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<document id="325C8D2ECE1673AE428610271740A831" ID-CLB-Dataset="42402" ID-DOI="10.11646/zootaxa.3872.5.9" ID-GBIF-Dataset="099868dc-c782-452c-aaa8-d3375dcbb364" ID-ISSN="1175-5326" ID-Zenodo-Dep="251251" ID-ZooBank="41BDCF15-0657-4CC8-82DC-37D04C5DDE38" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460933014911" checkinUser="plazi" docAuthor="Jin, Mengjie, Weir, Tom, Ślipiński, Adam &amp; Pang, Hong" docDate="2014" docId="214087F6FF8F0838FF262E77CA4E028E" docLanguage="en" docName="zt03872p600.pdf" docOrigin="Zootaxa 3872 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala katherine Jin, Weir, Slipinski" docType="treatment" docVersion="9" lastPageNumber="599" masterDocId="DD79FF8EFF8D0830FFB12D20C9180666" masterDocTitle="Description of a new species of Anomala Samouelle (Coleoptera: Scarabaeidae: Rutelinae) from Northern Territory, Australia" masterLastPageNumber="600" masterPageNumber="591" pageNumber="593" updateTime="1698582436333" updateUser="ExternalLinkService">
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<mods:title id="C148DB075785E6FD8D85465CFF495A8A">Description of a new species of Anomala Samouelle (Coleoptera: Scarabaeidae: Rutelinae) from Northern Territory, Australia</mods:title>
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<mods:namePart id="57864076DDEA26755C8AED930556404A">Jin, Mengjie</mods:namePart>
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<mods:namePart id="94D9E627B6BC5965A98FC050BCC092BE">Weir, Tom</mods:namePart>
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<mods:namePart id="BDA0339ADD54E3CD66CDFA80D2294E27">Ślipiński, Adam</mods:namePart>
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<mods:namePart id="B5334846A87DDA43C132B3F5A7902A98">Pang, Hong</mods:namePart>
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<treatment id="214087F6FF8F0838FF262E77CA4E028E" ID-DOI="http://doi.org/10.5281/zenodo.6141594" ID-GBIF-Taxon="119590228" ID-Zenodo-Dep="6141594" LSID="urn:lsid:plazi:treatment:214087F6FF8F0838FF262E77CA4E028E" httpUri="http://treatment.plazi.org/id/214087F6FF8F0838FF262E77CA4E028E" lastPageId="8" lastPageNumber="599" pageId="2" pageNumber="593">
<subSubSection id="E1F3656BFF8F0832FF262E77CB7D05F2" pageId="2" pageNumber="593" type="nomenclature">
<paragraph id="A95636E0FF8F0832FF262E77CA650514" blockId="2.[151,893,855,916]" box="[151,893,855,882]" pageId="2" pageNumber="593">
<heading id="F21E818CFF8F0832FF262E77CA650514" bold="true" box="[151,893,855,882]" fontSize="11" level="1" pageId="2" pageNumber="593" reason="1">
<emphasis id="9B9DEAF2FF8F0832FF262E77CA650514" bold="true" box="[151,893,855,882]" pageId="2" pageNumber="593">
<taxonomicName id="6EE94D63FF8F0832FF262E77CB6E0514" authority="Jin, Weir, Slipinski" authorityName="Jin, Weir, Slipinski" box="[151,630,855,882]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="katherine">
<emphasis id="9B9DEAF2FF8F0832FF262E77C8990517" bold="true" box="[151,385,855,881]" italics="true" pageId="2" pageNumber="593">Anomala katherine</emphasis>
Jin, Weir, Ślipiński
</taxonomicName>
, &amp; Pang,new species
</emphasis>
</heading>
</paragraph>
<paragraph id="A95636E0FF8F0832FF262E5BCB7D05F2" blockId="2.[151,893,855,916]" box="[151,613,891,916]" pageId="2" pageNumber="593">
(
<figureCitation id="31D22A65FF8F0832FF2E2E5BC83A05F5" box="[159,290,891,916]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="2" pageNumber="593">Figs. 1AL</figureCitation>
,
<figureCitation id="31D22A65FF8F0832FE9F2E5BC87305F5" box="[302,363,891,915]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="2" pageNumber="593">2AJ</figureCitation>
,
<figureCitation id="31D22A65FF8F0832FEC62E5BC8A805F5" box="[375,432,891,915]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="2" pageNumber="593">
<date id="DD571020FF8F0832FEC62E5BC89505F5" box="[375,397,891,915]" pageId="2" pageNumber="593">3I</date>
L
</figureCitation>
,
<figureCitation id="31D22A65FF8F0832FE0A2E5BCB5A05F5" box="[443,578,891,915]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="2" pageNumber="593">4CE, HK</figureCitation>
,
<figureCitation id="31D22A65FF8F0832FDFF2E5BCB4405F2" box="[590,604,891,916]" captionStart="FIGURE 5" captionStartId="7.[151,250,1672,1694]" captionTargetBox="[180,1403,697,1646]" captionTargetId="figure@7.[178,1409,697,1651]" captionTargetPageId="7" captionText="FIGURE 5. Distribution map: Anomala antiqua (Gyllenhal) (blue circles), Anomala katherine new species (red triangles)." httpUri="https://zenodo.org/record/251256/files/figure.png" pageId="2" pageNumber="593">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E1F3656BFF8F0832FF262EE3CDAE029A" pageId="2" pageNumber="593" type="diagnosis">
<paragraph id="A95636E0FF8F0832FF262EE3CDAE029A" blockId="2.[151,1437,963,2001]" pageId="2" pageNumber="593">
<emphasis id="9B9DEAF2FF8F0832FF262EE3C80905BD" bold="true" box="[151,273,963,988]" pageId="2" pageNumber="593">Diagnosis.</emphasis>
<taxonomicName id="6EE94D63FF8F0832FEAC2EE3C8E105BD" box="[285,505,963,987]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="katherine">
<emphasis id="9B9DEAF2FF8F0832FEAC2EE3C8E105BD" box="[285,505,963,987]" italics="true" pageId="2" pageNumber="593">Anomala katherine</emphasis>
</taxonomicName>
is distinguished from the similar species by the combination of the following characters: clypeus weakly reflexed along anterior margin (
<figureCitation id="31D22A65FF8F0832FC822EC6CA930599" box="[819,907,998,1023]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="2" pageNumber="593">Fig. 3L</figureCitation>
); maxilla with five sclerotised teeth on apex of galea (
<figureCitation id="31D22A65FF8F0832FF55292BC82A0245" box="[228,306,1035,1060]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="2" pageNumber="593">
Fig.
<date id="DD571020FF8F0832FEAB292BC82A0245" box="[282,306,1035,1059]" pageId="2" pageNumber="593">1I</date>
</figureCitation>
); protibia with single tooth along external margin (
<figureCitation id="31D22A65FF8F0832FC33292BCAC20245" box="[898,986,1035,1060]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="2" pageNumber="593">Fig. 2F</figureCitation>
); pronotum with marginal bead at base incomplete medially (
<figureCitation id="31D22A65FF8F0832FE25290EC8E90221" box="[404,497,1070,1095]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="2" pageNumber="593">Fig. 4C</figureCitation>
); abdominal ventrites with long and sparse setae forming transverse rows in the middle with most of the setae directed posteriorly and surpassing posterior margins of their ventrites; pygidium uniformly yellow or brown (
<figureCitation id="31D22A65FF8F0832FE662956CB3D02E9" box="[471,549,1142,1167]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="2" pageNumber="593">
Fig.
<date id="DD571020FF8F0832FDBD2957CB3D02E9" box="[524,549,1143,1167]" pageId="2" pageNumber="593">3I</date>
</figureCitation>
); phallobase longer than parameres with well sclerotised ventral plate (
<figureCitation id="31D22A65FF8F0832FAF42956CC9602E9" box="[1349,1422,1142,1167]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="2" pageNumber="593">Fig 3J</figureCitation>
), the plate does not extend to parameres; parameres fused at bases but broadly separated apically; endophallus with single setal patch. The new species appears to be very similar to
<taxonomicName id="6EE94D63FF8F0832FCDF299FCDF302B1" authority="Ohaus (Ohaus 1916)" authorityName="Ohaus (Ohaus" authorityYear="1916" box="[878,1259,1214,1239]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="crypsinoa">
<emphasis id="9B9DEAF2FF8F0832FCDF299FCAE402B1" box="[878,1020,1215,1239]" italics="true" pageId="2" pageNumber="593">A. crypsinoa</emphasis>
<bibRefCitation id="CD784B11FF8F0832FBB2299ECDF302B1" box="[1027,1259,1214,1239]" pageId="2" pageNumber="593" refString="Ohaus, F. (1916) Beitrag zur Kenntnis der Ruteliden (Col. lamell.). Stettiner Entomologische Zeitung, 77, 39 - 113." type="journal article">Ohaus (Ohaus 1916)</bibRefCitation>
</taxonomicName>
described from “Moa und
<collectingCountry id="D1FE7670FF8F0832FEBF29C4C849029D" box="[270,337,1252,1275]" name="Malaysia" pageId="2" pageNumber="593">Dama</collectingCountry>
Inseln bei Timor” but differs in having parameres broadly separated apically.
</paragraph>
</subSubSection>
<subSubSection id="E1F3656BFF8F0832FF762826C8280322" pageId="2" pageNumber="593" type="etymology">
<paragraph id="A95636E0FF8F0832FF762826C8280322" blockId="2.[151,1437,963,2001]" pageId="2" pageNumber="593">
<emphasis id="9B9DEAF2FF8F0832FF762826C8560379" bold="true" box="[199,334,1286,1311]" pageId="2" pageNumber="593">Etymology.</emphasis>
Named after its
<typeStatus id="76528842FF8F0832FDB62828CB2F0346" box="[519,567,1288,1312]" pageId="2" pageNumber="593">type</typeStatus>
locality
<taxonomicName id="6EE94D63FF8F0832FD292826CA1E0379" box="[664,774,1286,1311]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">Katherine</taxonomicName>
Gorge in the Northern Territory of
<collectingCountry id="D1FE7670FF8F0832FB232826CDE40379" box="[1170,1276,1286,1311]" name="Australia" pageId="2" pageNumber="593">Australia</collectingCountry>
; name is noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="E1F3656BFF8F0837FF76286EC8A30140" lastPageId="7" lastPageNumber="598" pageId="2" pageNumber="593" type="description">
<paragraph id="A95636E0FF8F0832FF76286ECA230301" blockId="2.[151,1437,963,2001]" box="[199,827,1358,1383]" pageId="2" pageNumber="593">
<emphasis id="9B9DEAF2FF8F0832FF76286EC8400301" bold="true" box="[199,344,1358,1383]" pageId="2" pageNumber="593">Description.</emphasis>
Length 11.013.0 mm; width 6.0
<quantity id="6E119B05FF8F0832FD6F286ECA2E0301" box="[734,822,1358,1383]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="593" unit="mm" value="7.5">7.5 mm</quantity>
.
</paragraph>
<paragraph id="A95636E0FF8F0832FF762853CDF403B2" blockId="2.[151,1437,963,2001]" pageId="2" pageNumber="593">
<emphasis id="9B9DEAF2FF8F0832FF762853C81303EA" bold="true" box="[199,267,1395,1420]" pageId="2" pageNumber="593">Male.</emphasis>
<emphasis id="9B9DEAF2FF8F0832FEA72853C87203ED" box="[278,362,1395,1419]" italics="true" pageId="2" pageNumber="593">Colour.</emphasis>
Dorsum light brown (
<figureCitation id="31D22A65FF8F0832FDC42853CBD003ED" box="[629,712,1395,1420]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="2" pageNumber="593">
Fig.
<date id="DD571020FF8F0832FD1F2853CBD003ED" box="[686,712,1395,1419]" pageId="2" pageNumber="593">3I</date>
</figureCitation>
), venter mostly yellowish brown. Elytra yellowish or reddish brown with metallic green or violet shine; head, anteromedian area and pronotal base, margins of scutellum and elytra usually darker to blackened; legs, particularly metalegs darker often with strong green shine.
</paragraph>
<paragraph id="A95636E0FF8F0832FF7628FFCBC300CA" blockId="2.[151,1437,963,2001]" pageId="2" pageNumber="593">
<emphasis id="9B9DEAF2FF8F0832FF7628FFC8130391" box="[199,267,1503,1527]" italics="true" pageId="2" pageNumber="593">Head.</emphasis>
Clypeus subrectangular with anterior and lateral margins weakly upturned (
<figureCitation id="31D22A65FF8F0832FBCE28FECDC10391" box="[1151,1241,1502,1527]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="2" pageNumber="593">Fig. 3L</figureCitation>
); frontal clypeus about as thick as labrum (
<figureCitation id="31D22A65FF8F0832FE062B23CB17007D" box="[439,527,1539,1564]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="2" pageNumber="593">Fig. 1C</figureCitation>
). Clypeus densely and coarsely punctate; punctures variable and often confluent; interspaces irregularly wrinkled and feebly shiny. Frontoclypeal suture distinct, straight. Frontal punctures similar in size and density to those on clypeus but punctures on vertex much sparser and smaller than frontal ones with smooth and more distinctly shiny interspaces. Ratio of frontal interocular distance to head width: 0.580.61. Antenna as in
<figureCitation id="31D22A65FF8F0832FE892BB3C89700CD" box="[312,399,1683,1708]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="2" pageNumber="593">Fig. 1H</figureCitation>
, club equal in length to shaft.
</paragraph>
<paragraph id="A95636E0FF8F0832FF762B98CA2E015A" blockId="2.[151,1437,963,2001]" pageId="2" pageNumber="593">
<emphasis id="9B9DEAF2FF8F0832FF762B98C82E00A9" box="[199,310,1720,1743]" italics="true" pageId="2" pageNumber="593">Pronotum</emphasis>
subtrapezoidial (
<figureCitation id="31D22A65FF8F0832FE482B96CB4C00A9" box="[505,596,1718,1743]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="2" pageNumber="593">Fig. 2A</figureCitation>
), widest at base, 0.60.7 times as long as broad, broadly curved anteriorly from a point shortly before middle. Anterior angles weakly acute; posterior angles obtuse. Pronotal base with marginal bead incomplete medially in front of scutellum. Pronotal punctures about as large as those on vertex, about 0.20.5 diameter apart; interspaces smooth and shiny.
</paragraph>
<paragraph id="A95636E0FF8F0832FF762A67CAD30139" blockId="2.[151,1437,963,2001]" box="[199,971,1862,1887]" pageId="2" pageNumber="593">
<emphasis id="9B9DEAF2FF8F0832FF762A67C82D0139" box="[199,309,1863,1887]" italics="true" pageId="2" pageNumber="593">Scutellum</emphasis>
(
<figureCitation id="31D22A65FF8F0832FEF42A66C8830139" box="[325,411,1862,1887]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="2" pageNumber="593">Fig. 4C</figureCitation>
) densely punctate, glabrous and rounded apically.
</paragraph>
<paragraph id="A95636E0FF8F0832FF762A4BCC0401B6" blockId="2.[151,1437,963,2001]" pageId="2" pageNumber="593">
<emphasis id="9B9DEAF2FF8F0832FF762A4BC80D01E5" box="[199,277,1899,1923]" italics="true" pageId="2" pageNumber="593">Elytra.</emphasis>
1.11.2 times as long and wide and 2.12.5 times as long as pronotum. Each elytron with 7 weakly defined punctate striae (
<figureCitation id="31D22A65FF8F0832FE1A2AB1CB1001CC" box="[427,520,1937,1962]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="2" pageNumber="593">Fig. 4D</figureCitation>
), punctures as large as those on pronotum; all intervals irregularly punctate, 2nd widest, 1st (sutural), 3rd and sometimes 5th weakly raised throughout, 4th broadly raised near elytral apex.
</paragraph>
<paragraph id="A95636E0FF8E0833FF762DB8CA8406B2" blockId="3.[151,1436,151,320]" pageId="3" pageNumber="594">
<emphasis id="9B9DEAF2FF8E0833FF762DB8C82A06D6" box="[199,306,152,176]" italics="true" pageId="3" pageNumber="594">Pygidium</emphasis>
with marginal bead complete but very fine with long, erect setae along margin. Surface fine and densely transversely rugose, feebly shiny, usually glabrous (
<figureCitation id="31D22A65FF8E0833FC892D9CCA8806B2" box="[824,912,188,213]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="3" pageNumber="594">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="A95636E0FF8E0833FF762DC1CB3E0759" blockId="3.[151,1436,151,320]" pageId="3" pageNumber="594">
<emphasis id="9B9DEAF2FF8E0833FF762DC1C808069E" box="[199,272,225,248]" italics="true" pageId="3" pageNumber="594">Venter.</emphasis>
Abdominal ventrites 25 with long, sparse setae forming transverse rows in the middle, directed posteriorly and surpassing the posterior margins of their ventrites. Ventrite 6 with apical fine line curved forward and enclosing a smooth apical area.
</paragraph>
<caption id="FD966668FF8E0833FF262ABCCAE20196" httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="3" pageNumber="594" targetBox="[162,1431,369,1909]" targetPageId="3">
<paragraph id="A95636E0FF8E0833FF262ABCCAE20196" blockId="3.[151,1436,1948,2032]" pageId="3" pageNumber="594">
<emphasis id="9B9DEAF2FF8E0833FF262ABCC84201D4" bold="true" box="[151,346,1948,1971]" pageId="3" pageNumber="594">FIGURES 1AM.</emphasis>
<taxonomicName id="6EE94D63FF8E0833FED32ABCC8A601D4" box="[354,446,1948,1970]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="9B9DEAF2FF8E0833FED32ABCC8A601D4" box="[354,446,1948,1970]" italics="true" pageId="3" pageNumber="594">Anomala</emphasis>
</taxonomicName>
spp. AL,
<taxonomicName id="6EE94D63FF8E0833FD842ABDCBAB01D4" box="[565,691,1948,1970]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">
<emphasis id="9B9DEAF2FF8E0833FD842ABDCBAB01D4" box="[565,691,1948,1970]" italics="true" pageId="3" pageNumber="594">A. katherine</emphasis>
</taxonomicName>
<taxonomicNameLabel id="80AE5789FF8E0833FD0A2ABDCA2201D4" box="[699,826,1948,1970]" pageId="3" pageNumber="594" rank="species">new species</taxonomicNameLabel>
: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M,
<taxonomicName id="6EE94D63FF8E0833FDD22AFBCA520196" authority="Gyllenhal" authorityName="Gyllenhal" box="[611,842,2010,2032]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="9B9DEAF2FF8E0833FDD22AFBCBD40196" box="[611,716,2011,2032]" italics="true" pageId="3" pageNumber="594">A. antiqua</emphasis>
(Gyllenhal)
</taxonomicName>
: maxilla, ventral.
</paragraph>
</caption>
<caption id="FD966668FF890834FF262A1CCA6E01E9" httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="4" pageNumber="595" targetBox="[178,1392,219,1788]" targetPageId="4">
<paragraph id="A95636E0FF890834FF262A1CCA6E01E9" blockId="4.[151,1436,1852,1935]" pageId="4" pageNumber="595">
<emphasis id="9B9DEAF2FF890834FF262A1CC8550137" bold="true" box="[151,333,1852,1875]" pageId="4" pageNumber="595">FIGURES 2AJ.</emphasis>
<taxonomicName id="6EE94D63FF890834FEE52A1CCB0D0134" box="[340,533,1852,1874]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">
<emphasis id="9B9DEAF2FF890834FEE52A1CCB0D0134" box="[340,533,1852,1874]" italics="true" pageId="4" pageNumber="595">Anomala katherine</emphasis>
</taxonomicName>
<taxonomicNameLabel id="80AE5789FF890834FDAD2A1DCB810134" box="[540,665,1852,1874]" pageId="4" pageNumber="595" rank="species">new species</taxonomicNameLabel>
, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal.
</paragraph>
</caption>
<caption id="FD966668FF880835FF262A17C9D601CC" httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="5" pageNumber="596" targetBox="[181,1436,219,1796]" targetPageId="5">
<paragraph id="A95636E0FF880835FF262A17C9D601CC" blockId="5.[151,1436,1847,1962]" pageId="5" pageNumber="596">
<emphasis id="9B9DEAF2FF880835FF262A17C84C012B" bold="true" box="[151,340,1847,1870]" pageId="5" pageNumber="596">FIGURES 3AL.</emphasis>
<taxonomicName id="6EE94D63FF880835FEED2A18C8A00128" box="[348,440,1848,1870]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="9B9DEAF2FF880835FEED2A18C8A00128" box="[348,440,1848,1870]" italics="true" pageId="5" pageNumber="596">Anomala</emphasis>
</taxonomicName>
spp. AD,
<taxonomicName id="6EE94D63FF880835FD852A19CA390128" authority="Gyllenhal" authorityName="Gyllenhal" box="[564,801,1848,1870]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="9B9DEAF2FF880835FD852A19CBB80128" box="[564,672,1849,1870]" italics="true" pageId="5" pageNumber="596">A. antiqua</emphasis>
(Gyllenhal)
</taxonomicName>
: A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. EH,
<taxonomicName id="6EE94D63FF880835FE692A77CBC3010A" authority="Fairmaire" authorityName="Fairmaire" box="[472,731,1878,1901]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="aeneotincta">
<emphasis id="9B9DEAF2FF880835FE692A77CB74010B" box="[472,620,1879,1901]" italics="true" pageId="5" pageNumber="596">A. aeneotincta</emphasis>
Fairmaire
</taxonomicName>
: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. IL,
<taxonomicName id="6EE94D63FF880835FEC72A56C8EA01ED" box="[374,498,1909,1931]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">
<emphasis id="9B9DEAF2FF880835FEC72A56C8EA01ED" box="[374,498,1909,1931]" italics="true" pageId="5" pageNumber="596">A. katherine</emphasis>
</taxonomicName>
<taxonomicNameLabel id="80AE5789FF880835FE4A2A56CB6201ED" box="[507,634,1909,1931]" pageId="5" pageNumber="596" rank="species">new species</taxonomicNameLabel>
: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head.
</paragraph>
</caption>
<caption id="FD966668FF8B0836FF262BFDCA560136" httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="6" pageNumber="597" targetBox="[189,1406,202,1733]" targetPageId="6">
<paragraph id="A95636E0FF8B0836FF262BFDCA560136" blockId="6.[151,1436,1757,1872]" pageId="6" pageNumber="597">
<emphasis id="9B9DEAF2FF8B0836FF262BFDC84C0095" bold="true" box="[151,340,1757,1780]" pageId="6" pageNumber="597">FIGURES 4AK.</emphasis>
<taxonomicName id="6EE94D63FF8B0836FEEA2BFEC8AF0092" box="[347,439,1758,1780]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="genus">
<emphasis id="9B9DEAF2FF8B0836FEEA2BFEC8AF0092" box="[347,439,1758,1780]" italics="true" pageId="6" pageNumber="597">Anomala</emphasis>
</taxonomicName>
spp. AC, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A,
<taxonomicName id="6EE94D63FF8B0836FB232A3CCC620157" authority="(Gyllenhal)" authorityName="Gyllenhal" baseAuthorityName="Gyllenhal" box="[1170,1402,1819,1841]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="9B9DEAF2FF8B0836FB232A3CCDE40157" box="[1170,1276,1820,1841]" italics="true" pageId="6" pageNumber="597">A. antiqua</emphasis>
(Gyllenhal)
</taxonomicName>
; B, F, G,
<taxonomicName id="6EE94D63FF8B0836FF7A2A1BC8D20136" authority="Fairmaire" authorityName="Fairmaire" box="[203,458,1850,1872]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="aeneotincta">
<emphasis id="9B9DEAF2FF8B0836FF7A2A1BC8450136" box="[203,349,1851,1872]" italics="true" pageId="6" pageNumber="597">A. aeneotincta</emphasis>
Fairmaire
</taxonomicName>
; CE, HK,
<taxonomicName id="6EE94D63FF8B0836FDFA2A1BCBDF0136" box="[587,711,1850,1872]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">
<emphasis id="9B9DEAF2FF8B0836FDFA2A1BCBDF0136" box="[587,711,1850,1872]" italics="true" pageId="6" pageNumber="597">A. katherine</emphasis>
</taxonomicName>
<taxonomicNameLabel id="80AE5789FF8B0836FD7C2A1BCA510136" box="[717,841,1850,1872]" pageId="6" pageNumber="597" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="A95636E0FF8B0836FF762A58CAEF01BE" blockId="6.[151,1436,1911,2008]" pageId="6" pageNumber="597">
<emphasis id="9B9DEAF2FF8B0836FF762A58C81B01E9" box="[199,259,1912,1935]" italics="true" pageId="6" pageNumber="597">Legs.</emphasis>
Protibia with blunt apical tooth and single sharp tooth on outer margin (
<figureCitation id="31D22A65FF8B0836FB9E2A57CD9C01F6" box="[1071,1156,1911,1936]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="6" pageNumber="597">Fig. 2F</figureCitation>
); larger claws of protarsi and mesotarsi apically, unequally bifurcate, those of protarsi enlarged (
<figureCitation id="31D22A65FF8B0836FC6A2ABBCD2201D2" box="[987,1082,1947,1972]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="6" pageNumber="597">Fig. 4H</figureCitation>
); mesotibia sinuate before the terminal tooth (
<figureCitation id="31D22A65FF8B0836FEF52A9FC88601BE" box="[324,414,1983,2008]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="6" pageNumber="597">Fig. 2G</figureCitation>
); apex of metatibia with short dense spines (
<figureCitation id="31D22A65FF8B0836FC212A9FCAF201BE" box="[912,1002,1983,2008]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="6" pageNumber="597">Fig. 2H</figureCitation>
).
</paragraph>
<paragraph id="A95636E0FF8A0837FF762DB8C8EE06C9" blockId="7.[151,1437,151,645]" box="[199,502,151,176]" pageId="7" pageNumber="598">
<emphasis id="9B9DEAF2FF8A0837FF762DB8C82C06D6" box="[199,308,152,176]" italics="true" pageId="7" pageNumber="598">Aedeagus</emphasis>
as in
<figureCitation id="31D22A65FF8A0837FEC72DB7C8EA06C9" box="[374,498,151,176]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="7" pageNumber="598">Figs. 3J, K</figureCitation>
.
</paragraph>
<paragraph id="A95636E0FF8A0837FF762D9CCD400691" blockId="7.[151,1437,151,645]" pageId="7" pageNumber="598">
<emphasis id="9B9DEAF2FF8A0837FF762D9CC83D06B3" bold="true" box="[199,293,188,213]" pageId="7" pageNumber="598">Female.</emphasis>
Externally similar to male but the split protarsal claws not enlarged (
<figureCitation id="31D22A65FF8A0837FBD02D9CCDAB06B2" box="[1121,1203,188,213]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="7" pageNumber="598">
Fig.
<date id="DD571020FF8A0837FB282D9CCDAB06B2" box="[1177,1203,188,212]" pageId="7" pageNumber="598">4I</date>
</figureCitation>
), antenna with club shorter than shaft and ventrite 6 with apex evenly curved, without smooth apical area.
</paragraph>
<paragraph id="A95636E0FF8A0837FF762C24CB0C04E2" blockId="7.[151,1437,151,645]" pageId="7" pageNumber="598">
<emphasis id="9B9DEAF2FF8A0837FF762C24C897077B" bold="true" box="[199,399,260,285]" pageId="7" pageNumber="598">
<typeStatus id="76528842FF8A0837FF762C24C814077B" box="[199,268,260,285]" pageId="7" pageNumber="598">Types</typeStatus>
.
<typeStatus id="76528842FF8A0837FE932C24C897077B" box="[290,399,260,285]" pageId="7" pageNumber="598" type="holotype">Holotype</typeStatus>
</emphasis>
male: Northern Territory: “
<geoCoordinate id="CCDD5027FF8A0837FD572C24CA58077A" box="[742,832,260,284]" direction="south" orientation="latitude" pageId="7" pageNumber="598" precision="925" value="-14.316667">14°19'S</geoCoordinate>
,
<geoCoordinate id="CCDD5027FF8A0837FCE52C24CAA6077B" box="[852,958,260,285]" direction="east" orientation="longitude" pageId="7" pageNumber="598" precision="925" value="132.41667">132°25'E</geoCoordinate>
,
<taxonomicName id="6EE94D63FF8A0837FC632C24CD58077B" box="[978,1088,260,285]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">Katherine</taxonomicName>
Gorge, N.T.,
<quantity id="6E119B05FF8A0837FB432C24CC59077B" box="[1266,1345,260,285]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="598" unit="km" value="24.0">24 km</quantity>
NE of
<taxonomicName id="6EE94D63FF8A0837FF262C07C8100726" box="[151,264,295,320]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">Katherine</taxonomicName>
,
<date id="DD571020FF8A0837FEA82C07C8770726" box="[281,367,295,320]" pageId="7" pageNumber="598" value="1972-10-16">16.x.72</date>
, M.S. Upton” (ANIC).
<emphasis id="9B9DEAF2FF8A0837FD3C2C07CA160726" bold="true" box="[653,782,295,320]" pageId="7" pageNumber="598">
<typeStatus id="76528842FF8A0837FD3C2C07CA110726" box="[653,777,295,320]" pageId="7" pageNumber="598" type="paratype">Paratypes</typeStatus>
:
</emphasis>
same data as the
<typeStatus id="76528842FF8A0837FC5E2C07CD490726" box="[1007,1105,295,320]" pageId="7" pageNumber="598" type="holotype">holotype</typeStatus>
(
<specimenCount id="BFEFFD69FF8A0837FBD62C07CD940726" box="[1127,1164,295,320]" pageId="7" pageNumber="598" type="generic">5♂</specimenCount>
, 3♀; ANIC); “
<geoCoordinate id="CCDD5027FF8A0837FAF22C07CC840726" box="[1347,1436,295,320]" direction="south" orientation="latitude" pageId="7" pageNumber="598" precision="925" value="-12.1">12°06'S</geoCoordinate>
<geoCoordinate id="CCDD5027FF8A0837FF262C6CC8180702" box="[151,256,332,356]" direction="east" orientation="longitude" pageId="7" pageNumber="598" precision="925" value="133.06667">133°04'E</geoCoordinate>
, Cooper Creek, N.T.,
<quantity id="6E119B05FF8A0837FE4A2C6CCB5D0703" box="[507,581,332,357]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="598" unit="km" value="19.0">19 km</quantity>
, E. by S. of Mt. Borradaile,
<date id="DD571020FF8A0837FC3B2C6CCAC20703" box="[906,986,332,357]" pageId="7" pageNumber="598" value="1972-11-02">2.xi.72</date>
, at light, E. Britton” (
<specimenCount id="BFEFFD69FF8A0837FB622C6CCC1F0702" box="[1235,1287,332,356]" pageId="7" pageNumber="598" type="generic">19♂</specimenCount>
, 9♀; ANIC); same data but
<date id="DD571020FF8A0837FEF52C4FC88C07EE" box="[324,404,367,392]" pageId="7" pageNumber="598" value="1972-11-09">9.xi.72</date>
, M.S. Upton” (
<specimenCount id="BFEFFD69FF8A0837FDFD2C50CB6907EE" box="[588,625,368,392]" pageId="7" pageNumber="598" type="generic">7♂</specimenCount>
, 1♀ANIC); same data but T.A. Weir &amp; T. Angeles (
<specimenCount id="BFEFFD69FF8A0837FB532C4FCC1E07EE" box="[1250,1286,367,392]" pageId="7" pageNumber="598" type="generic">5♂</specimenCount>
, 5♀; NTM); “
<geoCoordinate id="CCDD5027FF8A0837FF122CB4C9E307CA" box="[163,251,404,428]" direction="south" orientation="latitude" pageId="7" pageNumber="598" precision="925" value="-12.283334">12°17'S</geoCoordinate>
<geoCoordinate id="CCDD5027FF8A0837FEB22CB4C87507CA" box="[259,365,404,428]" direction="east" orientation="longitude" pageId="7" pageNumber="598" precision="925" value="133.33333">133°20'E</geoCoordinate>
, Cooper Creek, NT.,
<quantity id="6E119B05FF8A0837FDEF2CB4CBBD07CB" box="[606,677,404,429]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="598" unit="km" value="11.0">11 km</quantity>
S. by W. of Nimbuwah Rock,
<date id="DD571020FF8A0837FBB22CB4CD4B07CB" box="[1027,1107,404,429]" pageId="7" pageNumber="598" value="1972-11-01">1.xi.72</date>
, at light, E.B. Britton” (
<specimenCount id="BFEFFD69FF8A0837FAD52CB4CC8F07CA" box="[1380,1431,404,428]" pageId="7" pageNumber="598" type="generic">18♂</specimenCount>
, 8♀; ANIC); same data but T.A. Weir &amp; T. Angeles (
<specimenCount id="BFEFFD69FF8A0837FD472C98CA0407B6" box="[758,796,440,464]" pageId="7" pageNumber="598" type="generic">4♂</specimenCount>
, 11♀; NTM); “
<geoCoordinate id="CCDD5027FF8A0837FC7E2C97CD3F07B6" box="[975,1063,439,464]" direction="south" orientation="latitude" pageId="7" pageNumber="598" precision="925" value="-13.25">13°15'S</geoCoordinate>
<geoCoordinate id="CCDD5027FF8A0837FB802C97CD8207B6" box="[1073,1178,439,464]" direction="east" orientation="longitude" pageId="7" pageNumber="598" precision="925" value="131.1">131°06'E</geoCoordinate>
, Adelaide River, N.T.,
<date id="DD571020FF8A0837FF262CFCC9F50792" box="[151,237,476,500]" pageId="7" pageNumber="598" value="1972-10-17">17.x.72</date>
, M.S. Upton” (
<specimenCount id="BFEFFD69FF8A0837FE1B2CFCC8D70792" box="[426,463,476,500]" pageId="7" pageNumber="598" type="generic">9♂</specimenCount>
, 3♀; ANIC); “Tindal, N.T.,
<geoCoordinate id="CCDD5027FF8A0837FC992CFCCA980792" box="[808,896,476,500]" direction="south" orientation="latitude" pageId="7" pageNumber="598" precision="925" value="-14.516666">14°31'S</geoCoordinate>
<geoCoordinate id="CCDD5027FF8A0837FC3C2CFCCAEE0792" box="[909,1014,476,500]" direction="east" orientation="longitude" pageId="7" pageNumber="598" precision="925" value="132.36667">132°22'E</geoCoordinate>
,
<date id="DD571020FF8A0837FBB92CFCCDA30793" box="[1032,1211,476,501]" pageId="7" pageNumber="598" value="1967-12-01" valueMax="1967-12-20" valueMin="1967-12-01">120 Dec.1967</date>
, light trap, W.J.M. Vestjens” (1♀; ANIC); “
<geoCoordinate id="CCDD5027FF8A0837FE0B2F20CB0A047E" box="[442,530,512,536]" direction="south" orientation="latitude" pageId="7" pageNumber="598" precision="925" value="-12.383333">12°23'S</geoCoordinate>
<geoCoordinate id="CCDD5027FF8A0837FDAF2CDFCB90047E" box="[542,648,511,536]" direction="east" orientation="longitude" pageId="7" pageNumber="598" precision="925" value="132.93333">132°56'E</geoCoordinate>
,
<quantity id="6E119B05FF8A0837FD292F20CBCE047E" box="[664,726,511,536]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="598" unit="km" value="7.0">7 km</quantity>
NW by N of Cahills Crossing, N.T. (East Alligator River),
<date id="DD571020FF8A0837FF262F04C9FE045B" box="[151,230,548,573]" pageId="7" pageNumber="598" value="1972-11-04">4.xi.72</date>
, E. Britton” (1♀ ANIC); “
<geoCoordinate id="CCDD5027FF8A0837FD952F04CB64045B" box="[548,636,548,573]" direction="south" orientation="latitude" pageId="7" pageNumber="598" precision="925" value="-12.433333">12°26'S</geoCoordinate>
<geoCoordinate id="CCDD5027FF8A0837FD362F04CBE9045B" box="[647,753,548,573]" direction="east" orientation="longitude" pageId="7" pageNumber="598" precision="925" value="132.96666">132°58'E</geoCoordinate>
,
<quantity id="6E119B05FF8A0837FCB02F04CA26045B" box="[769,830,548,573]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="598" unit="km" value="1.0">1 km</quantity>
S of Cahills Crossing (East Alligator River), N.T.,
<date id="DD571020FF8A0837FF262F67C81B0406" box="[151,259,583,608]" pageId="7" pageNumber="598" value="1972-11-03">3.xi.1972</date>
, at light, E. Britton” (1♀; ANIC); “
<geoCoordinate id="CCDD5027FF8A0837FD142F67CBE60406" box="[677,766,583,608]" direction="south" orientation="latitude" pageId="7" pageNumber="598" precision="925" value="-12.1">12°06'S</geoCoordinate>
<geoCoordinate id="CCDD5027FF8A0837FCB92F67CA680406" box="[776,880,583,608]" direction="south" orientation="latitude" pageId="7" pageNumber="598" precision="925" value="-132.76666">132°46'S</geoCoordinate>
, Nourlangie Creek,
<quantity id="6E119B05FF8A0837FBEB2F67CD8E0406" box="[1114,1174,583,608]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="598" unit="km" value="6.0">6 km</quantity>
E of Mt. Cahill, N.T.,
<date id="DD571020FF8A0837FF262F4CC81104E2" box="[151,265,620,644]" pageId="7" pageNumber="598" value="1972-10-12">12.x.1972</date>
, T. Weir” (
<specimenCount id="BFEFFD69FF8A0837FE342F4CC8B104E2" box="[389,425,620,644]" pageId="7" pageNumber="598" type="generic">1♂</specimenCount>
, NTDA).
</paragraph>
<caption id="FD966668FF8A0837FF262BA8CC6900F8" box="[151,1393,1672,1694]" httpUri="https://zenodo.org/record/251256/files/figure.png" pageId="7" pageNumber="598" targetBox="[180,1403,697,1646]" targetPageId="7">
<paragraph id="A95636E0FF8A0837FF262BA8CC6900F8" blockId="7.[151,1393,1672,1694]" box="[151,1393,1672,1694]" pageId="7" pageNumber="598">
<emphasis id="9B9DEAF2FF8A0837FF262BA8C80A00FB" bold="true" box="[151,274,1672,1694]" pageId="7" pageNumber="598">FIGURE 5.</emphasis>
Distribution map:
<taxonomicName id="6EE94D63FF8A0837FE622BA8CBE400F8" authority="(Gyllenhal)" authorityName="Gyllenhal" baseAuthorityName="Gyllenhal" box="[467,764,1672,1694]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="9B9DEAF2FF8A0837FE622BA8CB9900F8" box="[467,641,1672,1694]" italics="true" pageId="7" pageNumber="598">Anomala antiqua</emphasis>
(Gyllenhal)
</taxonomicName>
(blue circles),
<taxonomicName id="6EE94D63FF8A0837FC252BA8CD4E00F8" box="[916,1110,1672,1694]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">
<emphasis id="9B9DEAF2FF8A0837FC252BA8CD4E00F8" box="[916,1110,1672,1694]" italics="true" pageId="7" pageNumber="598">Anomala katherine</emphasis>
</taxonomicName>
<taxonomicNameLabel id="80AE5789FF8A0837FBED2BA9CDCD00F8" box="[1116,1237,1672,1694]" pageId="7" pageNumber="598" rank="species">new species</taxonomicNameLabel>
(red triangles).
</paragraph>
</caption>
<paragraph id="A95636E0FF8A0837FF262BC9C8A30140" blockId="7.[151,489,1769,1830]" pageId="7" pageNumber="598">
<taxonomicName id="6EE94D63FF8A0837FF262BC9C8F10164" authority="Gyllenhal" authorityName="Gyllenhal" box="[151,489,1769,1794]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="9B9DEAF2FF8A0837FF262BC9C8790164" bold="true" box="[151,353,1769,1794]" pageId="7" pageNumber="598">Anomala antiqua</emphasis>
(Gyllenhal)
</taxonomicName>
(
<figureCitation id="31D22A65FF8A0837FF2E2A2DC8100140" box="[159,264,1805,1830]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="7" pageNumber="598">Figs. 1M</figureCitation>
,
<figureCitation id="31D22A65FF8A0837FEA22A2EC8710140" box="[275,361,1806,1830]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="7" pageNumber="598">3A3D</figureCitation>
,
<figureCitation id="31D22A65FF8A0837FEC52A2EC8800140" box="[372,408,1806,1830]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="7" pageNumber="598">4A</figureCitation>
,
<figureCitation id="31D22A65FF8A0837FE152A2DC8AA0140" box="[420,434,1805,1830]" captionStart="FIGURE 5" captionStartId="7.[151,250,1672,1694]" captionTargetBox="[180,1403,697,1646]" captionTargetId="figure@7.[178,1409,697,1651]" captionTargetPageId="7" captionText="FIGURE 5. Distribution map: Anomala antiqua (Gyllenhal) (blue circles), Anomala katherine new species (red triangles)." httpUri="https://zenodo.org/record/251256/files/figure.png" pageId="7" pageNumber="598">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E1F3656BFF8A0837FF262A75CB3B01CF" pageId="7" pageNumber="598" type="reference_group">
<paragraph id="A95636E0FF8A0837FF262A75CB5B010D" blockId="7.[151,982,1877,1961]" box="[151,579,1877,1899]" pageId="7" pageNumber="598">
<treatmentCitationGroup id="89F911CEFF8A0837FF262A75CB5B010D" box="[151,579,1877,1899]" pageId="7" pageNumber="598">
<taxonomicName id="6EE94D63FF8A0837FF262A75CB27010D" authority="Gyllenhal, 1817: 196" authorityName="Gyllenhal" authorityPageNumber="196" authorityYear="1817" box="[151,575,1877,1899]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="9B9DEAF2FF8A0837FF262A75C847010D" box="[151,351,1877,1899]" italics="true" pageId="7" pageNumber="598">Melolontha antiqua</emphasis>
<treatmentCitation id="284810F1FF8A0837FED42A75CB27010D" author="Gyllenhal" box="[357,575,1877,1899]" page="196" pageId="7" pageNumber="598" year="1817">
<bibRefCitation id="CD784B11FF8A0837FED42A75CB14010D" author="Gyllenhal" box="[357,524,1877,1899]" pageId="7" pageNumber="598" refString="Gyllenhal, L. (1817) [new taxa]. In: C. J. Schonherr; Synonymia Insectorum, oder Versuch einer Synonymie aller bisher bekannten Insecten; nach Fabricii Systema Elautheratorum etc. geordnet. Erster Band. Eleutherata oder Kafer. Dritter Theil. Hispa --- Molorchus. Em. Brucelius, Uppsala, Sweden, 506 pp. + Appendix: Descriptiones novarum specierum, 266 pp." type="book" year="1817">Gyllenhal, 1817</bibRefCitation>
: 196
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A95636E0FF8A0837FF262A54CB3B01CF" blockId="7.[151,982,1877,1961]" pageId="7" pageNumber="598">
<treatmentCitationGroup id="89F911CEFF8A0837FF262A54CB3B01CF" pageId="7" pageNumber="598">
<taxonomicName id="6EE94D63FF8A0837FF262A54CB1D01EC" authority="Carne, 1958: 170" authorityName="Carne" authorityPageNumber="170" authorityYear="1958" box="[151,517,1908,1930]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="9B9DEAF2FF8A0837FF262A54C85E01EC" box="[151,326,1908,1930]" italics="true" pageId="7" pageNumber="598">Anomala antiqua</emphasis>
:
<treatmentCitation id="284810F1FF8A0837FEE22A54CB1D01EC" author="Carne" box="[339,517,1908,1930]" page="170" pageId="7" pageNumber="598" year="1958">
<bibRefCitation id="CD784B11FF8A0837FEE22A54C8CB01EC" author="Carne" box="[339,467,1908,1930]" pageId="7" pageNumber="598" refString="Carne, P. B. (1958) A review of the Australian Rutelinae (Coleoptera: Scarabaeidae). Australian Journal of Zoology, 6, 162 - 240. http: // dx. doi. org / 10.1071 / zo 9580162" type="journal article" year="1958">Carne, 1958</bibRefCitation>
: 170
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="284810F1FF8A0837FDA12A54CA0E01EC" author="Cassis" box="[528,790,1908,1930]" page="363" pageId="7" pageNumber="598" year="1992">
<bibRefCitation id="CD784B11FF8A0837FDA12A54CBFB01EC" author="Cassis" box="[528,739,1908,1930]" pageId="7" pageNumber="598" refString="Cassis, G. &amp; Weir, T. A. (1992) Rutelinae. In: Houston, W. W. K. &amp; Weir, T. A. (Eds.), Zoological Catalogue of Australia. Vol. 9. Coleoptera: Scarabaeidae. Australian Government Publishing Service, Canberra, pp. 359 - 382. [Australia]" type="book" year="1992">Cassis &amp; Weir, 1992</bibRefCitation>
: 363
</treatmentCitation>
;
<treatmentCitation id="284810F1FF8A0837FC912A55CACA01EC" author="Zorn" box="[800,978,1908,1930]" page="257" pageId="7" pageNumber="598" year="2006">
<bibRefCitation id="CD784B11FF8A0837FC912A55CA8701EC" author="Zorn" box="[800,927,1908,1930]" pageId="7" pageNumber="598" refString="Zorn, C. (2006 a) Rutelinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Apollo Books, Stenstrup, pp. 248 - 277. [Denmark]" type="book" year="2006" yearSuffix="a">Zorn, 2006a</bibRefCitation>
: 257
</treatmentCitation>
. For full synonmy see
<treatmentCitation id="284810F1FF8A0837FEC52AB4CB0701CF" author="Zorn" box="[372,543,1939,1961]" page="257" pageId="7" pageNumber="598" year="2006">
<bibRefCitation id="CD784B11FF8A0837FEC52AB4C8F401CF" author="Zorn" box="[372,492,1939,1961]" pageId="7" pageNumber="598" refString="Zorn, C. (2006 a) Rutelinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Apollo Books, Stenstrup, pp. 248 - 277. [Denmark]" type="book" year="2006" yearSuffix="a">Zorn 2006a</bibRefCitation>
: 257
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1F3656BFF8A0838FF262AF5CDEA07EE" lastPageId="8" lastPageNumber="599" pageId="7" pageNumber="598" type="diagnosis">
<paragraph id="A95636E0FF8A0838FF262AF5CB320703" blockId="7.[151,1436,2005,2030]" lastBlockId="8.[151,1437,151,1256]" lastPageId="8" lastPageNumber="599" pageId="7" pageNumber="598">
<emphasis id="9B9DEAF2FF8A0837FF262AF5C8090188" bold="true" box="[151,273,2005,2030]" pageId="7" pageNumber="598">Diagnosis.</emphasis>
<taxonomicName id="6EE94D63FF8A0837FEA62AF5C8C0018B" box="[279,472,2005,2029]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="598" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="9B9DEAF2FF8A0837FEA62AF5C8C0018B" box="[279,472,2005,2029]" italics="true" pageId="7" pageNumber="598">Anomala antiqua</emphasis>
</taxonomicName>
is distinguished from the other Australian species by the combination of the following characters: clypeus strongly reflexed along anterior margin (
<figureCitation id="31D22A65FF850838FCF72DB7CABA06D6" box="[838,930,151,176]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="8" pageNumber="599">Fig. 3D</figureCitation>
); maxilla with 7 heavily sclerotised teeth on apex of galea (
<figureCitation id="31D22A65FF850838FEF32D9CC8BA06B2" box="[322,418,188,213]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="8" pageNumber="599">Fig. 1M</figureCitation>
); protibia with 2 teeth along external margin (
<figureCitation id="31D22A65FF850838FC082D9CCD0D06B2" box="[953,1045,188,213]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="8" pageNumber="599">Fig. 3A</figureCitation>
); pronotum with marginal bead at base complete medially (
<figureCitation id="31D22A65FF850838FE762DFFCB31069E" box="[455,553,223,248]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="8" pageNumber="599">Fig. 4A</figureCitation>
); pygidium uniformly brown or green; phallobase distinctly shorter than parameres with well sclerotised and apically pointed ventral plate (
<figureCitation id="31D22A65FF850838FC092C24CD17077A" box="[952,1039,260,285]" captionStart="FIGURES 3 A L" captionStartId="5.[151,264,1847,1870]" captionTargetBox="[181,1436,219,1796]" captionTargetId="figure@5.[166,1449,213,1806]" captionTargetPageId="5" captionText="FIGURES 3 A L. Anomala spp. A D, A. antiqua (Gyllenhal): A, dorsal view; B, aedeagus, inner; C, aedeagus, lateral; D, anterior portion of head. E H, A. aeneotincta Fairmaire: E, dorsal view; F, aedeagus, inner; G, aedeagus, lateral; H, anterior portion of head. I L, A. katherine new species: I, dorsal view; J, aedeagus, inner; K, aedeagus, lateral; L, anterior portion of head." httpUri="https://zenodo.org/record/251254/files/figure.png" pageId="8" pageNumber="599">Fig 3B</figureCitation>
), the plate extends to parameres; parameres fused at bases but broadly separated apically. This species is also on average larger and has distinctive green colouration of the dorsal side.
</paragraph>
<paragraph id="A95636E0FF850838FF762C4FCDEA07EE" blockId="8.[151,1437,151,1256]" box="[199,1266,367,392]" pageId="8" pageNumber="599">
<emphasis id="9B9DEAF2FF850838FF762C4FC83007EE" bold="true" box="[199,296,367,392]" pageId="8" pageNumber="599">
<typeStatus id="76528842FF850838FF762C4FC81407EE" box="[199,268,367,392]" pageId="8" pageNumber="599">Types</typeStatus>
. “
</emphasis>
2793 T N.T. [on mounting card]/
<taxonomicName id="6EE94D63FF850838FD2F2C4FCAE907EE" authority="Australasia, Blackb." authorityName="Australasia, Blackb." box="[670,1009,367,392]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="599" phylum="Arthropoda" rank="genus">Anomala Australasia, Blackb.</taxonomicName>
” (1, BMNH;
<typeStatus id="76528842FF850838FB3B2C50CDFE07EE" box="[1162,1254,368,392]" pageId="8" pageNumber="599" type="syntype">syntype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="E1F3656BFF850838FF762CB4CD110253" pageId="8" pageNumber="599" type="materials_examined">
<paragraph id="A95636E0FF850838FF762CB4CD110253" blockId="8.[151,1437,151,1256]" pageId="8" pageNumber="599">
<emphasis id="9B9DEAF2FF850838FF762CB4C8AB07CA" bold="true" box="[199,435,404,429]" pageId="8" pageNumber="599">Material examined.</emphasis>
Specimens from
<collectionCode id="CFF8AE25FF850838FDCE2CB4CBDB07CA" LSID="urn:lsid:biocol.org:col:32981" box="[639,707,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="8" pageNumber="599">ANIC</collectionCode>
(455), QM (37), AM (6),
<collectionCode id="CFF8AE25FF850838FC462CB4CD5A07CA" box="[1015,1090,404,428]" httpUri="http://grbio.org/cool/r09q-tgas" name="Agricultural Scientific Collections Unit" pageId="8" pageNumber="599">ASCU</collectionCode>
(2).
<emphasis id="9B9DEAF2FF850838FBCD2CB4CC1107CB" bold="true" box="[1148,1289,404,429]" pageId="8" pageNumber="599">Queensland</emphasis>
:
<quantity id="6E119B05FF850838FAAD2CB4CC7C07CB" box="[1308,1380,404,429]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="599" unit="km" value="11.0">11 km</quantity>
NW Mareeba, Southedge
<collectingCountry id="D1FE7670FF850838FE362C97C8FD07B6" box="[391,485,439,464]" name="Trinidad and Tobago" pageId="8" pageNumber="599">Tobacco</collectingCountry>
Research Station;
<quantity id="6E119B05FF850838FD0A2C98CA1C07B6" box="[699,772,439,464]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="599" unit="km" value="12.0">12 km</quantity>
SSE Heathlands;
<quantity id="6E119B05FF850838FC632C98CD0307B6" box="[978,1051,439,464]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="599" unit="km" value="12.0">12 km</quantity>
E by N Heathlands;
<quantity id="6E119B05FF850838FABA2C98CC4C07B6" box="[1291,1364,439,464]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="599" unit="km" value="12.0">12 km</quantity>
SE of Daintree;
<quantity id="6E119B05FF850838FEB72CFCC8550793" box="[262,333,476,501]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="599" unit="km" value="14.0">14 km</quantity>
ENE Heathlands;
<quantity id="6E119B05FF850838FDAE2CFCCB7E0793" box="[543,614,476,501]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="599" unit="km" value="15.0">15 km</quantity>
NE by E Heathlands;
<quantity id="6E119B05FF850838FCD32CFCCA830793" box="[866,923,476,501]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="599" unit="km" value="1.0">1 km</quantity>
S Heathlands;
<quantity id="6E119B05FF850838FBF42CFCCD940793" box="[1093,1164,476,501]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="599" unit="km" value="20.0">20 km</quantity>
N of Mt. Carbine;
<quantity id="6E119B05FF850838FAD22CFCCC840793" box="[1379,1436,476,501]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="599" unit="km" value="2.0">2 km</quantity>
NNW Jowalbinna;
<quantity id="6E119B05FF850838FEC72CDFC8AB047E" box="[374,435,511,536]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="599" unit="km" value="6.0">6 km</quantity>
NNE Coen;
<quantity id="6E119B05FF850838FDFE2CDFCB93047E" box="[591,651,511,536]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="599" unit="km" value="6.0">6 km</quantity>
SE of Chillagoe; Airville; Archer River; Atherton; Ayr; Bald Hills Station,
<quantity id="6E119B05FF850838FF422F04C821045B" box="[243,313,548,573]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="599" unit="km" value="30.0">30 km</quantity>
N of Cooktown;
<collectingCountry id="D1FE7670FF850838FE4A2F04CB49045B" box="[507,593,548,573]" name="Netherlands" pageId="8" pageNumber="599">Batavia</collectingCountry>
Downs; Bucasia; Burdekin Rural Education Centre
<quantity id="6E119B05FF850838FB262F04CDC6045B" box="[1175,1246,548,573]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="599" unit="km" value="30.0">30 km</quantity>
SW Ayr; Cairns; Cairns area, near Lake Placid; Cardwell; Conway Beach,
<quantity id="6E119B05FF850838FC9D2F68CA6D0406" box="[812,885,583,608]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="599" unit="km" value="27.0">27 km</quantity>
SE Proserpine; Cooktown, Walker Bay; Crystal Creek,
<quantity id="6E119B05FF850838FF5B2F4CC82904E3" box="[234,305,620,645]" metricMagnitude="4" metricUnit="m" metricValue="3.7014912" pageId="8" pageNumber="599" unit="mi" value="23.0">23 mi</quantity>
. SSE of Ingham; Cunnamulla; Davies Creek (10.6 road km from Kennedy Hwy),
<quantity id="6E119B05FF850838FB5A2F4CCC2D04E3" box="[1259,1333,620,645]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="599" unit="km" value="17.0">17 km</quantity>
EbyS of Mareeba; Deeral; Fletcher Creek rest area; Four Mile Beach, Port Douglas; Funnel Creek; Gap Creek; Gordonvale; Green Hills; Gulf Development Road between Mount Surprise and Newcastle Range; Gympie; Hann River; Heathlands; Helenslee Stn., Homestead,
<quantity id="6E119B05FF850838FDDB2FF8CBAC0496" box="[618,692,727,752]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="599" unit="km" value="48.0">48 km</quantity>
SW Charters Towers; Helenvale; Ipswich; Iron Range; Jourama Falls National Park; Kamerunga, Cairns; Kuranda; Lansdowne Station,
<quantity id="6E119B05FF850838FC602FDDCD140573" box="[977,1036,764,789]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="599" unit="km" value="7.0">7 km</quantity>
S of Woodstock; Laura; Mareeba; Mary Creek; Meringa; Mount Elliot National Park; Moreton; Mount Surprise; Mount Carbine; Mount Garnet; Mount Spec; Mount White; Palm Cove; Prince of
<collectingCountry id="D1FE7670FF850838FD582E64CA35053B" box="[745,813,836,861]" name="United Kingdom" pageId="8" pageNumber="599">Wales</collectingCountry>
Island, Gulf of Carpentaria; Rochford Scrub; Sarina; Shiptons Flat; Sliver Plains Homestead, Cape York; Station Creek,
<specimenCount id="BFEFFD69FF850838FC392E48CAD00519" box="[904,968,872,896]" pageId="8" pageNumber="599" type="generic">
<quantity id="6E119B05FF850838FC392E48CADB0519" box="[904,963,872,896]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="599" unit="m" value="11.0">11 m</quantity>
.
</specimenCount>
N Mount Molloy; The Boulders,
<quantity id="6E119B05FF850838FAF32E47CC6305E6" box="[1346,1403,871,896]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="599" unit="km" value="5.0">5 km</quantity>
W of Babinda; Thursday Island; Toomba; Townsville; W Claudie River, Iron Range; Weipa; Woodstock; Yeppoon.
<emphasis id="9B9DEAF2FF850838FF262E8FC86405AE" bold="true" box="[151,380,943,968]" pageId="8" pageNumber="599">Northern Territory</emphasis>
:
<quantity id="6E119B05FF850838FE3A2E90C8C905AE" box="[395,465,943,968]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="599" unit="km" value="14.0">14 km</quantity>
NW of Cape Crawford;
<quantity id="6E119B05FF850838FD552E90CA0505AE" box="[740,797,943,968]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="599" unit="km" value="3.0">3 km</quantity>
SSW of
<taxonomicName id="6EE94D63FF850838FC302E8FCAEB05AE" box="[897,1011,943,968]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="598" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">Katherine</taxonomicName>
; Adelaide River;
<taxonomicName id="6EE94D63FF850838FB072E8FCC6E05AE" authority="Beach" authorityName="Beach" box="[1206,1398,943,968]" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="599" phylum="Tracheophyta" rank="genus">Casuarina Beach</taxonomicName>
,
<quantity id="6E119B05FF850838FA312E90C9A3058B" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="599" unit="km" value="10.0">10 km</quantity>
NNE of Darwin; Daly River Mission; Holmes Jungle, Berrimah,
<quantity id="6E119B05FF850838FC2B2EF4CAF9058B" box="[922,993,980,1005]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="599" unit="km" value="10.0">10 km</quantity>
S Darwin;
<taxonomicName id="6EE94D63FF850838FBD02EF4CDD7058B" box="[1121,1231,980,1005]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="598" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">Katherine</taxonomicName>
Gorge,
<quantity id="6E119B05FF850838FA982EF4CC68058B" box="[1321,1392,980,1005]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="599" unit="km" value="24.0">24 km</quantity>
NE of
<taxonomicName id="6EE94D63FF850838FF042ED7C83F0276" box="[181,295,1015,1040]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="598" phylum="Arthropoda" rank="species" species="katherine" status="sp. nov.">Katherine</taxonomicName>
; McArthur River,
<quantity id="6E119B05FF850838FE442ED8CB240276" box="[501,572,1015,1040]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="599" unit="km" value="14.0">14 km</quantity>
S by W of Cape Crawford; Tindal.
<emphasis id="9B9DEAF2FF850838FC7C2ED7CDB00276" bold="true" box="[973,1192,1015,1040]" pageId="8" pageNumber="599">
Western
<collectingCountry id="D1FE7670FF850838FB892ED7CDB00276" box="[1080,1192,1015,1040]" name="Australia" pageId="8" pageNumber="599">Australia</collectingCountry>
</emphasis>
: Kimberley Research Station via Wyndham; Kununurra; Wyndham.
<emphasis id="9B9DEAF2FF850838FD10293CCA690253" bold="true" box="[673,881,1052,1077]" pageId="8" pageNumber="599">
New South
<collectingCountry id="D1FE7670FF850838FC9B293CCA690253" box="[810,881,1052,1077]" name="United Kingdom" pageId="8" pageNumber="599">Wales</collectingCountry>
</emphasis>
: Cabramatta.
</paragraph>
</subSubSection>
<subSubSection id="E1F3656BFF850838FF76291FCA4E028E" pageId="8" pageNumber="599" type="distribution">
<paragraph id="A95636E0FF850838FF76291FCB8202C6" blockId="8.[151,1437,151,1256]" pageId="8" pageNumber="599">
<emphasis id="9B9DEAF2FF850838FF76291FC878023E" bold="true" box="[199,352,1087,1112]" pageId="8" pageNumber="599">Distribution.</emphasis>
Widely distributed in Oriental and southeastern part of the Palaearctic Regions from
<collectingCountry id="D1FE7670FF850838FA8B291FCC65023E" box="[1338,1405,1087,1112]" name="China" pageId="8" pageNumber="599">China</collectingCountry>
to
<collectingCountry id="D1FE7670FF850838FF262944C9CC021B" box="[151,212,1124,1149]" name="India" pageId="8" pageNumber="599">India</collectingCountry>
. Introduced to
<collectingCountry id="D1FE7670FF850838FE212944C8E3021B" box="[400,507,1124,1149]" name="Australia" pageId="8" pageNumber="599">Australia</collectingCountry>
: Northern Territory, Western
<collectingCountry id="D1FE7670FF850838FCEF2944CADF021B" box="[862,967,1124,1149]" name="Australia" pageId="8" pageNumber="599">Australia</collectingCountry>
, Queensland, (
<figureCitation id="31D22A65FF850838FBCB2944CDDE021B" box="[1146,1222,1124,1149]" captionStart="FIGURE 5" captionStartId="7.[151,250,1672,1694]" captionTargetBox="[180,1403,697,1646]" captionTargetId="figure@7.[178,1409,697,1651]" captionTargetPageId="7" captionText="FIGURE 5. Distribution map: Anomala antiqua (Gyllenhal) (blue circles), Anomala katherine new species (red triangles)." httpUri="https://zenodo.org/record/251256/files/figure.png" pageId="8" pageNumber="599">Fig. 5</figureCitation>
). The Cabramatta specimen probably represents an interception.
</paragraph>
<paragraph id="A95636E0FF850838FF76298CCA4E028E" blockId="8.[151,1437,151,1256]" pageId="8" pageNumber="599">
<emphasis id="9B9DEAF2FF850838FF76298CC80902A3" bold="true" box="[199,273,1196,1221]" pageId="8" pageNumber="599">Notes.</emphasis>
The earliest Australian records that we have been able to find are: QLD: Cairns 1890, Cape York 1909; NT: Adelaide River 1892, Darwin 1916; WA: Wyndham 1929.
</paragraph>
</subSubSection>
</treatment>
</document>