treatments-xml/data/03/DE/36/03DE3646FF93FFC3FF43FA4FFA9BFC67.xml
2024-06-21 12:22:17 +02:00

294 lines
34 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="7C8C7D71467750A88824C7488D82D436" ID-DOI="10.11646/phytotaxa.515.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8061143" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1687282820830" checkinUser="plazi" docAuthor="Beek, Abraham Van De" docDate="2021" docId="03DE3646FF93FFC3FF43FA4FFA9BFC67" docLanguage="en" docName="Phytotaxa.515.1.1.pdf" docOrigin="Phytotaxa 515 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.515.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Rubus leptodytus A. Beek 2021, sp. nov." docType="treatment" docVersion="2" lastPageNumber="31" masterDocId="FFE74E3EFF8FFFDDFFCBFFE1FF96FF93" masterDocTitle="Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa" masterLastPageNumber="71" masterPageNumber="1" pageNumber="29" updateTime="1687379631607" updateUser="ExternalLinkService">
<mods:mods id="0D5EA7E1FA89DC89361FBFD647C2E249" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D095E5FB75B73C18FB94E5BF7F9E74A4">
<mods:title id="1831972EE05EE184AF9455B68391EDB8">Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa</mods:title>
</mods:titleInfo>
<mods:name id="90B7DB7736B181D5902A03A5049F6996" type="personal">
<mods:role id="A33E6FAC89BB0C1905C76673CD25E00D">
<mods:roleTerm id="55BB3F25BDD1040B76EC708DB6F32F19">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B2DD60EA8847AD40445BB9AD4D92159">Beek, Abraham Van De</mods:namePart>
</mods:name>
<mods:typeOfResource id="7DF2D545CDA549F29ABEECEF277E1A10">text</mods:typeOfResource>
<mods:relatedItem id="86DCFBFA8D4FF96B4531BF8CAB84E98F" type="host">
<mods:titleInfo id="43D9F6662F0BE7D66BA92FA38D270E28">
<mods:title id="F93AE45AD878DFEFBA63FB06D3F64F01">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="98F37584281844ADBF1208EAE7472F5C">
<mods:date id="33648C93DF1E6214E6FA52E8E28F3238">2021</mods:date>
<mods:detail id="B57195A6BF683E496CF6982ACEB44757" type="pubDate">
<mods:number id="40101FB985017B89F27D7B277C040416">2021-08-19</mods:number>
</mods:detail>
<mods:detail id="1591D3E4ACCC791CF32CA960E6757B9E" type="volume">
<mods:number id="913198B38C43E664FE7C53FCC440A353">515</mods:number>
</mods:detail>
<mods:detail id="DDD18AE818606890A7A28F421D17319A" type="issue">
<mods:number id="28E378599C16C8A967B6263D8E6CCE4C">1</mods:number>
</mods:detail>
<mods:extent id="DED7B4E289055CF242AC0F182FA53D72" unit="page">
<mods:start id="16A1747B7F3E05DADBC1F3F4DF1DE625">1</mods:start>
<mods:end id="B22850F0A33F23F2CA4D334B1F4DB8B0">71</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="E1B244E35B8861A66C3A522491A00041">
<mods:url id="A89B30F6997108B6278219A28A9A1DCD">http://dx.doi.org/10.11646/phytotaxa.515.1.1</mods:url>
</mods:location>
<mods:classification id="B9E8F351E316E0A2087380C031758309">journal article</mods:classification>
<mods:identifier id="839534F16E884085EE9E12361B5E857B" type="DOI">10.11646/phytotaxa.515.1.1</mods:identifier>
<mods:identifier id="879A535CC09C141F06AEB5DF58E0CB12" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="320B38B86C0110FE239AA41E7A3C22D6" type="Zenodo-Dep">8061143</mods:identifier>
</mods:mods>
<treatment id="03DE3646FF93FFC3FF43FA4FFA9BFC67" ID-DOI="http://doi.org/10.5281/zenodo.8067193" ID-Zenodo-Dep="8067193" LSID="urn:lsid:plazi:treatment:03DE3646FF93FFC3FF43FA4FFA9BFC67" httpUri="http://treatment.plazi.org/id/03DE3646FF93FFC3FF43FA4FFA9BFC67" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="C36DD4DBFF93FFC1FF43FA4FFDDBFA5B" box="[136,589,1454,1481]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BC88750FF93FFC1FF43FA4FFDDBFA5B" blockId="28.[136,1451,1454,1517]" box="[136,589,1454,1481]" pageId="28" pageNumber="29">
<emphasis id="B9035B42FF93FFC1FF43FA4FFF3DFA5B" bold="true" box="[136,171,1454,1480]" pageId="28" pageNumber="29">10.</emphasis>
<taxonomicName id="4C77FCD3FF93FFC1FF73FA4FFE73FA5B" ID-CoL="8RTRL" authority="A.Beek" authorityName="A. Beek" authorityYear="2021" box="[184,485,1454,1481]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="leptodytus" status="sp. nov.">
<emphasis id="B9035B42FF93FFC1FF73FA4FFE13FA5A" bold="true" box="[184,389,1454,1481]" italics="true" pageId="28" pageNumber="29">Rubus leptodytus</emphasis>
A.Beek
</taxonomicName>
<emphasis id="B9035B42FF93FFC1FE39FA4EFDDBFA5B" box="[498,589,1455,1480]" italics="true" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A230E639FF93FFC1FE39FA4EFDDBFA5B" box="[498,589,1455,1480]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF93FFC1FD91FA4FFC22FA7F" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BC88750FF93FFC1FD91FA4FFC22FA7F" blockId="28.[136,1451,1454,1517]" pageId="28" pageNumber="29">
<materialsCitation id="3B1F8D0DFF93FFC1FD91FA4FFC22FA7F" collectingDate="2019-11-20" collectionCode="L" collectorName="Beek" country="South Africa" location="Karnemelksvlei" municipality="Citrusdal" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
Type:—
<collectingCountry id="F360C7C0FF93FFC1FD7FFA4FFC1EFA5B" box="[692,904,1454,1480]" name="South Africa" pageId="28" pageNumber="29">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FF93FFC1FC53FA4EFBA9FA5B" box="[920,1087,1454,1481]" country="South Africa" name="Western Cape" pageId="28" pageNumber="29">Western Cape</collectingRegion>
:
<collectingMunicipality id="6BAC1D2AFF93FFC1FB84FA4FFB2FFA5B" box="[1103,1209,1454,1480]" pageId="28" pageNumber="29">Citrusdal</collectingMunicipality>
,
<location id="8EA8D18BFF93FFC1FB01FA4FFAE8FA5B" LSID="urn:lsid:plazi:treatment:03DE3646FF93FFC3FF43FA4FFA9BFC67:8EA8D18BFF93FFC1FB01FA4FFAE8FA5B" box="[1226,1406,1454,1480]" country="South Africa" county="Type" municipality="Citrusdal" name="Karnemelksvlei" pageId="28" pageNumber="29" stateProvince="Western Cape">Karnemelksvlei</location>
,
<date id="FFC9A190FF93FFC1FA44FA4FFED6FA7F" pageId="28" pageNumber="29" value="2019-11-20">
<collectingDate id="EF8D5878FF93FFC1FA44FA4FFED6FA7F" pageId="28" pageNumber="29" value="2019-11-20">20 November 2019</collectingDate>
</date>
,
<emphasis id="B9035B42FF93FFC1FE80FA33FE72FA7F" box="[331,484,1490,1516]" italics="true" pageId="28" pageNumber="29">
<collectorName id="2682E286FF93FFC1FE80FA33FE17FA7F" box="[331,385,1490,1516]" pageId="28" pageNumber="29">Beek</collectorName>
2019.93
</emphasis>
(
<typeStatus id="54CC39F2FF93FFC1FE39FA33FDC0FA7F" box="[498,598,1490,1516]" pageId="28" pageNumber="29">holotype</typeStatus>
<collectionCode id="ED661F95FF93FFC1FD96FA32FDE7FA7E" box="[605,625,1491,1517]" pageId="28" pageNumber="29">L</collectionCode>
; isotype NBG). (
<figureCitation id="134C9BD5FF93FFC1FCE4FA33FC3AFA7F" box="[815,940,1490,1516]" captionStart="FIGURE 8" captionStartId="29.[136,229,1750,1772]" captionTargetBox="[151,1436,190,1726]" captionTargetId="figure-18@29.[151,1436,190,1726]" captionTargetPageId="29" captionText="FIGURE 8. Rubus leptodytus. A. Holotype, primocane. B. Holotype, primocane. C. Holotype, inflorescence. D. Plant of the type, habit. E. Primocane. F. Primocane. G. Leaves. H. Flowering branch. I. Inflorescence. J. Pedicels. K. Flowers. L. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061165" httpUri="https://zenodo.org/record/8061165/files/figure.png" pageId="28" pageNumber="29">Figs 8AC</figureCitation>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF93FFC1FF43F9FBFAEEF957" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BC88750FF93FFC1FF43F9FBFAEEF957" blockId="28.[136,1452,1562,2093]" pageId="28" pageNumber="29">
<emphasis id="B9035B42FF93FFC1FF43F9FBFE93F9A7" bold="true" box="[136,261,1562,1588]" pageId="28" pageNumber="29">Diagnosis:</emphasis>
—A taxon with characteristics that are different from both
<taxonomicName id="4C77FCD3FF93FFC1FC63F9FAFB8EF9A7" authorityName="Smith" authorityYear="1819" box="[936,1048,1562,1588]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="rigidus">
<emphasis id="B9035B42FF93FFC1FC63F9FAFB8EF9A7" box="[936,1048,1562,1588]" italics="true" pageId="28" pageNumber="29">R. rigidus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C77FCD3FF93FFC1FB98F9FAFB5BF9A7" baseAuthorityName="Focke" baseAuthorityYear="1874" box="[1107,1229,1562,1588]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="ecklonii">
<emphasis id="B9035B42FF93FFC1FB98F9FAFB5BF9A7" box="[1107,1229,1562,1588]" italics="true" pageId="28" pageNumber="29">R. ecklonii</emphasis>
</taxonomicName>
occurs in the more northern regions of the
<collectingRegion id="49B349B2FF93FFC1FE59F9DEFDA7F9CB" box="[402,561,1598,1625]" country="South Africa" name="Western Cape" pageId="28" pageNumber="29">Western Cape</collectingRegion>
, from Citrusdal to Oudtshoorn. It has, in contrast to
<taxonomicName id="4C77FCD3FF93FFC1FB4EF9DEFB68F9CB" baseAuthorityName="Focke" baseAuthorityYear="1874" box="[1157,1278,1598,1624]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="ecklonii">
<emphasis id="B9035B42FF93FFC1FB4EF9DEFB68F9CB" box="[1157,1278,1598,1624]" italics="true" pageId="28" pageNumber="29">R. ecklonii</emphasis>
</taxonomicName>
, many stipitate glands on the primocane and some in the inflorescences like
<taxonomicName id="4C77FCD3FF93FFC1FCF1F982FC3FF9EF" authorityName="Smith" authorityYear="1819" box="[826,937,1634,1660]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="rigidus">
<emphasis id="B9035B42FF93FFC1FCF1F982FC3FF9EF" box="[826,937,1634,1660]" italics="true" pageId="28" pageNumber="29">R. rigidus</emphasis>
</taxonomicName>
, but the leaflets are much broader than in the latter, the serrature is shallower and the leaves have abaxially a very short tomentum (
<taxonomicName id="4C77FCD3FF93FFC1FBA5F967FB77F933" authorityName="A. Beek" authorityYear="2021" box="[1134,1249,1670,1696]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="leptodytus">
<emphasis id="B9035B42FF93FFC1FBA5F967FB77F933" box="[1134,1249,1670,1696]" italics="true" pageId="28" pageNumber="29">leptodytus</emphasis>
</taxonomicName>
means with thin clothes). Because it is very characteristic and has a rather wide distribution it is described here as a new species.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF93FFC3FF76F92EFC93FE27" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="description">
<paragraph id="8BC88750FF93FFC3FF76F92EFC93FE27" blockId="28.[136,1452,1562,2093]" lastBlockId="30.[136,1452,158,1013]" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<emphasis id="B9035B42FF93FFC1FF76F92EFEAEF97B" box="[189,312,1743,1768]" italics="true" pageId="28" pageNumber="29">Primocane</emphasis>
(
<figureCitation id="134C9BD5FF93FFC1FE8CF92FFE57F97B" box="[327,449,1742,1768]" captionStart="FIGURE 8" captionStartId="29.[136,229,1750,1772]" captionTargetBox="[151,1436,190,1726]" captionTargetId="figure-18@29.[151,1436,190,1726]" captionTargetPageId="29" captionText="FIGURE 8. Rubus leptodytus. A. Holotype, primocane. B. Holotype, primocane. C. Holotype, inflorescence. D. Plant of the type, habit. E. Primocane. F. Primocane. G. Leaves. H. Flowering branch. I. Inflorescence. J. Pedicels. K. Flowers. L. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061165" httpUri="https://zenodo.org/record/8061165/files/figure.png" pageId="28" pageNumber="29">Figs 8DF</figureCitation>
) suberect to high arching, when supported in shrubs up to more than
<quantity id="4C8F2AB5FF93FFC1FB08F92FFB78F97A" box="[1219,1262,1742,1769]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="m" value="3.0">3 m</quantity>
high, sometimes finally tip rooting, diam.
<quantity id="4C8F2AB5FF93FFC1FE56F913FD9FF89E" box="[413,521,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="28" pageNumber="29" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">510 mm</quantity>
, furrowed, densely very short stellate hairy, with numerous stipitate glands; prickles 417 per
<quantity id="4C8F2AB5FF93FFC1FF24F8F7FEBEF8A2" box="[239,296,1814,1841]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="28" pageNumber="29" unit="cm" value="5.0">5 cm</quantity>
, very unequal, with
<quantity id="4C8F2AB5FF93FFC1FDCDF8F7FDF4F8A2" box="[518,610,1814,1841]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="28" pageNumber="29" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
broad base, compressed, curved or hooked, up to 35(6) mm long; stipules linear
<quantity id="4C8F2AB5FF93FFC1FF1BF8DBFEA8F8C6" box="[208,318,1850,1877]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="28" pageNumber="29" unit="mm" value="11.5" valueMax="15.0" valueMin="8.0">815 mm</quantity>
, short hairy, with stipitate glands.
<emphasis id="B9035B42FF93FFC1FD0DF8DAFC82F8C7" box="[710,788,1851,1876]" italics="true" pageId="28" pageNumber="29">Leaves</emphasis>
(
<figureCitation id="134C9BD5FF93FFC1FCEFF8DBFCEAF8C7" box="[804,892,1850,1876]" captionStart="FIGURE 8" captionStartId="29.[136,229,1750,1772]" captionTargetBox="[151,1436,190,1726]" captionTargetId="figure-18@29.[151,1436,190,1726]" captionTargetPageId="29" captionText="FIGURE 8. Rubus leptodytus. A. Holotype, primocane. B. Holotype, primocane. C. Holotype, inflorescence. D. Plant of the type, habit. E. Primocane. F. Primocane. G. Leaves. H. Flowering branch. I. Inflorescence. J. Pedicels. K. Flowers. L. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061165" httpUri="https://zenodo.org/record/8061165/files/figure.png" pageId="28" pageNumber="29">Fig. 8G</figureCitation>
) 3-foliolate, rarely pedate 5-foliolate, or 2-jugate pinnate, adaxially glabrous, abaxially very short white tomentose, sometime becoming almost green; serrature fine to mediocre, obtuse, shallow, short obtuse periodical, with very short mucros; petiole
<quantity id="4C8F2AB5FF93FFC1FB9FF863FB3DF80E" box="[1108,1195,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="28" pageNumber="29" unit="cm" value="6.0" valueMax="8.0" valueMin="4.0">48 cm</quantity>
, very short tomentose, glandular, with 816 curved or hooked prickles; central leaflet of 3-foliolate or pedate leaves
<quantity id="4C8F2AB5FF93FFC1FB51F847FA83F852" box="[1178,1301,1958,1985]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" metricValueMax="7.5" metricValueMin="4.7" pageId="28" pageNumber="29" unit="mm" value="61.0" valueMax="75.0" valueMin="47.0">4775 mm</quantity>
, with cordate or rarely roundish base, suborbicular or almost quadrangular, gradually attenuate, acute or with broad short tip, or obtuse, usually lobate, widthlength index 0.761.14, petiolule
<quantity id="4C8F2AB5FF93FFC1FC9AF80FFC5BF79A" box="[849,973,2030,2057]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="4.3" metricValueMin="2.0" pageId="28" pageNumber="29" unit="mm" value="31.5" valueMax="43.0" valueMin="20.0">2043 mm</quantity>
; petiolule of basal leaves
<quantity id="4C8F2AB5FF93FFC1FB25F80FFACEF79A" box="[1262,1368,2030,2057]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="28" pageNumber="29" unit="mm" value="12.5" valueMax="20.0" valueMin="5.0">520 mm</quantity>
; central leaflets of pinnate leaves
<quantity id="4C8F2AB5FF93FFC1FE5DF7F3FD9AF7BE" box="[406,524,2066,2093]" metricMagnitude="-2" metricUnit="m" metricValue="5.15" metricValueMax="6.1" metricValueMin="4.2" pageId="28" pageNumber="29" unit="mm" value="51.5" valueMax="61.0" valueMin="42.0">4261 mm</quantity>
, with attenuate to cordate base, ovate, triangular, roundish or (elliptical-)obovate, acute or obtuse, widthlength index 0.690.96, petiolule
<quantity id="4C8F2AB5FF92FFC0FD09F866FCABF832" box="[706,829,1927,1953]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.2" metricValueMin="1.0" pageId="29" pageNumber="30" unit="mm" value="16.0" valueMax="22.0" valueMin="10.0">1022 mm</quantity>
.
<emphasis id="B9035B42FF92FFC0FC8CF866FB84F832" box="[839,1042,1927,1953]" italics="true" pageId="29" pageNumber="30">Flowering branch</emphasis>
(
<figureCitation id="134C9BD5FF92FFC0FBEAF866FBE1F832" box="[1057,1143,1927,1953]" captionStart="FIGURE 8" captionStartId="29.[136,229,1750,1772]" captionTargetBox="[151,1436,190,1726]" captionTargetId="figure-18@29.[151,1436,190,1726]" captionTargetPageId="29" captionText="FIGURE 8. Rubus leptodytus. A. Holotype, primocane. B. Holotype, primocane. C. Holotype, inflorescence. D. Plant of the type, habit. E. Primocane. F. Primocane. G. Leaves. H. Flowering branch. I. Inflorescence. J. Pedicels. K. Flowers. L. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061165" httpUri="https://zenodo.org/record/8061165/files/figure.png" pageId="29" pageNumber="30">Fig. 8H</figureCitation>
) furrowed, with dense very short stellate hairs and few to numerous stipitate glands; prickles 27 per
<quantity id="4C8F2AB5FF92FFC0FC13F84BFB87F856" box="[984,1041,1962,1989]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="30" unit="cm" value="5.0">5 cm</quantity>
from
<quantity id="4C8F2AB5FF92FFC0FB91F84BFB2FF856" box="[1114,1209,1962,1989]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="29" pageNumber="30" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">25 mm</quantity>
broad base (slightly) compressed, curved or hooked, up to
<quantity id="4C8F2AB5FF92FFC0FDF0F82EFD0CF87A" box="[571,666,1999,2025]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="29" pageNumber="30" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
long.
<emphasis id="B9035B42FF92FFC0FD2EF82EFCA5F87B" box="[741,819,1999,2024]" italics="true" pageId="29" pageNumber="30">Leaves</emphasis>
3-foliolate, or upper ones sometimes simple, adaxially glabrous or rarely with some appressed hairs, abaxially appressed grey tomentose or basal ones green; serrature usually shallow, fine to mediocre, with very short mucros, very shallowly to rather pronouncedly periodical.
<emphasis id="B9035B42FF91FFC3FB10FF7FFAE7FF2B" box="[1243,1393,158,184]" italics="true" pageId="30" pageNumber="31">Inflorescence</emphasis>
(
<figureCitation id="134C9BD5FF91FFC3FAB5FF7FFF36FF4F" captionStart="FIGURE 8" captionStartId="29.[136,229,1750,1772]" captionTargetBox="[151,1436,190,1726]" captionTargetId="figure-18@29.[151,1436,190,1726]" captionTargetPageId="29" captionText="FIGURE 8. Rubus leptodytus. A. Holotype, primocane. B. Holotype, primocane. C. Holotype, inflorescence. D. Plant of the type, habit. E. Primocane. F. Primocane. G. Leaves. H. Flowering branch. I. Inflorescence. J. Pedicels. K. Flowers. L. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061165" httpUri="https://zenodo.org/record/8061165/files/figure.png" pageId="30" pageNumber="31">Fig. 8I</figureCitation>
) cylindrical-pyramidal, with straight or somewhat flexuous axis, with 17 axial branches with sharp angle, short tomentose, with some to many stipitate glands, few weak curved or declining pricklets; peduncles often divided from base, longest with 416 flowers, tomentose, with some short stipitate glands; pedicels (
<figureCitation id="134C9BD5FF91FFC3FBA0FEEBFB2CFEB7" box="[1131,1210,266,292]" captionStart="FIGURE 8" captionStartId="29.[136,229,1750,1772]" captionTargetBox="[151,1436,190,1726]" captionTargetId="figure-18@29.[151,1436,190,1726]" captionTargetPageId="29" captionText="FIGURE 8. Rubus leptodytus. A. Holotype, primocane. B. Holotype, primocane. C. Holotype, inflorescence. D. Plant of the type, habit. E. Primocane. F. Primocane. G. Leaves. H. Flowering branch. I. Inflorescence. J. Pedicels. K. Flowers. L. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061165" httpUri="https://zenodo.org/record/8061165/files/figure.png" pageId="30" pageNumber="31">Fig. 8J</figureCitation>
)
<quantity id="4C8F2AB5FF91FFC3FB07FEEBFAADFEB6" box="[1228,1339,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="14.0" metricValueMin="2.0" pageId="30" pageNumber="31" unit="mm" value="8.0" valueMax="14.0" valueMin="2.0">214 mm</quantity>
.
<emphasis id="B9035B42FF91FFC3FA83FEEBFA32FEB7" box="[1352,1444,266,292]" italics="true" pageId="30" pageNumber="31">Flowers</emphasis>
: sepals patent to erect, 24 ×
<quantity id="4C8F2AB5FF91FFC3FE19FECFFDA5FEDA" box="[466,563,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="30" pageNumber="31" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
, ovate, with short tip, short grey tomentose and with few subsessile and short stipitate glands; petals (
<figureCitation id="134C9BD5FF91FFC3FE46FEB3FE72FEFF" box="[397,484,338,364]" captionStart="FIGURE 8" captionStartId="29.[136,229,1750,1772]" captionTargetBox="[151,1436,190,1726]" captionTargetId="figure-18@29.[151,1436,190,1726]" captionTargetPageId="29" captionText="FIGURE 8. Rubus leptodytus. A. Holotype, primocane. B. Holotype, primocane. C. Holotype, inflorescence. D. Plant of the type, habit. E. Primocane. F. Primocane. G. Leaves. H. Flowering branch. I. Inflorescence. J. Pedicels. K. Flowers. L. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061165" httpUri="https://zenodo.org/record/8061165/files/figure.png" pageId="30" pageNumber="31">Fig. 8K</figureCitation>
) pinkish red, orbicular or obovate, 4.56.0 ×
<quantity id="4C8F2AB5FF91FFC3FC11FEB3FBAFFEFE" box="[986,1081,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="30" pageNumber="31" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
; stamens deep purple red, shorter than or equalling styles; anthers and ovaries glabrous; styles deep purple red; receptacle with some hairs. Ripe
<emphasis id="B9035B42FF91FFC3FAB0FE97FA3DFE03" box="[1403,1451,374,400]" italics="true" pageId="30" pageNumber="31">fruit</emphasis>
(
<figureCitation id="134C9BD5FF91FFC3FF5BFE7BFF73FE27" box="[144,229,410,436]" captionStart="FIGURE 8" captionStartId="29.[136,229,1750,1772]" captionTargetBox="[151,1436,190,1726]" captionTargetId="figure-18@29.[151,1436,190,1726]" captionTargetPageId="29" captionText="FIGURE 8. Rubus leptodytus. A. Holotype, primocane. B. Holotype, primocane. C. Holotype, inflorescence. D. Plant of the type, habit. E. Primocane. F. Primocane. G. Leaves. H. Flowering branch. I. Inflorescence. J. Pedicels. K. Flowers. L. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061165" httpUri="https://zenodo.org/record/8061165/files/figure.png" pageId="30" pageNumber="31">Fig. 8L</figureCitation>
) pale yellow to brownish orange, not very juicy.
</paragraph>
</subSubSection>
<caption id="DF08D7D8FF92FFC0FF43F937FF4BF8A7" ID-DOI="http://doi.org/10.5281/zenodo.8061165" ID-Zenodo-Dep="8061165" httpUri="https://zenodo.org/record/8061165/files/figure.png" pageId="29" pageNumber="30" startId="29.[136,229,1750,1772]" targetBox="[151,1436,190,1726]" targetPageId="29" targetType="figure">
<paragraph id="8BC88750FF92FFC0FF43F937FF4BF8A7" blockId="29.[136,1452,1750,1844]" pageId="29" pageNumber="30">
<emphasis id="B9035B42FF92FFC0FF43F937FF68F978" bold="true" box="[136,254,1750,1772]" pageId="29" pageNumber="30">FIGURE 8.</emphasis>
<taxonomicName id="4C77FCD3FF92FFC0FECFF937FE3FF978" authorityName="A. Beek" authorityYear="2021" box="[260,425,1750,1771]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="leptodytus">
<emphasis id="B9035B42FF92FFC0FECFF937FE3FF978" box="[260,425,1750,1771]" italics="true" pageId="29" pageNumber="30">Rubus leptodytus</emphasis>
</taxonomicName>
.
<emphasis id="B9035B42FF92FFC0FE7EF937FE50F978" bold="true" box="[437,454,1750,1771]" pageId="29" pageNumber="30">A</emphasis>
. Holotype, primocane.
<emphasis id="B9035B42FF92FFC0FD61F937FD2CF978" bold="true" box="[682,698,1750,1771]" pageId="29" pageNumber="30">B</emphasis>
. Holotype, primocane.
<emphasis id="B9035B42FF92FFC0FC56F937FC38F97F" bold="true" box="[925,942,1750,1772]" pageId="29" pageNumber="30">C</emphasis>
. Holotype, inflorescence.
<emphasis id="B9035B42FF92FFC0FB66F937FB28F978" bold="true" box="[1197,1214,1750,1771]" pageId="29" pageNumber="30">D</emphasis>
. Plant of the type, habit.
<emphasis id="B9035B42FF92FFC0FF43F91BFF0EF89C" bold="true" box="[136,152,1786,1807]" pageId="29" pageNumber="30">E</emphasis>
. Primocane.
<emphasis id="B9035B42FF92FFC0FED2F91BFEBEF89C" bold="true" box="[281,296,1786,1807]" pageId="29" pageNumber="30">F</emphasis>
. Primocane.
<emphasis id="B9035B42FF92FFC0FE63F91BFE2DF883" bold="true" box="[424,443,1786,1808]" pageId="29" pageNumber="30">G</emphasis>
. Leaves.
<emphasis id="B9035B42FF92FFC0FDD3F91BFDBDF89C" bold="true" box="[536,555,1786,1807]" pageId="29" pageNumber="30">H</emphasis>
. Flowering branch.
<emphasis id="B9035B42FF92FFC0FD3BF91BFD6FF89C" bold="true" box="[752,761,1786,1807]" pageId="29" pageNumber="30">I</emphasis>
. Inflorescence.
<emphasis id="B9035B42FF92FFC0FC5FF91BFC36F89C" bold="true" box="[916,928,1786,1807]" pageId="29" pageNumber="30">J</emphasis>
. Pedicels.
<emphasis id="B9035B42FF92FFC0FBC1F91BFB8BF89C" bold="true" box="[1034,1053,1786,1807]" pageId="29" pageNumber="30">K</emphasis>
. Flowers.
<emphasis id="B9035B42FF92FFC0FB4FF91BFB02F89C" bold="true" box="[1156,1172,1786,1807]" pageId="29" pageNumber="30">L</emphasis>
. Fruits. Photographs: A. van de Beek.
</paragraph>
</caption>
<subSubSection id="C36DD4DBFF91FFC3FF76FE5FFC2AFE4B" box="[189,956,446,472]" pageId="30" pageNumber="31" type="biology_ecology">
<paragraph id="8BC88750FF91FFC3FF76FE5FFC2AFE4B" blockId="30.[136,1452,158,1013]" box="[189,956,446,472]" pageId="30" pageNumber="31">
<emphasis id="B9035B42FF91FFC3FF76FE5FFEB0FE4B" bold="true" box="[189,294,446,472]" pageId="30" pageNumber="31">Ecology:</emphasis>
—Humid places along ditches, in fields, hedges, roadsides.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF91FFC3FF76FE03FD8BFDFB" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BC88750FF91FFC3FF76FE03FD8BFDFB" blockId="30.[136,1452,158,1013]" pageId="30" pageNumber="31">
<emphasis id="B9035B42FF91FFC3FF76FE03FECFFE6F" bold="true" box="[189,345,482,508]" pageId="30" pageNumber="31">Distribution:</emphasis>
<taxonomicName id="4C77FCD3FF91FFC3FEBEFE03FDA3FE6F" authorityName="A. Beek" authorityYear="2021" box="[373,565,482,508]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="leptodytus">
<emphasis id="B9035B42FF91FFC3FEBEFE03FDA3FE6F" box="[373,565,482,508]" italics="true" pageId="30" pageNumber="31">Rubus leptodytus</emphasis>
</taxonomicName>
has a rather wide distribution in the
<collectingRegion id="49B349B2FF91FFC3FC1AFE02FBF9FE6F" box="[977,1135,482,509]" country="South Africa" name="Western Cape" pageId="30" pageNumber="31">Western Cape</collectingRegion>
,
<collectingCountry id="F360C7C0FF91FFC3FBB0FE03FA9BFE6F" box="[1147,1293,482,508]" name="South Africa" pageId="30" pageNumber="31">South Africa</collectingCountry>
; it is common south of Citrusdal near the Theerivier, and at several places around
<collectingCountry id="F360C7C0FF91FFC3FC4FFDE7FC4AFDB3" box="[900,988,518,544]" name="Algeria" pageId="30" pageNumber="31">Algeria</collectingCountry>
; more to the south near Tulbagh; further, in Oudtshoorn (Cango Valley), along the road to the Gamkaskloof, and near Caledon; specimens from Avontuur and Swellendam might also belong to it.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF91FFC3FF76FD93FA9BFC67" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BC88750FF91FFC3FF76FD93FA9BFC67" blockId="30.[136,1452,158,1013]" pageId="30" pageNumber="31">
<emphasis id="B9035B42FF91FFC3FF76FD93FD6AFD1F" bold="true" box="[189,764,626,653]" pageId="30" pageNumber="31">
<collectingRegion id="49B349B2FF91FFC3FF76FD93FEA9FD1F" box="[189,319,626,652]" name="Specimens" pageId="30" pageNumber="31">Specimens</collectingRegion>
:—
<collectingCountry id="F360C7C0FF91FFC3FEAAFD93FDABFD1F" box="[353,573,626,652]" name="South Africa" pageId="30" pageNumber="31">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FF91FFC3FD85FD93FD6AFD1F" box="[590,764,626,652]" country="South Africa" name="Western Cape" pageId="30" pageNumber="31">Western Cape</collectingRegion>
</emphasis>
: Theerivier, Citrusdal,
<date id="FFC9A190FF91FFC3FBD9FD92FB7BFD1F" box="[1042,1261,626,653]" pageId="30" pageNumber="31" value="1968-09-07">7 September 1968</date>
,
<emphasis id="B9035B42FF91FFC3FB36FD93FA3DFD1F" box="[1277,1451,626,652]" italics="true" pageId="30" pageNumber="31">Hanekom 1183</emphasis>
(PRE); Citrusdal, south of Theerivier,
<date id="FFC9A190FF91FFC3FD85FD77FCA5FD23" box="[590,819,662,688]" pageId="30" pageNumber="31" value="2019-11-20">20 November 2019</date>
,
<emphasis id="B9035B42FF91FFC3FC88FD77FC76FD23" box="[835,992,662,688]" italics="true" pageId="30" pageNumber="31">Beek 2019.94</emphasis>
(L); Citrusdal, south of Theerivier,
<date id="FFC9A190FF91FFC3FA44FD77FED4FD47" pageId="30" pageNumber="31" value="2019-11-20">20 November 2019</date>
,
<emphasis id="B9035B42FF91FFC3FE84FD5BFE7FFD47" box="[335,489,698,724]" italics="true" pageId="30" pageNumber="31">Beek 2019.95</emphasis>
(L); Algeria, along the road ±
<quantity id="4C8F2AB5FF91FFC3FC87FD5BFC02FD47" box="[844,916,698,724]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="10.0">10 km</quantity>
south of the Forest station, in shadow of rock,
<date id="FFC9A190FF91FFC3FF43FD3FFEF0FD6B" box="[136,358,734,760]" pageId="30" pageNumber="31" value="2019-11-20">20 November 2019</date>
,
<emphasis id="B9035B42FF91FFC3FEB8FD3FFD9BFD6B" box="[371,525,734,760]" italics="true" pageId="30" pageNumber="31">Beek 2019.96</emphasis>
(L); Algeria Forest Station,
<date id="FFC9A190FF91FFC3FC98FD3FFBA4FD6B" box="[851,1074,734,760]" pageId="30" pageNumber="31" value="1996-11-16">16 November 1996</date>
,
<emphasis id="B9035B42FF91FFC3FBF5FD3EFF24FC8F" italics="true" pageId="30" pageNumber="31">Van Rooyen, Steyn &amp; de Villiers 207</emphasis>
(NBG); Tulbagh, Winterhoek road,
<date id="FFC9A190FF91FFC3FD86FCE3FCBFFC8F" box="[589,809,770,796]" pageId="30" pageNumber="31" value="2019-11-21">21 November 2019</date>
,
<emphasis id="B9035B42FF91FFC3FCFEFCE3FC58FC8F" box="[821,974,770,796]" italics="true" pageId="30" pageNumber="31">Beek 2019.99</emphasis>
(L); Seweweekspoort, in Klein Swartberg near Ladismith, at the farm Aristata,
<emphasis id="B9035B42FF91FFC3FDF2FCC7FD63FCD3" box="[569,757,806,832]" italics="true" pageId="30" pageNumber="31">Henderson 1109</emphasis>
(PRE); Oudtshoorn, Cango Valley, after the off road to the Swartbergpass just before the Grobbelaars River, east side of the road,
<date id="FFC9A190FF91FFC3FC5EFCABFBD8FCF7" box="[917,1102,842,869]" pageId="30" pageNumber="31" value="2020-01-21">21 January 2020</date>
,
<emphasis id="B9035B42FF91FFC3FB93FCABFB79FCF7" box="[1112,1263,842,868]" italics="true" pageId="30" pageNumber="31">Beek 2020.12</emphasis>
(L); Boomplaas, Cango Valley,
<date id="FFC9A190FF91FFC3FEE4FC8FFE78FC1A" box="[303,494,878,905]" pageId="30" pageNumber="31" value="1975-02-05">5 February 1975</date>
,
<emphasis id="B9035B42FF91FFC3FE30FC8FFDE0FC1B" box="[507,630,878,904]" italics="true" pageId="30" pageNumber="31">Moffet 608</emphasis>
(NBG); Caledon, Riviersonderend Valley, near the farm Terra Bona,
<date id="FFC9A190FF91FFC3FA44FC8FFEA6FC3F" pageId="30" pageNumber="31" value="1992-02-18">18 February 1992</date>
,
<emphasis id="B9035B42FF91FFC3FEF0FC73FE62FC3F" box="[315,500,914,940]" italics="true" pageId="30" pageNumber="31">Henderson 1152</emphasis>
(PRE); Rivierzonder Einde, N. Ecksteen, November,
<emphasis id="B9035B42FF91FFC3FB9FFC73FB77FC3F" box="[1108,1249,914,940]" italics="true" pageId="30" pageNumber="31">Zeyher 2450</emphasis>
[?] (NBG,
<taxonomicName id="4C77FCD3FF91FFC3FA94FC73FA3AFC3F" authorityName="Linnaeus" authorityYear="1753" box="[1375,1452,914,940]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">Rubus</taxonomicName>
chrysocarpus); Baviaansrivier near Gnadendal and Waterfall near Tulbagh, November/December,
<emphasis id="B9035B42FF91FFC3FB22FC57FF56FC67" italics="true" pageId="30" pageNumber="31">Ecklon &amp; Zeyher 1708</emphasis>
(NBG,
<taxonomicName id="4C77FCD3FF91FFC3FED5FC3BFEFDFC67" authorityName="Linnaeus" authorityYear="1753" box="[286,363,986,1012]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">Rubus</taxonomicName>
chrysocarpus, it is not clear if this specimen is from Gnadendal or from Tulbagh).
</paragraph>
</subSubSection>
</treatment>
</document>