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<document id="17B43C4807333235F413ED0CC5FE885C" ID-DOI="10.11646/zootaxa.4012.1.4" ID-GBIF-Dataset="d32dd252-5a36-4368-a636-6e688bc4ed13" ID-ISSN="1175-5326" ID-Zenodo-Dep="232879" ID-ZooBank="C3B3A06D-C18C-4B66-83E9-51B605E11862" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461226104448" checkinUser="plazi" docAuthor="Kobieluszova, Lucie &amp; Kundrata, Robin" docDate="2015" docId="C3140C0EFF8BFFF03DBFB5FE5E8EC21C" docLanguage="en" docName="zt04012p096.pdf" docOrigin="Zootaxa 4012 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drilus robustus Kobieluszova &amp; Kundrata, 2015, sp. nov." docType="treatment" docVersion="8" masterDocId="3F2D7476FF82FFFD3D28B31B5D1EC058" masterDocTitle="Taxonomic review of Drilus Olivier, 1790 (Elateridae: Agrypninae: Drilini) from Asia Minor, with descriptions of seven new species and comments on the female antennal morphology in Drilini" updateTime="1698604902572" updateUser="plazi">
<mods:mods id="04CB458CD4FE5225E585E7EB6702160E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E9961A0176ED7A04FAE41B7B5EDA9A7D">
<mods:title id="9D73B6BDD5E745319ED0951B304E834B">Taxonomic review of Drilus Olivier, 1790 (Elateridae: Agrypninae: Drilini) from Asia Minor, with descriptions of seven new species and comments on the female antennal morphology in Drilini</mods:title>
</mods:titleInfo>
<mods:name id="9F577DCF76F32D2D48C80C26BF03A2E9" type="personal">
<mods:role id="258B8E4F25335C3C952901AB9389F992">
<mods:roleTerm id="F077859179F541A329E6086A6A4AF498">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF766F2DF4D04B152B5F72B8A5040CAE">Kobieluszova, Lucie</mods:namePart>
</mods:name>
<mods:name id="79CEFBA9DB0862E42C37DA60CF976BE6" type="personal">
<mods:role id="F9406FFD02AF56FA5221B4BCEA49CC8D">
<mods:roleTerm id="E6CF2242034818C1E1046B8694DE1224">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05B5C23802D2DA7B7CEB412987113614">Kundrata, Robin</mods:namePart>
</mods:name>
<mods:typeOfResource id="1BAB1E7D7B8F6AA43A989D8CFC5C01EC">text</mods:typeOfResource>
<mods:relatedItem id="20E591A5E3E8D26B05BD3EA4DA12B934" type="host">
<mods:titleInfo id="857CC47E173D285FFB2A2E4260344536">
<mods:title id="8521FD3EBB87BCCD14754CEA269FAD3E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F552C561A519417EE9EFB7AE0154A631">
<mods:date id="95694A7266ED52C85EE34D55281DCC83">2015</mods:date>
<mods:detail id="AB42800FD33DD41EC3721D45A6B7ABE5" type="volume">
<mods:number id="A251B3D12D60E69E56969906AC4D1037">4012</mods:number>
</mods:detail>
<mods:detail id="7661F5FC3324DC421A11F745C9F73B94" type="issue">
<mods:number id="EE597414F11F6EA9995555B39169858F">1</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="0EB8A4C18ABBACB49BB2CD334E3397E0">journal volume</mods:classification>
<mods:identifier id="B6B4EE94BBE18659B94DDA19C512F52E" type="DOI">10.11646/zootaxa.4012.1.4</mods:identifier>
<mods:identifier id="0E2245A5DF979A6971B13084FE4CB6B4" type="GBIF-Dataset">d32dd252-5a36-4368-a636-6e688bc4ed13</mods:identifier>
<mods:identifier id="458D161BB3F2EE39B1FD5311211675D3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2D2FB402E92F835FAFFF08644F7226F7" type="Zenodo-Dep">232879</mods:identifier>
<mods:identifier id="52B5731EC6D953F49EC51A4F430C4346" type="ZooBank">C3B3A06D-C18C-4B66-83E9-51B605E11862</mods:identifier>
</mods:mods>
<treatment id="C3140C0EFF8BFFF03DBFB5FE5E8EC21C" ID-DOI="http://doi.org/10.5281/zenodo.6112434" ID-GBIF-Taxon="119632738" ID-Zenodo-Dep="6112434" LSID="urn:lsid:plazi:treatment:C3140C0EFF8BFFF03DBFB5FE5E8EC21C" httpUri="http://treatment.plazi.org/id/C3140C0EFF8BFFF03DBFB5FE5E8EC21C" lastPageId="13" pageId="9">
<subSubSection id="03A7EE93FF8BFFF43DBFB5FE5F8DC778" pageId="9" type="nomenclature">
<paragraph id="4B02BD18FF8BFFF43DBFB5FE5CA6C6A7" blockId="9.[151,659,1765,1824]" box="[151,440,1765,1791]" pageId="9">
<heading id="104A0A74FF8BFFF43DBFB5FE5CA6C6A7" bold="true" box="[151,440,1765,1791]" fontSize="11" level="1" pageId="9" reason="1">
<emphasis id="79C9610AFF8BFFF43DBFB5FE5CA6C6A7" bold="true" box="[151,440,1765,1791]" pageId="9">
<taxonomicName id="8CBDC69BFF8BFFF43DBFB5FE5C4DC6A7" box="[151,339,1765,1791]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="79C9610AFF8BFFF43DBFB5FE5C4DC6A7" bold="true" box="[151,339,1765,1791]" italics="true" pageId="9">Drilus robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="62FADC71FF8BFFF43C72B5FE5CA6C6A7" box="[346,440,1765,1791]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4B02BD18FF8BFFF43DBFB41C5F8DC778" blockId="9.[151,659,1765,1824]" box="[151,659,1799,1824]" pageId="9">
(
<figureCitation id="D386A19DFF8BFFF43DB7B41C5DF9C778" box="[159,231,1799,1824]" captionStart="FIGURES 1 – 12" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[181,1396,203,1850]" captionTargetId="figure@2.[172,1408,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/232880/files/figure.png" pageId="9">Figs 8</figureCitation>
,
<figureCitation id="D386A19DFF8BFFF43DDBB4135C0FC778" box="[243,273,1800,1824]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="9">18</figureCitation>
,
<figureCitation id="D386A19DFF8BFFF43C35B4135C8EC778" box="[285,400,1800,1824]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="9">28, 32–33</figureCitation>
,
<figureCitation id="D386A19DFF8BFFF43CB3B4135CA4C778" box="[411,442,1800,1824]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="9">40</figureCitation>
,
<figureCitation id="D386A19DFF8BFFF43CEDB41C5F11C778" box="[453,527,1799,1824]" captionStart="FIGURES 46 – 50" captionStartId="10.[151,264,1240,1263]" captionTargetBox="[168,1420,385,1218]" captionTargetId="figure@10.[151,1436,373,1220]" captionTargetPageId="10" captionText="FIGURES 46 – 50. Drilus robustus sp. nov., female. 46, habitus; 47, genitalia; 48, antenna. Larval cerci. 49, D. robustus sp. nov.; 50, Drilus sp. (Turkey, Muğla Prov., Knidos). Scale bars = 2.0 mm (Fig. 46), 1.0 mm (Figs 47, 49 – 50), 0.5 mm (Fig. 48)." httpUri="https://zenodo.org/record/232884/files/figure.png" pageId="9">46–49</figureCitation>
,
<figureCitation id="D386A19DFF8BFFF43F31B41C5F95C778" box="[537,651,1799,1824]" captionStart="FIGURES 51 – 54" captionStartId="11.[151,264,1732,1755]" captionTargetBox="[151,1436,517,1711]" captionTargetId="figure@11.[151,1436,517,1711]" captionTargetPageId="11" captionText="FIGURES 51 – 54. Drilus robustus sp. nov. 51, larva; 52, mating couple; 53, the type locality (Turkey, Kemer env.); 54, larval skins inside the shells of Rumina decollata (Linnaeus, 1758)." httpUri="https://zenodo.org/record/232885/files/figure.png" pageId="9">51–52, 54</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8BFFF73DBFB4545926C118" lastPageId="10" pageId="9" type="materials_examined">
<paragraph id="4B02BD18FF8BFFF43DBFB4545FF7C73F" blockId="9.[151,1437,1871,2005]" box="[151,745,1871,1896]" pageId="9">
<emphasis id="79C9610AFF8BFFF43DBFB4545C2BC730" bold="true" box="[151,309,1871,1896]" pageId="9">
<typeStatus id="940603BAFF8BFFF43DBFB4545DCCC730" box="[151,210,1871,1896]" pageId="9">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="33AAFD88FF8BFFF43C14B4545C93C730" box="[316,397,1871,1896]" name="Turkey" pageId="9">Turkey</collectingCountry>
, Antalya province, Kemer env.
</paragraph>
<paragraph id="4B02BD18FF8BFFF73DEFB46F5876C0A0" blockId="9.[151,1437,1871,2005]" lastBlockId="10.[151,1436,151,320]" lastPageId="10" pageId="9">
<emphasis id="79C9610AFF8BFFF43DEFB46F5C66C7D5" bold="true" box="[199,376,1908,1933]" pageId="9">
<typeStatus id="940603BAFF8BFFF43DEFB46F5C1CC7D5" box="[199,258,1908,1933]" pageId="9">Type</typeStatus>
material.
</emphasis>
<typeStatus id="940603BAFF8BFFF43CABB46F5CF0C7D5" box="[387,494,1908,1933]" pageId="9" type="holotype">Holotype</typeStatus>
, male: &quot;
<collectingCountry id="33AAFD88FF8BFFF43F66B46F5FBFC7D5" box="[590,673,1908,1933]" name="Turkey" pageId="9">Turkey</collectingCountry>
mer. occ., Kemer env., 1.–
<date id="3F039BD8FF8BFFF43EF1B46F594EC7D5" box="[985,1104,1908,1933]" pageId="9" value="2011-11-06">11.6.2011</date>
, J. Bednář lgt.&quot; (
<collectionCode id="2DAC25DDFF8BFFF43832B46F5874C7D4" box="[1306,1386,1908,1932]" pageId="9">PCRK</collectionCode>
); 81
<typeStatus id="940603BAFF8BFFF43DBFB4825C19C7E9" box="[151,263,1945,1969]" pageId="9" type="paratype">paratypes</typeStatus>
:
<specimenCount id="5DBB7691FF8BFFF43C39B4835C75C7E9" box="[273,363,1944,1969]" pageId="9" type="male">5 males</specimenCount>
, the same data as for the
<typeStatus id="940603BAFF8BFFF43FA1B4835FF5C7E9" box="[649,747,1944,1969]" pageId="9" type="holotype">holotype</typeStatus>
(
<collectionCode id="2DAC25DDFF8BFFF43FD2B4835E20C7E8" box="[762,830,1944,1968]" pageId="9">PCJB</collectionCode>
,
<collectionCode id="2DAC25DDFF8BFFF43E62B4835E86C7E8" box="[842,920,1944,1968]" pageId="9">PCRK</collectionCode>
);
<specimenCount id="5DBB7691FF8BFFF43E84B483591BC7E9" box="[940,1029,1944,1969]" pageId="9" type="male">4 males</specimenCount>
, &quot;
<collectingCountry id="33AAFD88FF8BFFF43932B4835973C7E9" box="[1050,1133,1944,1969]" name="Turkey" pageId="9">Turkey</collectingCountry>
mer. occ., Kemer env., 1.–
<date id="3F039BD8FF8BFFF43DBFB4A75C16C78D" box="[151,264,1980,2005]" pageId="9" value="2011-11-06">11.6.2011</date>
, J. Brádka lgt.&quot; (
<collectionCode id="2DAC25DDFF8BFFF43CEEB4A75F12C78C" box="[454,524,1980,2004]" pageId="9">PCJR</collectionCode>
);
<specimenCount id="5DBB7691FF8BFFF43F08B4A75F67C78D" box="[544,633,1980,2005]" pageId="9" type="male">3 males</specimenCount>
, &quot;
<collectingCountry id="33AAFD88FF8BFFF43FA6B4A75FFFC78D" box="[654,737,1980,2005]" name="Turkey" pageId="9">Turkey</collectingCountry>
mer. occ., Kemer env., 1.–
<date id="3F039BD8FF8BFFF43923B4A7599CC78D" box="[1035,1154,1980,2005]" pageId="9" value="2011-11-06">11.6.2011</date>
, J. Januš lgt.&quot; (
<collectionCode id="2DAC25DDFF8BFFF43806B4A75872C78C" box="[1326,1388,1980,2004]" pageId="9">PCJJ</collectionCode>
);
<specimenCount id="5DBB7691FF8BFFF738A8B4A75DC5C0E8" lastPageId="10" pageId="9" type="male">62 males</specimenCount>
,
<specimenCount id="5DBB7691FF88FFF73DC3B3835C4DC0E8" box="[235,339,151,176]" pageId="10" type="female">1 female</specimenCount>
, &quot;
<collectingCountry id="33AAFD88FF88FFF73C43B38C5CA1C0E8" box="[363,447,151,176]" name="Turkey" pageId="10">Turkey</collectingCountry>
, Antalya pr., Kemer env.,
<quantity id="8C4510FDFF88FFF73FD1B3835E33C0E8" box="[761,813,152,176]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" unit="m" value="10.0">10m</quantity>
, 26.–
<date id="3F039BD8FF88FFF73E44B38C5EFCC0E8" box="[876,994,151,176]" pageId="10" value="2012-05-28">28.5.2012</date>
, Krol, Fusek &amp; Masek leg.&quot; (
<collectionCode id="2DAC25DDFF88FFF7386CB3835886C0E8" box="[1348,1432,152,176]" pageId="10">LMBC</collectionCode>
,
<collectionCode id="2DAC25DDFF88FFF73DBFB3A75DFBC08C" box="[151,229,188,212]" pageId="10">PCRK</collectionCode>
);
<specimenCount id="5DBB7691FF88FFF73DD0B3A75C4EC08D" box="[248,336,188,213]" pageId="10" type="male">5 males</specimenCount>
, &quot;
<collectingCountry id="33AAFD88FF88FFF73C4BB3A75CA9C08D" box="[355,439,188,213]" name="Turkey" pageId="10">Turkey</collectingCountry>
, Antalya pr., Kemer env.,
<quantity id="8C4510FDFF88FFF73FF1B3A75E29C08D" box="[729,823,188,213]" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="10" unit="m" value="30.0" valueMax="50.0" valueMin="10.0">10–50m</quantity>
, 26.–
<date id="3F039BD8FF88FFF73E59B3A75EF9C08D" box="[881,999,188,213]" pageId="10" value="2012-05-28">28.5.2012</date>
, Krol, Fusek &amp; Masek leg.&quot; (
<collectionCode id="2DAC25DDFF88FFF73804B3A75862C08C" box="[1324,1404,188,212]" pageId="10">PCRK</collectionCode>
);
<specimenCount id="5DBB7691FF88FFF738A6B3A75DCEC0A0" pageId="10" type="male">1 male</specimenCount>
, &quot;
<collectingCountry id="33AAFD88FF88FFF73DCCB3C45C26C0A0" box="[228,312,223,248]" name="Turkey" pageId="10">Turkey</collectingCountry>
, Antalya pr., Kemer env.,
<quantity id="8C4510FDFF88FFF73F74B3C45F8EC0A0" box="[604,656,223,248]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" unit="m" value="50.0">50m</quantity>
, forest, in flight,
<date id="3F039BD8FF88FFF73E79B3C45EDAC0A0" box="[849,964,223,248]" pageId="10" value="2012-05-28">28.5.2012</date>
, Krol, Fusek &amp; Masek leg.&quot; (
<collectionCode id="2DAC25DDFF88FFF73824B3FB5845C0A0" box="[1292,1371,224,248]" pageId="10">PCRK</collectionCode>
).
</paragraph>
<paragraph id="4B02BD18FF88FFF73DEFB21F5926C118" blockId="10.[151,1436,151,320]" pageId="10">
<emphasis id="79C9610AFF88FFF73DEFB21F5CF9C145" bold="true" box="[199,487,260,285]" pageId="10">Other material studied.</emphasis>
<specimenCount id="5DBB7691FF88FFF73CDAB21F5F5AC145" box="[498,580,260,285]" pageId="10" type="male">1 male</specimenCount>
, &quot;
<collectingCountry id="33AAFD88FF88FFF73F74B21F5FB2C145" box="[604,684,260,285]" name="Turkey" pageId="10">Türkei</collectingCountry>
, Catallar,
<date id="3F039BD8FF88FFF73E0DB21F5E87C145" box="[805,921,260,285]" pageId="10" value="1997-06-06">6.VI.1997</date>
, leg. H. Schmid&quot; (
<collectionCode id="2DAC25DDFF88FFF7395FB21F59DCC144" box="[1143,1218,260,284]" pageId="10">PCRS</collectionCode>
); 1 larva, &quot;
<collectingCountry id="33AAFD88FF88FFF7386CB21F5886C145" box="[1348,1432,260,285]" name="Turkey" pageId="10">Turkey</collectingCountry>
, Antalya pr., Kemer env.,
<quantity id="8C4510FDFF88FFF73C98B2335CFAC118" box="[432,484,296,320]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" unit="m" value="10.0">10m</quantity>
, 26.–
<date id="3F039BD8FF88FFF73F36B23C5F8AC118" box="[542,660,295,320]" pageId="10" value="2012-05-28">28.5.2012</date>
, Krol, Fusek &amp; Masek leg.&quot; (
<collectionCode id="2DAC25DDFF88FFF73EF4B2335932C118" box="[988,1068,296,320]" pageId="10">PCRK</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="1FC2ED90FF88FFF73DBFB7C35884C556" httpUri="https://zenodo.org/record/232884/files/figure.png" pageId="10" targetBox="[168,1420,385,1218]" targetPageId="10">
<paragraph id="4B02BD18FF88FFF73DBFB7C35884C556" blockId="10.[151,1436,1240,1294]" pageId="10">
<emphasis id="79C9610AFF88FFF73DBFB7C35C48C4B6" bold="true" box="[151,342,1240,1263]" pageId="10">FIGURES 46–50.</emphasis>
<taxonomicName id="8CBDC69BFF88FFF73C75B7C25CE2C4B6" box="[349,508,1241,1262]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="79C9610AFF88FFF73C75B7C25CE2C4B6" box="[349,508,1241,1262]" italics="true" pageId="10">Drilus robustus</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF88FFF73F2CB7C25F49C4B6" bold="true" box="[516,599,1241,1262]" pageId="10">
<taxonomicNameLabel id="62FADC71FF88FFF73F2CB7C25F49C4B6" box="[516,599,1241,1262]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 46, habitus; 47, genitalia; 48, antenna. Larval cerci. 49,
<taxonomicName id="8CBDC69BFF88FFF739D6B7C1586BC4B6" box="[1278,1397,1241,1263]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="79C9610AFF88FFF739D6B7C1580EC4B7" box="[1278,1296,1242,1263]" italics="true" pageId="10">D</emphasis>
.
<emphasis id="79C9610AFF88FFF73837B7C2586BC4B6" box="[1311,1397,1241,1262]" italics="true" pageId="10">robustus</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF88FFF73855B7C25DDDC555" bold="true" pageId="10">
<taxonomicNameLabel id="62FADC71FF88FFF73855B7C25DDDC555" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 50,
<taxonomicName id="8CBDC69BFF88FFF73DDEB7E35C29C555" box="[246,311,1272,1293]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="genus">
<emphasis id="79C9610AFF88FFF73DDEB7E35C29C555" box="[246,311,1272,1293]" italics="true" pageId="10">Drilus</emphasis>
</taxonomicName>
sp. (Turkey, Muğla Prov., Knidos). Scale bars = 2.0 mm (Fig. 46), 1.0 mm (Figs 47, 49–50), 0.5 mm (Fig. 48).
</paragraph>
</caption>
<subSubSection id="03A7EE93FF88FFF73DEFB6215CD3C673" pageId="10" type="diagnosis">
<paragraph id="4B02BD18FF88FFF73DEFB6215CD3C673" blockId="10.[151,1437,1338,2011]" pageId="10">
<emphasis id="79C9610AFF88FFF73DEFB6215C5FC50B" bold="true" box="[199,321,1338,1363]" pageId="10">Diagnosis.</emphasis>
The male of
<taxonomicName id="8CBDC69BFF88FFF73CCBB6205F8AC50B" box="[483,660,1339,1363]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="79C9610AFF88FFF73CCBB6205F8AC50B" box="[483,660,1339,1363]" italics="true" pageId="10">Drilus robustus</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF88FFF73FB7B6215FE2C50B" bold="true" box="[671,764,1338,1363]" pageId="10">
<taxonomicNameLabel id="62FADC71FF88FFF73FB7B6215FE2C50B" box="[671,764,1338,1363]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily distinguished from its congeners from
<collectingCountry id="33AAFD88FF88FFF73841B6215DC3C52F" name="Turkey" pageId="10">Asia Minor</collectingCountry>
by the following combination of characters: robust general habitus, reddish brown body coloration with bicolored elytra (
<figureCitation id="D386A19DFF88FFF73C48B6995CB6C5C3" box="[352,424,1410,1435]" captionStart="FIGURES 1 – 12" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[181,1396,203,1850]" captionTargetId="figure@2.[172,1408,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/232880/files/figure.png" pageId="10">Fig. 8</figureCitation>
), strongly pectinate antennnae (
<figureCitation id="D386A19DFF88FFF73E09B6995E67C5C3" box="[801,889,1410,1435]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="10">Fig. 28</figureCitation>
), and pronotum widest slightly before middle, constricted anteriorly (
<figureCitation id="D386A19DFF88FFF73CB2B6BD5CEEC5E7" box="[410,496,1446,1471]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="10">Fig. 18</figureCitation>
). This species is very similar to
<taxonomicName id="8CBDC69BFF88FFF73E42B6BC598FC5E7" authority="Olivier, 1913" authorityName="Olivier" authorityYear="1913" box="[874,1169,1446,1471]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="bleusei">
<emphasis id="79C9610AFF88FFF73E42B6BC5E60C5E6" box="[874,894,1447,1470]" italics="true" pageId="10">D</emphasis>
.
<emphasis id="79C9610AFF88FFF73EA6B6BD5EC3C5E6" box="[910,989,1446,1470]" italics="true" pageId="10">bleusei</emphasis>
(
<bibRefCitation id="2F2CC0E9FF88FFF73EC7B6BD5996C5E7" author="Olivier" box="[1007,1160,1446,1471]" pageId="10" refString="Olivier, E. (1913) Description d'une nouvelle espece de Drilidae [Col.]. Bulletin de la Societe Entomologique de France, 1913, 92 - 93." type="journal article" year="1913">Olivier, 1913</bibRefCitation>
)
</taxonomicName>
from Rhodes, but they differ in e.g. the coloration of elytra (only apical 2/5 of elytra black in
<taxonomicName id="8CBDC69BFF88FFF73EECB6D05928C5BB" box="[964,1078,1483,1507]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="bleusei">
<emphasis id="79C9610AFF88FFF73EECB6D05EC6C5BA" box="[964,984,1483,1506]" italics="true" pageId="10">D</emphasis>
.
<emphasis id="79C9610AFF88FFF73ECFB6D05928C5BB" box="[999,1078,1483,1507]" italics="true" pageId="10">bleusei</emphasis>
</taxonomicName>
), the shape of pronotum (more constricted anteriorly in
<taxonomicName id="8CBDC69BFF88FFF73C80B6F45F07C65E" box="[424,537,1518,1542]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="bleusei">
<emphasis id="79C9610AFF88FFF73C80B6F45CA2C65E" box="[424,444,1519,1542]" italics="true" pageId="10">D</emphasis>
.
<emphasis id="79C9610AFF88FFF73CE2B6F55F07C65E" box="[458,537,1518,1542]" italics="true" pageId="10">bleusei</emphasis>
</taxonomicName>
) and male genitalia (wider paramera and relatively shorter phallus in
<taxonomicName id="8CBDC69BFF88FFF73809B6F4588CC65E" box="[1313,1426,1518,1542]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="bleusei">
<emphasis id="79C9610AFF88FFF73809B6F4582BC65E" box="[1313,1333,1519,1542]" italics="true" pageId="10">D</emphasis>
.
<emphasis id="79C9610AFF88FFF7386BB6F5588CC65E" box="[1347,1426,1518,1542]" italics="true" pageId="10">bleusei</emphasis>
</taxonomicName>
) (
<bibRefCitation id="2F2CC0E9FF88FFF73DB6B5095CDFC673" author="Trllova" box="[158,449,1554,1579]" pageId="10" refString="Trllova, S. &amp; Kundrata, R. (2015) A review of the genus Selasia (Elateridae: Agrypninae: Drilini) in the Palaearctic Region. Zootaxa, 3920 (4), 563 - 571. http: // dx. doi. org / 10.11646 / zootaxa. 3920.4.5" type="journal article" year="2015">Trllova &amp; Kundrata 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF88FFF13DEFB52D5FB2C2DC" lastPageId="12" pageId="10" type="description">
<paragraph id="4B02BD18FF88FFF63DEFB52D5E2EC188" blockId="10.[151,1437,1338,2011]" lastBlockId="11.[151,1437,151,464]" lastPageId="11" pageId="10">
<emphasis id="79C9610AFF88FFF73DEFB52D5C47C617" bold="true" box="[199,345,1590,1615]" pageId="10">Description.</emphasis>
Male (
<typeStatus id="940603BAFF88FFF73C8FB52D5F11C617" box="[423,527,1590,1615]" pageId="10" type="holotype">holotype</typeStatus>
). Body 2.6 times longer than width at humeri, yellowish brown to reddish brown, antennae slightly darker, elytra with black coloration beginning in humeral part and continuing obliquely to the inner elytral margins; body covered with yellowish pubescence (
<figureCitation id="D386A19DFF88FFF73EACB5655ED2C6CF" box="[900,972,1662,1687]" captionStart="FIGURES 1 – 12" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[181,1396,203,1850]" captionTargetId="figure@2.[172,1408,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/232880/files/figure.png" pageId="10">Fig. 8</figureCitation>
). Head including eyes about as wide as anterior margin of pronotum (
<figureCitation id="D386A19DFF88FFF73CCBB5B95F2AC6E3" box="[483,564,1698,1723]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="10">Fig. 18</figureCitation>
), dorsally with shallow depression, sparsely covered with fine punctures, with sparse erect pubescence; clypeal margin widely concave. Eyes medium-sized, their frontal distance 2.3 times eye diameter. Mandibles robust, long, curved, shiny, incisor margin with stout tooth medially. Labrum covered with sparse long setae; maxillary palpi with ultimate palpomere obliquely cut, flattened apically. Antennae pectinate, reaching one third of elytral length, antennomeres 3–10 pectinate, apical antennomere simple, longest, more than 2 times longer than stem of penultimate antennomere (
<figureCitation id="D386A19DFF88FFF73FCCB44D5E26C737" box="[740,824,1878,1903]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="10">Fig. 28</figureCitation>
). Pronotum slightly convex, transverse, widest slightly before middle, 1.5 times wider posteriorly than length at midline. Anterior margin straight, lateral margins deeply emarginate at basal third, widened at apical two thirds, posterior margin slightly convex, bisinuate. Anterior angles inconspicuous, posterior angles rectangular, slightly turned upward (
<figureCitation id="D386A19DFF88FFF73EBEB4D95EF7C783" box="[918,1001,1986,2011]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="10">Fig. 18</figureCitation>
); surface of disc sparsely covered with fine punctures, with sparse erected setae, pubescence denser at angles and lateral margins. Scutellum flat, triangleshaped, apical part rounded. Prosternum transverse, with long transverse groove near frontal margin, frontal margin almost straight, prosternal process short, elevated, gradually declivitous. Mesoventrite widely v-shaped. Elytra subparallel-sided, with longitudinal keel running from humeri toward apex, finely punctured, covered with semierect pubescence, sparser basally, denser at lateral margins and apically. Legs slender, slightly compressed, with sparse, long, semierect setae, tarsomeres 1–4 gradually shortened, tarsomere 4 shortest, apical tarsomere apparently longest, almost as long as combined lengths of tarsomeres 2–4 (
<figureCitation id="D386A19DFF89FFF63EF6B274592CC1D0" box="[990,1074,367,392]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="11">Fig. 32</figureCitation>
). Last abdominal segments as in
<figureCitation id="D386A19DFF89FFF63DBFB28F5DF5C1F4" box="[151,235,404,429]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="11">Fig. 33</figureCitation>
. Aedeagus with phallus considerably curved, longer than paramera; paramera long, with basal inner parts strengthened; phallobase robust, widely v-shaped (
<figureCitation id="D386A19DFF89FFF63FE7B2AC5E3DC188" box="[719,803,439,464]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="11">Fig. 40</figureCitation>
).
</paragraph>
<caption id="1FC2ED90FF89FFF63DBFB5DF5E1CC6A1" httpUri="https://zenodo.org/record/232885/files/figure.png" pageId="11" targetBox="[151,1436,517,1711]" targetPageId="11">
<paragraph id="4B02BD18FF89FFF63DBFB5DF5E1CC6A1" blockId="11.[151,1436,1732,1785]" pageId="11">
<emphasis id="79C9610AFF89FFF63DBFB5DF5C4AC682" bold="true" box="[151,340,1732,1755]" pageId="11">FIGURES 51–54.</emphasis>
<taxonomicName id="8CBDC69BFF89FFF63C73B5DE5CE6C682" box="[347,504,1733,1754]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="79C9610AFF89FFF63C73B5DE5CE6C682" box="[347,504,1733,1754]" italics="true" pageId="11">Drilus robustus</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF89FFF63CD7B5DE5F4EC682" bold="true" box="[511,592,1733,1754]" pageId="11">
<taxonomicNameLabel id="62FADC71FF89FFF63CD7B5DE5F4EC682" box="[511,592,1733,1754]" pageId="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
51, larva; 52, mating couple; 53, the type locality (Turkey, Kemer env.); 54, larval skins inside the shells of
<taxonomicName id="8CBDC69BFF89FFF63CBCB5FF5FE0C6A1" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[404,766,1763,1785]" class="Insecta" family="Chrysomelidae" genus="Rumina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="decollata">
<emphasis id="79C9610AFF89FFF63CBCB5FF5F56C6A0" box="[404,584,1763,1785]" italics="true" pageId="11">Rumina decollata</emphasis>
(Linnaeus, 1758)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="4B02BD18FF89FFF13DEFB43D5F1CC0A0" blockId="11.[151,1436,1830,2035]" lastBlockId="12.[151,1437,151,896]" lastPageId="12" pageId="11">
Female. Larviform, body elongate. All body parts yellowish to light brown, head orange, dorsal surface of thoracic and abdominal segments with dark brown to black markings; body covered with short yellowish pubescence, sparse medially, dense at margins (
<figureCitation id="D386A19DFF89FFF63F98B4755E1BC7DF" box="[688,773,1902,1927]" captionStart="FIGURES 46 – 50" captionStartId="10.[151,264,1240,1263]" captionTargetBox="[168,1420,385,1218]" captionTargetId="figure@10.[151,1436,373,1220]" captionTargetPageId="10" captionText="FIGURES 46 – 50. Drilus robustus sp. nov., female. 46, habitus; 47, genitalia; 48, antenna. Larval cerci. 49, D. robustus sp. nov.; 50, Drilus sp. (Turkey, Muğla Prov., Knidos). Scale bars = 2.0 mm (Fig. 46), 1.0 mm (Figs 47, 49 – 50), 0.5 mm (Fig. 48)." httpUri="https://zenodo.org/record/232884/files/figure.png" pageId="11">Fig. 46</figureCitation>
). Head prognathous, well sclerotized, small, narrower than pronotal anterior margin. Eyes small, their frontal distance more than 5 times eye diameter. Antennae 11- segmented, filiform, covered with sparse semierect setae; apical antennomere minute, narrowed (
<figureCitation id="D386A19DFF89FFF639F2B4AD5833C797" box="[1242,1325,1974,1999]" captionStart="FIGURES 46 – 50" captionStartId="10.[151,264,1240,1263]" captionTargetBox="[168,1420,385,1218]" captionTargetId="figure@10.[151,1436,373,1220]" captionTargetPageId="10" captionText="FIGURES 46 – 50. Drilus robustus sp. nov., female. 46, habitus; 47, genitalia; 48, antenna. Larval cerci. 49, D. robustus sp. nov.; 50, Drilus sp. (Turkey, Muğla Prov., Knidos). Scale bars = 2.0 mm (Fig. 46), 1.0 mm (Figs 47, 49 – 50), 0.5 mm (Fig. 48)." httpUri="https://zenodo.org/record/232884/files/figure.png" pageId="11">Fig. 48</figureCitation>
, Table 1). Mandibles robust, shiny, considerably curved, incisor margin medially with small tooth. Pronotum widest at posterior two thirds; thoracic and abdominal tergites wrinkled, mainly medially. Abdomen with nine visible segments, apical segment much narrower and smaller. Ovipositor with fused valviferes and coxites. Vagina simple, slender, short, sac-like (
<figureCitation id="D386A19DFF8EFFF13C89B3C45CEBC0A0" box="[417,501,223,248]" captionStart="FIGURES 46 – 50" captionStartId="10.[151,264,1240,1263]" captionTargetBox="[168,1420,385,1218]" captionTargetId="figure@10.[151,1436,373,1220]" captionTargetPageId="10" captionText="FIGURES 46 – 50. Drilus robustus sp. nov., female. 46, habitus; 47, genitalia; 48, antenna. Larval cerci. 49, D. robustus sp. nov.; 50, Drilus sp. (Turkey, Muğla Prov., Knidos). Scale bars = 2.0 mm (Fig. 46), 1.0 mm (Figs 47, 49 – 50), 0.5 mm (Fig. 48)." httpUri="https://zenodo.org/record/232884/files/figure.png" pageId="12">Fig. 47</figureCitation>
).
</paragraph>
<paragraph id="4B02BD18FF8EFFF13DEFB21E5F6EC1AC" blockId="12.[151,1437,151,896]" pageId="12">
Larva. Body elongate, slightly widened towards apex. Body yellowish, head, dorsal surface of thoracic and abdominal segments, lateral pleural processes and legs brown; each tergite with a pair of black spots; body covered with long yellowish or brown pubescence (
<figureCitation id="D386A19DFF8EFFF13FBBB2575FF4C13D" box="[659,746,332,357]" captionStart="FIGURES 51 – 54" captionStartId="11.[151,264,1732,1755]" captionTargetBox="[151,1436,517,1711]" captionTargetId="figure@11.[151,1436,517,1711]" captionTargetPageId="11" captionText="FIGURES 51 – 54. Drilus robustus sp. nov. 51, larva; 52, mating couple; 53, the type locality (Turkey, Kemer env.); 54, larval skins inside the shells of Rumina decollata (Linnaeus, 1758)." httpUri="https://zenodo.org/record/232885/files/figure.png" pageId="12">Fig. 51</figureCitation>
). Head prognathous, well sclerotized, small, narrower than frontal pronotal margin. Mandibles sclerotized, narrow and falcate. Abdomen with sclerotized and pigmented tergites, with long setae, especially dorsally (
<figureCitation id="D386A19DFF8EFFF13FBCB28F5FF9C1F4" box="[660,743,404,429]" captionStart="FIGURES 46 – 50" captionStartId="10.[151,264,1240,1263]" captionTargetBox="[168,1420,385,1218]" captionTargetId="figure@10.[151,1436,373,1220]" captionTargetPageId="10" captionText="FIGURES 46 – 50. Drilus robustus sp. nov., female. 46, habitus; 47, genitalia; 48, antenna. Larval cerci. 49, D. robustus sp. nov.; 50, Drilus sp. (Turkey, Muğla Prov., Knidos). Scale bars = 2.0 mm (Fig. 46), 1.0 mm (Figs 47, 49 – 50), 0.5 mm (Fig. 48)." httpUri="https://zenodo.org/record/232884/files/figure.png" pageId="12">Fig. 49</figureCitation>
,
<figureCitation id="D386A19DFF8EFFF13FDCB28F5E21C1F5" box="[756,831,404,429]" captionStart="FIGURES 51 – 54" captionStartId="11.[151,264,1732,1755]" captionTargetBox="[151,1436,517,1711]" captionTargetId="figure@11.[151,1436,517,1711]" captionTargetPageId="11" captionText="FIGURES 51 – 54. Drilus robustus sp. nov. 51, larva; 52, mating couple; 53, the type locality (Turkey, Kemer env.); 54, larval skins inside the shells of Rumina decollata (Linnaeus, 1758)." httpUri="https://zenodo.org/record/232885/files/figure.png" pageId="12">51, 54</figureCitation>
). Lateral pleural and tergal processes well developed. Last segment small, slender. Cerci slightly bent upward, densely covered with long pubescence, cerci apically acuminate; with long apical part (
<figureCitation id="D386A19DFF8EFFF13F27B2C75F7DC1AC" box="[527,611,476,501]" captionStart="FIGURES 46 – 50" captionStartId="10.[151,264,1240,1263]" captionTargetBox="[168,1420,385,1218]" captionTargetId="figure@10.[151,1436,373,1220]" captionTargetPageId="10" captionText="FIGURES 46 – 50. Drilus robustus sp. nov., female. 46, habitus; 47, genitalia; 48, antenna. Larval cerci. 49, D. robustus sp. nov.; 50, Drilus sp. (Turkey, Muğla Prov., Knidos). Scale bars = 2.0 mm (Fig. 46), 1.0 mm (Figs 47, 49 – 50), 0.5 mm (Fig. 48)." httpUri="https://zenodo.org/record/232884/files/figure.png" pageId="12">Fig. 49</figureCitation>
).
</paragraph>
<paragraph id="4B02BD18FF8EFFF13DEFB2E45FB2C2DC" blockId="12.[151,1437,151,896]" pageId="12">
<emphasis id="79C9610AFF8EFFF13DEFB2E45C63C240" bold="true" box="[199,381,511,536]" pageId="12">Measurements.</emphasis>
<typeStatus id="940603BAFF8EFFF13CACB2E45CEEC240" box="[388,496,511,536]" pageId="12" type="holotype">Holotype</typeStatus>
. BL
<quantity id="8C4510FDFF8EFFF13F0EB11B5F61C24F" box="[550,639,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="12" unit="mm" value="7.1">7.1 mm</quantity>
, EL
<quantity id="8C4510FDFF8EFFF13F9DB2E45E10C24F" box="[693,782,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="12" unit="mm" value="5.2">5.2 mm</quantity>
, WHe
<quantity id="8C4510FDFF8EFFF13E75B11B5EABC24F" box="[861,949,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" unit="mm" value="1.8">1.8 mm</quantity>
, WHum
<quantity id="8C4510FDFF8EFFF13933B11B596AC24F" box="[1051,1140,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="12" unit="mm" value="2.7">2.7 mm</quantity>
, PL
<quantity id="8C4510FDFF8EFFF13980B11B581FC24F" box="[1192,1281,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="12" unit="mm" value="1.4">1.4 mm</quantity>
, PW
<quantity id="8C4510FDFF8EFFF13817B11B5886C24F" box="[1343,1432,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="12" unit="mm" value="2.2">2.2 mm</quantity>
, Edist
<quantity id="8C4510FDFF8EFFF13DF0B13F5C2EC264" box="[216,304,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="12" unit="mm" value="1.3">1.3 mm</quantity>
, Ediam
<quantity id="8C4510FDFF8EFFF13CA4B13F5CFBC264" box="[396,485,548,573]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="12" unit="mm" value="0.6">0.6 mm</quantity>
.
<typeStatus id="940603BAFF8EFFF13CD8B13E5F7EC265" box="[496,608,549,573]" pageId="12" type="paratype">Paratypes</typeStatus>
, males. BL
<quantity id="8C4510FDFF8EFFF13FCEB13F5E71C264" box="[742,879,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="9.5" metricValueMin="6.2" pageId="12" unit="mm" value="7.85" valueMax="9.5" valueMin="6.2">6.2–9.5 mm</quantity>
, EL
<quantity id="8C4510FDFF8EFFF13E8DB13F5930C264" box="[933,1070,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="6.6" metricValueMin="4.5" pageId="12" unit="mm" value="5.55" valueMax="6.6" valueMin="4.5">4.5–6.6 mm</quantity>
, WHe
<quantity id="8C4510FDFF8EFFF13954B13F5818C264" box="[1148,1286,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="12" unit="mm" value="1.95" valueMax="2.3" valueMin="1.6">1.6–2.3 mm</quantity>
, WHum
<quantity id="8C4510FDFF8EFFF13843B13F5DF1C207" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.6" metricValueMin="2.4" pageId="12" unit="mm" value="3.0" valueMax="3.6" valueMin="2.4">2.4– 3.6 mm</quantity>
, PL
<quantity id="8C4510FDFF8EFFF13C0BB1535CB2C207" box="[291,428,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="12" unit="mm" value="1.55" valueMax="1.8" valueMin="1.3">1.3–1.8 mm</quantity>
, PW 2.0–
<quantity id="8C4510FDFF8EFFF13F34B1535F6DC207" box="[540,627,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="12" unit="mm" value="2.8">2.8 mm</quantity>
, Edist
<quantity id="8C4510FDFF8EFFF13FE8B15C5E57C207" box="[704,841,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="12" unit="mm" value="1.35" valueMax="1.6" valueMin="1.1">1.1–1.6 mm</quantity>
, Ediam
<quantity id="8C4510FDFF8EFFF13E8EB15C592EC207" box="[934,1072,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="12" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
.
<typeStatus id="940603BAFF8EFFF13913B15359BEC238" box="[1083,1184,584,608]" pageId="12" type="paratype">Paratype</typeStatus>
, female. BL
<quantity id="8C4510FDFF8EFFF13819B15C5886C207" box="[1329,1432,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="12" unit="mm" value="20.5">20.5 mm</quantity>
, BW
<quantity id="8C4510FDFF8EFFF13DE3B1775C3AC2DC" box="[203,292,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="12" unit="mm" value="4.7">4.7 mm</quantity>
. Larva. BL 18.0 mm, BW
<quantity id="8C4510FDFF8EFFF13F66B1775FB6C2DC" box="[590,680,620,645]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="12" unit="mm" value="3.5">3.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8EFFF13DEFB1945FE7C2F0" box="[199,761,655,680]" pageId="12" type="distribution">
<paragraph id="4B02BD18FF8EFFF13DEFB1945FE7C2F0" blockId="12.[151,1437,151,896]" box="[199,761,655,680]" pageId="12">
<emphasis id="79C9610AFF8EFFF13DEFB1945C7EC2F0" bold="true" box="[199,352,655,680]" pageId="12">Distribution.</emphasis>
<collectingCountry id="33AAFD88FF8EFFF13C4FB1945CA9C2F0" box="[359,439,655,680]" name="Turkey" pageId="12">Turkey</collectingCountry>
(Antalya Province;
<figureCitation id="D386A19DFF8EFFF13FB1B1945FF3C2F0" box="[665,749,655,680]" captionStart="FIGURE 55" captionStartId="12.[151,250,1911,1933]" captionTargetBox="[160,1421,963,1875]" captionTargetId="figure@12.[151,1436,951,1889]" captionTargetPageId="12" captionText="FIGURE 55. The distribution of Drilus species in Asia Minor." httpUri="https://zenodo.org/record/232886/files/figure.png" pageId="12">Fig. 55</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8EFFF13DEFB1AF590EC295" box="[199,1040,692,717]" pageId="12" type="etymology">
<paragraph id="4B02BD18FF8EFFF13DEFB1AF590EC295" blockId="12.[151,1437,151,896]" box="[199,1040,692,717]" pageId="12">
<emphasis id="79C9610AFF8EFFF13DEFB1AF5C50C295" bold="true" box="[199,334,692,717]" pageId="12">Etymology.</emphasis>
The specific epithet refers to the robust habitus of this species.
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF8EFFF03DEFB1CC5E8EC21C" lastPageId="13" pageId="12" type="biology_ecology">
<paragraph id="4B02BD18FF8EFFF13DEFB1CC5CAAC3D8" blockId="12.[151,1437,151,896]" pageId="12">
<emphasis id="79C9610AFF8EFFF13DEFB1CC5CA2C2A8" bold="true" box="[199,444,727,752]" pageId="12">Biology and ecology.</emphasis>
<taxonomicName id="8CBDC69BFF8EFFF13CEBB1C35F6CC2A8" box="[451,626,728,752]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="79C9610AFF8EFFF13CEBB1C35F6CC2A8" box="[451,626,728,752]" italics="true" pageId="12">Drilus robustus</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF8EFFF13F51B1CC5FCDC2A8" bold="true" box="[633,723,727,752]" pageId="12">
<taxonomicNameLabel id="62FADC71FF8EFFF13F51B1CC5FCDC2A8" box="[633,723,727,752]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
occurs in a lowland forest; most specimens were collected at a forest edge (
<figureCitation id="D386A19DFF8EFFF13C09B1E75C6AC34D" box="[289,372,764,789]" captionStart="FIGURES 51 – 54" captionStartId="11.[151,264,1732,1755]" captionTargetBox="[151,1436,517,1711]" captionTargetId="figure@11.[151,1436,517,1711]" captionTargetPageId="11" captionText="FIGURES 51 – 54. Drilus robustus sp. nov. 51, larva; 52, mating couple; 53, the type locality (Turkey, Kemer env.); 54, larval skins inside the shells of Rumina decollata (Linnaeus, 1758)." httpUri="https://zenodo.org/record/232885/files/figure.png" pageId="12">Fig. 53</figureCitation>
). Larvae prey on the snail
<taxonomicName id="8CBDC69BFF8EFFF13FB4B1E65936C34D" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[668,1064,764,789]" class="Insecta" family="Chrysomelidae" genus="Rumina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="decollata">
<emphasis id="79C9610AFF8EFFF13FB4B1E65E7CC34C" box="[668,866,764,788]" italics="true" pageId="12">Rumina decollata</emphasis>
(
<bibRefCitation id="2F2CC0E9FF8EFFF13E58B1E75901C34D" author="Linnaeus" box="[880,1055,764,789]" pageId="12" refString="Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth Edition, Laurentii Salvii, Stockholm, 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
(Achatinoidea:
<taxonomicName id="8CBDC69BFF8EFFF139F3B1E7587BC34D" box="[1243,1381,764,789]" class="Gastropoda" family="Subulinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" phylum="Mollusca" rank="family">Subulinidae</taxonomicName>
;
<figureCitation id="D386A19DFF8EFFF13858B1E75DA9C360" captionStart="FIGURES 51 – 54" captionStartId="11.[151,264,1732,1755]" captionTargetBox="[151,1436,517,1711]" captionTargetId="figure@11.[151,1436,517,1711]" captionTargetPageId="11" captionText="FIGURES 51 – 54. Drilus robustus sp. nov. 51, larva; 52, mating couple; 53, the type locality (Turkey, Kemer env.); 54, larval skins inside the shells of Rumina decollata (Linnaeus, 1758)." httpUri="https://zenodo.org/record/232885/files/figure.png" pageId="12">Fig. 54</figureCitation>
). Adults can be found from late May to early June. After pupation, males sit on grass and tree branches waiting for the females which they mate almost immediately after their emergence (L. Fusek, J. Bednář, pers. communication;
<figureCitation id="D386A19DFF8EFFF13C7CB07C5CB9C3D8" box="[340,423,871,896]" captionStart="FIGURES 51 – 54" captionStartId="11.[151,264,1732,1755]" captionTargetBox="[151,1436,517,1711]" captionTargetId="figure@11.[151,1436,517,1711]" captionTargetPageId="11" captionText="FIGURES 51 – 54. Drilus robustus sp. nov. 51, larva; 52, mating couple; 53, the type locality (Turkey, Kemer env.); 54, larval skins inside the shells of Rumina decollata (Linnaeus, 1758)." httpUri="https://zenodo.org/record/232885/files/figure.png" pageId="12">Fig. 52</figureCitation>
).
</paragraph>
<caption id="1FC2ED90FF8EFFF13DBFB46C5E0AC7D5" box="[151,788,1911,1933]" httpUri="https://zenodo.org/record/232886/files/figure.png" pageId="12" targetBox="[160,1421,963,1875]" targetPageId="12">
<paragraph id="4B02BD18FF8EFFF13DBFB46C5E0AC7D5" blockId="12.[151,788,1911,1933]" box="[151,788,1911,1933]" pageId="12">
<emphasis id="79C9610AFF8EFFF13DBFB46C5C01C7D4" bold="true" box="[151,287,1911,1933]" pageId="12">FIGURE 55.</emphasis>
The distribution of
<taxonomicName id="8CBDC69BFF8EFFF13CC2B46C5F35C7D4" box="[490,555,1911,1932]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
<emphasis id="79C9610AFF8EFFF13CC2B46C5F35C7D4" box="[490,555,1911,1932]" italics="true" pageId="12">Drilus</emphasis>
</taxonomicName>
species in Asia Minor.
</paragraph>
</caption>
<paragraph id="4B02BD18FF8FFFF03D5FB10A5E8EC21C" blockId="13.[119,1888,524,580]" pageId="13">
<emphasis id="79C9610AFF8FFFF03D5FB10A5DD0C27F" bold="true" box="[119,206,529,551]" pageId="13">Table 1.</emphasis>
Numbers of antennomeres in females of different
<taxonomicName id="8CBDC69BFF8FFFF03FE6B10A5E0FC27F" box="[718,785,529,551]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="tribe" tribe="Drilini">Drilini</taxonomicName>
species. 1 xx+1 – the apical antennomere is minute;? – one of the antennomeres might in fact consist of two fused antennomeres; pers. com. – personal communication; det. – determined.
</paragraph>
</subSubSection>
</treatment>
</document>
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