treatments-xml/data/5B/56/E8/5B56E872FF954A5DFF328808FECDEC4E.xml

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<document id="807B7F0235C28FF584EF3DC0FF08ABE2" ID-CLB-Dataset="26920" ID-DOI="10.11646/zootaxa.4599.1.1" ID-GBIF-Dataset="ea727a9b-324b-4c6b-a625-bc7cdd204473" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668058" ID-ZooBank="7CDD24FF-5BA3-4D06-898D-A3E220515D4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127209543" checkinUser="plazi" docAuthor="Longino, John T." docDate="2019" docId="5B56E872FF954A5DFF328808FECDEC4E" docLanguage="en" docName="zootaxa.4599.1.1.pdf.imf" docOrigin="Zootaxa 4599 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole andersoni Longino 2019, new species" docType="treatment" docVersion="23" lastPageNumber="23" masterDocId="A76F900AFF804A4BFFA58B6FFFF4EF5B" masterDocTitle="Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest" masterLastPageNumber="126" masterPageNumber="1" pageNumber="22" updateTime="1732580243751" updateUser="ExternalLinkService">
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<mods:title id="B42B2B013A1420F64822C2A5555479BB">Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest</mods:title>
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<mods:namePart id="B77CF1222AD45F81F9ED5F2895160B42">Longino, John T.</mods:namePart>
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<mods:date id="BCBECAF19BF66E32F388B9C10090DA4F">2019</mods:date>
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<mods:number id="4090ECB640B21598B0E0A2098944F987">2019-05-06</mods:number>
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<treatment id="5B56E872FF954A5DFF328808FECDEC4E" ID-DOI="http://doi.org/10.5281/zenodo.5586163" ID-GBIF-Taxon="157062586" ID-Zenodo-Dep="5586163" LSID="urn:lsid:plazi:treatment:5B56E872FF954A5DFF328808FECDEC4E" httpUri="http://treatment.plazi.org/id/5B56E872FF954A5DFF328808FECDEC4E" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="9BE50AEFFF954A5EFF328808FEF1ECF8" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="D3405964FF954A5EFF328808FDECECD9" blockId="21.[151,536,871,931]" box="[151,536,871,898]" pageId="21" pageNumber="22">
<heading id="8808EE08FF954A5EFF328808FDECECD9" bold="true" box="[151,536,871,898]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="E18B8576FF954A5EFF328808FDECECD9" bold="true" box="[151,536,871,898]" pageId="21" pageNumber="22">
<taxonomicName id="14FF22E7FF954A5EFF328808FE74ECDA" ID-CoL="CMKTT" authority="Longino, 2019" authorityName="Longino" authorityYear="2019" box="[151,384,871,897]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="andersoni" status="sp. nov.">
<emphasis id="E18B8576FF954A5EFF328808FE74ECDA" bold="true" box="[151,384,871,897]" italics="true" pageId="21" pageNumber="22">Pheidole andersoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAB8380DFF954A5EFE2D8807FDECECD9" box="[392,536,872,898]" pageId="21" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D3405964FF954A5EFF3288E5FEF1ECF8" blockId="21.[151,536,871,931]" box="[151,261,906,931]" pageId="21" pageNumber="22">
(
<figureCitation id="4BC445E1FF954A5EFF3B88E5FF08ECF8" box="[158,252,906,931]" captionStart="PLATE 12" captionStartId="78.[151,238,1954,1977]" captionTargetId="figure@78.[202,1384,194,1933]" captionTargetPageId="78" captionText="PLATE 12." httpUri="https://zenodo.org/record/2668253/files/figure.png" pageId="21" pageNumber="22">Plate 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF954A5EFF3288BDFC6FECB0" box="[151,923,978,1003]" pageId="21" pageNumber="22" type="description">
<paragraph id="D3405964FF954A5EFF3288BDFC6FECB0" blockId="21.[151,1437,978,2011]" box="[151,923,978,1003]" pageId="21" pageNumber="22">
<taxonomicName id="14FF22E7FF954A5EFF3288BDFF0FECB0" authorityName="Biolleyi Forel" authorityYear="1908" box="[151,251,978,1003]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="E18B8576FF954A5EFF3288BDFF0FECB0" box="[151,251,978,1003]" italics="true" pageId="21" pageNumber="22">Pheidole</emphasis>
</taxonomicName>
JTL-230: morphospecies code previously used on AntWeb.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF954A5EFF628898FCF1EBC4" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="D3405964FF954A5EFF628898FEA8EB20" blockId="21.[151,1437,978,2011]" pageId="21" pageNumber="22">
<materialsCitation id="63975339FF954A5EFF628898FE87EB0C" ID-GBIF-Occurrence="2237851471" collectingDate="2007-07-15" collectionCode="UNAM" collectorName="J. Longino" country="Mexico" elevation="0" location="Tonala" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<typeStatus id="0C44E7C6FF954A5EFF628898FEABEB54" box="[199,351,1015,1039]" pageId="21" pageNumber="22" type="holotype">HOLOTYPE</typeStatus>
: 1 major worker,
<collectingCountry id="ABE819F4FF954A5EFD848899FD8DEB54" box="[545,633,1014,1039]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
,
<collectingRegion id="113B9786FF954A5EFD218899FD17EB54" box="[644,739,1014,1039]" country="Mexico" name="Chiapas" pageId="21" pageNumber="22">Chiapas</collectingRegion>
:
<quantity id="1407F481FF954A5EFD488899FCD8EB54" box="[749,812,1014,1039]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="22" unit="km" value="19.0">19km</quantity>
ENE
<location id="D6200FBFFF954A5EFCD58899FC4BEB54" LSID="urn:lsid:plazi:treatment:5B56E872FF954A5DFF328808FECDEC4E:D6200FBFFF954A5EFCD58899FC4BEB54" box="[880,959,1014,1039]" country="Mexico" name="Tonala" pageId="21" pageNumber="22" stateProvince="Chiapas">Tonalá</location>
, 16.15630 -93.59920 ±
<elevation id="58D2BE57FF954A5EFB648898FA98EB55" box="[1217,1388,1014,1039]" isEnumeration="true" pageId="21" pageNumber="22">
<quantity id="1407F481FF954A5EFB648898FAFEEB55" box="[1217,1290,1015,1039]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="m" value="100.0">100 m</quantity>
,
<quantity id="1407F481FF954A5EFAB08899FA98EB55" box="[1301,1388,1014,1039]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="21" pageNumber="22" unit="m" value="1250.0">1250 m</quantity>
</elevation>
,
<date id="A7417FA4FF954A5EFAD28899FF0FEB68" pageId="21" pageNumber="22" value="2007-07-15">
<collectingDate id="B705864CFF954A5EFAD28899FF0FEB68" pageId="21" pageNumber="22" value="2007-07-15">15- Jul-2007</collectingDate>
</date>
, second growth wet forest, nest in clay soil (
<collectorName id="7E0A3CB2FF954A5EFD5A8F74FC8FEB68" box="[767,891,1050,1075]" pageId="21" pageNumber="22">J. Longino</collectorName>
, JTL6054) [
<collectionCode id="B5EEC1A1FF954A5EFBA08F74FB94EB68" box="[1029,1120,1051,1075]" country="Mexico" name="Universidad Nacional Autonoma de Mexico" pageId="21" pageNumber="22">UNAM</collectionCode>
, unique specimen identifier CASENT0645179]
</materialsCitation>
.
<materialsCitation id="63975339FF954A5EFE248F50FEADEB20" ID-GBIF-Occurrence="2237851436" collectingDate="2007-07-15" collectionCode="CAS, DZUP, JTLC, MCZC, UNAM, USNM" collectorName="J. Longino" country="Mexico" elevation="0" location="Tonala" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<typeStatus id="0C44E7C6FF954A5EFE248F50FDD4EB0C" box="[385,544,1087,1111]" pageId="21" pageNumber="22" type="paratype">PARATYPES</typeStatus>
: major, minor workers: same data as holotype [
<collectionCode id="B5EEC1A1FF954A5EFBF48F50FB78EB0C" box="[1105,1164,1087,1111]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="21" pageNumber="22" type="Museum">CAS</collectionCode>
,
<collectionCode id="B5EEC1A1FF954A5EFB398F50FB11EB0D" box="[1180,1253,1087,1110]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="21" pageNumber="22" type="University or college">DZUP</collectionCode>
,
<collectionCode id="B5EEC1A1FF954A5EFB518F50FAC3EB0C" box="[1268,1335,1087,1111]" pageId="21" pageNumber="22">JTLC</collectionCode>
,
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,
<collectionCode id="B5EEC1A1FF954A5EFF328F0CFF1BEB20" box="[151,239,1123,1147]" country="Mexico" name="Universidad Nacional Autonoma de Mexico" pageId="21" pageNumber="22">UNAM</collectionCode>
,
<collectionCode id="B5EEC1A1FF954A5EFF5F8F0CFEA5EB20" box="[250,337,1123,1147]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22" type="Museum">USNM</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="D3405964FF954A5EFF628FE9FCF1EBC4" blockId="21.[151,1437,978,2011]" box="[199,773,1158,1183]" pageId="21" pageNumber="22">
<emphasis id="E18B8576FF954A5EFF628FE9FE51EBC4" bold="true" box="[199,421,1158,1183]" pageId="21" pageNumber="22">Geographic range.</emphasis>
<collectingCountry id="ABE819F4FF954A5EFE098FE9FDF5EBC4" box="[428,513,1158,1183]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
(
<collectingRegion id="113B9786FF954A5EFDB48FE9FD85EBC4" box="[529,625,1158,1183]" country="Mexico" name="Chiapas" pageId="21" pageNumber="22">Chiapas</collectingRegion>
),
<collectingCountry id="ABE819F4FF954A5EFD218FE9FCF5EBC4" box="[644,769,1158,1183]" name="Guatemala" pageId="21" pageNumber="22">Guatemala</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF954A5EFF628FC5FD51E95C" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="D3405964FF954A5EFF628FC5FD51E95C" blockId="21.[151,1437,978,2011]" pageId="21" pageNumber="22">
<emphasis id="E18B8576FF954A5EFF628FC5FE6FEB98" bold="true" box="[199,411,1194,1219]" pageId="21" pageNumber="22">
Diagnosis.
<emphasis id="E18B8576FF954A5EFEF78FC5FE6FEB98" bold="true" box="[338,411,1194,1219]" italics="true" pageId="21" pageNumber="22">Minor</emphasis>
</emphasis>
: face and mesosoma foveolate, with foveolation faint to absent on lower half of katepisternum; promesonotal groove absent; propodeal spines one fourth to one fifth length of posterior face of propodeum; gaster smooth and shining; abundant erect setae on mesosomal dorsum and gaster; tibiae with 2-3 erect setae; color orange.
<emphasis id="E18B8576FF954A5EFE6D8E79FDFBEA74" bold="true" box="[456,527,1302,1327]" italics="true" pageId="21" pageNumber="22">Major</emphasis>
: inner hypostomal teeth distinct, closely spaced; scape base terete; face with distinct scrobes, delimited dorsally by frontal carinae and forming concave trough below them, ventral and posterior margins less delimited, surface of scrobe smooth and shiny; head weakly depressed posteriorly; face surface generally shiny, space between frontal carinae smooth, space between eye and antennal fossa and vertex lobes with widely separated, subparallel, longitudinal carinae; propodeal spines one half length of posterior face of propodeum; gastral dorsum smooth and shining; sides of head with abundant erect setae; abundant flexuous erect setae on mesosomal dorsum, tibiae, and gaster.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF954A5EFF628D7DFD9BE9CC" pageId="21" pageNumber="22" type="description">
<paragraph id="D3405964FF954A5EFF628D7DFD9BE914" blockId="21.[151,1437,978,2011]" pageId="21" pageNumber="22">
<emphasis id="E18B8576FF954A5EFF628D7DFDD9E970" bold="true" box="[199,557,1554,1579]" pageId="21" pageNumber="22">Measurements, minor worker</emphasis>
: HW 0.45, HL 0.49, SL 0.41, EL 0.10, WL 0.56, PSL 0.03, PTW 0.08, PPW 0.14, CI 91, SI 93, PSLI 6, PPI 168 (n=5).
</paragraph>
<paragraph id="D3405964FF954A5EFF628D35FD9BE9CC" blockId="21.[151,1437,978,2011]" pageId="21" pageNumber="22">
<emphasis id="E18B8576FF954A5EFF628D35FDD9E928" bold="true" box="[199,557,1626,1651]" pageId="21" pageNumber="22">Measurements, major worker</emphasis>
: HW 0.81, HL 0.98, SL 0.44, EL 0.14, WL 0.82, PSL 0.07, PTW 0.19, PPW 0.33, CI 83, SI 54, PSLI 8, PPI 176 (n=5).
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF954A5EFF628DCDFB2BE810" pageId="21" pageNumber="22" type="biology_ecology">
<paragraph id="D3405964FF954A5EFF628DCDFB2BE810" blockId="21.[151,1437,978,2011]" pageId="21" pageNumber="22">
<emphasis id="E18B8576FF954A5EFF628DCDFED3E9E0" bold="true" box="[199,295,1698,1723]" pageId="21" pageNumber="22">Biology.</emphasis>
This species occurs in cloud forest habitats. Minor and major workers recruit to baits on the forest floor and occur in Winkler samples. Two nests have been observed. One was in a vertical clay bank at a trail edge. A small cylindrical clay turret extended horizontally from the bank,
<quantity id="1407F481FF954A5EFC158D85FBEDE859" box="[944,1049,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="21" pageNumber="22" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5-10 mm</quantity>
long and
<quantity id="1407F481FF954A5EFB348D84FB18E859" box="[1169,1260,1771,1795]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="21" pageNumber="22" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
wide. A major worker was at the entrance. A partial excavation revealed a chamber a few cm deep with many minor and major workers. A second nest was in bare soil of a road in a coffee farm (no further observation details).
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF954A5DFF628C39FDBBED96" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="discussion">
<paragraph id="D3405964FF954A5DFF628C39FDA4EE3E" blockId="21.[151,1437,978,2011]" lastBlockId="22.[151,1437,151,789]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="E18B8576FF954A5EFF628C39FEBCE834" bold="true" box="[199,328,1878,1903]" pageId="21" pageNumber="22">Comments</emphasis>
. A set of species occurs from the southern
<collectingCountry id="ABE819F4FF954A5EFC988C38FC7EE834" box="[829,906,1879,1903]" name="United States of America" pageId="21" pageNumber="22">U.S.A.</collectingCountry>
to
<collectingCountry id="ABE819F4FF954A5EFC178C38FBDAE834" box="[946,1070,1878,1903]" name="Costa Rica" pageId="21" pageNumber="22">Costa Rica</collectingCountry>
that share a habitus. The minor workers of most species are similar to the common
<taxonomicName id="14FF22E7FF954A5EFD4C8C13FCA1E8C8" authorityName="Roger" authorityYear="1863" box="[745,853,1914,1939]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="flavens">
<emphasis id="E18B8576FF954A5EFD4C8C13FCA1E8C8" box="[745,853,1914,1939]" italics="true" pageId="21" pageNumber="22">P. flavens</emphasis>
</taxonomicName>
and thus easily escape detection in mixed samples and samples without major workers. The major workers have subrectangular heads, HW is in the 0.7-1.0 range, the antennae are short (SI 40-55), the antennal scrobes are well marked, the frontal carinae are expanded to form the pronounced dorsal margin of the scrobes, the face is generally shiny with widely separated longitudinal carinae distributed in various ways, the vertex lobes are pronounced and with a deep posterior sinus separating them, and in profile the posterior part of the head is often compressed. Most species are orange, but
<taxonomicName id="14FF22E7FF964A5DFB3F8B8EFAF0EFA3" authorityName="Longino" authorityYear="2019" box="[1178,1284,225,248]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="zannia">
<emphasis id="E18B8576FF964A5DFB3F8B8EFAF0EFA3" box="[1178,1284,225,248]" italics="true" pageId="22" pageNumber="23">P. zannia</emphasis>
</taxonomicName>
in
<collectingCountry id="ABE819F4FF964A5DFA8A8BB0FA68EFA3" box="[1327,1436,223,248]" name="Honduras" pageId="22" pageNumber="23">Honduras</collectingCountry>
appears to be in a mimicry complex and has divergent harlequin color, with black head and gaster and sharply contrasting orange mesosoma. This species also has the minor worker face smooth and shiny rather than foveolate. All of the species appear to nest in soil.
</paragraph>
<paragraph id="D3405964FF964A5DFF628A00FDBBED96" blockId="22.[151,1437,151,789]" pageId="22" pageNumber="23">
The species are distributed in a mosaic across Middle America, with various degrees of allopatry, parapatry, and narrow sympatry.
<taxonomicName id="14FF22E7FF964A5DFE358AFBFDC1EEF7" authorityName="Wilson" authorityYear="2003" box="[400,565,404,429]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mera">
<emphasis id="E18B8576FF964A5DFE358AFBFDC1EEF7" box="[400,565,404,429]" italics="true" pageId="22" pageNumber="23">Pheidole mera</emphasis>
</taxonomicName>
is known from south
<collectingRegion id="113B9786FF964A5DFC8D8AFAFC9DEEF7" box="[808,873,405,428]" country="United States of America" name="Texas" pageId="22" pageNumber="23">Texas</collectingRegion>
in the
<collectingCountry id="ABE819F4FF964A5DFC138AFBFBF7EEF7" box="[950,1027,404,428]" name="United States of America" pageId="22" pageNumber="23">U.S.A.</collectingCountry>
<taxonomicName id="14FF22E7FF964A5DFBAC8AFBFB10EEF6" authorityName="Longino" authorityYear="2019" box="[1033,1252,404,429]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="E18B8576FF964A5DFBAC8AFBFB10EEF6" box="[1033,1252,404,429]" italics="true" pageId="22" pageNumber="23">Pheidole andersoni</emphasis>
</taxonomicName>
is a cloud forest species in the Sierra Madre de Chiapas and adjacent parts of
<collectingCountry id="ABE819F4FF964A5DFCF28AD8FC20EE8B" box="[855,980,439,464]" name="Guatemala" pageId="22" pageNumber="23">Guatemala</collectingCountry>
.
<taxonomicName id="14FF22E7FF964A5DFC458AD8FB8AEE8B" authorityName="Longino" authorityYear="2019" box="[992,1150,439,464]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tikal">
<emphasis id="E18B8576FF964A5DFC458AD8FB8AEE8B" box="[992,1150,439,464]" italics="true" pageId="22" pageNumber="23">Pheidole tikal</emphasis>
</taxonomicName>
is in the
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region of
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.
<taxonomicName id="14FF22E7FF964A5DFE808AB3FE15EEAF" authorityName="Longino" authorityYear="2019" box="[293,481,476,501]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="zannia">
<emphasis id="E18B8576FF964A5DFE808AB3FE15EEAF" box="[293,481,476,501]" italics="true" pageId="22" pageNumber="23">Pheidole zannia</emphasis>
</taxonomicName>
occurs in cloud forests of central
<collectingCountry id="ABE819F4FF964A5DFC2B8AB3FC0BEEAE" box="[910,1023,476,501]" name="Honduras" pageId="22" pageNumber="23">Honduras</collectingCountry>
.
<taxonomicName id="14FF22E7FF964A5DFBB58AB3FB3BEEAE" authorityName="Longino" authorityYear="2019" box="[1040,1231,476,501]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="natalie">
<emphasis id="E18B8576FF964A5DFBB58AB3FB3BEEAE" box="[1040,1231,476,501]" italics="true" pageId="22" pageNumber="23">Pheidole natalie</emphasis>
</taxonomicName>
has a somewhat disjointed distribution, with morphologically very similar populations in cloud forests of southern
<collectingCountry id="ABE819F4FF964A5DFAE28A90FA68ED43" box="[1351,1436,511,536]" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingCountry>
(
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), northern
<collectingCountry id="ABE819F4FF964A5DFEDC894BFE12ED66" box="[377,486,548,573]" name="Honduras" pageId="22" pageNumber="23">Honduras</collectingCountry>
(Cusuco), and
<collectingCountry id="ABE819F4FF964A5DFD38894BFCE0ED66" box="[669,788,548,573]" name="Nicaragua" pageId="22" pageNumber="23">Nicaragua</collectingCountry>
.
<taxonomicName id="14FF22E7FF964A5DFC87894BFBD4ED66" authorityName="Longino" authorityYear="2019" box="[802,1056,548,573]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="corniclypeus">
<emphasis id="E18B8576FF964A5DFC87894BFBD4ED66" box="[802,1056,548,573]" italics="true" pageId="22" pageNumber="23">Pheidole corniclypeus</emphasis>
</taxonomicName>
occurs in lowland rainforests in eastern
<collectingCountry id="ABE819F4FF964A5DFF4B8928FEAAED3B" box="[238,350,583,608]" name="Honduras" pageId="22" pageNumber="23">Honduras</collectingCountry>
,
<collectingCountry id="ABE819F4FF964A5DFECC8928FE2BED3B" box="[361,479,583,608]" name="Nicaragua" pageId="22" pageNumber="23">Nicaragua</collectingCountry>
, and possibly into northeastern
<collectingCountry id="ABE819F4FF964A5DFCE08927FC30ED3B" box="[837,964,583,608]" name="Costa Rica" pageId="22" pageNumber="23">Costa Rica</collectingCountry>
.
<taxonomicName id="14FF22E7FF964A5DFC6B8928FB38ED3B" authorityName="Longino" authorityYear="2019" box="[974,1228,583,608]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="E18B8576FF964A5DFC6B8928FB38ED3B" box="[974,1228,583,608]" italics="true" pageId="22" pageNumber="23">Pheidole costaricensis</emphasis>
</taxonomicName>
occurs in lowland to lower montane wet forest in
<collectingCountry id="ABE819F4FF964A5DFE5B8903FD88EDDE" box="[510,636,620,645]" name="Costa Rica" pageId="22" pageNumber="23">Costa Rica</collectingCountry>
, and there is the potential for another lowland Pacific slope species in
<collectingCountry id="ABE819F4FF964A5DFF3289FFFEE2EDF3" box="[151,278,655,680]" name="Costa Rica" pageId="22" pageNumber="23">Costa Rica</collectingCountry>
. The one well-documented case of sympatry is the co-occurrence of
<taxonomicName id="14FF22E7FF964A5DFB8289FEFB66EDF3" authorityName="Longino" authorityYear="2019" box="[1063,1170,655,680]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="natalie">
<emphasis id="E18B8576FF964A5DFB8289FEFB66EDF3" box="[1063,1170,655,680]" italics="true" pageId="22" pageNumber="23">P. natalie</emphasis>
</taxonomicName>
and
<taxonomicName id="14FF22E7FF964A5DFB6E89FEFA83EDF3" authorityName="Longino" authorityYear="2019" box="[1227,1399,655,680]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="corniclypeus">
<emphasis id="E18B8576FF964A5DFB6E89FEFA83EDF3" box="[1227,1399,655,680]" italics="true" pageId="22" pageNumber="23">P. corniclypeus</emphasis>
</taxonomicName>
on one mountainside in central
<collectingCountry id="ABE819F4FF964A5DFE7189DBFDBFED96" box="[468,587,692,717]" name="Nicaragua" pageId="22" pageNumber="23">Nicaragua</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF964A5DFF6289B8FECDEC4E" pageId="22" pageNumber="23" type="etymology">
<paragraph id="D3405964FF964A5DFF6289B8FECDEC4E" blockId="22.[151,1437,151,789]" pageId="22" pageNumber="23">
<emphasis id="E18B8576FF964A5DFF6289B8FEBCEDAB" bold="true" box="[199,328,727,752]" pageId="22" pageNumber="23">Etymology</emphasis>
: In honor of Bob Anderson, Coleopterist, who has contributed so much to Neotropical myrmecology.
</paragraph>
</subSubSection>
</treatment>
</document>