187 lines
23 KiB
XML
187 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.984.53587" ID-GBIF-Dataset="9d291585-91c0-45e5-94d9-ee28795291ba" ID-PMC="PMC7658186" ID-Pensoft-Pub="1313-2970-984-83" ID-Pensoft-UUID="1448149CA44B51ED9EE57DC3A924C430" ID-PubMed="33223870" ID-ZooBank="D8B945AE620949578BB1D96DDBB5AA44" ModsDocID="1313-2970-984-83" checkinTime="1604561368050" checkinUser="pensoft" docAuthor="Irwin, Michael E., Winterton, Shaun L. & Metz, Mark A." docDate="2020" docId="21FE030021BE59BCB84E85A8F1669B25" docLanguage="en" docName="ZooKeys 984: 83-132" docOrigin="ZooKeys 984" docSource="http://dx.doi.org/10.3897/zookeys.984.53587" docTitle="Calophytus schlingeri Irwin & Winterton & Metz 2020, sp. nov." docType="treatment" docUuid="76F032BE-4F50-46B1-908E-146A0BB5E1EE" docUuidSource="ZooBank" docVersion="4" id="1448149CA44B51ED9EE57DC3A924C430" lastPageNumber="83" masterDocId="1448149CA44B51ED9EE57DC3A924C430" masterDocTitle="New stiletto flies from New Caledonia (Therevidae, Agapophytinae)" masterLastPageNumber="132" masterPageNumber="83" pageNumber="83" updateTime="1668169746090" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New stiletto flies from New Caledonia (Therevidae, Agapophytinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Irwin, Michael E.</mods:namePart>
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<mods:affiliation>Emeritus, University of Illinois, Urbana-Champaign, Illinois, USA & University of Arizona, Tucson, USA</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Winterton, Shaun L.</mods:namePart>
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<mods:affiliation>California State Collection of Arthropods, California Department of Food & Agriculture, Sacramento, California, USA</mods:affiliation>
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<mods:nameIdentifier type="email">wintertonshaun@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Metz, Mark A.</mods:namePart>
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<mods:affiliation>USDA, ARS, Systematic Entomology Laboratory, Beltsville, Maryland, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3535-535X</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>984</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>83</mods:start>
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<mods:end>132</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.984.53587</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.984.53587</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-984-83</mods:identifier>
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<mods:identifier type="ZooBank">D8B945AE620949578BB1D96DDBB5AA44</mods:identifier>
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<mods:identifier type="Pensoft-UUID">1448149CA44B51ED9EE57DC3A924C430</mods:identifier>
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<treatment ID-GBIF-Taxon="168756108" LSID="urn:lsid:zoobank.org:act:76F032BE-4F50-46B1-908E-146A0BB5E1EE" httpUri="http://treatment.plazi.org/id/21FE030021BE59BCB84E85A8F1669B25" lastPageNumber="83" pageId="0" pageNumber="83">
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<subSubSection pageId="0" pageNumber="83" type="nomenclature">
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<paragraph pageId="0" pageNumber="83">
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<taxonomicName LSID="21FE0300-21BE-59BC-B84E-85A8F1669B25" authority="Irwin & Winterton & Metz, 2020" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Agapophytinae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus schlingeri" order="Therevidae" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="schlingeri" status="sp. nov.">Calophytus schlingeri</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="83">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lateral view of antennae of Calophytus gen. nov. A C. chazeaui sp. nov. B C. grandiosus sp. nov. C C. schlingeri sp. nov. D C. matilei sp. nov. E C. webbi sp. nov. (figures not to scale) (drawings by J. Marie Metz)." figureDoi="10.3897/zookeys.984.53587.figure5" httpUri="https://binary.pensoft.net/fig/471321" pageId="0" pageNumber="83">Figs 5C</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scutum and scutellum of Calophytus gen. nov. and Jeanchazeauia gen. nov. A C. schlingeri sp. nov. B C. grandiosus sp. nov. C C. matilei sp. nov. D C. webbi sp. nov. E J. nubilosus sp. nov. (figures not to scale) (drawings by J. Marie Metz)." figureDoi="10.3897/zookeys.984.53587.figure6" httpUri="https://binary.pensoft.net/fig/471322" pageId="0" pageNumber="83">, 6A</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Wings of Calophytus gen. nov. and Jeanchazeauia gen. nov. A C. chazeaui sp. nov. B C. schlingeri sp. nov. C C. grandiosus sp. nov. D C. matilei sp. nov. E C. webbi sp. nov. F C. monteithi sp. nov. G J. amoa sp. nov. H J. nubilosus sp. nov. (female) (figures not to scale)." figureDoi="10.3897/zookeys.984.53587.figure7" httpUri="https://binary.pensoft.net/fig/471313" pageId="0" pageNumber="83">, 7B</figureCitation>
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Calophytus schlingeri sp. nov. A adult male (MEI 030158), anterolateral view B same, oblique view C adult female (MEI 030172), lateral view D same, oblique view. Body length: male: 8.6 mm; female: 9.8 mm." figureDoi="10.3897/zookeys.984.53587.figure16" httpUri="https://binary.pensoft.net/fig/471326" pageId="0" pageNumber="83">, 16</figureCitation>
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Calophytus schlingeri sp. nov., cleared male genitalia A epandrium B gonocoxites and aedeagus, ventral view C aedeagus, dorsal view D gonocoxites, ventral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.984.53587.figure17" httpUri="https://binary.pensoft.net/fig/471327" pageId="0" pageNumber="83">, 17</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Calophytus schlingeri sp. nov., genitalia A epandrium B gonocoxites, ventral view C same, lateral view D aedeagus, lateral view E same, dorsal view F female genitalia, lateral view, internal structures shown. Scale line: 0.2 mm. Abbreviations: ag, accessory gland; d, distiphallus; da, dorsal apodeme of parameral sheath; ea, ejaculatory apodeme; c, cercus; A 1, primary acanthophorite spines; A 2 secondary acanthophorite spines; ga, gonocoxal apodeme; gs, gonostylus; igp, inner gonocoxal process (articulated); lea, lateral ejaculatory apodeme; ogp, outer gonocoxal process; s 8, sternite VIII; s 10, sternite X; s, spermatheca; ss, spermathecal sac; ssd, spermathecal sac duct; sd, spermathecal duct; vl, ventral lobe." figureDoi="10.3897/zookeys.984.53587.figure18" httpUri="https://binary.pensoft.net/fig/471328" pageId="0" pageNumber="83">, 18</figureCitation>
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Distribution of Calophytus spp. collecting records throughout New Caledonia. Provinces are labelled and ecoregions delineated by colour." figureDoi="10.3897/zookeys.984.53587.figure22" httpUri="https://binary.pensoft.net/fig/471332" pageId="0" pageNumber="83">, 22</figureCitation>
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Female abdominal tergites VI-VIII A Calophytus schlingeri sp. nov. B Jeanchazeauia amoa sp. nov. C Jeanchazeauia nubilosus sp. nov. D Jeanchazeauia rufinatus sp. nov. Scale line: 0.2 mm." figureDoi="10.3897/zookeys.984.53587.figure23" httpUri="https://binary.pensoft.net/fig/471333" pageId="0" pageNumber="83">, 23A</figureCitation>
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<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Natural habitats of New Caledonian agapophytine stiletto flies A Riviere Bleue Reserve, sclerophyll forest and maquis scrub, habitat of C. chazeaui sp. nov., C. grandiosus sp. nov., C. schlingeri sp. nov., and J. nubilosus sp. nov. B Pindai Peninsula, tropical dry forest, habitat of C. webbi sp. nov. C Mount Dzumac, rainforest, habitat of J. nubilosus sp. nov. D Mount Dore, rainforest and dry sclerophyll forest E Mount Dore, Malaise trap in flyway (photographs by Michael E. Irwin)." figureDoi="10.3897/zookeys.984.53587.figure31" httpUri="https://binary.pensoft.net/fig/471341" pageId="0" pageNumber="83">, 31A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="83" type="diagnosis">
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<paragraph pageId="0" pageNumber="83">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="83">Flagellum shorter than scape; single row of postocular setae dorsally; scutum dark except for orange suffusion on notopleural callus; legs yellow except for tarsi dark apically; wing with dark infuscation, fenestrate apically; male abdomen with silver velutum pubescence.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="83" type="description">
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<paragraph pageId="0" pageNumber="83">Description.</paragraph>
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<paragraph pageId="0" pageNumber="83">
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Length 8.5-10.7 mm.
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<emphasis italics="true" pageId="0" pageNumber="83">Head</emphasis>
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. Glossy black. Frons smooth, raised around base of antenna. Eyes separated by slightly more than 2
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<normalizedToken originalValue="×">x</normalizedToken>
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width of median ocellus. Occiput black with silver or gold pubescence laterally, extending ventrally onto gena, admixed laterally and ventrally with light brown to white, fine setae; parafacial silver pubescent as elongate wedge shape, extending ventrally along margin of buccal area. Postocular macrosetae on occiput few in number, arranged dorsally in single row in both sexes, typically black, sometimes with orange suffusion. Scape 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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head length; yellow basally, black apically. Pedicel short, 1/8 length of scape, black, sparsely yellow pubescent, with numerous short, brown setae. Basal flagellomere
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<normalizedToken originalValue="½×">1/2x</normalizedToken>
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length of scape; elongate, gradually tapering to a blunt point apically with short, black setae covering entire length. Second flagellomere cylindrical. Third flagellomere subequal in length of second. Style subequal in length to third flagellomere, spiculate. Palpus more or less cylindrical, slightly capitate at apex; yellow basally, black at apex; yellow pubescent basally, black pubescent at apex; yellow setose basally, black setose at apex. Mouthparts yellow.
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<emphasis italics="true" pageId="0" pageNumber="83">Thorax</emphasis>
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. Dark brown to black, except prothorax, postpronotal lobe, and notopleuron yellow; prosternum, ventral proepimeron, posterodorsal katepisternum, posterior scutum, scutellum, subscutellum, metanepisternum, metepimeron, meron and metakatepisternum overlain with silver pubescence; pronotum, anatergite, and dorsal proepimeron gold-yellow pubescent; scutum with faint dorsocentral silver pubescent stripes; macrosetae black (np: 2; sa: 1; pa: 1, dc: 0, sc: 1); scutum with relatively short setae.
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<emphasis italics="true" pageId="0" pageNumber="83">Legs</emphasis>
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. Yellow, except for black suffusion on foretarsus and hind coxa, legs with pale setae except dark setae apically on femora and on all tarsi; coxae silver pubescent, especially on hind coxa; hind femur with posteroventral macroseta yellow; hind basal tarsomere with admix of white and gold setae ventrally and medially. Coxae with yellow setae anteriorly, hind coxa with two black lateral setae. Forecoxa with four marginal and one submarginal anteroventral, black and yellow macrosetae. Midcoxa with two marginal and three submarginal, anteroventral, black and yellow macrosetae. Hind coxa with anteroventral margin extended ventrally in a short point with three marginal, black or yellow macrosetae.
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<emphasis italics="true" pageId="0" pageNumber="83">Wing</emphasis>
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. Strongly marked, variable in shape, irregular. Costal cell with pale yellow infuscation; brown infuscation in cells sc, r adjacent to pterostigma, basal portions of r1 and r2+3, and r4+5, d, bm, basal portions of m3 and cua1, cup, and entire wing apex; infuscated areas also with dark microtrichia, hyaline areas of membrane largely void of microtrichia; pterostigma brown. Venation brown, except costa and R at base of wing gold; cell m3 widely open at wing margin. Haltere yellow.
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<emphasis italics="true" pageId="0" pageNumber="83">Abdomen</emphasis>
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. Segments mostly dark brown, males with tergites I-III and sternites II-V with portion of medial area lighter, light brown to dark yellow; female usually uniform dark brown although sometimes with lighter brown to dark yellow areas laterally on tergite II; male with tergites II-VI with silver velutum pubescence; tergites I-III with sparse short, pale, fine setae, longer laterally; tergite IV admixed with gold and black setae; remaining tergites with black setae.
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<emphasis italics="true" pageId="0" pageNumber="83">Genitalia</emphasis>
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. Male: epandrium narrowed posteriorly, with uniform scattered, dark setae. Cercus relatively narrow, rounded posteriorly. Subepandrial sclerite narrow, 1/3 width of epandrium; partially sclerotised, lateral margins more strongly so. Gonocoxites with scattered elongate brown setae; slightly denser ventrally; outer gonocoxal process sub-triangular in profile; inner gonocoxal process uniformly narrow, curved medially, spatulate apically. Parameral sheath of aedeagus with
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<normalizedToken originalValue="‘T’">'T'</normalizedToken>
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-shaped dorsal apodeme, arms slightly bifurcated, ventral apodeme fork relatively short, lobes rounded, distiphallus straight; basiphallus bulbous, lateral ejaculatory apodeme narrow, band-like with lateral process; Ejaculatory apodeme robust. Female: tergite VIII wider than long; anteromedial, projection long and narrow, 1/3
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<normalizedToken originalValue="×">x</normalizedToken>
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length of tergite VIII; posterior margin emarginated. Acanthophorite brown; acanthophorite spines brown admixed with short, brown setae. Sternite X sharply acute posteriorly. Spermathecal duct equal to four furcal lengths. Spermathecal sac duct equal to two furcal lengths; spermathecal sac ovoid, longer than wide; 1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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length of spermathecal sac duct.
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<emphasis italics="true" pageId="0" pageNumber="83">Variation</emphasis>
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. Some paratypes with frons sparsely pubescent; ventral parafacial variably silver and/or gold pubescent; all postocular macrosetae black; additional postocular macrosetae ventral to single dorsal row; apical 3/4 scape brown; basal flagellomere with covering of short, black, filiform setae variable from basal 1/4 to entire; notopleuron variably coloured light to dark brown; midcoxae variably gold to brown; coxae with anteroventral, marginal macrosetae number variable from 3-5 and variably coloured gold to black; hind coxa lateral macrosetae number variable, 0-2; wing infuscation variable in intensity and distribution.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="83" type="etymology">
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<paragraph pageId="0" pageNumber="83">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="83">Named in honour of the late Dr Evert I. Schlinger, who participated in the New Caledonian expeditions, from which much of the material for this paper was collected.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="83" type="comments">
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<paragraph pageId="0" pageNumber="83">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="83">
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<taxonomicName authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus schlingeri" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="schlingeri">
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<emphasis italics="true" pageId="0" pageNumber="83">Calophytus schlingeri</emphasis>
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</taxonomicName>
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sp. nov. is very similar to
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<taxonomicName lsidName="C. matilei" pageId="0" pageNumber="83" rank="species" species="matilei">
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<emphasis italics="true" pageId="0" pageNumber="83">C. matilei</emphasis>
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</taxonomicName>
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sp. nov. and can be distinguished based on the pattern of silver pubescence on the face, pattern of wing infuscation and whether the mid- and hind coxae are largely dark yellow or brown. These are the only known species in the genus with silver velutum pubescence on the male abdomen. Specimens of
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<taxonomicName authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus schlingeri" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="schlingeri">
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<emphasis italics="true" pageId="0" pageNumber="83">Calophytus schlingeri</emphasis>
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</taxonomicName>
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sp. nov. were collected in a variety of habitats including dry sclerophyll forest, rainforest and maquis scrub (Fig.
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<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Natural habitats of New Caledonian agapophytine stiletto flies A Riviere Bleue Reserve, sclerophyll forest and maquis scrub, habitat of C. chazeaui sp. nov., C. grandiosus sp. nov., C. schlingeri sp. nov., and J. nubilosus sp. nov. B Pindai Peninsula, tropical dry forest, habitat of C. webbi sp. nov. C Mount Dzumac, rainforest, habitat of J. nubilosus sp. nov. D Mount Dore, rainforest and dry sclerophyll forest E Mount Dore, Malaise trap in flyway (photographs by Michael E. Irwin)." figureDoi="10.3897/zookeys.984.53587.figure31" httpUri="https://binary.pensoft.net/fig/471341" pageId="0" pageNumber="83">31A</figureCitation>
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). GenBank sequences for this species (see
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<bibRefCitation DOI="https://doi.org/10.1111/syen.12147" author="Winterton, SL" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="83" pagination="144 - 161" refId="B18" refString="Winterton, SL, Hardy, NB, Gaimari, SD, Hauser, M, Hill, HN, Holston, KC, Irwin, ME, Lambkin, CL, Metz, MA, Turco, F, Webb, D, Yang, L, Yeates, DK, Wiegmann, BM, 2016. The phylogeny of stiletto flies (Diptera: Therevidae). Systematic Entomology 41: 144 - 161, DOI: https://doi.org/10.1111/syen.12147" title="The phylogeny of stiletto flies (Diptera: Therevidae)." url="https://doi.org/10.1111/syen.12147" volume="41" year="2016">Winterton et al. 2016</bibRefCitation>
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: table S1): KM885001 (28S rDNA), KM879117 (EF1a).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="83" type="materials_examined">
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<paragraph pageId="0" pageNumber="83">Specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="83">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Holotype</emphasis>
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male, New Caledonia: Province Sud: Plaine des Lacs, 5 km E of Grand Lac, hand netted [
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<geoCoordinate degrees="22.267" direction="south" orientation="latitude" precision="55" value="-22.267">-22.267</geoCoordinate>
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,
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<geoCoordinate degrees="166.967" direction="east" orientation="longitude" precision="55" value="166.967">166.967</geoCoordinate>
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], 300 m, 22-25.I.1984, M. Pogue, M. Epstein (MEI030164, MNHN).
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</paragraph>
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<paragraph pageId="0" pageNumber="83">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="83">Paratypes</emphasis>
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. New Caledonia: Province Sud: 1 male, 2 females, Plaine des Lacs, 5 km E of Grand Lac, black light (UV) [
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<geoCoordinate degrees="22.267" direction="south" orientation="latitude" precision="55" value="-22.267">-22.267</geoCoordinate>
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,
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<geoCoordinate degrees="166.967" direction="east" orientation="longitude" precision="55" value="166.967">166.967</geoCoordinate>
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], 300 m, 22-25.I.1984, M. Pogue, M. Epstein (MEI030165, 03173-4, UMSP); 4 males, 3 females, Plaine des Lacs, 5 km E of Grand Lac, hand netted [
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<geoCoordinate degrees="22.267" direction="south" orientation="latitude" precision="55" value="-22.267">-22.267</geoCoordinate>
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,
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<geoCoordinate degrees="166.967" direction="east" orientation="longitude" precision="55" value="166.967">166.967</geoCoordinate>
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], 300 m, 22-25.I.1984, M. Pogue, M. Epstein, male, (MEI030161-3, 030166, 030175-7, FSCA, CASC); 1 female, Mt. Koghis, Malaise trap in forest [
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<geoCoordinate degrees="22.167" direction="south" orientation="latitude" precision="55" value="-22.167">-22.167</geoCoordinate>
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,
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<geoCoordinate degrees="166.533" direction="east" orientation="longitude" precision="55" value="166.533">166.533</geoCoordinate>
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], 600 m, 2-5.XII.2001, T. Pape, B. Viklund (MEI135023, NHRS); 1 male,
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<normalizedToken originalValue="Rivière">Riviere</normalizedToken>
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des Pirogues black light (UV) [approximated as
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<geoCoordinate degrees="22.275" direction="south" orientation="latitude" precision="55" value="-22.275">-22.275</geoCoordinate>
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,
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<geoCoordinate degrees="166.687" direction="east" orientation="longitude" precision="55" value="166.687">166.687</geoCoordinate>
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], 7-9.II.1984, M. Pogue, M. Epstein (MEI030167, UMSP); 2 males, 4 females,
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<normalizedToken originalValue="Rivière">Riviere</normalizedToken>
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Bleue Provincial Park, 25.8 km
|
||
<normalizedToken originalValue="Rivière">Riviere</normalizedToken>
|
||
Bleue Road, Malaise trap across forest path [
|
||
<geoCoordinate degrees="22.117" direction="south" orientation="latitude" precision="55" value="-22.117">-22.117</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="166.658" direction="east" orientation="longitude" precision="55" value="166.658">166.658</geoCoordinate>
|
||
], 213 m, various dates: 5-28.XI.1992, D.W. Webb, E.I. & M. Schlinger (MEI030158-9, 030170-2, CSCA); 1 male,
|
||
<normalizedToken originalValue="Rivière">Riviere</normalizedToken>
|
||
Bleue Provincial Park,
|
||
<normalizedToken originalValue="Rivière">Riviere</normalizedToken>
|
||
Bleue Road, Malaise trap across forest path [
|
||
<geoCoordinate degrees="22.117" direction="south" orientation="latitude" precision="55" value="-22.117">-22.117</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="166.658" direction="east" orientation="longitude" precision="55" value="166.658">166.658</geoCoordinate>
|
||
], 290 m, 5-16.XI.1992, D.W. Webb, male, (MEI030160, CSCA); 1 female, 17 km NNE
|
||
<normalizedToken originalValue="Nouméa">Noumea</normalizedToken>
|
||
, Mt. Koghis, [Koghis stream waterfall] Malaise trap in tropical forest [
|
||
<geoCoordinate degrees="22.171" direction="south" orientation="latitude" precision="55" value="-22.171">-22.171</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="166.512" direction="east" orientation="longitude" precision="55" value="166.512">166.512</geoCoordinate>
|
||
], 500 m, 21-29.XI.1992, D.W. Webb, (MEI030169, CSCA).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |