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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.239.3734" ID-GBIF-Dataset="58b3b015-a919-456b-9175-f96878341e31" ID-PMC="PMC3497256" ID-Pensoft-Pub="1313-2970-239-1" ID-PubMed="23226711" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-239-1" ModsDocOrigin="ZooKeys 239" ModsDocTitle="Polyaxone monaxonids: revision of raspailiid sponges with polyactine megascleres ( Cyamon and Trikentrion)" checkinTime="1451248578713" checkinUser="pensoft" docAuthor="Soest, Rob van, Carballo, Jose Luis &amp; Hooper, John" docDate="2012" docId="2203E72F032E81B6D7151497FD5EF7DC" docLanguage="en" docName="ZooKeys 239: 1-70" docOrigin="ZooKeys 239" docSource="http://dx.doi.org/10.3897/zookeys.239.3734" docTitle="Trikentrion catalina Sim &amp; Bakus 1986" docType="treatment" docVersion="3" lastPageNumber="46" masterDocId="3639FF9B0F54FFEACF6A573ADC3CDF3D" masterDocTitle="Polyaxone monaxonids: revision of raspailiid sponges with polyactine megascleres (Cyamon and Trikentrion)" masterLastPageNumber="70" masterPageNumber="1" pageNumber="44" updateTime="1668154678110" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Polyaxone monaxonids: revision of raspailiid sponges with polyactine megascleres (Cyamon and Trikentrion)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Soest, Rob van</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Carballo, Jose Luis</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hooper, John</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>239</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>70</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.239.3734</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.239.3734</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-239-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152037784" LSID="urn:lsid:plazi:treatment:2203E72F032E81B6D7151497FD5EF7DC" httpUri="http://treatment.plazi.org/id/2203E72F032E81B6D7151497FD5EF7DC" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44">
<subSubSection pageId="43" pageNumber="44" type="nomenclature">
<paragraph pageId="43" pageNumber="44">
<taxonomicName LSID="http://species-id.net/wiki/Trikentrion_catalina" authority="Sim &amp; Bakus, 1986" authorityName="Sim &amp; Bakus" authorityYear="1986" class="Demospongiae" family="Raspailiidae" genus="Trikentrion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trikentrion catalina" order="Axinellida" pageId="43" pageNumber="44" phylum="Porifera" rank="species" species="catalina">Trikentrion catalina (Sim &amp; Bakus, 1986)</taxonomicName>
Figs 25
<normalizedToken originalValue="AF">A-F</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="reference_group">
<paragraph pageId="43" pageNumber="44">
<taxonomicName class="Demospongiae" family="Raspailiidae" genus="Cyamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyamon catalina" order="Axinellida" pageId="43" pageNumber="44" phylum="Porifera" rank="species" species="catalina">Cyamon catalina</taxonomicName>
<bibRefCitation author="Sim, CJ" journalOrPublisher="Occasional Papers of the Allan Hancock Foundation" pageId="51" pageNumber="52" pagination="1 - 23" title="Marine sponges of Santa Catalina Island, California." volume="5" year="1986">Sim and Bakus 1986</bibRefCitation>
: 18, fig. 4;
<bibRefCitation pageId="43" pageNumber="44">Lee et al. 2007</bibRefCitation>
: 211 (California).
</paragraph>
<paragraph pageId="43" pageNumber="44">
<taxonomicName lsidName="Not (Trikentrion) catalina" pageId="43" pageNumber="44" rank="species" species="catalina" subGenus="Trikentrion">Not: Trikentrion catalina</taxonomicName>
;
<bibRefCitation author="Gomez, P" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="51" pageNumber="52" pagination="223 - 237" title="New records for the sponge fauna (Porifea: Demospongiae) of the Pacific coast of Mexico (eastern Pacific Ocean)." volume="115" year="2002">
<normalizedToken originalValue="Gómez">Gomez</normalizedToken>
et al. 2002
</bibRefCitation>
=
<taxonomicName class="Demospongiae" family="Raspailiidae" genus="Trikentrion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trikentrion helium" order="Axinellida" pageId="43" pageNumber="44" phylum="Porifera" rank="species" species="helium">Trikentrion helium</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="material examined">
<paragraph pageId="43" pageNumber="44">Material examined.</paragraph>
<paragraph pageId="43" pageNumber="44">
Holotype USNM 33631, preserved in alcohol, California, Santa Catalina Island, Bird Rock,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="33.45">33.45°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-118.4833">118.4833°W</geoCoordinate>
, on rocky cliff at 50 m depth, coll. K. McCleneghan.
</paragraph>
<paragraph pageId="43" pageNumber="44">Not examined: paratype BMNH 1985 (reg. nr. unknown), Santa Catalina Island, Ship Rock, on rock at 46 m depth, coll. R. Given.</paragraph>
</subSubSection>
<subSubSection lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44" type="description">
<paragraph pageId="43" pageNumber="44">Description.</paragraph>
<paragraph pageId="43" pageNumber="44">
Flabelliform sponge (Fig. 25A), measuring 15
<normalizedToken originalValue="×">x</normalizedToken>
8 by 0.4 cm, attached to rocks by a 3
<normalizedToken originalValue="×">x</normalizedToken>
0.6 cm stalk. Surface hispid. No oscules apparent. Consistency firm and leathery. Color reddish orange (alive), pale beige (alcohol).
</paragraph>
<paragraph pageId="43" pageNumber="44">Skeleton (Fig. 25B): choanosome densely packed with three-claded polyactines; ectosome with long, relatively thick styles surrounded by dense bouquets of short thin styles; trichodragmata commonly observed especially in the peripheral parts.</paragraph>
<paragraph pageId="43" pageNumber="44">Spicules: long (thin) styles, short thin styles, polyactines, trichodragmas.</paragraph>
<paragraph pageId="43" pageNumber="44">
Long (thin) styles (Fig. 25C), usually broken and only a few could be measured: 1400-5400
<normalizedToken originalValue="×">x</normalizedToken>
8-40
<normalizedToken originalValue="µm">µm</normalizedToken>
, so not really thin.
</paragraph>
<paragraph pageId="44" pageNumber="45">
<pageBreakToken pageId="44" pageNumber="45" start="start">Short</pageBreakToken>
thin styles (Fig. 25D), 130
<normalizedToken originalValue="611.3">-611.3-</normalizedToken>
730
<normalizedToken originalValue="×">x</normalizedToken>
3
<normalizedToken originalValue="5.6">-5.6-</normalizedToken>
8
<normalizedToken originalValue="µm">µm</normalizedToken>
,
</paragraph>
<paragraph pageId="44" pageNumber="45">
Polyactines (Figs 25E, E1), predominantly three-claded, with spined shorter basal cladi (Fig. 25E1), occasionally with few or no spines on the basal cladi, and smooth, longer, relatively pointed lateral cladi; occasionally four-claded; size basal cladi 78
<normalizedToken originalValue="98.7">-98.7-</normalizedToken>
126
<normalizedToken originalValue="×">x</normalizedToken>
16
<normalizedToken originalValue="25.3">-25.3-</normalizedToken>
31
<normalizedToken originalValue="µm">µm</normalizedToken>
, lateral cladi 156
<normalizedToken originalValue="197.7">-197.7-</normalizedToken>
236
<normalizedToken originalValue="×">x</normalizedToken>
18
<normalizedToken originalValue="24.4">-24.4-</normalizedToken>
29
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
<paragraph pageId="45" pageNumber="46">
<pageBreakToken pageId="45" pageNumber="46" start="start">Trichodragmas</pageBreakToken>
: straight, with lightly spined raphides (Figs 25F, F1), 63
<normalizedToken originalValue="79.3">-79.3-</normalizedToken>
88
<normalizedToken originalValue="×">x</normalizedToken>
7
<normalizedToken originalValue="10.2">-10.2-</normalizedToken>
13
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="distribution">
<paragraph pageId="45" pageNumber="46">Distribution.</paragraph>
<paragraph pageId="45" pageNumber="46">Santa Catalina Island, Southern California.</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="ecology">
<paragraph pageId="45" pageNumber="46">Ecology.</paragraph>
<paragraph pageId="45" pageNumber="46">On rocks, from 46-50 m depth.</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="discussion">
<paragraph pageId="45" pageNumber="46">Discussion.</paragraph>
<paragraph pageId="45" pageNumber="46">
This species is assigned to
<taxonomicName class="Demospongiae" family="Raspailiidae" genus="Trikentrion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trikentrion" order="Axinellida" pageId="45" pageNumber="46" phylum="Porifera" rank="genus">Trikentrion</taxonomicName>
, because of the flabellate shape resembling
<taxonomicName class="Demospongiae" family="Raspailiidae" genus="Trikentrion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trikentrion flabelliforme" order="Axinellida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="flabelliforme">Trikentrion flabelliforme</taxonomicName>
<bibRefCitation author="Hentschel, E" journalOrPublisher="Abhandlungen herausgegeben von der Senckenbergischen naturforschenden Gesellschaft" pageId="51" pageNumber="52" pagination="293 - 448" title="Kiesel- und Hornschwaemme der Aru- und Kei-Inseln." volume="34" year="1912">Hentschel (1912)</bibRefCitation>
, the sagittal polyactines, three-claded with spines only on the basal cladus, and the possession of trichodragmas. It is similar to
<taxonomicName class="Demospongiae" family="Raspailiidae" genus="Trikentrion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trikentrion helium" order="Axinellida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="helium">Trikentrion helium</taxonomicName>
in the lack of genuine choanosomal oxeas, and the short basal cladi of the polyactines. Remarkably, when describing
<taxonomicName class="Demospongiae" family="Raspailiidae" genus="Trikentrion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trikentrion catalina" order="Axinellida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="catalina">Trikentrion catalina</taxonomicName>
, Sim &amp; Bakus (1986) did not notice - they did not discuss
<taxonomicName class="Demospongiae" family="Raspailiidae" genus="Trikentrion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trikentrion helium" order="Axinellida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="helium">Trikentrion helium</taxonomicName>
- the similarities with their species. Specimens assigned to
<taxonomicName class="Demospongiae" family="Raspailiidae" genus="Trikentrion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trikentrion catalina" order="Axinellida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="catalina">Trikentrion catalina</taxonomicName>
by
<bibRefCitation author="Gomez, P" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="51" pageNumber="52" pagination="223 - 237" title="New records for the sponge fauna (Porifea: Demospongiae) of the Pacific coast of Mexico (eastern Pacific Ocean)." volume="115" year="2002">
<normalizedToken originalValue="Gómez">Gomez</normalizedToken>
et al. (2002)
</bibRefCitation>
are considered to be long to
<taxonomicName class="Demospongiae" family="Raspailiidae" genus="Trikentrion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trikentrion helium" order="Axinellida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="helium">Trikentrion helium</taxonomicName>
(see above).
</paragraph>
<caption pageId="45" pageNumber="46">
<paragraph pageId="45" pageNumber="46">
Figure 25.
<taxonomicName class="Demospongiae" family="Raspailiidae" genus="Trikentrion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trikentrion catalina" order="Axinellida" pageId="45" pageNumber="46" phylum="Porifera" rank="species" species="catalina">Trikentrion catalina</taxonomicName>
(Sim &amp; Bakus, 1986), holotype USNM 33631, A shape of holotype specimen (scale bar = 1 cm) B cross section of skeleton C detail of head of long thin style D details of short thin style E three- and four-claded polyactines E1 detail of spined basal clade of polyactine F raphide F1 detail of raphide showing spination.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>