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<document id="344D5CBD394971E87D04822B10A7752B" ID-CLB-Dataset="20607" ID-DOI="10.11646/zootaxa.5100.1.6" ID-GBIF-Dataset="4e55483a-a5d2-48e6-b868-524c84158851" ID-ISSN="1175-5326" ID-Zenodo-Dep="6127852" ID-ZooBank="3ED947F2-7E12-4585-BE63-967785BAB369" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645085107942" checkinUser="plazi" docAuthor="Hutchinson, Paul M. &amp; Allsopp, Peter G." docDate="2022" docId="038EED58C7164C3CFF77FD8CFF64FB58" docLanguage="en" docName="zootaxa.5100.1.6.pdf" docOrigin="Zootaxa 5100 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systellopini Von Dalla Torre 1912" docType="treatment" docVersion="8" lastPageNumber="120" masterDocId="FFB79520C7174C3DFFE0FF94FF84FFE2" masterDocTitle="Liomenochilus ongi (Coleoptera: Scarabaeidae: Melolonthinae: Systellopini), a new genus and new species from northern Western Australia" masterLastPageNumber="128" masterPageNumber="119" pageNumber="120" updateTime="1699314375074" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B91F370011D3B0A01E7C9DBBD169DF70" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="272B7AB769E70817C4251E5BF8526CC8">
<mods:title id="78BD43E01C0F2698EBE9FF0856C670A2">Liomenochilus ongi (Coleoptera: Scarabaeidae: Melolonthinae: Systellopini), a new genus and new species from northern Western Australia</mods:title>
</mods:titleInfo>
<mods:name id="76C619CBE7ED999FF717D1CDC2137658" type="personal">
<mods:role id="5F176E16769538C974982CB9AB2AE63D">
<mods:roleTerm id="256F27969C06A57321C2F41ED97293CD">Author</mods:roleTerm>
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<mods:namePart id="CB8F6D4260BC419F60289DE732D4D829">Hutchinson, Paul M.</mods:namePart>
<mods:nameIdentifier id="D2D28E83EEBBEC16891186E4CF8A3B82" type="ORCID">0000-0001-8747-1320</mods:nameIdentifier>
<mods:affiliation id="D81D777DD5A2E2E5397AD85E3447B72C">Department of Primary Industries and Regional Development (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia 6986, Australia. paul. hutchinson @ dpird. wa. gov. au; https: // orcid. org / 0000 - 0001 - 8747 - 1320</mods:affiliation>
<mods:nameIdentifier id="01129921399568B06BB5495E71C64972" type="email">paul.hutchinson@dpird.wa.gov.au</mods:nameIdentifier>
</mods:name>
<mods:name id="FFDD8E3D69122C71F61A3C6E14DFEB61" type="personal">
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<mods:namePart id="3EBA4D8A130EE060824F62ECF955367E">Allsopp, Peter G.</mods:namePart>
<mods:affiliation id="88EBDFE64C49D3C44B2C2200299BFBAB">Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia.</mods:affiliation>
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<mods:title id="09DF7AFA27C600A5BCFFCE2F0C24B9E5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="348F73DDCFDB99988BFC6B729D639038">
<mods:date id="A9C34468142E216780603F4D7551C047">2022</mods:date>
<mods:detail id="BC98D11F7A001E2C412B39C5517155ED" type="pubDate">
<mods:number id="9F45703B5969D95F890DFCB713729BF5">2022-02-17</mods:number>
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<mods:detail id="CE695BBE71444FBBB621723968E4491A" type="volume">
<mods:number id="C4D6431D6D421431325920E4C67E3C5F">5100</mods:number>
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<mods:classification id="709C09794E0CAA4B11BE37634F263886">journal article</mods:classification>
<mods:identifier id="231B90C33A1F46807264C4B89C1223EF" type="CLB-Dataset">20607</mods:identifier>
<mods:identifier id="CE6B05476DAA4AC4F0AB58269F83FBB4" type="DOI">10.11646/zootaxa.5100.1.6</mods:identifier>
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<treatment id="038EED58C7164C3CFF77FD8CFF64FB58" ID-DOI="http://doi.org/10.5281/zenodo.6312409" ID-GBIF-Taxon="193389249" ID-Zenodo-Dep="6312409" LSID="urn:lsid:plazi:treatment:038EED58C7164C3CFF77FD8CFF64FB58" httpUri="http://treatment.plazi.org/id/038EED58C7164C3CFF77FD8CFF64FB58" lastPageNumber="120" pageId="1" pageNumber="120">
<subSubSection id="C33D0FC5C7164C3CFF77FD8CFEF7FDD0" box="[151,371,536,562]" pageId="1" pageNumber="120" type="nomenclature">
<paragraph id="8B985C4EC7164C3CFF77FD8CFEF7FDD0" blockId="1.[151,371,536,562]" box="[151,371,536,562]" pageId="1" pageNumber="120">
<heading id="D0D0EB22C7164C3CFF77FD8CFEF7FDD0" bold="true" box="[151,371,536,562]" fontSize="11" level="1" pageId="1" pageNumber="120" reason="1">
<emphasis id="B953805CC7164C3CFF77FD8CFEF7FDD0" bold="true" box="[151,371,536,562]" pageId="1" pageNumber="120">
Tribe
<taxonomicName id="4C2727CDC7164C3CFF02FD8CFEF7FDD0" ID-CoL="L9G" authorityName="Von Dalla Torre" authorityYear="1912" box="[226,371,536,562]" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" rank="tribe" tribe="Systellopini">Systellopini</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D0FC5C7164C3CFF77FDF4FF64FB58" pageId="1" pageNumber="120" type="diagnosis">
<paragraph id="8B985C4EC7164C3CFF77FDF4FD68FC00" blockId="1.[151,1437,608,1210]" pageId="1" pageNumber="120">
<emphasis id="B953805CC7164C3CFF77FDF4FD16FD98" bold="true" box="[151,658,608,634]" pageId="1" pageNumber="120">
Revised diagnosis (
<figureCitation id="131C40CBC7164C3CFE60FDF4FE74FD98" box="[384,496,608,634]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1940,1964]" captionTargetBox="[151,1436,214,1915]" captionTargetId="figure-19@2.[151,1436,179,1915]" captionTargetPageId="2" captionText="FIGURES 1–7. Liomenochilus ongi Hutchinson &amp; Allsopp, new species, holotype male. 1, dorsal view; 2, ventral view; 3, lateral view; 4, head, dorsal view; 5, head and pronotum, oblique view; 6, pygidium; 7, label." figureDoi="http://doi.org/10.5281/zenodo.6127854" httpUri="https://zenodo.org/record/6127854/files/figure.png" pageId="1" pageNumber="120">Figs. 1–6</figureCitation>
,
<figureCitation id="131C40CBC7164C3CFDE0FDF4FDAFFD98" box="[512,555,608,634]" captionStart="FIGURES 8–9" captionStartId="3.[151,264,1220,1244]" captionTargetBox="[151,1436,417,1197]" captionTargetId="figure-66@3.[151,1436,417,1197]" captionTargetPageId="3" captionText="FIGURES 8–9. Liomenochilus ongi Hutchinson &amp; Allsopp, new species, holotype male, aedeagus. 8, dorsal view; 9, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6127856" httpUri="https://zenodo.org/record/6127856/files/figure.png" pageId="1" pageNumber="120">8–9</figureCitation>
,
<figureCitation id="131C40CBC7164C3CFDDCFDF4FD01FD98" box="[572,645,608,634]" captionStart-0="FIGURES 12–18. 12–17" captionStart-1="FIGURES 19–24" captionStartId-0="6.[151,264,1816,1840]" captionStartId-1="7.[151,264,860,884]" captionTargetBox-0="[157,1416,216,1776]" captionTargetBox-1="[151,1436,181,837]" captionTargetId-0="figure-19@6.[151,1436,181,1792]" captionTargetId-1="figure-17@7.[151,1436,181,837]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 12–18. 12–17. Representative males of each of six systellopine genera, dorsal view. 12, Chilodiplus weiri Allsopp, 1993 (photograph by CSIRO). 13, Enamillus pilicollis (Sharp, 1877). 14, Prochelyna heterodoxa Burmeister, 1855. 15, Sarothromerus sharpi (Blackburn, 1905). 16, Sphyrocallus brunneus Sharp, 1877. 17, Systellopus obtusus Sharp, 1877. 18, Antennal club of male Sarothromerus sharpi (Blackburn, 1905)." captionText-1="FIGURES 19–24. Representative males of each of six systellopine genera, head and pronotum, oblique view. 19 Chilodiplus weiri Allsopp, 1993 (photograph by CSIRO). 20, Enamillus pilicollis (Sharp, 1877). 21 Prochelyna heterodoxa Burmeister, 1855. 22, Sarothromerus sharpi (Blackburn, 1905). 23, Sphyrocallus brunneus Sharp, 1877. 24, Systellopus obtusus Sharp, 1877." figureDoi-0="http://doi.org/10.5281/zenodo.6127862" figureDoi-1="http://doi.org/10.5281/zenodo.6127864" httpUri-0="https://zenodo.org/record/6127862/files/figure.png" httpUri-1="https://zenodo.org/record/6127864/files/figure.png" pageId="1" pageNumber="120">12–24</figureCitation>
).
</emphasis>
Modified from
<bibRefCitation id="EFB621BFC7164C3CFCAFFDF4FC7BFD98" author="Allsopp, P. G." box="[847,1023,608,634]" pageId="1" pageNumber="120" pagination="197 - 227" refId="ref4361" refString="Allsopp, P. G. (1990) Revision of the Systellopini (Coleoptera: Scarabaeidae: Melolonthinae). Invertebrate Taxonomy, 3, 197 - 227. https: // doi. org / 10.1071 / IT 9890197" type="journal article" year="1990">Allsopp (1990)</bibRefCitation>
. Male. Form normal, not unusually depressed; surface of body and legs dull coloured, not metallic. Labrum large, extended in front of clypeus and in same general plane, either clearly differentiated by a complete suture dorsally or suture only evident on lateral declivity. Clypeus without a fringe of erect, stout-pointed spines along anterior margin, lateral edges usually deeply emarginate before ocular canthus. Antennal club with 3, 5, 6 or 7 lamellae. Pronotum with defined lateral margins and edges. Mesosternum and metasternum without a trace of a median process. Anterior leg with coxae projecting to a distance not greater than width of coxal cavity; tibiae each with 1 or 2 teeth on outer edge in addition to apical tooth. Metacoxae inflated, usually without defined lateral margins; metafemora inflated; metatibia very short and stout, ratio length:width 0.9–2.2:1, complete transverse, setose ridge on middle of outer side, 2 distal spurs, 1 above and 1 below articulation of tarsus. All claws simple and equal. Abdomen without longitudinal ridge along each side. Parameres long and slender, simple and symmetrical.
</paragraph>
<paragraph id="8B985C4EC7164C3CFF27FC78FF64FB58" blockId="1.[151,1437,608,1210]" pageId="1" pageNumber="120">
The tribe contains
<taxonomicName id="4C2727CDC7164C3CFE42FC78FD3EFBE4" authority="Sharp, 1877" authorityName="Sharp" authorityYear="1877" box="[418,698,1004,1030]" class="Insecta" family="Melolonthidae" genus="Chilodiplus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B953805CC7164C3CFE42FC78FDA1FBE4" box="[418,549,1004,1030]" italics="true" pageId="1" pageNumber="120">Chilodiplus</emphasis>
<bibRefCitation id="EFB621BFC7164C3CFDCFFC78FD3EFBE4" author="Sharp, D." box="[559,698,1004,1030]" pageId="1" pageNumber="120" pagination="311 - 320" refId="ref4956" refString="Sharp, D. (1877) Descriptions of some new forms of aberrant Melolonthini from Australia forming a distinct subtribe (Systellopides). Annali del Museo Civico di Storia Naturale Giacomo Doria, 9, 311 - 320." type="journal article" year="1877">Sharp, 1877</bibRefCitation>
</taxonomicName>
(4 species),
<taxonomicName id="4C2727CDC7164C3CFCB1FC78FBD3FBE4" authority="Sharp, 1877" authorityName="Sharp" authorityYear="1877" box="[849,1111,1004,1030]" class="Insecta" family="Melolonthidae" genus="Enamillus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B953805CC7164C3CFCB1FC78FC47FBE4" box="[849,963,1004,1030]" italics="true" pageId="1" pageNumber="120">Enamillus</emphasis>
<bibRefCitation id="EFB621BFC7164C3CFC2DFC78FBD3FBE4" author="Sharp, D." box="[973,1111,1004,1030]" pageId="1" pageNumber="120" pagination="311 - 320" refId="ref4956" refString="Sharp, D. (1877) Descriptions of some new forms of aberrant Melolonthini from Australia forming a distinct subtribe (Systellopides). Annali del Museo Civico di Storia Naturale Giacomo Doria, 9, 311 - 320." type="journal article" year="1877">Sharp, 1877</bibRefCitation>
</taxonomicName>
(8 species),
<taxonomicName id="4C2727CDC7164C3CFB0FFC78FEACFBC8" authority="Erichson, 1847" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Melolonthidae" genus="Prochelyna" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B953805CC7164C3CFB0FFC78FAEBFBE4" box="[1263,1391,1004,1030]" italics="true" pageId="1" pageNumber="120">Prochelyna</emphasis>
Erichson, 1847
</taxonomicName>
(3 species),
<taxonomicName id="4C2727CDC7164C3CFE5CFB84FCACFBC8" authority="Blackburn, 1907" authorityName="Blackburn" authorityYear="1907" box="[444,808,1040,1066]" class="Insecta" family="Melolonthidae" genus="Sarothromerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B953805CC7164C3CFE5CFB84FDE6FBC8" box="[444,610,1040,1066]" italics="true" pageId="1" pageNumber="120">Sarothromerus</emphasis>
Blackburn, 1907
</taxonomicName>
(3 species),
<taxonomicName id="4C2727CDC7164C3CFC5CFB84FB64FBC8" authority="Sharp, 1877" authorityName="Sharp" authorityYear="1877" box="[956,1248,1040,1066]" class="Insecta" family="Melolonthidae" genus="Sphyrocallus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B953805CC7164C3CFC5CFB84FBC9FBC8" box="[956,1101,1040,1066]" italics="true" pageId="1" pageNumber="120">Sphyrocallus</emphasis>
<bibRefCitation id="EFB621BFC7164C3CFBB6FB84FB64FBC8" author="Sharp, D." box="[1110,1248,1040,1066]" pageId="1" pageNumber="120" pagination="311 - 320" refId="ref4956" refString="Sharp, D. (1877) Descriptions of some new forms of aberrant Melolonthini from Australia forming a distinct subtribe (Systellopides). Annali del Museo Civico di Storia Naturale Giacomo Doria, 9, 311 - 320." type="journal article" year="1877">Sharp, 1877</bibRefCitation>
</taxonomicName>
(3 species), and
<taxonomicName id="4C2727CDC7164C3CFF77FBA0FE20FBAC" authority="Sharp, 1877" authorityName="Sharp" authorityYear="1877" box="[151,420,1076,1102]" class="Insecta" family="Melolonthidae" genus="Systellopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B953805CC7164C3CFF77FBA0FE91FBAC" box="[151,277,1076,1102]" italics="true" pageId="1" pageNumber="120">Systellopus</emphasis>
<bibRefCitation id="EFB621BFC7164C3CFEFDFBA0FE20FBAC" author="Sharp, D." box="[285,420,1076,1102]" pageId="1" pageNumber="120" pagination="311 - 320" refId="ref4956" refString="Sharp, D. (1877) Descriptions of some new forms of aberrant Melolonthini from Australia forming a distinct subtribe (Systellopides). Annali del Museo Civico di Storia Naturale Giacomo Doria, 9, 311 - 320." type="journal article" year="1877">Sharp, 1877</bibRefCitation>
</taxonomicName>
(1 species) (
<bibRefCitation id="EFB621BFC7164C3CFDD4FBA0FD54FBAC" author="Allsopp, P. G." box="[564,720,1076,1102]" pageId="1" pageNumber="120" pagination="197 - 227" refId="ref4361" refString="Allsopp, P. G. (1990) Revision of the Systellopini (Coleoptera: Scarabaeidae: Melolonthinae). Invertebrate Taxonomy, 3, 197 - 227. https: // doi. org / 10.1071 / IT 9890197" type="journal article" year="1990">Allsopp 1990</bibRefCitation>
,
<bibRefCitation id="EFB621BFC7164C3CFD3CFBA0FC93FBAC" author="Allsopp, P. G." box="[732,791,1076,1102]" pageId="1" pageNumber="120" pagination="49 - 55" refId="ref4402" refString="Allsopp, P. G. (1993) Three new species of Chilodiplus Sharp (Coleoptera: Scarabaeidae: Melolonthinae) from northern Australia. Australian Entomologist, 20, 49 - 55." type="journal article" year="1993">1993</bibRefCitation>
;
<bibRefCitation id="EFB621BFC7164C3CFCC2FBA0FC5FFBAC" author="Weir, T. A. &amp; Lawrence, J. F. &amp; Lemann, C. &amp; Gunter, N. L." box="[802,987,1076,1102]" pageId="1" pageNumber="120" pagination="1819" refId="ref5152" refString="Weir, T. A., Lawrence, J. F., Lemann, C. &amp; Gunter, N. L. (2019) Scarabaeidae: Melolonthinae Leach 1819. In: Slipinski, A. &amp; Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 447 - 507." type="book chapter" year="2019">
Weir
<emphasis id="B953805CC7164C3CFCBFFBA1FC1CFBAC" box="[863,920,1076,1102]" italics="true" pageId="1" pageNumber="120">et al.</emphasis>
2019
</bibRefCitation>
), and
<taxonomicName id="4C2727CDC7164C3CFBFFFBA0FF48FB91" authority="Hutchinson &amp; Allsopp" authorityName="Hutchinson &amp; Allsopp" authorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Liomenochilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B953805CC7164C3CFBFFFBA0FB40FBAC" box="[1055,1220,1076,1102]" italics="true" pageId="1" pageNumber="120">Liomenochilus</emphasis>
Hutchinson &amp; Allsopp
</taxonomicName>
<emphasis id="B953805CC7164C3CFF32FBCCFEC9FB90" bold="true" box="[210,333,1112,1138]" pageId="1" pageNumber="120">
<taxonomicNameLabel id="A2603D27C7164C3CFF32FBCCFEC9FB90" box="[210,333,1112,1138]" pageId="1" pageNumber="120" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
(1 species). A further species,
<taxonomicName id="4C2727CDC7164C3CFD42FBCCFBABFB90" authority="Westwood, 1842" authorityName="Westwood" authorityYear="1842" box="[674,1071,1112,1138]" class="Insecta" family="Melolonthidae" genus="Metascelis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="flexilis">
<emphasis id="B953805CC7164C3CFD42FBCCFCEFFB90" box="[674,875,1112,1138]" italics="true" pageId="1" pageNumber="120">Metascelis flexilis</emphasis>
Westwood, 1842
</taxonomicName>
, was included in the
<taxonomicName id="4C2727CDC7164C3CFAF6FBCCFA18FB90" authorityName="Von Dalla Torre" authorityYear="1912" box="[1302,1436,1112,1138]" pageId="1" pageNumber="120" rank="tribe" tribe="Systellopini">Systellopini</taxonomicName>
(
<bibRefCitation id="EFB621BFC7164C3CFF7FFBE8FEE7FB74" author="Dalla Torre, K. W. von" box="[159,355,1148,1175]" pageId="1" pageNumber="120" pagination="1 - 84" refId="ref4699" refString="Dalla Torre, K. W. von. (1912) Scarabaeidae: Melolonthinae. I. Coleoptorum Catalogus, 45, 1 - 84." type="journal article" year="1912">Dalla Torre 1912</bibRefCitation>
), but the only known specimen is from the India-Nepal border and is not a systellopine (
<bibRefCitation id="EFB621BFC7164C3CFAA2FBE8FF57FB58" author="Allsopp, P. G." pageId="1" pageNumber="120" pagination="197 - 227" refId="ref4361" refString="Allsopp, P. G. (1990) Revision of the Systellopini (Coleoptera: Scarabaeidae: Melolonthinae). Invertebrate Taxonomy, 3, 197 - 227. https: // doi. org / 10.1071 / IT 9890197" type="journal article" year="1990">Allsopp 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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