37" authorityName="Jara-Muñoz &amp; Moonlight &amp; Zabala-Rivera" authorityYear="2023" box="[976,1123,446,472]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="species" species="betancurii">
<emphasis id="799CE776FF9A7E46FC2BFE04FBFEFE37" box="[976,1123,446,472]" italics="true" pageId="4" pageNumber="173">B. betancurii</emphasis>
</taxonomicName>
and the most similar species in sect. 
<taxonomicName id="8CE840E7FF9A7E46FF1FFE58FEE3FE13" box="[228,382,483,508]" pageId="4" pageNumber="173" rank="section" section="Ruizopavonia">
<emphasis id="799CE776FF9A7E46FF1FFE58FEE3FE13" box="[228,382,483,508]" italics="true" pageId="4" pageNumber="173">Ruizopavonia</emphasis>
</taxonomicName>
, and one species in sect. 
<taxonomicName id="8CE840E7FF9A7E46FD50FE59FCD7FE13" authorityName="Klotzsch (1855: 220) De Candolle" authorityYear="1864" box="[683,842,482,508]" class="Magnoliopsida" family="Begoniaceae" genus="Cyathocnemis" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="genus">
<emphasis id="799CE776FF9A7E46FD50FE59FCD7FE13" box="[683,842,482,508]" italics="true" pageId="4" pageNumber="173">Cyathocnemis</emphasis>
</taxonomicName>
(
<taxonomicName id="8CE840E7FF9A7E46FCA6FE58FB95FE13" authorityName="Smith &amp; Schubert" authorityYear="1946" box="[861,1032,482,508]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="species" species="magdalenae">
<emphasis id="799CE776FF9A7E46FCA6FE58FB95FE13" box="[861,1032,482,508]" italics="true" pageId="4" pageNumber="173">B. magdalenae</emphasis>
</taxonomicName>
). Our placement of 
<taxonomicName id="8CE840E7FF9A7E46FB0DFE58FA16FE13" authorityName="Jara-Muñoz &amp; Moonlight &amp; Zabala-Rivera" authorityYear="2023" box="[1270,1419,482,508]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="species" species="betancurii">
<emphasis id="799CE776FF9A7E46FB0DFE58FA16FE13" box="[1270,1419,482,508]" italics="true" pageId="4" pageNumber="173">B. betancurii</emphasis>
</taxonomicName>
in section 
<taxonomicName id="8CE840E7FF9A7E46FF24FDBCFEE4FDCF" box="[223,377,519,544]" pageId="4" pageNumber="173" rank="section" section="Ruizopavonia">
<emphasis id="799CE776FF9A7E46FF24FDBCFEE4FDCF" box="[223,377,519,544]" italics="true" pageId="4" pageNumber="173">Ruizopavonia</emphasis>
</taxonomicName>
is based on i) the presence of four staminate and three pistillate tepals (less than five), which is the common condition among the section, ii) oblong or elliptic anthers longer than the filaments, and iii) the raised secondary veins on the underside, which distinguish this and other sections in Neotropical clade II (NC-ii) 
<emphasis id="799CE776FF9A7E46FA96FDF4FA36FD87" box="[1389,1451,591,616]" italics="true" pageId="4" pageNumber="173">sensu</emphasis>
<bibRefCitation id="2F794695FF9A7E46FF73FDC9FE08FD63" author="Moonlight, P. W. &amp; Ardi, W. H. &amp; Arroyo Padilla, L. &amp; Chung, K. - F. &amp; Fuller, D. &amp; Girmansyah, D. &amp; Hollands, R. &amp; Jara-Munoz, O. A. &amp; Kiew, R. &amp; Leong, W. - C. &amp; Liu, Y. &amp; Mahardika, A. &amp; Marasinghe, L. D. K. &amp; O'Connor, M. &amp; Peng, C. &amp; Perez, A. J. &amp; Phutthai, T. &amp; Pullan, M. &amp; Rajbhandary, S. &amp; Reynel, C. &amp; Rubite, R. R. &amp; Sang, J. &amp; Scherberich, D. &amp; Shui, Y. - M. &amp; Tebbitt, M. C. &amp; Thomas, D. C. &amp; Wilson, H. P. &amp; Zaini, N. H. &amp; Hughes, M." box="[136,405,626,652]" pageId="4" pageNumber="173" pagination="267 - 323" refId="ref3808" refString="Moonlight, P. W., Ardi, W. H., Arroyo Padilla, L., Chung, K. - F., Fuller, D., Girmansyah, D., Hollands, R., Jara-Munoz, O. A., Kiew, R., Leong, W. - C., Liu, Y., Mahardika, A., Marasinghe, L. D. K., O'Connor, M., Peng, C. - I, Perez, A. J., Phutthai, T., Pullan, M., Rajbhandary, S., Reynel, C., Rubite, R. R., Sang, J., Scherberich, D., Shui, Y. - M., Tebbitt, M. C., Thomas, D. C., Wilson, H. P., Zaini, N. H. &amp; Hughes, M. (2018) Dividing and conquering the fastest-growing genus: towards a natural sectional classification of the mega-diverse genus Begonia (Begoniaceae). Taxon 67: 267 - 323. https: // doi. org / 10.12705 / 672.3" type="journal article" year="2018">
Moonlight 
<emphasis id="799CE776FF9A7E46FEFCFDC8FEDCFD63" box="[263,321,626,652]" italics="true" pageId="4" pageNumber="173">et al.</emphasis>
(2018)
</bibRefCitation>
from section 
<taxonomicName id="8CE840E7FF9A7E46FDCEFDC8FD1DFD63" authorityName="Klotzsch (1854: 123) De Candolle" authorityYear="1859" box="[565,640,627,652]" class="Magnoliopsida" family="Begoniaceae" genus="Lepsia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="173" phylum="Tracheophyta" rank="genus">
<emphasis id="799CE776FF9A7E46FDCEFDC8FD1DFD63" box="[565,640,627,652]" italics="true" pageId="4" pageNumber="173">Lepsia</emphasis>
</taxonomicName>
. The larger size of the tepals is the most distinctive feature of the new species among species in the section 
<taxonomicName id="8CE840E7FF9A7E46FDDCFD2CFD5CFD5F" box="[551,705,663,688]" pageId="4" pageNumber="173" rank="section" section="Ruizopavonia">
<emphasis id="799CE776FF9A7E46FDDCFD2CFD5CFD5F" box="[551,705,663,688]" italics="true" pageId="4" pageNumber="173">Ruizopavonia</emphasis>
</taxonomicName>
and members of section 
<taxonomicName id="8CE840E7FF9A7E46FC21FD2DFBE2FD5F" box="[986,1151,662,688]" pageId="4" pageNumber="173" rank="section" section="Cyathocmemis">
<emphasis id="799CE776FF9A7E46FC21FD2DFBE2FD5F" box="[986,1151,662,688]" italics="true" pageId="4" pageNumber="173">Cyathocmemis</emphasis>
</taxonomicName>
from the northern Andes.
</paragraph>
</subSubSection>
</treatment>
</document>