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<mods:title id="214747A06BC37ABBEE700128DE8D303C">The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha)</mods:title>
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Metriorhynchinae
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(Zoological Code)
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Lydekker 1888
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A: 91, converted clade name (PhyloCode)
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RegNum registration number1018.
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genus:
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<emphasis id="B90DE802FF85730A7B010DBF0FC0FA38" box="[609,762,1455,1479]" italics="true" pageId="53" pageNumber="600">Metriorhynchus</emphasis>
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.
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<emphasis id="B90DE802FF85730A78E30E150CD3F9E0" box="[387,489,1541,1567]" italics="true" pageId="53" pageNumber="600">Etymology</emphasis>
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Pertaining to
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. The stem
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- is based on the genus
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. -
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, is a Latin feminine plural suffix for pertaining to. Under Article 29.2 of the Zoological Code, the suffix -
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denotes a subfamily rank within the family-group.
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<emphasis id="B90DE802FF85730A78090EF10F39F904" box="[361,515,1761,1787]" italics="true" pageId="53" pageNumber="600">Geological range</emphasis>
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Middle Jurassic (middle Callovian) to Early Cretaceous (early Valanginian) (
<bibRefCitation id="EFE849E1FF85730A78680F370C98F8C0" author="Andrews CW" box="[264,418,1831,1855]" pageId="53" pageNumber="600" refId="ref48442" refString="Andrews CW. A Descriptive Catalogue of the Marine Reptiles of the Oxford Clay, Part Two. London: British Museum (Natural History), 1913." type="book" year="1913">Andrews 1913</bibRefCitation>
,
<bibRefCitation id="EFE849E1FF85730A78D30F370F4BF8C0" author="Young MT &amp; Brusatte SL &amp; Ruta M" box="[435,625,1831,1855]" pageId="53" pageNumber="600" pagination="801 - 59" refId="ref60618" refString="Young MT, Brusatte SL, Ruta M et al. The evolution of Metriorhynchoidea (Mesoeucrocodylia, Thalattosuchia): an integrated approach using geometrics morphometrics, analysis of disparity and biomechanics. Zoological Journal of the Linnean Society 2010; 158: 801 - 59." type="journal article" year="2010">
Young
<emphasis id="B90DE802FF85730A789E0F370F16F8C0" box="[510,556,1831,1855]" italics="true" pageId="53" pageNumber="600">et al</emphasis>
. 2010
</bibRefCitation>
,
<bibRefCitation id="EFE849E1FF85730A7BE20F370D9FF8A1" author="Sachs S &amp; Young MT &amp; Hornung J" pageId="53" pageNumber="600" pagination="104479" refId="ref58635" refString="Sachs S, Young MT, Hornung J. The enigma of Enaliosuchus, and a reassessment of the Early Cretaceous fossil record of Metriorhynchidae. Cretaceous Research 2020; 114: 104479." type="journal article" year="2020">
Sachs
<emphasis id="B90DE802FF85730A7BA60F370FCEF8C0" box="[710,756,1831,1855]" italics="true" pageId="53" pageNumber="600">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC63410FF85730A78790F690F69F86C" blockId="53.[113,762,1913,2007]" box="[281,595,1913,1939]" pageId="53" pageNumber="600">
<emphasis id="B90DE802FF85730A78790F690F69F86C" box="[281,595,1913,1939]" italics="true" pageId="53" pageNumber="600">PhyloCode phylogenetic definition</emphasis>
</paragraph>
<paragraph id="8BC63410FF85730A79110F8F0E55FF38" blockId="53.[113,762,1913,2007]" lastBlockId="53.[810,1459,144,199]" pageId="53" pageNumber="600">
The smallest clade within
<taxonomicName id="4C794F93FF85730A78A70F8F0F4EF848" authorityName="sensu Young and Andrade" authorityYear="2009" box="[455,628,1951,1975]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="family">
<emphasis id="B90DE802FF85730A78A70F8F0F4EF848" box="[455,628,1951,1975]" italics="true" pageId="53" pageNumber="600">Metriorhynchidae</emphasis>
</taxonomicName>
containing
<taxonomicName id="4C794F93FF85730A79110FAF0F7FF828" authority="von Meyer, 1831" authorityName="von Meyer" authorityYear="1831" box="[113,581,1983,2007]" class="Reptilia" family="Metriorhynchidae" genus="Rhacheosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B90DE802FF85730A79110FAF0C65F828" box="[113,351,1983,2007]" italics="true" pageId="53" pageNumber="600">Rhacheosaurus gracilis</emphasis>
<bibRefCitation id="EFE849E1FF85730A78150FAF0F7FF828" author="von Meyer H" box="[373,581,1983,2007]" pageId="53" pageNumber="600" pagination="171 - 200" refId="ref56665" refString="von Meyer H. Neue fossile Reptilien aus der Ordnung der Saurier. Nova Acta physico-medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosum 1831; 15: 171 - 200." type="journal article" year="1831">von Meyer, 1831</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C794F93FF85730A7B010FAF0911FF57" authority="(Holl 1829)" authorityName="Metriorhynchoidea" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[609,1067,144,2007]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="species" species="brevirostris">
<emphasis id="B90DE802FF85730A7B010FAF0EAFFF57" box="[609,917,144,2007]" italics="true" pageId="53" pageNumber="600">Metriorhynchus brevirostris</emphasis>
(
<bibRefCitation id="EFE849E1FF85730A7ACF0880091AFF57" author="Holl F" box="[943,1056,144,168]" pageId="53" pageNumber="600" refId="ref53683" refString="Holl F. Handbuch der Petrefactenkunde Teil 1. Dresden: Hilscher, 1829, 416." type="book" year="1829">Holl 1829</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C794F93FF85730A7D1408800E50FF38" authority="(Andrews 1913)" baseAuthorityName="Andrews" baseAuthorityYear="1913" class="Reptilia" family="Metriorhynchidae" genus="Gracilineustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="species" species="leedsi">
<emphasis id="B90DE802FF85730A7D1408800804FF57" box="[1140,1342,144,168]" italics="true" pageId="53" pageNumber="600">Gracilineustes leedsi</emphasis>
(
<bibRefCitation id="EFE849E1FF85730A7C3808800E64FF38" author="Andrews CW" pageId="53" pageNumber="600" refId="ref48442" refString="Andrews CW. A Descriptive Catalogue of the Marine Reptiles of the Oxford Clay, Part Two. London: British Museum (Natural History), 1913." type="book" year="1913">Andrews 1913</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BC63410FF85730A7D7208F709F6FEFE" blockId="53.[1042,1228,232,256]" box="[1042,1228,231,257]" pageId="53" pageNumber="600">
<emphasis id="B90DE802FF85730A7D7208F709F6FEFE" box="[1042,1228,231,257]" italics="true" pageId="53" pageNumber="600">Reference phylogeny</emphasis>
</paragraph>
<paragraph id="8BC63410FF85730A7A4A091D0E52FEDA" blockId="53.[810,872,270,293]" box="[810,872,269,293]" pageId="53" pageNumber="600">
<figureCitation id="13422895FF85730A7A4A091D0E58FEDA" box="[810,866,269,293]" captionStart="Figure 3" captionStartId="19.[113,178,1185,1209]" captionTargetBox="[117,1455,147,1154]" captionTargetId="figure-283@19.[114,1458,144,1157]" captionTargetPageId="19" captionText="Figure 3. The equal weights, no constraints strict consensus topology (C0). Length = 4164, CI = 0.303, RI = 0.831, RC = 0.251, HI = 0.697, number of most parsimonious cladograms hit the hold limit.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S1 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians.Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/." figureDoi="http://doi.org/10.5281/zenodo.11241251" httpUri="https://zenodo.org/record/11241251/files/figure.png" pageId="53" pageNumber="600">Fig. 3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C363679BFF85730A7A5B09550EEBFD62" pageId="53" pageNumber="600" type="diagnosis">
<paragraph id="8BC63410FF85730A7A5B09550898FEA0" blockId="53.[810,1459,325,669]" box="[827,1442,325,351]" pageId="53" pageNumber="600">
<emphasis id="B90DE802FF85730A7A5B09550898FEA0" box="[827,1442,325,351]" italics="true" pageId="53" pageNumber="600">Zoological Code diagnosis and PhyloCode diagnostic apomorphies</emphasis>
</paragraph>
<paragraph id="8BC63410FF85730A7A4A097B0EEBFD62" blockId="53.[810,1459,325,669]" pageId="53" pageNumber="600">
Metriorhynchid crocodylomorphs with the following unique combination of characters (4): reduction in external ornamentation on the maxilla (shared with the South American
<taxonomicName id="4C794F93FF85730A7A5009D90EF3FE1E" authorityName="von Meyer" authorityYear="1832" box="[816,969,457,481]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7A5009D90EF3FE1E" box="[816,969,457,481]" italics="true" pageId="53" pageNumber="600">Metriorhynchus</emphasis>
</taxonomicName>
species,
<taxonomicName id="4C794F93FF85730A7D4809DA099FFE1E" box="[1064,1189,458,481]" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7D4809DA099FFE1E" box="[1064,1189,458,481]" italics="true" pageId="53" pageNumber="600">Torvoneustes</emphasis>
</taxonomicName>
, the E-clade,
<taxonomicName id="4C794F93FF85730A7C5609D90897FE1E" box="[1334,1453,457,481]" class="Reptilia" family="Metriorhynchidae" genus="Plesiosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7C5609D90897FE1E" box="[1334,1453,457,481]" italics="true" pageId="53" pageNumber="600">Plesiosuchus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C794F93FF85730A7A4A09F90EAAFDFF" authorityName="Bonaparte" authorityYear="1850" box="[810,912,489,512]" class="Reptilia" family="Metriorhynchidae" genus="Geosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7A4A09F90EAAFDFF" box="[810,912,489,512]" italics="true" pageId="53" pageNumber="600">Geosaurus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C794F93FF85730A7AAA09F9098BFDFE" authorityName="Vignaud &amp; Gasparini" authorityYear="1996" box="[970,1201,489,513]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="species" species="andiniensis">
<emphasis id="B90DE802FF85730A7AAA09F9098BFDFE" box="[970,1201,489,513]" italics="true" pageId="53" pageNumber="600">Dakosaurus andiniensis</emphasis>
</taxonomicName>
) (18.02); lack of ornamentation on the intertemporal bar (shared with
<taxonomicName id="4C794F93FF85730A7C500A190897FDDF" box="[1328,1453,521,544]" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7C500A190897FDDF" box="[1328,1453,521,544]" italics="true" pageId="53" pageNumber="600">Torvoneustes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C794F93FF85730A7A4A0A380EAAFDC0" authorityName="Bonaparte" authorityYear="1850" box="[810,912,552,575]" class="Reptilia" family="Metriorhynchidae" genus="Geosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7A4A0A380EAAFDC0" box="[810,912,552,575]" italics="true" pageId="53" pageNumber="600">Geosaurus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C794F93FF85730A7AA90A370904FDC0" box="[969,1086,551,575]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7AA90A370904FDC0" box="[969,1086,551,575]" italics="true" pageId="53" pageNumber="600">Dakosaurus</emphasis>
</taxonomicName>
) (24.1); posterodorsal retraction of the posterior margin of the narial fossa (shared with
<taxonomicName id="4C794F93FF85730A7C560A570889FDA1" box="[1334,1459,583,606]" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7C560A570889FDA1" box="[1334,1459,583,606]" italics="true" pageId="53" pageNumber="600">Torvoneustes</emphasis>
</taxonomicName>
and the E-clade) (104.26); the humerus is shorter than the scapula (749.2).
</paragraph>
</subSubSection>
<subSubSection id="C363679BFF8573077D520AAD0FBDFB1C" lastPageId="56" lastPageNumber="603" pageId="53" pageNumber="600" type="discussion">
<paragraph id="8BC63410FF85730A7D520AAD0991FD28" blockId="53.[810,1459,701,795]" box="[1074,1195,701,727]" pageId="53" pageNumber="600">
<emphasis id="B90DE802FF85730A7D520AAD0991FD28" box="[1074,1195,701,727]" italics="true" pageId="53" pageNumber="600">Composition</emphasis>
</paragraph>
<paragraph id="8BC63410FF85730A7A4A0AF30EFCFCE5" blockId="53.[810,1459,701,795]" pageId="53" pageNumber="600">
<taxonomicName id="4C794F93FF85730A7A4A0AF30E8BFD04" box="[810,945,739,763]" class="Reptilia" family="Metriorhynchidae" genus="Gracilineustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7A4A0AF30E8BFD04" box="[810,945,739,763]" italics="true" pageId="53" pageNumber="600">Gracilineustes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C794F93FF85730A7ADB0AF30961FD04" authorityName="Parrilla-Bel, Young, Moreno-Azanza &amp; Canudo" authorityYear="2013" box="[955,1115,739,763]" family="Metriorhynchidae" genus="Maledictosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7ADB0AF30961FD04" box="[955,1115,739,763]" italics="true" pageId="53" pageNumber="600">Maledictosuchus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C794F93FF85730A7D050AF309C4FD04" authorityName="von Meyer" authorityYear="1832" box="[1125,1278,739,763]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7D050AF309C4FD04" box="[1125,1278,739,763]" italics="true" pageId="53" pageNumber="600">Metriorhynchus</emphasis>
</taxonomicName>
, and the subclade
<taxonomicName id="4C794F93FF85730A7A4A0B130EFAFCE4" box="[810,960,771,795]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="582" phylum="Chordata" rank="tribe" tribe="Rhacheosaurini">
<emphasis id="B90DE802FF85730A7A4A0B130EFAFCE4" box="[810,960,771,795]" italics="true" pageId="53" pageNumber="600">Rhacheosaurini</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC63410FF85730A7D5B0B2B099BFCAA" blockId="53.[810,1459,827,1108]" box="[1083,1185,827,853]" pageId="53" pageNumber="600">
<emphasis id="B90DE802FF85730A7D5B0B2B099BFCAA" box="[1083,1185,827,853]" italics="true" pageId="53" pageNumber="600">Comments</emphasis>
</paragraph>
<paragraph id="8BC63410FF85730A7A4A0B710ED3FBAC" blockId="53.[810,1459,827,1108]" pageId="53" pageNumber="600">
<emphasis id="B90DE802FF85730A7A4A0B710EA4FC86" box="[810,926,865,889]" italics="true" pageId="53" pageNumber="600">Authorship:</emphasis>
While the nomen
<taxonomicName id="4C794F93FF85730A7D060B71081AFC86" authorityName="Lydekker" authorityYear="1888" box="[1126,1312,865,889]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="subFamily" subFamily="Metriorhynchinae">Metriorhynchinae</taxonomicName>
was first used by
<bibRefCitation id="EFE849E1FF85730A7A270B90093AFC67" author="Lydekker R" box="[839,1024,896,920]" pageId="53" pageNumber="600" refId="ref56097" refString="Lydekker R. Catalogue of the fossil Reptilia and Amphibia in the British Museum (Natural History), Cromwell Road, S. W., Part 1. Containing the orders Ornithosauria, Crocodilia, Dinosauria, Squamata, Rhynchocephalia, and Proterosauria. London: British Museum of Natural History. 1888 a, 309." type="book" year="1888">Lydekker (1888a)</bibRefCitation>
, under the Zoological Code
<bibRefCitation id="EFE849E1FF85730A7C7F0B900889FC67" author="Fitzinger L" box="[1311,1459,896,920]" pageId="53" pageNumber="600" refId="ref51847" refString="Fitzinger L. Systema Reptilium: Amblyglossae. Fasciculus primus, Part 1. Vienna: Braumuller et Seidel, 1843, 106." type="book" year="1843">Fitzinger 1843</bibRefCitation>
is the nominal authority. The nominal author of a family-group is the author who first erected a family-group taxon that is valid (in fulfilment of Article 11), and in accordance with the Zoological Code Principle of Coordination applied to family-group names (Article 36.1). Under the PhyloCode, the nominal authority is
<bibRefCitation id="EFE849E1FF85730A7A4A0C2C0ED9FBAB" author="Lydekker R" box="[810,995,1084,1108]" pageId="53" pageNumber="600" refId="ref56097" refString="Lydekker R. Catalogue of the fossil Reptilia and Amphibia in the British Museum (Natural History), Cromwell Road, S. W., Part 1. Containing the orders Ornithosauria, Crocodilia, Dinosauria, Squamata, Rhynchocephalia, and Proterosauria. London: British Museum of Natural History. 1888 a, 309." type="book" year="1888">Lydekker (1888a)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC63410FF85730A7A4A0C6609D3F9FA" blockId="53.[810,1460,1141,1541]" pageId="53" pageNumber="600">
<emphasis id="B90DE802FF85730A7A4A0C660977FB72" box="[810,1101,1141,1165]" italics="true" pageId="53" pageNumber="600">Prior phylogenetic definition:</emphasis>
<bibRefCitation id="EFE849E1FF85730A7D390C6608BDFB72" author="Young MT &amp; Andrade MB" box="[1113,1415,1141,1166]" pageId="53" pageNumber="600" pagination="551 - 85" refId="ref60527" refString="Young MT, Andrade MB. What is Geosaurus? Redescription of Geosaurus giganteus (Thalattosuchia, Metriorhynchidae) from the Upper Jurassic of Bayern, Germany. Zoological Journal of the Linnean Society 2009; 157: 551 - 85." type="journal article" year="2009">Young and Andrade (2009)</bibRefCitation>
defined
<taxonomicName id="4C794F93FF85730A7A090C85092FFB52" authorityName="Lydekker" authorityYear="1888" box="[873,1045,1173,1197]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="subFamily" subFamily="Metriorhynchinae">
<emphasis id="B90DE802FF85730A7A090C85092FFB52" box="[873,1045,1173,1197]" italics="true" pageId="53" pageNumber="600">Metriorhynchinae</emphasis>
</taxonomicName>
as the most inclusive clade including
<taxonomicName id="4C794F93FF85730A7A4A0CA409DEFB33" authority="von Meyer, 1832" authorityName="von Meyer" authorityYear="1832" box="[810,1252,1204,1228]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="species" species="geoffroyii">
<emphasis id="B90DE802FF85730A7A4A0CA4091FFB33" box="[810,1061,1204,1228]" italics="true" pageId="53" pageNumber="600">Metriorhynchus geoffroyii</emphasis>
<bibRefCitation id="EFE849E1FF85730A7D4F0CA409DEFB33" author="von Meyer H" box="[1071,1252,1204,1228]" pageId="53" pageNumber="600" refId="ref56694" refString="von Meyer H. Palaeologica zur Geschichte der Erde und ihrer Geschopfe. Frankfurt am Main: Siegmund Schmerber, 1832, 560." type="book" year="1832">von Meyer, 1832</bibRefCitation>
</taxonomicName>
, but not
<taxonomicName id="4C794F93FF85730A7C2D0CA409AFFB14" authority="(von Sommerring 1816)" baseAuthorityName="von Sommerring" baseAuthorityYear="1816" class="Reptilia" family="Metriorhynchidae" genus="Geosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="species" species="giganteus">
<emphasis id="B90DE802FF85730A7C2D0CA40EBEFB14" italics="true" pageId="53" pageNumber="600">Geosaurus giganteus</emphasis>
(
<bibRefCitation id="EFE849E1FF85730A7AF80CC409B3FB14" author="von Sommerring ST" box="[920,1161,1235,1259]" pageId="53" pageNumber="600" pagination="37 - 58" refId="ref59308" refString="von Sommerring ST. Ueber die Lacerta gigantea der Vorwelt. Denkschriften der Koniglichen Akademie der Wissenschaften zu Munchen Classe der Mathematik und Naturwissenschaften 1816; 6: 37 - 58." type="journal article" year="1816">von Sömmerring 1816</bibRefCitation>
)
</taxonomicName>
. Here we have decided to choose a more stable clade to attach the nomen
<taxonomicName id="4C794F93FF85730A7C660CE30888FAF4" authorityName="Lydekker" authorityYear="1888" box="[1286,1458,1267,1291]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="subFamily" subFamily="Metriorhynchinae">
<emphasis id="B90DE802FF85730A7C660CE30888FAF4" box="[1286,1458,1267,1291]" italics="true" pageId="53" pageNumber="600">Metriorhynchinae</emphasis>
</taxonomicName>
to, to ensure future ergotaxonomic and nomenclatural stability. In some datasets
<taxonomicName id="4C794F93FF85730A7A880D2109C1FAB6" box="[1000,1275,1329,1353]" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="53" pageNumber="600" phylum="Chordata" rank="species" species="superciliosus">
<emphasis id="B90DE802FF85730A7A880D2109C1FAB6" box="[1000,1275,1329,1353]" italics="true" pageId="53" pageNumber="600">Thalattosuchus superciliosus</emphasis>
</taxonomicName>
is the basalmost member of
<taxonomicName id="4C794F93FF85730A7AC60D400968FA97" authorityName="Lydekker" authorityYear="1888" box="[934,1106,1360,1384]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="subFamily" subFamily="Metriorhynchinae">
<emphasis id="B90DE802FF85730A7AC60D400968FA97" box="[934,1106,1360,1384]" italics="true" pageId="53" pageNumber="600">Metriorhynchinae</emphasis>
</taxonomicName>
(e.g. Young
<emphasis id="B90DE802FF85730A7DB90D41083EFA97" box="[1241,1284,1360,1384]" italics="true" pageId="53" pageNumber="600">et al</emphasis>
. 2021), whereas in other datasets
<taxonomicName id="4C794F93FF85730A7AB70D600952FA77" authorityName="Young, Brignon, Sachs, Hornung, Foffa, Kitson, Johnson &amp; Steel" authorityYear="2020" box="[983,1128,1392,1416]" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7AB70D600952FA77" box="[983,1128,1392,1416]" italics="true" pageId="53" pageNumber="600">Thalattosuchus</emphasis>
</taxonomicName>
falls outside of
<taxonomicName id="4C794F93FF85730A7C660D6009B8FA58" authority="sensu Young and Andrade (2009)" authorityName="sensu Young and Andrade" authorityYear="2009" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="family">
<emphasis id="B90DE802FF85730A7C660D600889FA77" box="[1286,1459,1392,1416]" italics="true" pageId="53" pageNumber="600">Metriorhynchidae</emphasis>
sensu
<bibRefCitation id="EFE849E1FF85730A7A080D8009B8FA58" author="Young MT &amp; Andrade MB" box="[872,1154,1423,1448]" pageId="53" pageNumber="600" pagination="551 - 85" refId="ref60527" refString="Young MT, Andrade MB. What is Geosaurus? Redescription of Geosaurus giganteus (Thalattosuchia, Metriorhynchidae) from the Upper Jurassic of Bayern, Germany. Zoological Journal of the Linnean Society 2009; 157: 551 - 85." type="journal article" year="2009">Young and Andrade (2009)</bibRefCitation>
</taxonomicName>
(see:
<bibRefCitation id="EFE849E1FF85730A7DDE0D9F086CFA58" author="Wilberg EW" box="[1214,1366,1423,1447]" pageId="53" pageNumber="600" pagination="621 - 37" refId="ref59841" refString="Wilberg EW. What's in an outgroup? The impact of outgroup choice on the phylogenetic position of Thalattosuchia (Crocodylomorpha) and the origin of Crocodyliformes. Systematic Biology 2015 b; 64: 621 - 37. https: // doi. org / 10.1093 / sysbio / syv 020" type="journal article" year="2015">
Wilberg 2015
<bibRefCitation id="EFE849E1FF85730A7C290D9F086CFA58" author="Hicham B &amp; Nehili A &amp; Ouzzaouit LA" box="[1353,1366,1423,1447]" pageId="53" pageNumber="600" pagination="104804" refId="ref53643" refString="Hicham B, Nehili A, Ouzzaouit LA et al. Discovery of the teleosauroid crocodylomorph from the early Jurassic of Chaara cave, Middle Atlas of Morocco. Journal of African Earth Sciences 2023; 198: 104804." type="journal article" year="2023">b</bibRefCitation>
</bibRefCitation>
, Wilberg
<emphasis id="B90DE802FF85730A7A4A0DBF0E68FA39" box="[810,850,1454,1478]" italics="true" pageId="53" pageNumber="600">et al</emphasis>
. 2023). Given the instability of
<taxonomicName id="4C794F93FF85730A7DF20DBE0819FA39" authorityName="Young, Brignon, Sachs, Hornung, Foffa, Kitson, Johnson &amp; Steel" authorityYear="2020" box="[1170,1315,1454,1478]" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7DF20DBE0819FA39" box="[1170,1315,1454,1478]" italics="true" pageId="53" pageNumber="600">Thalattosuchus</emphasis>
</taxonomicName>
, and how well supported the
<taxonomicName id="4C794F93FF85730A7AA40DDE0971FA19" authority="Rhacheosaurini" authorityName="Rhacheosaurini" box="[964,1099,1486,1510]" class="Reptilia" family="Metriorhynchidae" genus="Gracilineustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7AA40DDE0971FA19" box="[964,1099,1486,1510]" italics="true" pageId="53" pageNumber="600">Gracilineustes</emphasis>
</taxonomicName>
+
<taxonomicName id="4C794F93FF85730A7D070DDE09C7FA19" box="[1127,1277,1486,1510]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="582" phylum="Chordata" rank="tribe" tribe="Rhacheosaurini">
<emphasis id="B90DE802FF85730A7D070DDE09C7FA19" box="[1127,1277,1486,1510]" italics="true" pageId="53" pageNumber="600">Rhacheosaurini</emphasis>
</taxonomicName>
clade is, we have decided to make the latter
<taxonomicName id="4C794F93FF85730A7D570DFD09D9F9FA" authorityName="Lydekker" authorityYear="1888" box="[1079,1251,1517,1541]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="subFamily" subFamily="Metriorhynchinae">
<emphasis id="B90DE802FF85730A7D570DFD09D9F9FA" box="[1079,1251,1517,1541]" italics="true" pageId="53" pageNumber="600">Metriorhynchinae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC63410FF8573077A4A0E370FBDFB1C" blockId="53.[810,1460,1574,2006]" lastBlockId="56.[129,778,1196,1251]" lastPageId="56" lastPageNumber="603" pageId="53" pageNumber="600">
<emphasis id="B90DE802FF85730A7A4A0E370EA2F9C1" box="[810,920,1575,1598]" italics="true" pageId="53" pageNumber="600">Discussion:</emphasis>
The most recent list of characters used to diagnose
<taxonomicName id="4C794F93FF85730A7A4A0E560EDEF9A1" authorityName="Lydekker" authorityYear="1888" box="[810,996,1606,1630]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="subFamily" subFamily="Metriorhynchinae">Metriorhynchinae</taxonomicName>
comes from
<bibRefCitation id="EFE849E1FF85730A7D1E0E560894F9A1" author="Young MT &amp; Andrade MB" box="[1150,1454,1606,1630]" pageId="53" pageNumber="600" pagination="551 - 85" refId="ref60527" refString="Young MT, Andrade MB. What is Geosaurus? Redescription of Geosaurus giganteus (Thalattosuchia, Metriorhynchidae) from the Upper Jurassic of Bayern, Germany. Zoological Journal of the Linnean Society 2009; 157: 551 - 85." type="journal article" year="2009">Young and Andrade (2009)</bibRefCitation>
. However, our knowledge of thalattosuchians has expanded since then, and only three of those characters can define a clade: atlas intercentrum subequal in length to atlas centrum (675.1); posterior cervical vertebra centra length is less than centrum height (697.2); tibia being shorter than the femur (less than 40%) (805.4). Unfortunately, these characters are not well sampled, and the final character also unites
<taxonomicName id="4C794F93FF85730A7D9A0F320846F8C6" authorityName="Arldt" authorityYear="1907" box="[1274,1404,1826,1849]" class="Reptilia" family="Metriorhynchidae" genus="Neustosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7D9A0F320846F8C6" box="[1274,1404,1826,1849]" italics="true" pageId="53" pageNumber="600">Neustosaurus</emphasis>
</taxonomicName>
with
<taxonomicName id="4C794F93FF85730A7A4A0F500EDCF8A7" authorityName="Lydekker" authorityYear="1888" box="[810,998,1856,1880]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="subFamily" subFamily="Metriorhynchinae">Metriorhynchinae</taxonomicName>
. The Valanginian genera
<taxonomicName id="4C794F93FF85730A7C610F500845F8A7" authorityName="Koken" authorityYear="1883" box="[1281,1407,1856,1880]" class="Reptilia" family="Metriorhynchidae" genus="Enaliosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7C610F500845F8A7" box="[1281,1407,1856,1880]" italics="true" pageId="53" pageNumber="600">Enaliosuchus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C794F93FF85730A7A4A0F700E96F888" authorityName="Arldt" authorityYear="1907" box="[810,940,1888,1911]" class="Reptilia" family="Metriorhynchidae" genus="Neustosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="53" pageNumber="600" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF85730A7A4A0F700E96F888" box="[810,940,1888,1911]" italics="true" pageId="53" pageNumber="600">Neustosaurus</emphasis>
</taxonomicName>
are extremely poorly understood (
<bibRefCitation id="EFE849E1FF85730A7C440F700897F887" author="Raspail E" box="[1316,1453,1888,1912]" pageId="53" pageNumber="600" refId="ref58136" refString="Raspail E. Observations sur un Nouveau Genre de Saurien Fossile, le Neustosaurus Gigondarum. Paris: Raspail, 1842, 56." type="book" year="1842">Raspail 1842</bibRefCitation>
,
<bibRefCitation id="EFE849E1FF85730A7A4A0F6F0E9DF868" author="Koken E" box="[810,935,1919,1943]" pageId="53" pageNumber="600" pagination="735 - 827" refId="ref55297" refString="Koken E. Die Reptilian der norddeutschen unteren Kreide. Zeitschrift deutschen Geologischen Gesellschaft 1883; 35: 735 - 827." type="journal article" year="1883">Koken 1883</bibRefCitation>
,
<bibRefCitation id="EFE849E1FF85730A7AD20F6F0978F868" author="Piveteau J" box="[946,1090,1919,1943]" pageId="53" pageNumber="600" pagination="30 - 47" refId="ref57800" refString="Piveteau J. Le reptile de Gigondas et l'evolution des Metriorhynchides. Annales de Paleontologie 1928; 17: 30 - 47." type="journal article" year="1928">Piveteau 1928</bibRefCitation>
,
<bibRefCitation id="EFE849E1FF85730A7D2D0F6F09C9F868" author="Sachs S &amp; Young MT &amp; Hornung J" box="[1101,1267,1919,1943]" pageId="53" pageNumber="600" pagination="104479" refId="ref58635" refString="Sachs S, Young MT, Hornung J. The enigma of Enaliosuchus, and a reassessment of the Early Cretaceous fossil record of Metriorhynchidae. Cretaceous Research 2020; 114: 104479." type="journal article" year="2020">
Sachs
<emphasis id="B90DE802FF85730A7DEC0F90098EF868" box="[1164,1204,1919,1943]" italics="true" pageId="53" pageNumber="600">et al</emphasis>
. 2020
</bibRefCitation>
) and them being a clade of early diverging metriorhynchines creates a ghost lineage spanning from the Valanginian into the Bathonian. Moreover, the position of
<taxonomicName id="4C794F93FF87730878740F650F1EF872" box="[276,548,1909,1933]" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="602" phylum="Chordata" rank="species" species="superciliosus">
<emphasis id="B90DE802FF87730878740F650F1EF872" box="[276,548,1909,1933]" italics="true" pageId="55" pageNumber="602">Thalattosuchus superciliosus</emphasis>
</taxonomicName>
is unstable between different datasets. Sometimes
<taxonomicName id="4C794F93FF87730878C50F840F0CF853" authorityName="Young, Brignon, Sachs, Hornung, Foffa, Kitson, Johnson &amp; Steel" authorityYear="2020" box="[421,566,1940,1964]" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="602" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF87730878C50F840F0CF853" box="[421,566,1940,1964]" italics="true" pageId="55" pageNumber="602">Thalattosuchus</emphasis>
</taxonomicName>
is recovered as the first early diverging metriorhynchine (e.g. Young
<emphasis id="B90DE802FF8773087B1A0FA40F9CF834" box="[634,678,1971,1995]" italics="true" pageId="55" pageNumber="602">et al</emphasis>
. 2021), whereas in other datasets
<taxonomicName id="4C794F93FF8773087D200F6509EBF872" authorityName="Young, Brignon, Sachs, Hornung, Foffa, Kitson, Johnson &amp; Steel" authorityYear="2020" box="[1088,1233,1909,1933]" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="602" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF8773087D200F6509EBF872" box="[1088,1233,1909,1933]" italics="true" pageId="55" pageNumber="602">Thalattosuchus</emphasis>
</taxonomicName>
is the sister-taxon to
<taxonomicName id="4C794F93FF8773087A4A0F840EDEF853" authorityName="Lydekker" authorityYear="1888" box="[810,996,1940,1964]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="602" phylum="Chordata" rank="subFamily" subFamily="Metriorhynchinae">Metriorhynchinae</taxonomicName>
+
<taxonomicName id="4C794F93FF8773087D610F8409B8F853" authorityName="Bonaparte" authorityYear="1845" box="[1025,1154,1940,1964]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="55" pageNumber="602" phylum="Chordata" rank="subFamily" subFamily="Geosaurinae">Geosaurinae</taxonomicName>
(
<bibRefCitation id="EFE849E1FF8773087DF70F84080FF853" author="Wilberg EW" box="[1175,1333,1940,1964]" pageId="55" pageNumber="602" pagination="621 - 37" refId="ref59841" refString="Wilberg EW. What's in an outgroup? The impact of outgroup choice on the phylogenetic position of Thalattosuchia (Crocodylomorpha) and the origin of Crocodyliformes. Systematic Biology 2015 b; 64: 621 - 37. https: // doi. org / 10.1093 / sysbio / syv 020" type="journal article" year="2015">
Wilberg 2015
<bibRefCitation id="EFE849E1FF8773087C480F84080FF853" author="Hicham B &amp; Nehili A &amp; Ouzzaouit LA" box="[1320,1333,1940,1964]" pageId="55" pageNumber="602" pagination="104804" refId="ref53643" refString="Hicham B, Nehili A, Ouzzaouit LA et al. Discovery of the teleosauroid crocodylomorph from the early Jurassic of Chaara cave, Middle Atlas of Morocco. Journal of African Earth Sciences 2023; 198: 104804." type="journal article" year="2023">b</bibRefCitation>
</bibRefCitation>
, Wilberg
<emphasis id="B90DE802FF8773087CC20F850E06F834" italics="true" pageId="55" pageNumber="602">et al</emphasis>
. 2023). Given the instability of
<taxonomicName id="4C794F93FF8773087DEB0FA30826F834" authorityName="Young, Brignon, Sachs, Hornung, Foffa, Kitson, Johnson &amp; Steel" authorityYear="2020" box="[1163,1308,1971,1995]" family="Metriorhynchidae" genus="Thalattosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="55" pageNumber="602" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF8773087DEB0FA30826F834" box="[1163,1308,1971,1995]" italics="true" pageId="55" pageNumber="602">Thalattosuchus</emphasis>
</taxonomicName>
, and how well supported the
<taxonomicName id="4C794F93FF88730778770CBC0CA4FB3B" authority="Rhacheosaurini" authorityName="Rhacheosaurini" box="[279,414,1196,1220]" class="Reptilia" family="Metriorhynchidae" genus="Gracilineustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="603" phylum="Chordata" rank="genus">
<emphasis id="B90DE802FF88730778770CBC0CA4FB3B" box="[279,414,1196,1220]" italics="true" pageId="56" pageNumber="603">Gracilineustes</emphasis>
</taxonomicName>
+
<taxonomicName id="4C794F93FF88730778DA0CBC0F60FB3B" box="[442,602,1196,1220]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="582" phylum="Chordata" rank="tribe" tribe="Rhacheosaurini">Rhacheosaurini</taxonomicName>
clade is, we have decided to make the latter clade
<taxonomicName id="4C794F93FF88730778A70CDB0FB9FB1C" authorityName="Lydekker" authorityYear="1888" box="[455,643,1227,1251]" class="Reptilia" family="Metriorhynchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="603" phylum="Chordata" rank="subFamily" subFamily="Metriorhynchinae">Metriorhynchinae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>