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<document id="DEA390F4C4257B4E85B13004B20BC305" ID-DOI="10.11646/zootaxa.4012.1.4" ID-GBIF-Dataset="d32dd252-5a36-4368-a636-6e688bc4ed13" ID-ISSN="1175-5326" ID-Zenodo-Dep="232879" ID-ZooBank="C3B3A06D-C18C-4B66-83E9-51B605E11862" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461226104448" checkinUser="plazi" docAuthor="Kobieluszova, Lucie &amp; Kundrata, Robin" docDate="2015" docId="C3140C0EFF8CFFF33DBFB38C5883C576" docLanguage="en" docName="zt04012p096.pdf" docOrigin="Zootaxa 4012 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drilus sanliurfensis Kobieluszova &amp; Kundrata, 2015, sp. nov." docType="treatment" docUuid="A14767A9-E162-4BB5-A4A1-E0305EFC6D67" docUuidSource="ZooBank" docVersion="9" masterDocId="3F2D7476FF82FFFD3D28B31B5D1EC058" masterDocTitle="Taxonomic review of Drilus Olivier, 1790 (Elateridae: Agrypninae: Drilini) from Asia Minor, with descriptions of seven new species and comments on the female antennal morphology in Drilini" updateTime="1698604902572" updateUser="plazi">
<mods:mods id="027E1307F317B3C0B922649852D63F84" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5E05C537FA3933B451D33B3E0B58D304">
<mods:title id="E74D3DFF33CFD9C5819E1B53D4CCA25D">Taxonomic review of Drilus Olivier, 1790 (Elateridae: Agrypninae: Drilini) from Asia Minor, with descriptions of seven new species and comments on the female antennal morphology in Drilini</mods:title>
</mods:titleInfo>
<mods:name id="AF5BDEC3C691185BA160538DE867CBF4" type="personal">
<mods:role id="005679ABD2E6F273878177AB6AB02273">
<mods:roleTerm id="7A2082035A1B98BD02168AD7D2414496">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D94A4F464D37CB95D6890A068E450EB">Kobieluszova, Lucie</mods:namePart>
</mods:name>
<mods:name id="51C4C2B89B29F3BC6EF69CE88703BC98" type="personal">
<mods:role id="B1DE62BE9633102AD421813652E9E477">
<mods:roleTerm id="DDB5E740AFF32B36F32FC439DE4CC812">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DA02F749FBE7B5F4D44EA6737615FE60">Kundrata, Robin</mods:namePart>
</mods:name>
<mods:typeOfResource id="48D4D0425FA54CA217B9955DD9755C4A">text</mods:typeOfResource>
<mods:relatedItem id="25D972667858B35E253C9479D698F54A" type="host">
<mods:titleInfo id="BD60E5C316FF57B430A4D2177CA616BF">
<mods:title id="684D27A5A2BC2EC8D5ECD3D3ABD59EBD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F256AA989D944BF8DF20650D4DB89773">
<mods:date id="134EB553DD7C58343D55B7D5E9165076">2015</mods:date>
<mods:detail id="0F44573FC88E26815D30F72095F1477F" type="volume">
<mods:number id="64C51B6C08ACFE4DC6D1DF2D8F8B1279">4012</mods:number>
</mods:detail>
<mods:detail id="6B901D1FB814D892681CCF72C38D24E4" type="issue">
<mods:number id="E972A363FD8FB8F5FFC62F8FC6A1BAE7">1</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="EE33F3266C3C804DDE8DEEFE88206CA3">journal volume</mods:classification>
<mods:identifier id="BABF99DE403812559B835D8FF7ACEC97" type="DOI">10.11646/zootaxa.4012.1.4</mods:identifier>
<mods:identifier id="12EB0CCDBE8F6C65676398E790CD3C1F" type="GBIF-Dataset">d32dd252-5a36-4368-a636-6e688bc4ed13</mods:identifier>
<mods:identifier id="1C901B44943D57776248519269EB3178" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D9DA5C870D2DB3DCB63EB85AF54C82B8" type="Zenodo-Dep">232879</mods:identifier>
<mods:identifier id="5FFA743F617CF95C408EFAEF1BD5A9B2" type="ZooBank">C3B3A06D-C18C-4B66-83E9-51B605E11862</mods:identifier>
</mods:mods>
<treatment id="C3140C0EFF8CFFF33DBFB38C5883C576" ID-DOI="http://doi.org/10.5281/zenodo.6112436" ID-GBIF-Taxon="119632734" ID-Zenodo-Dep="6112436" LSID="urn:lsid:zoobank.org:act:A14767A9-E162-4BB5-A4A1-E0305EFC6D67" httpUri="http://treatment.plazi.org/id/C3140C0EFF8CFFF33DBFB38C5883C576" pageId="14">
<subSubSection id="03A7EE93FF8CFFF33DBFB38C5C72C08B" pageId="14" type="nomenclature">
<paragraph id="4B02BD18FF8CFFF33DBFB38C5CF2C0EA" blockId="14.[151,492,151,211]" box="[151,492,151,178]" pageId="14">
<heading id="104A0A74FF8CFFF33DBFB38C5CF2C0EA" bold="true" box="[151,492,151,178]" fontSize="11" level="1" pageId="14" reason="1">
<emphasis id="79C9610AFF8CFFF33DBFB38C5CF2C0EA" bold="true" box="[151,492,151,178]" pageId="14">
<taxonomicName id="8CBDC69BFF8CFFF33DBFB38C5C99C0E9" box="[151,391,151,177]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="species" species="sanliurfensis" status="sp. nov.">
<emphasis id="79C9610AFF8CFFF33DBFB38C5C99C0E9" bold="true" box="[151,391,151,177]" italics="true" pageId="14">Drilus sanliurfensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="62FADC71FF8CFFF33CA6B3835CF2C0EA" box="[398,492,152,178]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4B02BD18FF8CFFF33DBFB3A15C72C08B" blockId="14.[151,492,151,211]" box="[151,364,186,211]" pageId="14">
(
<figureCitation id="D386A19DFF8CFFF33DB7B3A15DF9C08B" box="[159,231,186,211]" captionStart="FIGURES 1 – 12" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[181,1396,203,1850]" captionTargetId="figure@2.[172,1408,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/232880/files/figure.png" pageId="14">Figs 9</figureCitation>
,
<figureCitation id="D386A19DFF8CFFF33DDBB3A05C0FC08B" box="[243,273,187,211]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="14">19</figureCitation>
,
<figureCitation id="D386A19DFF8CFFF33C35B3A05C25C08B" box="[285,315,187,211]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="14">29</figureCitation>
,
<figureCitation id="D386A19DFF8CFFF33C6FB3A05C7AC08B" box="[327,356,187,211]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="14">41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8CFFF33DBFB21959DFC1DF" pageId="14" type="materials_examined">
<paragraph id="4B02BD18FF8CFFF33DBFB2195C19C13B" blockId="14.[151,1437,258,1327]" pageId="14">
<emphasis id="79C9610AFF8CFFF33DBFB2195C56C143" bold="true" box="[151,328,258,283]" pageId="14">
<typeStatus id="940603BAFF8CFFF33DBFB2195DCCC143" box="[151,210,258,283]" pageId="14">Type</typeStatus>
material.
</emphasis>
<typeStatus id="940603BAFF8CFFF33C7BB2195CA0C143" box="[339,446,258,283]" pageId="14" type="holotype">Holotype</typeStatus>
, male, &quot;
<collectingCountry id="33AAFD88FF8CFFF33F36B2195F6CC143" box="[542,626,258,283]" name="Turkey" pageId="14">Turkey</collectingCountry>
, Prov. Urfa, Halfeti, valley of Euphrat,
<quantity id="8C4510FDFF8CFFF3396FB219598AC142" box="[1095,1172,258,283]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="14" unit="m" value="500.0">500 m</quantity>
,
<geoCoordinate id="2E89DBDFFF8CFFF3398BB219580BC143" box="[1187,1301,258,283]" direction="east" orientation="longitude" pageId="14" precision="92" value="37.875">37°52.5'E</geoCoordinate>
,
<geoCoordinate id="2E89DBDFFF8CFFF3380CB2195886C143" box="[1316,1432,258,283]" direction="north" orientation="latitude" pageId="14" precision="92" value="37.241665">37°14.5'N</geoCoordinate>
,
<date id="3F039BD8FF8CFFF33DBFB23C5C52C167" box="[151,332,295,319]" pageId="14" value="1989-04-28" valueMax="1989-05-01" valueMin="1989-04-28">28.IV–1.V.1989</date>
, leg. Fabián, Ronkay &amp; Ronkay&quot; (
<collectionCode id="2DAC25DDFF8CFFF33FFDB23C5E2FC167" LSID="urn:lsid:biocol.org:col:34991" box="[725,817,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="14">HNHM</collectionCode>
); 3
<typeStatus id="940603BAFF8CFFF33E73B23C5ED7C167" box="[859,969,295,319]" pageId="14" type="paratype">paratypes</typeStatus>
, males, the same data as for the
<typeStatus id="940603BAFF8CFFF33812B23D5882C167" box="[1338,1436,294,319]" pageId="14" type="holotype">holotype</typeStatus>
(
<collectionCode id="2DAC25DDFF8CFFF33DB7B2505DE5C13B" LSID="urn:lsid:biocol.org:col:34991" box="[159,251,331,355]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="14">HNHM</collectionCode>
).
</paragraph>
<paragraph id="4B02BD18FF8CFFF33DEFB27559DFC1DF" blockId="14.[151,1437,258,1327]" box="[199,1217,366,391]" pageId="14">
<emphasis id="79C9610AFF8CFFF33DEFB2755C7BC1DF" bold="true" box="[199,357,366,391]" pageId="14">
<typeStatus id="940603BAFF8CFFF33DEFB2755C1CC1DF" box="[199,258,366,391]" pageId="14">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="33AAFD88FF8CFFF33C44B2755CA3C1DF" box="[364,445,366,391]" name="Turkey" pageId="14">Turkey</collectingCountry>
, Prov. Urfa, Halfeti, valley of Euphrat,
<quantity id="8C4510FDFF8CFFF33E5FB2755EA1C1DE" box="[887,959,366,391]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="14" unit="m" value="500.0">500 m</quantity>
,
<geoCoordinate id="2E89DBDFFF8CFFF33EE3B2755921C1DF" box="[971,1087,366,391]" direction="north" orientation="latitude" pageId="14" precision="92" value="37.241665">37°14.5'N</geoCoordinate>
,
<geoCoordinate id="2E89DBDFFF8CFFF33963B27559A3C1DF" box="[1099,1213,366,391]" direction="east" orientation="longitude" pageId="14" precision="92" value="37.875">37°52.5'E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8CFFF33DEFB2895EE1C197" pageId="14" type="diagnosis">
<paragraph id="4B02BD18FF8CFFF33DEFB2895EE1C197" blockId="14.[151,1437,258,1327]" pageId="14">
<emphasis id="79C9610AFF8CFFF33DEFB2895C5FC1F3" bold="true" box="[199,321,402,427]" pageId="14">Diagnosis.</emphasis>
This species is similar in morphology and coloration to
<taxonomicName id="8CBDC69BFF8CFFF33EE4B2885959C1F3" box="[972,1095,403,427]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="79C9610AFF8CFFF33EE4B2885EFEC1F2" box="[972,992,403,426]" italics="true" pageId="14">D</emphasis>
.
<emphasis id="79C9610AFF8CFFF33ED8B2885959C1F3" box="[1008,1095,403,427]" italics="true" pageId="14">mertliki</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF8CFFF33978B28959B2C1F3" bold="true" box="[1104,1196,402,427]" pageId="14">
<taxonomicNameLabel id="62FADC71FF8CFFF33978B28959B2C1F3" box="[1104,1196,402,427]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but differs from the latter by the shape of pronotum (
<figureCitation id="D386A19DFF8CFFF33F2CB2AD5F9AC197" box="[516,644,438,463]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="14">Figs 16, 19</figureCitation>
) and male genitalia (
<figureCitation id="D386A19DFF8CFFF33E58B2AD5EEFC197" box="[880,1009,438,463]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="14">Figs 37, 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8CFFF33DEFB2C15ED2C4BE" pageId="14" type="description">
<paragraph id="4B02BD18FF8CFFF33DEFB2C1590DC423" blockId="14.[151,1437,258,1327]" pageId="14">
<emphasis id="79C9610AFF8CFFF33DEFB2C15C47C1AB" bold="true" box="[199,345,474,499]" pageId="14">Description.</emphasis>
Male (
<typeStatus id="940603BAFF8CFFF33C8EB2C15F10C1AB" box="[422,526,474,499]" pageId="14" type="holotype">holotype</typeStatus>
). Body 3.4 times longer than width at humeri. Head black, elytra and ventral parts dark brown, pronotum and scutellum lighter, antennomeres 3–11 brown, mouthparts, scape, pedicel, legs and last abdominal ventrite yellowish brown; body covered with yellowish to light brown pubescence (
<figureCitation id="D386A19DFF8CFFF339D5B1395858C263" box="[1277,1350,546,571]" captionStart="FIGURES 1 – 12" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[181,1396,203,1850]" captionTargetId="figure@2.[172,1408,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/232880/files/figure.png" pageId="14">Fig. 9</figureCitation>
). Head including eyes about as wide as anterior margin of pronotum (
<figureCitation id="D386A19DFF8CFFF33E45B15D5EDDC207" box="[877,963,582,607]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="14">Fig. 19</figureCitation>
), dorsally with wide shallow depression, lateral margins above antennal sockets elevated, covered with large shallow punctures, with sparse semierect or erect pubescence; clypeal margin widely concave. Eyes medium-sized, their frontal distance 1.9 times eye diameter. Mandibles long, curved; incisor margin with short robust tooth in middle part. Labrum transverse; maxillary palpi slender, apical palpomere obliquely widely cut. Antennae pectinate, reaching one third of elytral length, antennomere 3 serrate, antennomeres 4–10 pectinate, apical antennomere simple, longest (
<figureCitation id="D386A19DFF8CFFF3381CB1E1588EC34B" box="[1332,1424,762,787]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="14">Fig. 29</figureCitation>
). Pronotum slightly convex, transverse, widest at middle, 1.5 times wider posteriorly than length at midline. Anterior margin almost straight, lateral margins sinuate, posterior margin convex, bisinuate. Both anterior and posterior angles rectangular (
<figureCitation id="D386A19DFF8CFFF33C50B07D5CD3C327" box="[376,461,870,895]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="14">Fig. 19</figureCitation>
); surface of disc sparsely punctured, with sparse erected setae, pubescence denser at angles. Prosternum transverse, slightly wrinkled, frontal margin almost straight, prosternal process minute, gradually declivitous. Mesoventrite v-shaped, anteromedially depressed. Elytra elongate, subparallel-sided, with longitudinal keel running from humeri toward apex, inconspicuous at apical 1/3 of elytra, slightly wrinkled, sparsely punctured, covered with semierect pubescence. Legs slender, slightly compressed, with sparse, long, semierect setae, tarsomeres 1–4 gradually shortened, tarsomere 4 shortest, apical tarsomere long, slender, slightly longer than combined lengths of tarsomeres 3 and 4. Aedeagus with robust, considerably curved phallus; paramera long; phallobase v-shaped (
<figureCitation id="D386A19DFF8CFFF33CE3B7795F00C423" box="[459,542,1122,1147]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="14">Fig. 41</figureCitation>
). Female and immature stages are unknown.
</paragraph>
<paragraph id="4B02BD18FF8CFFF33DEFB79D5ED2C4BE" blockId="14.[151,1437,258,1327]" pageId="14">
<emphasis id="79C9610AFF8CFFF33DEFB79D5C63C4C7" bold="true" box="[199,381,1158,1183]" pageId="14">Measurements.</emphasis>
<typeStatus id="940603BAFF8CFFF33CACB79D5CEEC4C7" box="[388,496,1158,1183]" pageId="14" type="holotype">Holotype</typeStatus>
. BL
<quantity id="8C4510FDFF8CFFF33F0EB79D5F61C4C6" box="[550,639,1158,1183]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="14" unit="mm" value="5.5">5.5 mm</quantity>
, EL
<quantity id="8C4510FDFF8CFFF33F9DB79C5E10C4C6" box="[693,782,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="14" unit="mm" value="4.1">4.1 mm</quantity>
, WHe
<quantity id="8C4510FDFF8CFFF33E75B79C5EABC4C6" box="[861,949,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="14" unit="mm" value="1.2">1.2 mm</quantity>
, WHum
<quantity id="8C4510FDFF8CFFF33933B79D596AC4C6" box="[1051,1140,1158,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="14" unit="mm" value="1.6">1.6 mm</quantity>
, PL
<quantity id="8C4510FDFF8CFFF33980B79C581FC4C6" box="[1192,1281,1159,1183]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="14" unit="mm" value="0.9">0.9 mm</quantity>
, PW
<quantity id="8C4510FDFF8CFFF33817B79C5886C4C6" box="[1343,1432,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="14" unit="mm" value="1.3">1.3 mm</quantity>
, Edist
<quantity id="8C4510FDFF8CFFF33DFFB7B05C30C49A" box="[215,302,1195,1219]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="14" unit="mm" value="0.7">0.7 mm</quantity>
, Ediam
<quantity id="8C4510FDFF8CFFF33CA1B7B05CFCC49A" box="[393,482,1195,1219]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" unit="mm" value="0.4">0.4 mm</quantity>
.
<typeStatus id="940603BAFF8CFFF33CC4B7B05F43C49B" box="[492,605,1195,1219]" pageId="14" type="paratype">Paratypes</typeStatus>
. BL
<quantity id="8C4510FDFF8CFFF33FB9B7B15E07C49A" box="[657,793,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" metricValueMax="6.4" metricValueMin="4.5" pageId="14" unit="mm" value="5.45" valueMax="6.4" valueMin="4.5">4.5–6.4 mm</quantity>
, EL
<quantity id="8C4510FDFF8CFFF33E65B7B15EC8C49A" box="[845,982,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.8" metricValueMin="3.5" pageId="14" unit="mm" value="4.15" valueMax="4.8" valueMin="3.5">3.5–4.8 mm</quantity>
, WHe 1.0–
<quantity id="8C4510FDFF8CFFF3397CB7B059B5C49A" box="[1108,1195,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="14" unit="mm" value="1.4">1.4 mm</quantity>
, WHum 1.3–2.0 mm, PL 0.7–1.0 mm, PW
<quantity id="8C4510FDFF8CFFF33CACB7D55F13C4BE" box="[388,525,1230,1255]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="14" unit="mm" value="1.35" valueMax="1.6" valueMin="1.1">1.1–1.6 mm</quantity>
, Edist
<quantity id="8C4510FDFF8CFFF33F71B7D55FFCC4BE" box="[601,738,1230,1255]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="14" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
, Ediam
<quantity id="8C4510FDFF8CFFF33E16B7D55ED6C4BE" box="[830,968,1230,1255]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="14" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8CFFF33DEFB7E95E19C553" box="[199,775,1266,1291]" pageId="14" type="distribution">
<paragraph id="4B02BD18FF8CFFF33DEFB7E95E19C553" blockId="14.[151,1437,258,1327]" box="[199,775,1266,1291]" pageId="14">
<emphasis id="79C9610AFF8CFFF33DEFB7E95C7EC553" bold="true" box="[199,352,1266,1291]" pageId="14">Distribution.</emphasis>
<collectingCountry id="33AAFD88FF8CFFF33C4FB7E95CA9C553" box="[359,439,1266,1291]" name="Turkey" pageId="14">Turkey</collectingCountry>
(Şanlıurfa Province;
<figureCitation id="D386A19DFF8CFFF33F8FB7E95FE4C553" box="[679,762,1266,1291]" captionStart="FIGURE 55" captionStartId="12.[151,250,1911,1933]" captionTargetBox="[160,1421,963,1875]" captionTargetId="figure@12.[151,1436,951,1889]" captionTargetPageId="12" captionText="FIGURE 55. The distribution of Drilus species in Asia Minor." httpUri="https://zenodo.org/record/232886/files/figure.png" pageId="14">Fig. 55</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8CFFF33DEFB60D5883C576" box="[199,1437,1302,1327]" pageId="14" type="etymology">
<paragraph id="4B02BD18FF8CFFF33DEFB60D5883C576" blockId="14.[151,1437,258,1327]" box="[199,1437,1302,1327]" pageId="14">
<emphasis id="79C9610AFF8CFFF33DEFB60D5C50C577" bold="true" box="[199,334,1302,1327]" pageId="14">Etymology.</emphasis>
The specific epithet is derived from the name of the Turkish province in which this species occurs.
</paragraph>
</subSubSection>
</treatment>
</document>
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