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<document ID-DOI="http://doi.org/10.5281/zenodo.3792378" ID-GBIF-Dataset="125703f8-ddaa-4f28-9673-97b7c469b05c" ID-GBIF-Taxon="164126587" ID-ISSN="1313–2970" ID-Zenodo-Dep="3792378" checkinTime="1588014657767" checkinUser="plazi" docAuthor="Chamorro-Lacayo, Maria &amp; Konstantinov, Alexander" docDate="2009" docId="552287F6FFA9FFB1FF538D04FBC76B1E" docLanguage="en" docName="ZK_article_1978.pdf" docOrigin="ZooKeys 8 (8)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@60a754c6" docStyleName="zookeys.2008.journal_article" docTitle="Chlamisini Gressitt 1946" docType="key" docVersion="6" lastPageId="22" lastPageNumber="85" masterDocId="A91BFF8EFFBCFFA7FFDF8F7BFF946F13" masterDocTitle="Synopsis of warty leaf beetle genera of the world (Coleoptera, Chrysomelidae, Cryptocephalinae, Chlamisini)" masterLastPageNumber="88" masterPageNumber="63" pageId="21" pageNumber="84" updateTime="1636502541170" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Synopsis of warty leaf beetle genera of the world (Coleoptera, Chrysomelidae, Cryptocephalinae, Chlamisini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chamorro-Lacayo, Maria</mods:namePart>
<mods:affiliation>National Museum of Natural History, Washington D. C. ,,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Konstantinov, Alexander</mods:namePart>
<mods:affiliation>Systematic Entomology Laboratory, MRC- 168 Washington, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-04-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>8</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>8</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>63</mods:start>
<mods:end>88</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.8.90</mods:identifier>
<mods:identifier type="GBIF-Dataset">125703f8-ddaa-4f28-9673-97b7c469b05c</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576444</mods:identifier>
</mods:mods>
<subSection lastPageId="22" lastPageNumber="85" pageId="21" pageNumber="84" type="key">
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792378" ID-GBIF-Taxon="164126587" ID-Zenodo-Dep="3792378" LSID="urn:lsid:plazi:treatment:552287F6FFA9FFB1FF538D04FBC76B1E" httpUri="http://treatment.plazi.org/id/552287F6FFA9FFB1FF538D04FBC76B1E" lastPageId="22" lastPageNumber="85" pageId="21" pageNumber="84">
<subSubSection box="[140,330,639,666]" pageId="21" pageNumber="84" type="nomenclature">
<paragraph blockId="21.[140,330,639,666]" box="[140,330,639,666]" pageId="21" pageNumber="84">
<heading bold="true" box="[140,330,639,666]" fontSize="11" level="1" pageId="21" pageNumber="84" reason="1">
<emphasis bold="true" box="[140,330,639,666]" pageId="21" pageNumber="84">
Key to
<taxonomicName authorityName="Gressitt" authorityYear="1946" box="[236,330,639,666]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="84" phylum="Arthropoda" rank="tribe" tribe="Chlamisini">genera</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="85" pageId="21" pageNumber="84" type="key">
<key lastPageId="22" lastPageNumber="85" pageId="21" pageNumber="84">
<keyStep pageId="21" pageNumber="84">
<paragraph blockId="21.[140,1108,695,1672]" pageId="21" pageNumber="84">
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1 Head with large laminar projection in occipital region. (
<figureCitation box="[876,1047,695,721]" captionStart="Figure 6" captionStartId="16.[140,212,1622,1644]" captionTargetBox="[140,1108,165,1584]" captionTargetId="figure@16.[140,1108,165,1603]" captionTargetPageId="16" captionText="Figure 6. Kakita monrosi (Bokermann). A, Ventral view. B, Lateral view. C, Antenna. D, Frontal view. E, Prosternum." figureDoi="http://doi.org/10.5281/zenodo.3788014" httpUri="https://zenodo.org/record/3788014/files/figure.png" pageId="21" pageNumber="84">Figs. 6 A, B, D</figureCitation>
)...... .............................
<emphasis bold="true" box="[467,1107,729,756]" pageId="21" pageNumber="84">
<taxonomicName authority="Chamorro-Lacayo &amp; Konstantinov" authorityName="Chamorro-Lacayo &amp; Konstantinov" authorityYear="2009" box="[467,975,729,756]" class="Insecta" family="Chrysomelidae" genus="Kakita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[467,550,729,756]" italics="true" pageId="21" pageNumber="84">Kakita</emphasis>
Chamorro-Lacayo &amp; Konstantinov
</taxonomicName>
, new name
</emphasis>
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</paragraph>
<paragraph blockId="21.[140,1108,695,1672]" box="[140,1107,765,792]" pageId="21" pageNumber="84">
<keyLead box="[140,1107,765,792]" pageId="21" pageNumber="84">
– Head without large laminar projection in occipital region ..........................
<emphasis bold="true" box="[1092,1107,765,791]" pageId="21" pageNumber="84">2</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="84">
<paragraph blockId="21.[140,1108,695,1672]" pageId="21" pageNumber="84">
<keyLead pageId="21" pageNumber="84">
2(1) Legs atypical; femora and tibiae flattened; tarsi reduced, retractile, able to fit into groove along apex of tibia (
<figureCitation box="[599,691,835,862]" captionStart="Figure 3" captionStartId="7.[140,212,1526,1548]" captionTargetBox="[148,1095,175,1500]" captionTargetId="figure@7.[136,1110,165,1507]" captionTargetPageId="7" captionText="Figure 3. Chlamisini, ventral view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770843" httpUri="https://zenodo.org/record/3770843/files/figure.png" pageId="21" pageNumber="84">Fig. 3 B</figureCitation>
) ...
<taxonomicName authority="Lacordaire, 1848" authorityName="Lacordaire" authorityYear="1848" box="[730,1107,835,862]" class="Insecta" family="Chrysomelidae" genus="Carcinobaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[730,1107,835,862]" pageId="21" pageNumber="84">
<emphasis bold="true" box="[730,897,835,862]" italics="true" pageId="21" pageNumber="84">Carcinobaena</emphasis>
<bibRefCitation author="Lacordaire" box="[905,1107,835,861]" firstAuthor="Lacordaire" pageId="21" pageNumber="84" refId="ref10277" refString="Lacordaire MT (1848) Monographie des Coleopteres subpentameres de la famille des Phytophages. Tome second. Bruxelles et Leipzig, 890 pp." type="book" year="1848">Lacordaire, 1848</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="21.[140,1108,695,1672]" box="[140,1107,870,897]" pageId="21" pageNumber="84">
<keyLead box="[140,1107,870,897]" pageId="21" pageNumber="84">
– Legs normal and not as above (
<figureCitation box="[595,724,871,897]" captionStart="Figure 3" captionStartId="7.[140,212,1526,1548]" captionTargetBox="[148,1095,175,1500]" captionTargetId="figure@7.[136,1110,165,1507]" captionTargetPageId="7" captionText="Figure 3. Chlamisini, ventral view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770843" httpUri="https://zenodo.org/record/3770843/files/figure.png" pageId="21" pageNumber="84">Fig. 3 E, H</figureCitation>
) ................................................
<emphasis bold="true" box="[1092,1107,870,896]" pageId="21" pageNumber="84">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="84">
<paragraph blockId="21.[140,1108,695,1672]" pageId="21" pageNumber="84">
<keyLead pageId="21" pageNumber="84">
3(1) Prosternum nearly rectangular, with posterior margin slightly narrower than anterior margin (
<figureCitation box="[438,537,941,968]" captionStart="Figure 3" captionStartId="7.[140,212,1526,1548]" captionTargetBox="[148,1095,175,1500]" captionTargetId="figure@7.[136,1110,165,1507]" captionTargetPageId="7" captionText="Figure 3. Chlamisini, ventral view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770843" httpUri="https://zenodo.org/record/3770843/files/figure.png" pageId="21" pageNumber="84">Fig. 3 H</figureCitation>
). Pronotal and elytral tubercules reduced.............. ....................................................................
<taxonomicName authority="Monros, 1948" authorityName="Monros" authorityYear="1948" box="[751,1107,975,1003]" class="Insecta" family="Chrysomelidae" genus="Melittochlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[751,1107,975,1003]" pageId="21" pageNumber="84">
<emphasis bold="true" box="[751,931,976,1003]" italics="true" pageId="21" pageNumber="84">Melittochlamys</emphasis>
<bibRefCitation author="Monros" box="[939,1107,975,1002]" firstAuthor="Monros" pageId="21" pageNumber="84" pagination="171 - 200" refId="ref10353" refString="Monros F (1948) Descripcion de diez nuevos Camptosoma neotropicales. Acta Zoologica Lilloana 6: 171 - 200." type="journal article" year="1948">Monrós, 1948</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="21.[140,1108,695,1672]" pageId="21" pageNumber="84">
<keyLead pageId="21" pageNumber="84">
– Prosternum varying in shape, triangular, or angulate between mesocoxae, but never rectangular, with posterior margin much narrower than anterior margin (
<figureCitation box="[297,424,1082,1108]" captionStart="Figure 3" captionStartId="7.[140,212,1526,1548]" captionTargetBox="[148,1095,175,1500]" captionTargetId="figure@7.[136,1110,165,1507]" captionTargetPageId="7" captionText="Figure 3. Chlamisini, ventral view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770843" httpUri="https://zenodo.org/record/3770843/files/figure.png" pageId="21" pageNumber="84">Figs. 3 A, I</figureCitation>
). Pronotal and elytral tubercules well developed ................
<emphasis bold="true" box="[1092,1107,1081,1107]" pageId="21" pageNumber="84">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="84">
<paragraph blockId="21.[140,1108,695,1672]" pageId="21" pageNumber="84">
<keyLead pageId="21" pageNumber="84">
4(3) Head with vertex longitudinally impressed. Tarsal claws simple. Body about
<quantity box="[246,328,1152,1179]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="84" unit="mm" value="10.0">10 mm</quantity>
long and longer, bright metallic color. Pronotal and elytral tubercules pronounced .................................................................
<taxonomicName authority="Voet, 1806" authorityName="Voet" authorityYear="1806" box="[869,1107,1187,1214]" class="Insecta" family="Chrysomelidae" genus="Fulcidax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[869,1107,1187,1214]" pageId="21" pageNumber="84">
<emphasis bold="true" box="[869,972,1187,1214]" italics="true" pageId="21" pageNumber="84">Fulcidax</emphasis>
<bibRefCitation author="Voet" box="[979,1107,1187,1213]" firstAuthor="Voet" pageId="21" pageNumber="84" refId="ref10750" refString="Voet JE (1806) Catalogus systematicus coleopterorum, vol. 2. La Haye, 82 pp." type="book" year="1806">Voet, 1806</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="21.[140,1108,695,1672]" pageId="21" pageNumber="84">
<keyLead pageId="21" pageNumber="84">
– Head with vertex entire or convex. Tarsal claws usually appendiculate, if simple, see
<taxonomicName authorityName="Lacordaire" authorityYear="1848" box="[334,407,1259,1284]" class="Insecta" family="Chrysomelidae" genus="Exema" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis box="[334,407,1259,1284]" italics="true" pageId="21" pageNumber="84">Exema</emphasis>
</taxonomicName>
. Body less than
<quantity box="[592,675,1258,1284]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="84" unit="mm" value="10.0">10 mm</quantity>
long ...............................................
<emphasis bold="true" box="[1092,1107,1258,1284]" pageId="21" pageNumber="84">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="84">
<paragraph blockId="21.[140,1108,695,1672]" pageId="21" pageNumber="84">
<keyLead pageId="21" pageNumber="84">
5(4) Prosternal process distally widened between midcoxae (
<figureCitation box="[859,956,1293,1320]" captionStart="Figure 3" captionStartId="7.[140,212,1526,1548]" captionTargetBox="[148,1095,175,1500]" captionTargetId="figure@7.[136,1110,165,1507]" captionTargetPageId="7" captionText="Figure 3. Chlamisini, ventral view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770843" httpUri="https://zenodo.org/record/3770843/files/figure.png" pageId="21" pageNumber="84">Fig. 3 D</figureCitation>
). Metascutellum broadly exposed (
<figureCitation box="[495,586,1328,1355]" captionStart="Figure 1" captionStartId="4.[140,212,1526,1548]" captionTargetBox="[166,1087,177,1506]" captionTargetId="figure@4.[137,1109,165,1507]" captionTargetPageId="4" captionText="Figure 1. Chlamisini, dorsal view. A, Aulacochlamys distincta (Achard). B, A. costicollis (Lacordaire. C, Carcinobaena pilula (Klug). D, Chlamisus foveolatus (Knoch). E, Diplacaspis prosternalis (Schaeffer). F, Exema elliptica Karren. G, Fulcidax coelestina (Lacordaire). H, Hymetes javana Lacordaire. I, Melittochlamys specula (Klug). Į, Neochlamisus insularis (Schaeffer). K, N. velutinus Karren. L, Pseudochlamys megalostomoides Lacordaire ♀." figureDoi="http://doi.org/10.5281/zenodo.3788012" httpUri="https://zenodo.org/record/3788012/files/figure.png" pageId="21" pageNumber="84">Fig. 1 E</figureCitation>
) ........................
<taxonomicName authority="Jacobson, 1924" authorityName="Jacobson" authorityYear="1924" box="[779,1107,1328,1355]" class="Insecta" family="Chrysomelidae" genus="Diplacaspis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[779,1107,1328,1355]" pageId="21" pageNumber="84">
<emphasis bold="true" box="[779,917,1328,1355]" italics="true" pageId="21" pageNumber="84">Diplacaspis</emphasis>
<bibRefCitation author="Jacobson" box="[924,1107,1328,1354]" firstAuthor="Jacobson" pageId="21" pageNumber="84" pagination="237 - 244" refId="ref9947" refString="Jacobson G (1924) Annotationes synonymicae et systematicae de Coleopteris. Revue Russe d'Entomologie (Russkoe entomologicheskoe obozrenie) 18: 237 - 244." type="journal article" year="1924">Jacobson, 1924</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="21.[140,1108,695,1672]" pageId="21" pageNumber="84">
<keyLead pageId="21" pageNumber="84">
– Sides of prosternal process convergent or serrate, if slightly widened between mesocoxae, then metascutellum not exposed (
<figureCitation box="[761,854,1399,1425]" captionStart="Figure 1" captionStartId="4.[140,212,1526,1548]" captionTargetBox="[166,1087,177,1506]" captionTargetId="figure@4.[137,1109,165,1507]" captionTargetPageId="4" captionText="Figure 1. Chlamisini, dorsal view. A, Aulacochlamys distincta (Achard). B, A. costicollis (Lacordaire. C, Carcinobaena pilula (Klug). D, Chlamisus foveolatus (Knoch). E, Diplacaspis prosternalis (Schaeffer). F, Exema elliptica Karren. G, Fulcidax coelestina (Lacordaire). H, Hymetes javana Lacordaire. I, Melittochlamys specula (Klug). Į, Neochlamisus insularis (Schaeffer). K, N. velutinus Karren. L, Pseudochlamys megalostomoides Lacordaire ♀." figureDoi="http://doi.org/10.5281/zenodo.3788012" httpUri="https://zenodo.org/record/3788012/files/figure.png" pageId="21" pageNumber="84">Fig. 1 A</figureCitation>
) ..............................
<emphasis bold="true" box="[1092,1107,1398,1424]" pageId="21" pageNumber="84">6</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="84">
<paragraph blockId="21.[140,1108,695,1672]" pageId="21" pageNumber="84">
<keyLead pageId="21" pageNumber="84">
6(5) Pronotum with 6 distinct, small, sharp, longitudinal carinae that converge medially near posterior margin (fan). Small size, elongate, uniform hue, generally black (
<figureCitation box="[388,523,1504,1531]" captionStart="Figure 1" captionStartId="4.[140,212,1526,1548]" captionTargetBox="[166,1087,177,1506]" captionTargetId="figure@4.[137,1109,165,1507]" captionTargetPageId="4" captionText="Figure 1. Chlamisini, dorsal view. A, Aulacochlamys distincta (Achard). B, A. costicollis (Lacordaire. C, Carcinobaena pilula (Klug). D, Chlamisus foveolatus (Knoch). E, Diplacaspis prosternalis (Schaeffer). F, Exema elliptica Karren. G, Fulcidax coelestina (Lacordaire). H, Hymetes javana Lacordaire. I, Melittochlamys specula (Klug). Į, Neochlamisus insularis (Schaeffer). K, N. velutinus Karren. L, Pseudochlamys megalostomoides Lacordaire ♀." figureDoi="http://doi.org/10.5281/zenodo.3788012" httpUri="https://zenodo.org/record/3788012/files/figure.png" pageId="21" pageNumber="84">Figs. 1 A, B</figureCitation>
) ..............................
<taxonomicName authority="Monros, 1952" authorityName="Monros" authorityYear="1952" box="[757,1107,1503,1531]" class="Insecta" family="Chrysomelidae" genus="Aulacochlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[757,1107,1503,1531]" pageId="21" pageNumber="84">
<emphasis bold="true" box="[757,931,1504,1531]" italics="true" pageId="21" pageNumber="84">Aulacochlamys</emphasis>
<bibRefCitation author="Monros" box="[939,1107,1503,1530]" firstAuthor="Monros" pageId="21" pageNumber="84" pagination="489 - 672" refId="ref10435" refString="Monros F (1952) Revision de las especies argentinas de Chlamisinae (Col., Chrysomelidae) (5 a contribucion al conocimiento de Chlamisinae). Acta Zoologica Lilloana 10: 489 - 672." type="journal article" year="1952">Monrós, 1952</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="21.[140,1108,695,1672]" pageId="21" pageNumber="84">
<keyLead pageId="21" pageNumber="84">
– Pronotum with or without bumps, but never with 6 longitudinal carinae that converge medially near posterior margin.....................................................
<emphasis bold="true" box="[1092,1107,1575,1601]" pageId="21" pageNumber="84">7</emphasis>
</keyLead>
</paragraph>
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<keyStep lastPageId="22" lastPageNumber="85" pageId="21" pageNumber="84">
<paragraph blockId="21.[140,1108,695,1672]" lastBlockId="22.[140,1108,164,1038]" lastPageId="22" lastPageNumber="85" pageId="21" pageNumber="84">
<keyLead lastPageId="22" lastPageNumber="85" pageId="21" pageNumber="84">
7(6) Head not completely retracted into the prothorax; mandibles in males larger than in females (
<figureCitation box="[437,568,1645,1672]" captionStart="Figure 4" captionStartId="10.[140,212,1526,1548]" captionTargetBox="[140,1106,199,1486]" captionTargetId="figure@10.[140,1108,199,1506]" captionTargetPageId="10" captionText="Figure 4. Chlamisini, frontal view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). J, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770847" httpUri="https://zenodo.org/record/3770847/files/figure.png" pageId="21" pageNumber="84">Fig. 4 K, L</figureCitation>
). Prosternum strongly and abruptly constricted beyond anterior margin; prosternal process more than ¾ as long as prosternum. (
<figureCitation box="[322,457,202,228]" captionStart="Figure 3" captionStartId="7.[140,212,1526,1548]" captionTargetBox="[148,1095,175,1500]" captionTargetId="figure@7.[136,1110,165,1507]" captionTargetPageId="7" captionText="Figure 3. Chlamisini, ventral view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770843" httpUri="https://zenodo.org/record/3770843/files/figure.png" pageId="22" pageNumber="85">Figs. 3 K, L</figureCitation>
) .................................
<taxonomicName authority="Lacordaire, 1848" authorityName="Lacordaire" authorityYear="1848" box="[721,1107,201,228]" class="Insecta" family="Chrysomelidae" genus="Pseudochlamys" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[721,1107,201,228]" pageId="22" pageNumber="85">
<emphasis bold="true" box="[721,898,201,228]" italics="true" pageId="22" pageNumber="85">Pseudochlamys</emphasis>
<bibRefCitation author="Lacordaire" box="[905,1107,201,227]" firstAuthor="Lacordaire" pageId="22" pageNumber="85" refId="ref10277" refString="Lacordaire MT (1848) Monographie des Coleopteres subpentameres de la famille des Phytophages. Tome second. Bruxelles et Leipzig, 890 pp." type="book" year="1848">Lacordaire, 1848</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="22.[140,1108,164,1038]" pageId="22" pageNumber="85">
<keyLead pageId="22" pageNumber="85">
– Head completely retracted into prothorax; mandibles in males as large as in females (
<figureCitation box="[380,482,272,298]" captionStart="Figure 4" captionStartId="10.[140,212,1526,1548]" captionTargetBox="[140,1106,199,1486]" captionTargetId="figure@10.[140,1108,199,1506]" captionTargetPageId="10" captionText="Figure 4. Chlamisini, frontal view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). J, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770847" httpUri="https://zenodo.org/record/3770847/files/figure.png" pageId="22" pageNumber="85">Fig. 4 A</figureCitation>
). Prosternum gradually constricted at about 2/3 of its length .........................................................................................................
<emphasis bold="true" box="[1092,1107,307,333]" pageId="22" pageNumber="85">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="22" pageNumber="85">
<paragraph blockId="22.[140,1108,164,1038]" pageId="22" pageNumber="85">
<keyLead pageId="22" pageNumber="85">
8(7) Metasternum anteriorly elongated into deltoid projection (
<figureCitation box="[909,1007,343,369]" captionStart="Figure 3" captionStartId="7.[140,212,1526,1548]" captionTargetBox="[148,1095,175,1500]" captionTargetId="figure@7.[136,1110,165,1507]" captionTargetPageId="7" captionText="Figure 3. Chlamisini, ventral view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770843" httpUri="https://zenodo.org/record/3770843/files/figure.png" pageId="22" pageNumber="85">Fig. 3 G</figureCitation>
). Antennae serrate beyond antennomere 5 ......................
<taxonomicName authority="Lacordaire, 1848" authorityName="Lacordaire" authorityYear="1848" box="[798,1107,377,404]" class="Insecta" family="Chrysomelidae" genus="Hymetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[798,1107,377,404]" pageId="22" pageNumber="85">
<emphasis bold="true" box="[798,898,378,404]" italics="true" pageId="22" pageNumber="85">Hymetes</emphasis>
<bibRefCitation author="Lacordaire" box="[905,1107,377,403]" firstAuthor="Lacordaire" pageId="22" pageNumber="85" refId="ref10277" refString="Lacordaire MT (1848) Monographie des Coleopteres subpentameres de la famille des Phytophages. Tome second. Bruxelles et Leipzig, 890 pp." type="book" year="1848">Lacordaire, 1848</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="22.[140,1108,164,1038]" pageId="22" pageNumber="85">
<keyLead pageId="22" pageNumber="85">
– Metasternum anteriorly concave with two denticles on sides of concavity (
<figureCitation box="[254,348,448,475]" captionStart="Figure 3" captionStartId="7.[140,212,1526,1548]" captionTargetBox="[148,1095,175,1500]" captionTargetId="figure@7.[136,1110,165,1507]" captionTargetPageId="7" captionText="Figure 3. Chlamisini, ventral view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770843" httpUri="https://zenodo.org/record/3770843/files/figure.png" pageId="22" pageNumber="85">Fig. 3 B</figureCitation>
). ...................................................................................................
<emphasis bold="true" box="[1092,1107,448,474]" pageId="22" pageNumber="85">9</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="22" pageNumber="85">
<paragraph blockId="22.[140,1108,164,1038]" pageId="22" pageNumber="85">
<keyLead pageId="22" pageNumber="85">
9(8) Body usually metallic in color. Frons on canthus of eye usually with yellow spots, or if without yellow spots, then elytron with two velvety spots. Male ejaculatory guide asymmetrical, with sheath (
<figureCitation box="[754,847,554,580]" captionStart="Figure 7" captionStartId="19.[140,212,1068,1090]" captionTargetBox="[249,998,170,1037]" captionTargetId="figure@19.[235,1013,165,1048]" captionTargetPageId="19" captionText="Figure 7. Genitalia of Chlamisini. A, Male ejaculatory guide of Neochlamisus velutinus Karren, dorsal and lateral views. B, Male ejaculatory guide of Chlamisus maculipes (Chevrolat), dorsal and lateral views. C, Spermatheca of Neochlamisus velutinus. D, Spermatheca of Chlamisus maculipes." figureDoi="http://doi.org/10.5281/zenodo.3770851" httpUri="https://zenodo.org/record/3770851/files/figure.png" pageId="22" pageNumber="85">Fig. 7 A</figureCitation>
).................................. ........................................................................
<taxonomicName authority="Karren, 1972" authorityName="Karren" authorityYear="1972" box="[780,1107,588,615]" class="Insecta" family="Chrysomelidae" genus="Neochlamisus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[780,1107,588,615]" pageId="22" pageNumber="85">
<emphasis bold="true" box="[780,941,588,615]" italics="true" pageId="22" pageNumber="85">Neochlamisus</emphasis>
<bibRefCitation author="Karren" box="[948,1107,589,615]" firstAuthor="Karren" pageId="22" pageNumber="85" pagination="875 - 988" refId="ref10110" refString="Karren JB (1972) A revision of the subfamily Chlamisinae of America north of Mexico (Coleoptera: Chrysomelidae). The University of Kansas Science Bulletin 49: 875 - 988." type="journal article" year="1972">Karren, 1972</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="22.[140,1108,164,1038]" pageId="22" pageNumber="85">
<keyLead pageId="22" pageNumber="85">
– Body usually not metallic in color. Frons on canthus of eye usually without yellow spots (if face entirely or partly yellow, with yellow area extending onto canthus, elytra without velvety spots). Male ejaculatory guide symmetrical, without sheath (
<figureCitation box="[429,522,730,756]" captionStart="Figure 7" captionStartId="19.[140,212,1068,1090]" captionTargetBox="[249,998,170,1037]" captionTargetId="figure@19.[235,1013,165,1048]" captionTargetPageId="19" captionText="Figure 7. Genitalia of Chlamisini. A, Male ejaculatory guide of Neochlamisus velutinus Karren, dorsal and lateral views. B, Male ejaculatory guide of Chlamisus maculipes (Chevrolat), dorsal and lateral views. C, Spermatheca of Neochlamisus velutinus. D, Spermatheca of Chlamisus maculipes." figureDoi="http://doi.org/10.5281/zenodo.3770851" httpUri="https://zenodo.org/record/3770851/files/figure.png" pageId="22" pageNumber="85">Fig. 7 B</figureCitation>
) ..........................................................................
<emphasis bold="true" box="[1077,1107,729,755]" pageId="22" pageNumber="85">10</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="22" pageNumber="85">
<paragraph blockId="22.[140,1108,164,1038]" pageId="22" pageNumber="85">
<keyLead pageId="22" pageNumber="85">
10(9) Males without spines or spinulae on first ventrite. Fifth antennomere nearly as large as 6
<superScript attach="none" box="[388,402,799,813]" fontSize="6" pageId="22" pageNumber="85">th</superScript>
. Sutural serration of elytra usually incomplete (suture entire immediately following mesoscutellum) (
<figureCitation box="[711,819,835,862]" captionStart="Figure 1" captionStartId="4.[140,212,1526,1548]" captionTargetBox="[166,1087,177,1506]" captionTargetId="figure@4.[137,1109,165,1507]" captionTargetPageId="4" captionText="Figure 1. Chlamisini, dorsal view. A, Aulacochlamys distincta (Achard). B, A. costicollis (Lacordaire. C, Carcinobaena pilula (Klug). D, Chlamisus foveolatus (Knoch). E, Diplacaspis prosternalis (Schaeffer). F, Exema elliptica Karren. G, Fulcidax coelestina (Lacordaire). H, Hymetes javana Lacordaire. I, Melittochlamys specula (Klug). Į, Neochlamisus insularis (Schaeffer). K, N. velutinus Karren. L, Pseudochlamys megalostomoides Lacordaire ♀." figureDoi="http://doi.org/10.5281/zenodo.3788012" httpUri="https://zenodo.org/record/3788012/files/figure.png" pageId="22" pageNumber="85">Fig. 1 D</figureCitation>
). Prosternum posteriorly pointed (narrowed), posteriorly much narrower than anterior margin........... .......................................................................
<taxonomicName authority="Rafinesque, 1815" authorityName="Rafinesque" authorityYear="1815" box="[769,1107,905,932]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[769,1107,905,932]" pageId="22" pageNumber="85">
<emphasis bold="true" box="[769,893,905,932]" italics="true" pageId="22" pageNumber="85">Chlamisus</emphasis>
<bibRefCitation author="Rafinesque" box="[900,1107,905,931]" firstAuthor="Rafinesque" pageId="22" pageNumber="85" refId="ref10566" refString="Rafinesque CS (1815) Analyse de la nature ou tableau de l'univers et des corps organises. Palermo, 224 pp." type="book" year="1815">Rafinesque, 1815</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="22.[140,1108,164,1038]" pageId="22" pageNumber="85">
<keyLead pageId="22" pageNumber="85">
– Males with spines or spinulae on first ventrite. Fifth antennomere much smaller than 6
<superScript attach="none" box="[412,426,975,989]" fontSize="6" pageId="22" pageNumber="85">th</superScript>
. Sutural serration of elytra complete (
<figureCitation box="[870,968,976,1003]" captionStart="Figure 1" captionStartId="4.[140,212,1526,1548]" captionTargetBox="[166,1087,177,1506]" captionTargetId="figure@4.[137,1109,165,1507]" captionTargetPageId="4" captionText="Figure 1. Chlamisini, dorsal view. A, Aulacochlamys distincta (Achard). B, A. costicollis (Lacordaire. C, Carcinobaena pilula (Klug). D, Chlamisus foveolatus (Knoch). E, Diplacaspis prosternalis (Schaeffer). F, Exema elliptica Karren. G, Fulcidax coelestina (Lacordaire). H, Hymetes javana Lacordaire. I, Melittochlamys specula (Klug). Į, Neochlamisus insularis (Schaeffer). K, N. velutinus Karren. L, Pseudochlamys megalostomoides Lacordaire ♀." figureDoi="http://doi.org/10.5281/zenodo.3788012" httpUri="https://zenodo.org/record/3788012/files/figure.png" pageId="22" pageNumber="85">Fig. 1 F</figureCitation>
). Pronotum without longitudinal carinae.................................
<taxonomicName authority="Lacordaire, 1848" authorityName="Lacordaire" authorityYear="1848" box="[818,1107,1011,1038]" class="Insecta" family="Chrysomelidae" genus="Exema" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[818,1107,1011,1038]" pageId="22" pageNumber="85">
<emphasis bold="true" box="[818,898,1012,1038]" italics="true" pageId="22" pageNumber="85">Exema</emphasis>
<bibRefCitation author="Lacordaire" box="[905,1107,1011,1037]" firstAuthor="Lacordaire" pageId="22" pageNumber="85" refId="ref10277" refString="Lacordaire MT (1848) Monographie des Coleopteres subpentameres de la famille des Phytophages. Tome second. Bruxelles et Leipzig, 890 pp." type="book" year="1848">Lacordaire, 1848</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>
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