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<document id="EF22BF86472B2D020D7A604DFF18A8B6" ID-DOI="10.11646/zootaxa.4012.1.4" ID-GBIF-Dataset="d32dd252-5a36-4368-a636-6e688bc4ed13" ID-ISSN="1175-5326" ID-Zenodo-Dep="232879" ID-ZooBank="C3B3A06D-C18C-4B66-83E9-51B605E11862" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461226104448" checkinUser="plazi" docAuthor="Kobieluszova, Lucie &amp; Kundrata, Robin" docDate="2015" docId="C3140C0EFF87FFFA3DBFB66659CCC145" docLanguage="en" docName="zt04012p096.pdf" docOrigin="Zootaxa 4012 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drilus huijbregtsi Kobieluszova &amp; Kundrata, 2015, sp. nov." docType="treatment" docVersion="8" masterDocId="3F2D7476FF82FFFD3D28B31B5D1EC058" masterDocTitle="Taxonomic review of Drilus Olivier, 1790 (Elateridae: Agrypninae: Drilini) from Asia Minor, with descriptions of seven new species and comments on the female antennal morphology in Drilini" updateTime="1698604902572" updateUser="plazi">
<mods:mods id="25F4379E6AA129F34B6FEB8CFE8986E7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0AE2AB01D231E8175F60CBE25F7020EA">
<mods:title id="182339102430C83B21F6DB8433C1FBC5">Taxonomic review of Drilus Olivier, 1790 (Elateridae: Agrypninae: Drilini) from Asia Minor, with descriptions of seven new species and comments on the female antennal morphology in Drilini</mods:title>
</mods:titleInfo>
<mods:name id="8FF26F40961E055006ED45E9CBEEEC8A" type="personal">
<mods:role id="AEA3B075DBACE1D4D2CFE50A70B0E426">
<mods:roleTerm id="703398A22512DBBB7D46736C747BEBE9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F626D48CD9E0D560A93CBAD13222E4D">Kobieluszova, Lucie</mods:namePart>
</mods:name>
<mods:name id="F7615DBCC0C87DD92DFE04AD0F9807DF" type="personal">
<mods:role id="DC9510596942087E362D577D60D6044D">
<mods:roleTerm id="9408BBF532ADAD7C1BD062A4884A5BF2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C81D745440C942C8A0C8C7D5D6FFE75">Kundrata, Robin</mods:namePart>
</mods:name>
<mods:typeOfResource id="FF643A0D1E0EC22D5F86FCA133527740">text</mods:typeOfResource>
<mods:relatedItem id="0AC0664435BF8F63760AE2B93E3C00F7" type="host">
<mods:titleInfo id="6942918279572B0C761C871808BDED29">
<mods:title id="77EDF168EA161A571B02F00198CC1DDD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8CC3BFDB1B71753D930BBC879EC5AEC7">
<mods:date id="9CCF66367508CA75971D3447A960143B">2015</mods:date>
<mods:detail id="D32940D88A22FFD46BD6718FE309E624" type="volume">
<mods:number id="E5F676A9B69A38713DD0F99F4E2545A9">4012</mods:number>
</mods:detail>
<mods:detail id="E98072847692C64931C10D1ECF1B4C32" type="issue">
<mods:number id="7CC9A6969B46715D109678E794503AFE">1</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="6E53642C94D9EA6AAEC0B5CF95583289">journal volume</mods:classification>
<mods:identifier id="6305BC7685F813BF0D51B410F4041154" type="DOI">10.11646/zootaxa.4012.1.4</mods:identifier>
<mods:identifier id="9C42913DBBA6E6D7BA9AC5620321B9EF" type="GBIF-Dataset">d32dd252-5a36-4368-a636-6e688bc4ed13</mods:identifier>
<mods:identifier id="B5BC3694709C21AD276DB70D980B6847" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="169348FE78FA92FF7D3E535F155EEE97" type="Zenodo-Dep">232879</mods:identifier>
<mods:identifier id="DCB9A5999046D0EB00AD30654E6EAA83" type="ZooBank">C3B3A06D-C18C-4B66-83E9-51B605E11862</mods:identifier>
</mods:mods>
<treatment id="C3140C0EFF87FFFA3DBFB66659CCC145" ID-DOI="http://doi.org/10.5281/zenodo.6112424" ID-GBIF-Taxon="119632740" ID-Zenodo-Dep="6112424" LSID="urn:lsid:plazi:treatment:C3140C0EFF87FFFA3DBFB66659CCC145" httpUri="http://treatment.plazi.org/id/C3140C0EFF87FFFA3DBFB66659CCC145" lastPageId="7" pageId="5">
<subSubSection id="03A7EE93FF87FFF83DBFB6665C72C5E0" pageId="5" type="nomenclature">
<paragraph id="4B02BD18FF87FFF83DBFB6665CCFC5CF" blockId="5.[151,465,1405,1464]" box="[151,465,1405,1431]" pageId="5">
<heading id="104A0A74FF87FFF83DBFB6665CCFC5CF" bold="true" box="[151,465,1405,1431]" fontSize="11" level="1" pageId="5" reason="1">
<emphasis id="79C9610AFF87FFF83DBFB6665CCFC5CF" bold="true" box="[151,465,1405,1431]" pageId="5">
<taxonomicName id="8CBDC69BFF87FFF83DBFB6665C73C5CF" box="[151,365,1405,1431]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="huijbregtsi" status="sp. nov.">
<emphasis id="79C9610AFF87FFF83DBFB6665C73C5CF" bold="true" box="[151,365,1405,1431]" italics="true" pageId="5">Drilus huijbregtsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="62FADC71FF87FFF83C5CB6665CCFC5CF" box="[372,465,1405,1431]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4B02BD18FF87FFF83DBFB6845C72C5E0" blockId="5.[151,465,1405,1464]" box="[151,364,1439,1464]" pageId="5">
(
<figureCitation id="D386A19DFF87FFF83DB7B6845DF9C5E0" box="[159,231,1439,1464]" captionStart="FIGURES 1 – 12" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[181,1396,203,1850]" captionTargetId="figure@2.[172,1408,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/232880/files/figure.png" pageId="5">Figs 3</figureCitation>
,
<figureCitation id="D386A19DFF87FFF83DDBB6BB5C0FC5E0" box="[243,273,1440,1464]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="5">14</figureCitation>
,
<figureCitation id="D386A19DFF87FFF83C35B6BB5C25C5E0" box="[285,315,1440,1464]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="5">23</figureCitation>
,
<figureCitation id="D386A19DFF87FFF83C6FB6845C7AC5E0" box="[327,356,1439,1464]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="5">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF87FFF83DBFB6FC5F32C67C" pageId="5" type="materials_examined">
<paragraph id="4B02BD18FF87FFF83DBFB6FC584AC658" blockId="5.[151,1437,1511,2005]" box="[151,1364,1511,1536]" pageId="5">
<emphasis id="79C9610AFF87FFF83DBFB6FC5C5BC658" bold="true" box="[151,325,1511,1536]" pageId="5">
<typeStatus id="940603BAFF87FFF83DBFB6FC5DCCC658" box="[151,210,1511,1536]" pageId="5">Type</typeStatus>
material.
</emphasis>
<typeStatus id="940603BAFF87FFF83C64B6FC5CA9C658" box="[332,439,1511,1536]" pageId="5" type="holotype">Holotype</typeStatus>
, male, &quot;Amasia, Coll. D. A. Reclaire,
<taxonomicName id="8CBDC69BFF87FFF83E4BB6F35972C5A7" authority="Ahr. E." authorityName="Ahr. E." box="[867,1132,1511,1536]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="79C9610AFF87FFF83E4BB6F35908C658" box="[867,1046,1512,1536]" italics="true" pageId="5">Drilus concolor</emphasis>
Ahr. E.
</taxonomicName>
le Moult&quot; (
<collectionCode id="2DAC25DDFF87FFF839C5B6F35856C658" LSID="urn:lsid:biocol.org:col:34992" box="[1261,1352,1512,1536]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="5">RMNH</collectionCode>
).
</paragraph>
<paragraph id="4B02BD18FF87FFF83DEFB5175F32C67C" blockId="5.[151,1437,1511,2005]" box="[199,556,1548,1573]" pageId="5">
<emphasis id="79C9610AFF87FFF83DEFB5175C7BC67D" bold="true" box="[199,357,1548,1573]" pageId="5">
<typeStatus id="940603BAFF87FFF83DEFB5175C1CC67D" box="[199,258,1548,1573]" pageId="5">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="33AAFD88FF87FFF83C44B5175CA3C67D" box="[364,445,1548,1573]" name="Turkey" pageId="5">Turkey</collectingCountry>
, Amasya.
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF87FFF83DEFB534597CC634" pageId="5" type="diagnosis">
<paragraph id="4B02BD18FF87FFF83DEFB534597CC634" blockId="5.[151,1437,1511,2005]" pageId="5">
<emphasis id="79C9610AFF87FFF83DEFB5345C5FC610" bold="true" box="[199,321,1583,1608]" pageId="5">Diagnosis.</emphasis>
<taxonomicName id="8CBDC69BFF87FFF83C60B52B5F10C610" box="[328,526,1584,1608]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="huijbregtsi" status="sp. nov.">
<emphasis id="79C9610AFF87FFF83C60B52B5F10C610" box="[328,526,1584,1608]" italics="true" pageId="5">Drilus huijbregtsi</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF87FFF83F3EB5345F71C610" bold="true" box="[534,623,1583,1608]" pageId="5">
<taxonomicNameLabel id="62FADC71FF87FFF83F3EB5345F71C610" box="[534,623,1583,1608]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar in habitus and coloration to
<taxonomicName id="8CBDC69BFF87FFF83936B52A5984C61F" box="[1054,1178,1584,1608]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="turcicus" status="sp. nov.">
<emphasis id="79C9610AFF87FFF83936B52A592CC610" box="[1054,1074,1585,1608]" italics="true" pageId="5">D</emphasis>
.
<emphasis id="79C9610AFF87FFF83968B52B5984C61F" box="[1088,1178,1584,1607]" italics="true" pageId="5">turcicus</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF87FFF83989B53459E4C610" bold="true" box="[1185,1274,1583,1608]" pageId="5">
<taxonomicNameLabel id="62FADC71FF87FFF83989B53459E4C610" box="[1185,1274,1583,1608]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
These species differ in the shapes of clypeus, antennae (
<figureCitation id="D386A19DFF87FFF83F4FB54F5FF9C634" box="[615,743,1620,1645]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="5">Figs 23, 31</figureCitation>
) and male genitalia (
<figureCitation id="D386A19DFF87FFF83EFBB54F594BC634" box="[979,1109,1620,1645]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="5">Figs 35, 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF87FFFA3DEFB56C5C60C08C" lastPageId="7" pageId="5" type="description">
<paragraph id="4B02BD18FF87FFFB3DEFB56C5CF0C188" blockId="5.[151,1437,1511,2005]" lastBlockId="6.[151,1437,151,464]" lastPageId="6" pageId="5">
<emphasis id="79C9610AFF87FFF83DEFB56C5C47C6C8" bold="true" box="[199,345,1655,1680]" pageId="5">Description.</emphasis>
Male (
<typeStatus id="940603BAFF87FFF83C82B56C5F0DC6C8" box="[426,531,1655,1680]" pageId="5" type="holotype">holotype</typeStatus>
). Body 3.3 times longer than width at humeri, dark brown to black, trochanters and tarsi brown; body covered with light brown pubescence (
<figureCitation id="D386A19DFF87FFF83E41B5875EAEC6EC" box="[873,944,1692,1717]" captionStart="FIGURES 1 – 12" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[181,1396,203,1850]" captionTargetId="figure@2.[172,1408,195,1856]" captionTargetPageId="2" captionText="FIGURES 1 – 12. Habitus images of Drilus species. 1, Drilus akbesianus (Fairmaire, 1895); 2, D. badius sp. nov.; 3, D. huijbregtsi sp. nov.; 4, D. latithorax Pic, 1902; 5, D. mertliki sp. nov.; 6, D. obscuricornis Pic, 1899; 7, D. rectus Schaufuss, 1867; 8, D. robustus sp. nov.; 9, D. sanliurfensis sp. nov.; 10, D. teunisseni sp. nov.; 11, D. turcicus sp. nov.; 12, D. iranicus Wittmer, 1967. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/232880/files/figure.png" pageId="5">Fig. 3</figureCitation>
). Head including eyes slightly wider than anterior margin of pronotum (
<figureCitation id="D386A19DFF87FFF83F2AB5A45F42C680" box="[514,604,1727,1752]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="5">Fig. 14</figureCitation>
), dorsally with wide shallow depression, covered with large shallow punctures, with sparse semierect to erect pubescence; clypeal margin widely concave. Eyes medium-sized, their frontal distance 2.0 times eye diameter. Mandibles curved, incisor margin with short conspicuous tooth medially. Labrum transverse; maxillary palpi slender, apical palpomere obliquely cut. Antennae pectinate (only two and eight antennomeres present of left and right antenna, respectively); antennomere 3 long, serrate, antennomeres 4–8 pectinate (
<figureCitation id="D386A19DFF87FFF83C23B46F5C41C7D4" box="[267,351,1908,1933]" captionStart="FIGURES 22 – 33" captionStartId="4.[151,264,1794,1817]" captionTargetBox="[155,1414,201,1764]" captionTargetId="figure@4.[151,1436,193,1773]" captionTargetPageId="4" captionText="FIGURES 22 – 33. Antennae of Drilus species. 22, Drilus badius sp. nov.; 23, D. huijbregtsi sp. nov.; 24, D. latithorax Pic, 1902; 25, D. mertliki sp. nov.; 26, D. obscuricornis Pic, 1899; 27, D. rectus Schaufuss, 1867; 28, D. robustus sp. nov.; 29, D. sanliurfensis sp. nov.; 30, D. teunisseni sp. nov.; 31, D. turcicus sp. nov. Drilus robustus sp. nov., male. 32, leg; 33, last abdominal segments. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232882/files/figure.png" pageId="5">Fig. 23</figureCitation>
). Pronotum slightly convex, transverse, widest at middle, 1.6 times wider than length at midline. Anterior margin almost straight, lateral margins convex, widened; posterior margin bisinuate. Anterior angles inconspicuous, posterior angles rectangular (
<figureCitation id="D386A19DFF87FFF83FA3B4A75FC0C78C" box="[651,734,1980,2005]" captionStart="FIGURES 13 – 21" captionStartId="3.[151,264,1423,1446]" captionTargetBox="[161,1430,204,1401]" captionTargetId="figure@3.[151,1436,193,1403]" captionTargetPageId="3" captionText="FIGURES 13 – 21. Pronota of Drilus species. 13, Drilus badius sp. nov.; 14, D. huijbregtsi sp. nov.; 15, D. latithorax Pic, 1902; 16, D. mertliki sp. nov.; 17, D. obscuricornis Pic, 1899; 18, D. robustus sp. nov.; 19, D. sanliurfensis sp. nov.; 20, D. teunisseni sp. nov.; 21, D. turcicus sp. nov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/232881/files/figure.png" pageId="5">Fig. 14</figureCitation>
); surface of disc sparsely covered with fine shallow punctures, with sparse semierect to erect pubescence, pubescence denser at angles. Scutellum flat, gradually narrowed, wrinkled; apex widely rounded. Prosternum transverse, slightly wrinkled, frontal margin almost straight, posterior margin widely convex, steeply declivitous, prosternal process inconspicuous. Mesoventrite widely v-shaped, with groove anteromedially. Elytra relatively long, subparallel-sided, slightly wrinkled, covered with semierect pubescence. Legs slender, slightly compressed, with sparse, long, semierect setae, tarsomeres 1–4 gradually shortened, tarsomere 4 shortest, apical tarsomere longest, slender, longer than combined lengths of tarsomeres 3 and 4. Penultimate ventrite medially with deep rounded emargination. Aedeagus with robust, considerably curved phallus; paramera rather short, slightly membranous apically; phallobase long, u-shaped (
<figureCitation id="D386A19DFF84FFFB398FB28F59E1C1F5" box="[1191,1279,404,429]" captionStart="FIGURES 34 – 45" captionStartId="6.[151,264,1845,1868]" captionTargetBox="[163,1430,529,1821]" captionTargetId="figure@6.[151,1436,517,1825]" captionTargetPageId="6" captionText="FIGURES 34 – 45. Male genitalia of Drilus species. 34, Drilus badius sp. nov.; 35, D. huijbregtsi sp. nov.; 36, D. latithorax Pic, 1902; 37, D. mertliki sp. nov.; 38, D. obscuricornis Pic, 1899; 39, D. rectus Schaufuss, 1867 (Turkey, Adana Prov.); 40, D. robustus sp. nov.; 41, D. sanliurfensis sp. nov.; 42, D. teunisseni sp. nov.; 43, D. turcicus sp. nov.; 44, D. iranicus Wittmer, 1967; 45, Drilus sp. (Abkhazia). Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="6">Fig. 35</figureCitation>
). Female and immature stages are unknown.
</paragraph>
<caption id="1FC2ED90FF84FFFB3DBFB42E5FD7C7F0" httpUri="https://zenodo.org/record/232883/files/figure.png" pageId="6" targetBox="[163,1430,529,1821]" targetPageId="6">
<paragraph id="4B02BD18FF84FFFB3DBFB42E5FD7C7F0" blockId="6.[151,1436,1845,1960]" pageId="6">
<emphasis id="79C9610AFF84FFFB3DBFB42E5C4BC713" bold="true" box="[151,341,1845,1868]" pageId="6">FIGURES 34–45.</emphasis>
Male genitalia of
<taxonomicName id="8CBDC69BFF84FFFB3F3DB42D5F48C713" box="[533,598,1846,1867]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
<emphasis id="79C9610AFF84FFFB3F3DB42D5F48C713" box="[533,598,1846,1867]" italics="true" pageId="6">Drilus</emphasis>
</taxonomicName>
species. 34,
<taxonomicName id="8CBDC69BFF84FFFB3FF4B42D5E77C713" box="[732,873,1846,1867]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="badius" status="sp. nov.">
<emphasis id="79C9610AFF84FFFB3FF4B42D5E77C713" box="[732,873,1846,1867]" italics="true" pageId="6">Drilus badius</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF84FFFB3E58B42D5EDCC713" bold="true" box="[880,962,1846,1867]" pageId="6">
<taxonomicNameLabel id="62FADC71FF84FFFB3E58B42D5EDCC713" box="[880,962,1846,1867]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 35,
<taxonomicName id="8CBDC69BFF84FFFB3ED1B42C5999C713" box="[1017,1159,1846,1868]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="huijbregtsi" status="sp. nov.">
<emphasis id="79C9610AFF84FFFB3ED1B42C5999C713" box="[1017,1159,1846,1868]" italics="true" pageId="6">D. huijbregtsi</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF84FFFB39A6B42D59FFC713" bold="true" box="[1166,1249,1846,1867]" pageId="6">
<taxonomicNameLabel id="62FADC71FF84FFFB39A6B42D59FFC713" box="[1166,1249,1846,1867]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 36,
<taxonomicName id="8CBDC69BFF84FFFB383FB42C5DE7C732" authority="Pic, 1902" authorityName="Pic" authorityYear="1902" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="latithorax">
<emphasis id="79C9610AFF84FFFB383FB42C5882C713" box="[1303,1436,1846,1868]" italics="true" pageId="6">D. latithorax</emphasis>
Pic, 1902
</taxonomicName>
; 37,
<taxonomicName id="8CBDC69BFF84FFFB3C01B44E5C89C732" box="[297,407,1877,1898]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="79C9610AFF84FFFB3C01B44E5C89C732" box="[297,407,1877,1898]" italics="true" pageId="6">D. mertliki</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF84FFFB3CB4B44E5CF3C732" bold="true" box="[412,493,1877,1898]" pageId="6">
<taxonomicNameLabel id="62FADC71FF84FFFB3CB4B44E5CF3C732" box="[412,493,1877,1898]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 38,
<taxonomicName id="8CBDC69BFF84FFFB3F08B44E5E2DC732" authority="Pic, 1899" authorityName="Pic" authorityYear="1899" box="[544,819,1876,1898]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="obscuricornis">
<emphasis id="79C9610AFF84FFFB3F08B44E5FD5C732" box="[544,715,1877,1898]" italics="true" pageId="6">D. obscuricornis</emphasis>
Pic, 1899
</taxonomicName>
; 39,
<taxonomicName id="8CBDC69BFF84FFFB3E4AB44E5974C732" authority="Schaufuss, 1867" authorityName="Schaufuss" authorityYear="1867" box="[866,1130,1876,1898]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="rectus">
<emphasis id="79C9610AFF84FFFB3E4AB44E5E6AC732" box="[866,884,1877,1898]" italics="true" pageId="6">D</emphasis>
.
<emphasis id="79C9610AFF84FFFB3EA9B44E5EA3C732" box="[897,957,1877,1898]" italics="true" pageId="6">rectus</emphasis>
Schaufuss, 1867
</taxonomicName>
(Turkey, Adana Prov.); 40,
<taxonomicName id="8CBDC69BFF84FFFB38ABB44E5DF3C7D0" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="79C9610AFF84FFFB38ABB44E588BC732" box="[1411,1429,1877,1898]" italics="true" pageId="6">D</emphasis>
.
<emphasis id="79C9610AFF84FFFB3DBFB4685DF3C7D0" box="[151,237,1907,1928]" italics="true" pageId="6">robustus</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF84FFFB3DDDB46F5C59C7D1" bold="true" box="[245,327,1908,1929]" pageId="6">
<taxonomicNameLabel id="62FADC71FF84FFFB3DDDB46F5C59C7D1" box="[245,327,1908,1929]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 41,
<taxonomicName id="8CBDC69BFF84FFFB3C55B46F5F3EC7D0" box="[381,544,1907,1929]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="sanliurfensis" status="sp. nov.">
<emphasis id="79C9610AFF84FFFB3C55B46F5C91C7D1" box="[381,399,1908,1929]" italics="true" pageId="6">D</emphasis>
.
<emphasis id="79C9610AFF84FFFB3CB6B4685F3EC7D0" box="[414,544,1907,1928]" italics="true" pageId="6">sanliurfensis</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF84FFFB3F0FB46F5F67C7D1" bold="true" box="[551,633,1908,1929]" pageId="6">
<taxonomicNameLabel id="62FADC71FF84FFFB3F0FB46F5F67C7D1" box="[551,633,1908,1929]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 42,
<taxonomicName id="8CBDC69BFF84FFFB3F98B46F5E2BC7D1" box="[688,821,1908,1929]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="teunisseni" status="sp. nov.">
<emphasis id="79C9610AFF84FFFB3F98B46F5FDCC7D1" box="[688,706,1908,1929]" italics="true" pageId="6">D</emphasis>
.
<emphasis id="79C9610AFF84FFFB3FF8B46F5E2BC7D1" box="[720,821,1908,1929]" italics="true" pageId="6">teunisseni</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF84FFFB3E15B46F5E8EC7D1" bold="true" box="[829,912,1908,1929]" pageId="6">
<taxonomicNameLabel id="62FADC71FF84FFFB3E15B46F5E8EC7D1" box="[829,912,1908,1929]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 43,
<taxonomicName id="8CBDC69BFF84FFFB3EEEB46F5926C7D1" box="[966,1080,1908,1929]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="turcicus" status="sp. nov.">
<emphasis id="79C9610AFF84FFFB3EEEB46F5EC6C7D1" box="[966,984,1908,1929]" italics="true" pageId="6">D</emphasis>
.
<emphasis id="79C9610AFF84FFFB3ECFB46F5926C7D1" box="[999,1080,1908,1929]" italics="true" pageId="6">turcicus</emphasis>
</taxonomicName>
<emphasis id="79C9610AFF84FFFB3917B46F598FC7D1" bold="true" box="[1087,1169,1908,1929]" pageId="6">
<taxonomicNameLabel id="62FADC71FF84FFFB3917B46F598FC7D1" box="[1087,1169,1908,1929]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 44,
<taxonomicName id="8CBDC69BFF84FFFB39E0B46F5DD3C7F0" authority="Wittmer, 1967" authorityName="Wittmer" authorityYear="1967" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="79C9610AFF84FFFB39E0B46F59C4C7D1" box="[1224,1242,1908,1929]" italics="true" pageId="6">D</emphasis>
.
<emphasis id="79C9610AFF84FFFB39C0B46F5825C7D1" box="[1256,1339,1908,1929]" italics="true" pageId="6">iranicus</emphasis>
Wittmer, 1967
</taxonomicName>
; 45,
<taxonomicName id="8CBDC69BFF84FFFB3DD5B4895C20C7FF" box="[253,318,1938,1959]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
<emphasis id="79C9610AFF84FFFB3DD5B4895C20C7FF" box="[253,318,1938,1959]" italics="true" pageId="6">Drilus</emphasis>
</taxonomicName>
sp. (Abkhazia). Scale bars = 0.25 mm.
</paragraph>
</caption>
<paragraph id="4B02BD18FF85FFFA3DEFB38C5C60C08C" blockId="7.[151,1436,151,285]" pageId="7">
<emphasis id="79C9610AFF85FFFA3DEFB38C5C63C0E8" bold="true" box="[199,381,151,176]" pageId="7">Measurements.</emphasis>
BL
<quantity id="8C4510FDFF85FFFA3C99B38C5F14C0F7" box="[433,522,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="7" unit="mm" value="6.4">6.4 mm</quantity>
, EL
<quantity id="8C4510FDFF85FFFA3F69B3835F84C0F7" box="[577,666,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="7" unit="mm" value="4.4">4.4 mm</quantity>
, WHe
<quantity id="8C4510FDFF85FFFA3FC2B3835E5DC0F7" box="[746,835,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" unit="mm" value="1.3">1.3 mm</quantity>
, WHum
<quantity id="8C4510FDFF85FFFA3E82B383591AC0F7" box="[938,1028,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="7" unit="mm" value="1.9">1.9 mm</quantity>
, PL 1.0 mm, PW
<quantity id="8C4510FDFF85FFFA39F9B38C5834C0F7" box="[1233,1322,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="7" unit="mm" value="1.6">1.6 mm</quantity>
, Edist
<quantity id="8C4510FDFF85FFFA3851B3835DDBC08C" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" unit="mm" value="0.9">0.9 mm</quantity>
, Ediam
<quantity id="8C4510FDFF85FFFA3C08B3A75C64C08C" box="[288,378,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="7" unit="mm" value="0.4">0.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF85FFFA3DEFB3C45E76C0A0" box="[199,872,223,248]" pageId="7" type="distribution">
<paragraph id="4B02BD18FF85FFFA3DEFB3C45E76C0A0" blockId="7.[151,1436,151,285]" box="[199,872,223,248]" pageId="7">
<emphasis id="79C9610AFF85FFFA3DEFB3C45C7EC0A0" bold="true" box="[199,352,223,248]" pageId="7">Distribution.</emphasis>
Northern
<collectingCountry id="33AAFD88FF85FFFA3CFBB3C45F3DC0A0" box="[467,547,223,248]" name="Turkey" pageId="7">Turkey</collectingCountry>
(Amasya Province;
<figureCitation id="D386A19DFF85FFFA3E20B3C45E42C0A0" box="[776,860,223,248]" captionStart="FIGURE 55" captionStartId="12.[151,250,1911,1933]" captionTargetBox="[160,1421,963,1875]" captionTargetId="figure@12.[151,1436,951,1889]" captionTargetPageId="12" captionText="FIGURE 55. The distribution of Drilus species in Asia Minor." httpUri="https://zenodo.org/record/232886/files/figure.png" pageId="7">Fig. 55</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="03A7EE93FF85FFFA3DEFB21F59CCC145" box="[199,1234,260,285]" pageId="7" type="etymology">
<paragraph id="4B02BD18FF85FFFA3DEFB21F59CCC145" blockId="7.[151,1436,151,285]" box="[199,1234,260,285]" pageId="7">
<emphasis id="79C9610AFF85FFFA3DEFB21F5C50C145" bold="true" box="[199,334,260,285]" pageId="7">Etymology.</emphasis>
This species is named after Dr. Hans Huijbregts (RMNH, Leiden,
<collectingCountry id="33AAFD88FF85FFFA3911B21F59D8C145" box="[1081,1222,260,285]" name="Netherlands" pageId="7">Netherlands</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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