treatments-xml/data/EB/1A/9B/EB1A9B52FFA4FFAD710A51DFFC93FDE3.xml
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<document id="8DBC5B8500295D82AF9D0064261081C2" ID-DOI="10.1080/00222933.2019.1596328" ID-GBIF-Dataset="31e9dec8-ac91-43e6-aef3-c43fb0ed82ad" ID-ISSN="1464-5262" ID-Zenodo-Dep="3675703" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582134628034" checkinUser="valdenar" docAuthor="Molina-Acevedo, Isabel C." docDate="2019" docId="EB1A9B52FFA4FFAD710A51DFFC93FDE3" docLanguage="en" docName="JNATHIST.53.910.517540.pdf.imf" docOrigin="Journal of Natural History 53 (9)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Treadwellphysa dartevellei Molina-Acevedo 2019" docType="treatment" docVersion="11" lastPageNumber="525" masterDocId="1723E32AFFA1FFA470405151FF2DFFA4" masterDocTitle="Redescription of two species previously regarded as Marphysa de Quatrefagesı 1865 ı with the description of new species of Treadwellphysa Molina-Acevedo and Carrera-Parraı 2017" masterLastPageNumber="540" masterPageNumber="517" pageNumber="521" updateTime="1698803554584" updateUser="ExternalLinkService">
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<mods:title id="F8120A5B86D7F2060D0CC125072F0BC4">Redescription of two species previously regarded as Marphysa de Quatrefagesı 1865 ı with the description of new species of Treadwellphysa Molina-Acevedo and Carrera-Parraı 2017</mods:title>
</mods:titleInfo>
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<mods:namePart id="E513FFA94EB604DE81842458AFFFC3EC">Molina-Acevedo, Isabel C.</mods:namePart>
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<mods:title id="4498CF22AF5574CA34FF4BBA393EB221">Journal of Natural History</mods:title>
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<mods:date id="54D9ED948EDBDD25E9DE749C758E4B4A">2019</mods:date>
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<mods:number id="7F37F09DB903D3A09447D9EFE961AD0B">2019-04-30</mods:number>
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<mods:number id="60EF729DA253EF52EFAD878710EC0079">53</mods:number>
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<treatment id="EB1A9B52FFA4FFAD710A51DFFC93FDE3" ID-DOI="http://doi.org/10.5281/zenodo.3680422" ID-GBIF-Taxon="162266730" ID-Zenodo-Dep="3680422" LSID="urn:lsid:plazi:treatment:EB1A9B52FFA4FFAD710A51DFFC93FDE3" httpUri="http://treatment.plazi.org/id/EB1A9B52FFA4FFAD710A51DFFC93FDE3" lastPageId="9" lastPageNumber="525" pageId="5" pageNumber="521">
<subSubSection id="2BA979CFFFA4FFA1710A51DFFDF3FF6E" pageId="5" pageNumber="521" type="key">
<subSubSection id="2BA979CFFFA4FFA1710A51DFFCF7FF03" box="[330,986,142,167]" pageId="5" pageNumber="521" type="nomenclature">
<paragraph id="630C2A44FFA4FFA1710A51DFFCF7FF03" blockId="5.[330,986,142,202]" box="[330,986,142,167]" pageId="5" pageNumber="521">
<heading id="38449D28FFA4FFA1710A51DFFCFBFF03" box="[330,982,142,167]" pageId="5" pageNumber="521">
<taxonomicName id="A4B351C7FFA4FFA1710A51DFFC65FF03" ID-CoL="85S5Q" authority="(Monroi 1936)" authorityName="Molina-Acevedo" authorityYear="2019" baseAuthorityName="Monroi" baseAuthorityYear="1936" box="[330,840,142,167]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="521" phylum="Annelida" rank="species" species="dartevellei">
<emphasis id="51C7F656FFA4FFA1710A51DFFDBBFF03" bold="true" box="[330,662,142,167]" italics="true" pageId="5" pageNumber="521">Treadwellphysa dartevellei</emphasis>
(Monro, 1936)
</taxonomicName>
, comb. nov
</heading>
.
</paragraph>
</subSubSection>
<subSection id="0A3C33AFFFA4FFA1720551E0FDF3FF6E" box="[581,734,177,202]" pageId="5" pageNumber="521" type="multiple">
<subSubSection id="2BA979CFFFA4FFA1720551E0FDF3FF6E" box="[581,734,177,202]" pageId="5" pageNumber="521" type="description">
<paragraph id="630C2A44FFA4FFA1720551E0FDF3FF6E" blockId="5.[330,986,142,202]" box="[581,734,177,202]" pageId="5" pageNumber="521">
<heading id="38449D28FFA4FFA1720551E0FDF3FF6E" box="[581,734,177,202]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="521" reason="5">
(
<figureCitation id="FB8836C1FFA4FFA1720D51E0FDFBFF6E" box="[589,726,177,202]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="6.[160,225,1420,1443]" captionStartId-1="7.[160,225,1307,1330]" captionTargetBox-0="[162,1153,143,1382]" captionTargetBox-1="[164,1151,143,1274]" captionTargetId-0="figure@6.[161,1154,142,1389]" captionTargetId-1="figure@7.[163,1153,142,1275]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." captionText-1="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi-0="http://doi.org/10.5281/zenodo.3675705" figureDoi-1="http://doi.org/10.5281/zenodo.3675707" httpUri-0="https://zenodo.org/record/3675705/files/figure.png" httpUri-1="https://zenodo.org/record/3675707/files/figure.png" pageId="5" pageNumber="521">
Figures 1
<emphasis id="51C7F656FFA4FFA172FA51E0FDE5FF6E" box="[698,712,177,202]" italics="true" pageId="5" pageNumber="521"></emphasis>
2
</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
</subSection>
</subSubSection>
<subSubSection id="2BA979CFFFA4FFA170E051B7FEE5FE84" pageId="5" pageNumber="521" type="reference_group">
<paragraph id="630C2A44FFA4FFA170E051B7FEE5FE84" blockId="5.[160,1088,230,288]" pageId="5" pageNumber="521">
<treatmentCitationGroup id="43A30D6AFFA4FFA170E051B7FEE5FE84" pageId="5" pageNumber="521">
<treatmentCitation id="E2120C55FFA4FFA170E051B7FDC2FF5A" box="[160,751,230,254]" pageId="5" pageNumber="521">
<taxonomicName id="A4B351C7FFA4FFA170E051B7FDA1FF5A" ID-CoL="855GV" authority="Monroi 1936: 246 - 248" authorityName="Monroi" authorityPageNumber="246 - 248" authorityYear="1936" box="[160,652,230,254]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="521" phylum="Annelida" rank="species" species="dartevellei">
<emphasis id="51C7F656FFA4FFA170E051B7FEAEFF5A" box="[160,387,230,254]" italics="true" pageId="5" pageNumber="521">Marphysa dartevellei</emphasis>
Monro, 1936: 246
<emphasis id="51C7F656FFA4FFA1721351B7FD4DFF5A" box="[595,608,230,254]" italics="true" pageId="5" pageNumber="521"></emphasis>
248
</taxonomicName>
, figs a
<emphasis id="51C7F656FFA4FFA1729451B7FDCCFF5A" box="[724,737,230,254]" italics="true" pageId="5" pageNumber="521"></emphasis>
f;
</treatmentCitation>
<treatmentCitation id="E2120C55FFA4FFA172B851B7FB11FF5A" box="[760,1084,230,254]" pageId="5" pageNumber="521">
<bibRefCitation id="072257B5FFA4FFA172B851B7FB11FF5A" author="Fauvel Pi Rullier F" box="[760,1084,230,254]" pageId="5" pageNumber="521" pagination="143 - 205" refId="ref10923" refString="Fauvel Pi Rullier F. 1959. Annelides Polychetes. Resultats scientifiques des Campagnes de Calypso 4. Ann Inst Oceanogr. 37: 143 - 205." type="journal article" year="1959">Fauvel and Rullier 1959: 169</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="E2120C55FFA4FFA170825059FEE5FE84" box="[194,456,264,288]" pageId="5" pageNumber="521">
<bibRefCitation id="072257B5FFA4FFA170825059FEE5FE84" author="Amoureux L" box="[194,456,264,288]" pageId="5" pageNumber="521" pagination="41 - 65" refId="ref10646" refString="Amoureux L. 1973. Quelques annelides polychetes de l ' Afrique occidentale et equatoriale. Cah Off Rech Sci Tech Outre-Meri Ser Oceanogr. 11 (1): 41 - 65." type="journal article" year="1973">
Amoureux 1973: 57
<emphasis id="51C7F656FFA4FFA171E05059FE80FE84" box="[416,429,264,288]" italics="true" pageId="5" pageNumber="521"></emphasis>
58
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</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFA4FFA170E05002FD12FD23" pageId="5" pageNumber="521" type="materials_examined">
<paragraph id="630C2A44FFA4FFA170E05002FEADFEC8" blockId="5.[160,1156,339,468]" box="[160,384,339,364]" pageId="5" pageNumber="521">
<emphasis id="51C7F656FFA4FFA170E05002FEADFEC8" bold="true" box="[160,384,339,364]" italics="true" pageId="5" pageNumber="521">Material examined</emphasis>
</paragraph>
<paragraph id="630C2A44FFA4FFA170E05024FD35FE70" blockId="5.[160,1156,339,468]" pageId="5" pageNumber="521">
<emphasis id="51C7F656FFA4FFA170E05024FE7FFE2A" bold="true" box="[160,338,373,398]" italics="true" pageId="5" pageNumber="521">Type material.</emphasis>
<materialsCitation id="D3DB2019FFA4FFA171225027FD35FE70" ID-GBIF-Occurrence="2570076585" collectionCode="BMNH" location="Africa" municipality="Angola" pageId="5" pageNumber="521" specimenCount="1" stateProvince="Cape" typeStatus="holotype">
<typeStatus id="BC0894E6FFA4FFA171225027FEE4FE2A" box="[354,457,374,398]" pageId="5" pageNumber="521" type="holotype">Holotype</typeStatus>
British Museum of Natural History (
<collectionCode id="05A2B281FFA4FFA173C65027FCFDFE2A" box="[902,976,374,398]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="5" pageNumber="521">BMNH</collectionCode>
) 1935.1.29.1
<emphasis id="51C7F656FFA4FFA174235027FB5DFE2A" box="[1123,1136,374,398]" italics="true" pageId="5" pageNumber="521"></emphasis>
2;
<quantity id="A44B87A1FFA4FFA170E050C8FFC4FE15" box="[160,233,409,433]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="521" unit="km" value="85.0">
<locationDeviation id="72BE3366FFA4FFA170E050C8FFC4FE15" box="[160,233,409,433]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="521" unit="km" value="85.0">85 km</locationDeviation>
</quantity>
. North of Banana,
<collectingRegion id="A177E4A6FFA4FFA1718050C8FED4FE15" box="[448,505,409,433]" country="South Africa" name="Western Cape" pageId="5" pageNumber="521">Cape</collectingRegion>
Malembo,
<collectingRegion id="A177E4A6FFA4FFA1723A50C8FDF4FE15" box="[634,729,409,433]" country="Angola" name="Cabinda" pageId="5" pageNumber="521">Cabinda</collectingRegion>
,
<collectingMunicipality id="8368B03EFFA4FFA172A550C8FC1AFE15" box="[741,823,409,433]" pageId="5" pageNumber="521">Angola</collectingMunicipality>
,
<location id="666C7C9FFFA4FFA1730450C8FCABFE15" LSID="urn:lsid:plazi:treatment:EB1A9B52FFA4FFAD710A51DFFC93FDE3:666C7C9FFFA4FFA1730450C8FCABFE15" box="[836,902,409,433]" municipality="Angola" name="Africa" pageId="5" pageNumber="521" stateProvince="Cape">Africa</location>
,
<location id="666C7C9FFFA4FFA173D250C8FB16FE15" LSID="urn:lsid:plazi:treatment:EB1A9B52FFA4FFAD710A51DFFC93FDE3:666C7C9FFFA4FFA173D250C8FB16FE15" box="[914,1083,409,433]" municipality="Angola" name="Atlantic Ocean" pageId="5" pageNumber="521" stateProvince="Cape">Atlantic Ocean</location>
; 1933; on rock bored by
<taxonomicName id="A4B351C7FFA4FFA1712F50EDFEC2FE70" box="[367,495,444,468]" class="Bivalvia" family="Mytilidae" genus="Lithodomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Mytilida" pageId="5" pageNumber="521" phylum="Mollusca" rank="genus">
<emphasis id="51C7F656FFA4FFA1712F50EDFEC2FE70" box="[367,495,444,468]" italics="true" pageId="5" pageNumber="521">
<location id="666C7C9FFFA4FFA1712F50EDFEC2FE70" LSID="urn:lsid:plazi:treatment:EB1A9B52FFA4FFAD710A51DFFC93FDE3:666C7C9FFFA4FFA1712F50EDFEC2FE70" box="[367,495,444,468]" municipality="Angola" name="Lithodomus" pageId="5" pageNumber="521" stateProvince="Cape">Lithodomus</location>
</emphasis>
</taxonomicName>
<specimenCount id="75B5E1CDFFA4FFA171B850EDFD35FE70" box="[504,536,444,468]" pageId="5" pageNumber="521" type="generic">sp.</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="630C2A44FFA4FFA170E05357FD12FD23" blockId="5.[160,1155,518,647]" pageId="5" pageNumber="521">
<emphasis id="51C7F656FFA4FFA170E05357FEBDFDBB" bold="true" box="[160,400,518,543]" italics="true" pageId="5" pageNumber="521">Additional material.</emphasis>
<materialsCitation id="D3DB2019FFA4FFA171E05356FD12FD23" ID-GBIF-Occurrence="2570076584" collectingDate="1964-09-18" collectionCode="BMNH, LACM-AHF" country="Angola" location="Africa" municipality="Ghana" pageId="5" pageNumber="521" specimenCode="BMNH 2013233, POLY 11098" specimenCount="1" stateProvince="Greater Accra Region">
<collectionCode id="05A2B281FFA4FFA171E05356FECAFDBB" box="[416,487,519,543]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="5" pageNumber="521">BMNH</collectionCode>
1939.7.17.23
<emphasis id="51C7F656FFA4FFA1723C5357FDA4FDBA" box="[636,649,518,542]" italics="true" pageId="5" pageNumber="521"></emphasis>
24 (3); Cacongo (formerly Landana),
<collectingCountry id="1BA46AD4FFA4FFA1746E5356FBADFDBB" box="[1070,1152,519,543]" name="Angola" pageId="5" pageNumber="521">Angola</collectingCountry>
, Africa.
<specimenCode id="3315823FFFA4FFA170AD5378FEB6FDE5" box="[237,411,553,577]" pageId="5" pageNumber="521">BMNH 2013233</specimenCode>
(1);
<collectingCountry id="1BA46AD4FFA4FFA1718C5378FD3AFDE5" box="[460,535,553,577]" name="Ghana" pageId="5" pageNumber="521">Ghana</collectingCountry>
, Africa.
<collectionCode id="05A2B281FFA4FFA1722E5378FDCAFDE5" box="[622,743,553,577]" pageId="5" pageNumber="521">LACM-AHF</collectionCode>
<specimenCode id="3315823FFFA4FFA172AE5378FC58FDE5" box="[750,885,553,577]" pageId="5" pageNumber="521">POLY 11098</specimenCode>
(3) Paradise beach, Ada Foah, east of Pranpram,
<collectingRegion id="A177E4A6FFA4FFA1718C531DFDE7FDC0" box="[460,714,588,612]" country="Ghana" name="Greater Accra" pageId="5" pageNumber="521">Greater Accra Region</collectingRegion>
;
<collectingMunicipality id="8368B03EFFA4FFA1729B531DFC0BFDC0" box="[731,806,588,612]" pageId="5" pageNumber="521">Ghana</collectingMunicipality>
,
<location id="666C7C9FFFA4FFA17377531DFC54FDC0" LSID="urn:lsid:plazi:treatment:EB1A9B52FFA4FFAD710A51DFFC93FDE3:666C7C9FFFA4FFA17377531DFC54FDC0" box="[823,889,588,612]" country="Angola" municipality="Ghana" name="Africa" pageId="5" pageNumber="521" stateProvince="Greater Accra Region">Africa</location>
;
<geoCoordinate id="06874C83FFA4FFA173CA531DFCCCFDC0" box="[906,993,588,612]" degrees="5.776" direction="north" orientation="latitude" pageId="5" pageNumber="521" value="5.776">5.776°N</geoCoordinate>
<geoCoordinate id="06874C83FFA4FFA173B0531DFB7BFDC0" box="[1008,1110,588,612]" degrees="-0.647" direction="west" orientation="longitude" pageId="5" pageNumber="521" value="-0.647">-0.647°W</geoCoordinate>
;
<date id="170D0C84FFA4FFA17428531DFE70FD23" pageId="5" pageNumber="521" value="1964-09-18">
<collectingDate id="0749F56CFFA4FFA17428531DFE70FD23" pageId="5" pageNumber="521" value="1964-09-18">18 September 1964</collectingDate>
</date>
; coll.
<location id="666C7C9FFFA4FFA171DF533EFD16FD23" LSID="urn:lsid:plazi:treatment:EB1A9B52FFA4FFAD710A51DFFC93FDE3:666C7C9FFFA4FFA171DF533EFD16FD23" box="[415,571,623,647]" country="Angola" municipality="Ghana" name="Bjorn Berland" pageId="5" pageNumber="521" stateProvince="Greater Accra Region">Björn Berland</location>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFA4FFAC70E053E8FED3FB7D" lastPageId="8" lastPageNumber="524" pageId="5" pageNumber="521" type="description">
<paragraph id="630C2A44FFA4FFA170E053E8FE0BFD76" blockId="5.[160,1157,697,1727]" box="[160,294,697,722]" pageId="5" pageNumber="521">
<emphasis id="51C7F656FFA4FFA170E053E8FE0BFD76" bold="true" box="[160,294,697,722]" italics="true" pageId="5" pageNumber="521">Description</emphasis>
</paragraph>
<paragraph id="630C2A44FFA4FFA170E0538DFEF5FCF8" blockId="5.[160,1157,697,1727]" pageId="5" pageNumber="521">
<typeStatus id="BC0894E6FFA4FFA170E0538DFE2AFD50" box="[160,263,732,756]" pageId="5" pageNumber="521" type="holotype">Holotype</typeStatus>
incomplete, laterally dissected, with 105 chaetigers, L10 = 10.2 mm, W10 = 5 mm, TL =
<quantity id="A44B87A1FFA4FFA171DA53AEFEDCFCB3" box="[410,497,767,791]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="5" pageNumber="521" unit="mm" value="66.0">66 mm</quantity>
. Anterior region of body with convex dorsum and flat ventrum, without groove; body depressed from chaetiger 10, widest at chaetiger 33, tapering after chaetiger 45.
</paragraph>
<paragraph id="630C2A44FFA4FFA170805236FEFAFBD6" blockId="5.[160,1157,697,1727]" pageId="5" pageNumber="521">
Prostomium bilobed,
<quantity id="A44B87A1FFA4FFA171FF5236FD34FCDB" box="[447,537,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="521" unit="mm" value="1.8">1.8 mm</quantity>
long,
<quantity id="A44B87A1FFA4FFA1722B5236FD9FFCDB" box="[619,690,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="521" unit="mm" value="3.0">3 mm</quantity>
wide; lobes frontally rounded; median sulcus shallow (
<figureCitation id="FB8836C1FFA4FFA1711352DBFE86FC06" box="[339,427,906,930]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(a
<emphasis id="51C7F656FFA4FFA1718452DBFEFCFC06" box="[452,465,906,930]" italics="true" pageId="5" pageNumber="521"></emphasis>
c)), ventral sulcus deep (
<figureCitation id="FB8836C1FFA4FFA172A852DBFC6FFC06" box="[744,834,906,930]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(b)). Prostomial appendages in a semicircle, median antenna isolated by a gap. Palps reaching second peristomial ring; lateral antennae reaching middle of first chaetiger; median antenna broken, in additional specimen (LACM-AHF
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, L10 =
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) reaching middle of second chaetiger. Palpophores and ceratophores ring-shaped, short, thick; palpostyles and ceratostyles tapering, slender, without articulation. Eyes rounded, brown, between palps and lateral antennae.
</paragraph>
<paragraph id="630C2A44FFA4FFA17080552DFBA9FB7E" blockId="5.[160,1157,697,1727]" pageId="5" pageNumber="521">
Peristomium larger than prostomium (
<quantity id="A44B87A1FFA4FFA17237552DFDE2FB30" box="[631,719,1148,1172]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="521" unit="mm" value="2.1">2.1 mm</quantity>
long,
<quantity id="A44B87A1FFA4FFA1735A552DFC5DFB30" box="[794,880,1148,1172]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="521" unit="mm" value="5.2">5.2 mm</quantity>
wide); first ring 2 times longer than second ring, separation between rings distinct on all sides (
<figureCitation id="FB8836C1FFA4FFA173A355CEFB11FB13" box="[995,1084,1183,1207]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(a
<emphasis id="51C7F656FFA4FFA1741555CEFB4FFB13" box="[1109,1122,1183,1207]" italics="true" pageId="5" pageNumber="521"></emphasis>
c)). Ventral lip with a slight central depression and a couple of shallow wrinkles (
<figureCitation id="FB8836C1FFA4FFA173BC5593FB79FB7E" box="[1020,1108,1218,1242]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(b)).
</paragraph>
<paragraph id="630C2A44FFA4FFA1708055B5FBA9F9DE" blockId="5.[160,1157,697,1727]" pageId="5" pageNumber="521">
Maxillary apparatus with MF = 1+1, 3+4, 4+0, 5+6, 1+1 (
<figureCitation id="FB8836C1FFA4FFA1737855B5FCBDFB58" box="[824,912,1252,1276]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(d)). Maxillary carriers 4.1 times shorter than length of MI. MI forceps-like; closing system 5.8 times shorter than length of MI; ligament between MI and MII sclerotised (
<figureCitation id="FB8836C1FFA4FFA17359547BFC5DFAE6" box="[793,880,1322,1346]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(d,e)). MII with recurved teeth; cavity opening oval, 2.9 times shorter than length of MII; ligament between MII and MIII and right MIV, sclerotised (
<figureCitation id="FB8836C1FFA4FFA17210543EFD86FA23" box="[592,683,1391,1415]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(d,e)). MIII short; with blunt teeth; with irregular attachment lamella, situated in centre of right edge of maxilla, sclerotised (
<figureCitation id="FB8836C1FFA4FFA170E854E5FE2CFA68" box="[168,257,1460,1484]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(d,e)). Left MIV with two larger lateral teeth; attachment lamella semicircular, slender, better developed on left side, situated 2/3 of anterior edge of maxilla (
<figureCitation id="FB8836C1FFA4FFA174665486FBA9FA4B" box="[1062,1156,1495,1519]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(d,e)). Right MIV with two larger lateral teeth; attachment lamella semicircular, wide, better developed on the right side, situated along posterior edge (
<figureCitation id="FB8836C1FFA4FFA17385574DFB0EF990" box="[965,1059,1564,1588]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(d)). MV rectangular, with a short rounded tooth (
<figureCitation id="FB8836C1FFA4FFA172C0576EFDF7F9F3" box="[640,730,1599,1623]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(d,e)). Mandibles brown (
<figureCitation id="FB8836C1FFA4FFA17440576EFB71F9F3" box="[1024,1116,1599,1623]" captionStart="Figure 1" captionStartId="6.[160,225,1420,1443]" captionTargetBox="[162,1153,143,1382]" captionTargetId="figure@6.[161,1154,142,1389]" captionTargetPageId="6" captionText="Figure 1. Treadwellphysa dartevellei (Monro, 1936) comb. nov., holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm." figureDoi="http://doi.org/10.5281/zenodo.3675705" httpUri="https://zenodo.org/record/3675705/files/figure.png" pageId="5" pageNumber="521">Figure 1</figureCitation>
(f)); calcareous cutting plates broken, cutting plates brown, oval, with up to 10 growth rings.
</paragraph>
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Branchiae palmate with up to four filaments, present from chaetiger 35 until last parapodium of fragment (
<figureCitation id="FB8836C1FFA4FFA1718557F6FD0EF91B" box="[453,547,1703,1727]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="5" pageNumber="521">Figure 2</figureCitation>
(d
<emphasis id="51C7F656FFA4FFA1727857F6FD68F91B" box="[568,581,1703,1727]" italics="true" pageId="5" pageNumber="521"></emphasis>
f)). One filament along chaetigers 35L to 38L; three filaments along chaetigers 39L to 53L; four filaments from chaetiger 54L to end of fragment. Branchial filaments longer than dorsal cirri (
<figureCitation id="FB8836C1FFA7FFA2734A5727FC44F92A" box="[778,873,1654,1678]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="6" pageNumber="522">Figure 2</figureCitation>
(d
<emphasis id="51C7F656FFA7FFA2733E5727FCA6F92A" box="[894,907,1654,1678]" italics="true" pageId="6" pageNumber="522"></emphasis>
f)).
</paragraph>
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<paragraph id="630C2A44FFA7FFA270E054DDFFD2F9BD" blockId="6.[160,1157,1420,1561]" pageId="6" pageNumber="522">
<emphasis id="51C7F656FFA7FFA270E054DDFFD3FA07" bold="true" box="[160,254,1420,1443]" pageId="6" pageNumber="522">Figure 1.</emphasis>
<taxonomicName id="A4B351C7FFA7FFA2714854DCFDB2FA00" authority="(Monro, 1936)" authorityName="Molina-Acevedo" authorityYear="2019" baseAuthorityName="Monroi" baseAuthorityYear="1936" box="[264,671,1421,1444]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="6" pageNumber="522" phylum="Annelida" rank="species" species="dartevellei" status="comb. nov.">
<emphasis id="51C7F656FFA7FFA2714854DCFD2AFA00" box="[264,519,1421,1444]" italics="true" pageId="6" pageNumber="522">Treadwellphysa dartevellei</emphasis>
(
<bibRefCitation id="072257B5FFA7FFA2725754DCFDB5FA00" author="Monro CCA" box="[535,664,1421,1444]" pageId="6" pageNumber="522" pagination="245 - 248" refId="ref11485" refString="Monro CCA. 1936. Notes on some Polychaeta from the Congo coast. Rev Zool Bot Africaines. 28: 245 - 248." type="journal article" year="1936">Monro, 1936</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFA7FFA272E954DCFC30FA00" box="[681,797,1421,1444]" pageId="6" pageNumber="522" rank="species">comb. nov.</taxonomicNameLabel>
, holotype BMNH 1935.1.29.12. (a) Anterior end, dorsal view; (b) anterior end, ventral view; (c) anterior end, lateral view; (d) maxillary apparatus, dorsal view, arrows indicate the elevation at the base of MII; (e) left MIMV, lateral view, arrows indicate ligaments between maxillae; (f) mandibles. Scale bars: ac = 2.55 mm; df = 0.85 mm.
</paragraph>
</caption>
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<paragraph id="630C2A44FFA6FFA370E0544AFBA8FA5B" blockId="7.[160,1157,1307,1535]" pageId="7" pageNumber="523">
<emphasis id="51C7F656FFA6FFA370E0544AFFD1FA96" bold="true" box="[160,252,1307,1330]" pageId="7" pageNumber="523">Figure 2.</emphasis>
<taxonomicName id="A4B351C7FFA6FFA37145544AFDA7FA96" authority="(Monro, 1936)" authorityName="Molina-Acevedo" authorityYear="2019" baseAuthorityName="Monroi" baseAuthorityYear="1936" box="[261,650,1307,1330]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="7" pageNumber="523" phylum="Annelida" rank="species" species="dartevellei" status="comb. nov.">
<emphasis id="51C7F656FFA6FFA37145544AFED6FA96" box="[261,507,1307,1330]" italics="true" pageId="7" pageNumber="523">Treadwellphysa dartevellei</emphasis>
(
<bibRefCitation id="072257B5FFA6FFA37247544AFDAEFA96" author="Monro CCA" box="[519,643,1307,1330]" pageId="7" pageNumber="523" pagination="245 - 248" refId="ref11485" refString="Monro CCA. 1936. Notes on some Polychaeta from the Congo coast. Rev Zool Bot Africaines. 28: 245 - 248." type="journal article" year="1936">Monro, 1936</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFA6FFA372D1544AFDD3FA96" box="[657,766,1307,1330]" pageId="7" pageNumber="523" rank="species">comb. nov.</taxonomicNameLabel>
(a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324. Scale bars: a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view.
</paragraph>
</caption>
<paragraph id="630C2A44FFA6FFAC7080576EFD62FE7A" blockId="7.[160,1156,1599,1727]" lastBlockId="8.[160,1156,142,1241]" lastPageId="8" lastPageNumber="524" pageId="7" pageNumber="523">
First three parapodia smaller; best developed in chaetigers 5
<emphasis id="51C7F656FFA6FFA37314576EFC4CF9F3" box="[852,865,1599,1623]" italics="true" pageId="7" pageNumber="523"></emphasis>
39, following ones becoming gradually smaller. Notopodial cirri conical, longer than ventral cirri in first 15 chaetigers, following ones gradually decreasing in size, shorter than ventral cirri (
<figureCitation id="FB8836C1FFA6FFA373A657D5FB6BF938" box="[998,1094,1668,1692]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="7" pageNumber="523">Figure 2</figureCitation>
(a
<emphasis id="51C7F656FFA6FFA3741A57D5FB4AF938" box="[1114,1127,1668,1692]" italics="true" pageId="7" pageNumber="523"></emphasis>
f)). Prechaetal lobes short, as a transverse fold in all chaetigers (
<figureCitation id="FB8836C1FFA6FFA3730B57F6FC8BF91B" box="[843,934,1703,1727]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="7" pageNumber="523">Figure 2</figureCitation>
(a
<emphasis id="51C7F656FFA6FFA373FB57F6FCE5F91B" box="[955,968,1703,1727]" italics="true" pageId="7" pageNumber="523"></emphasis>
f)). Chaetal lobes in chaetigers 1
<emphasis id="51C7F656FFA9FFAC710651DFFE7EFF02" box="[326,339,142,166]" italics="true" pageId="8" pageNumber="524"></emphasis>
33 rounded, of similar size to postchaetal lobes, with aciculae emerging dorsal to midline; from chaetiger 34 triangular, longer than other lobes, with acicula emerging in midline (
<figureCitation id="FB8836C1FFA9FFAC71D55185FEDDFF48" box="[405,496,212,236]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(a
<emphasis id="51C7F656FFA9FFAC72455182FD3FFF4F" box="[517,530,211,235]" italics="true" pageId="8" pageNumber="524"></emphasis>
f)). Postchaetal lobes poorly developed in chaetigers 1
<emphasis id="51C7F656FFA9FFAC74365182FBAEFF4F" box="[1142,1155,211,235]" italics="true" pageId="8" pageNumber="524"></emphasis>
25, conical in first four chaetigers, rounded from chaetiger five; progressively decreasing in size from chaetigers 8 to 25; following ones inconspicuous (
<figureCitation id="FB8836C1FFA9FFAC73045048FC8DFE95" box="[836,928,281,305]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(a
<emphasis id="51C7F656FFA9FFAC73F45048FCECFE95" box="[948,961,281,305]" italics="true" pageId="8" pageNumber="524"></emphasis>
f)). Ventral cirri in chaetigers 1
<emphasis id="51C7F656FFA9FFAC7166506AFE1EFEF7" box="[294,307,315,339]" italics="true" pageId="8" pageNumber="524"></emphasis>
4 digitiform; in chaetigers 5 to end of the fragment with a swollen base as a transverse welt with a short digitiform tip, gradually decreasing in size (
<figureCitation id="FB8836C1FFA9FFAC738C500FFB05FED2" box="[972,1064,350,374]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(a
<emphasis id="51C7F656FFA9FFAC747C500FFB64FED2" box="[1084,1097,350,374]" italics="true" pageId="8" pageNumber="524"></emphasis>
f)). In complete specimen (BMNH 1939.7.17.23
<emphasis id="51C7F656FFA9FFAC721850D0FD48FE3D" box="[600,613,385,409]" italics="true" pageId="8" pageNumber="524"></emphasis>
24, L10 = 7.2 mm, 158 chaetigers) in chaetigers 5
<emphasis id="51C7F656FFA9FFAC743650D0FBAEFE3D" box="[1142,1155,385,409]" italics="true" pageId="8" pageNumber="524"></emphasis>
91 with a swollen base as a transverse welt with short digitiform tip; from chaetiger 92 digitiform, gradually decreasing in size.
</paragraph>
<paragraph id="630C2A44FFA9FFAC708050B8FDFEFD28" blockId="8.[160,1156,142,1241]" pageId="8" pageNumber="524">
Aciculae blunt, reddish along most of the length, amber on the distal tip (
<figureCitation id="FB8836C1FFA9FFAC73A850B8FB6DFDA5" box="[1000,1088,489,513]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(a
<emphasis id="51C7F656FFA9FFAC741550B8FB4FFDA5" box="[1109,1122,489,513]" italics="true" pageId="8" pageNumber="524"></emphasis>
e)); in additional specimen (BMNH 1939.7.17.23
<emphasis id="51C7F656FFA9FFAC72D2535AFDB2FD87" box="[658,671,523,547]" italics="true" pageId="8" pageNumber="524"></emphasis>
24) last chaetigers with two aciculae, one amber, and the second darker (
<figureCitation id="FB8836C1FFA9FFAC7255537FFD58FDE2" box="[533,629,558,582]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(f)). First three chaetigers with three aciculae, chaetigers 4
<emphasis id="51C7F656FFA9FFAC71675300FE19FDCD" box="[295,308,593,617]" italics="true" pageId="8" pageNumber="524"></emphasis>
28 with four, chaetigers 29
<emphasis id="51C7F656FFA9FFAC721D5300FD47FDCD" box="[605,618,593,617]" italics="true" pageId="8" pageNumber="524"></emphasis>
33 with three, chaetigers 34
<emphasis id="51C7F656FFA9FFAC73E15300FC83FDCD" box="[929,942,593,617]" italics="true" pageId="8" pageNumber="524"></emphasis>
101 with two, from chaetiger 102 with only one acicula (
<figureCitation id="FB8836C1FFA9FFAC72765325FDBDFD28" box="[566,656,628,652]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(a
<emphasis id="51C7F656FFA9FFAC72E45322FD9CFD2F" box="[676,689,627,651]" italics="true" pageId="8" pageNumber="524"></emphasis>
e)).
</paragraph>
<paragraph id="630C2A44FFA9FFAC708053C7FED3FB7D" blockId="8.[160,1156,142,1241]" pageId="8" pageNumber="524">
Limbate chaetae of two lengths in the same chaetiger, longer in dorsal region of the parapodia, reduced in number in anterior chaetiger. Three
<typeStatus id="BC0894E6FFA9FFAC737753E8FC5EFD75" box="[823,883,697,721]" pageId="8" pageNumber="524">types</typeStatus>
of pectinate chaetae; in anterior chaetigers, isodont narrow, with short and slender teeth, with 1
<emphasis id="51C7F656FFA9FFAC73AF538AFCD1FD57" box="[1007,1020,731,755]" italics="true" pageId="8" pageNumber="524"></emphasis>
2 pectinate, with up to 16 teeth (
<figureCitation id="FB8836C1FFA9FFAC71C853AFFEC8FCB2" box="[392,485,766,790]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(g)); in median-posterior chaetigers, isodont wide, with long and slender teeth, with 5
<emphasis id="51C7F656FFA9FFAC71FE5270FEE6FC9D" box="[446,459,801,825]" italics="true" pageId="8" pageNumber="524"></emphasis>
8 pectinate, with up to 20
<emphasis id="51C7F656FFA9FFAC72B45270FC2CFC9D" box="[756,769,801,825]" italics="true" pageId="8" pageNumber="524"></emphasis>
22 teeth (
<figureCitation id="FB8836C1FFA9FFAC73315270FCE2FC9D" box="[881,975,801,825]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(h)); in posterior chaetigers, anodont wide, with long and thick teeth, with 3
<emphasis id="51C7F656FFA9FFAC73025212FC62FCFF" box="[834,847,835,859]" italics="true" pageId="8" pageNumber="524"></emphasis>
4 pectinate, with up to 14
<emphasis id="51C7F656FFA9FFAC74365212FBAEFCFF" box="[1142,1155,835,859]" italics="true" pageId="8" pageNumber="524"></emphasis>
15 teeth (
<figureCitation id="FB8836C1FFA9FFAC714D5237FE41FCDA" box="[269,364,870,894]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(i)). Compound spinifalciger absent. Compound spinigers present in all chaetigers; in anterior-median region with blades of two lengths, shorter ones (
<figureCitation id="FB8836C1FFA9FFAC746752D8FBA9FC05" box="[1063,1156,905,929]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(k)) more abundant than longer ones (
<figureCitation id="FB8836C1FFA9FFAC722952FDFDE7FC60" box="[617,714,940,964]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(j)); in posterior region with blades of similar size, decreasing in number, two or three per parapodium. Compound falcigers present from chaetiger 45 to last chaetiger of fragment; in median-posterior region, with blades of similar length (48 µm,
<figureCitation id="FB8836C1FFA9FFAC72495545FD45FB88" box="[521,616,1044,1068]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(l)), more abundant than spinigers; all with blunt teeth, of similar size, distal tooth directed upward, proximal tooth directed laterally. Subacicular hooks bidentate, reddish along most of its length, translucent on the distal tip, starting from chaetiger 35, one per chaetiger, except for replacement; with distal triangular tooth, smaller than proximal, directed upward, with proximal blunt tooth, directed laterally (
<figureCitation id="FB8836C1FFA9FFAC712E5590FEE0FB7D" box="[366,461,1217,1241]" captionStart="Figure 2" captionStartId="7.[160,225,1307,1330]" captionTargetBox="[164,1151,143,1274]" captionTargetId="figure@7.[163,1153,142,1275]" captionTargetPageId="7" captionText="Figure 2. Treadwellphysa dartevellei (Monro, 1936) comb. nov. (a) Parapodium 3; (b) parapodium 8; (c) parapodium 21; (d) parapodium 53; (e) parapodium 71; (f) parapodium 151; (g) chaetiger 21, isodont pectinate narrow with short and slender teeth; (h) chaetiger 93, isodont pectinate wide with long and slender teeth; (i) chaetiger 150, anodont pectinate wide with long and thick teeth; (j) chaetiger 8, compound spiniger with long blade; (k) chaetiger 8, compound spiniger with short blade; (l) chaetiger 71, compound falciger; (m) chaetiger 93, subacicular hook. ae, g, l from holotype BMNH 1935.1.29.12; f, hk, m from non-type BMNH 1939.7.17.2324.Scale bars:a = 0.15 mm; b = 0.29 mm; c = 0.42 mm; d = 0.3 mm; e = 0.6 mm; f = 0.2 mm; gi = 7 µm; jl = 17.5 µm; m = 60 µm. All parapodia in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3675707" httpUri="https://zenodo.org/record/3675707/files/figure.png" pageId="8" pageNumber="524">Figure 2</figureCitation>
(m)).
</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFA9FFAC70E05457FE72F91B" pageId="8" pageNumber="524" type="materials_examined">
<paragraph id="630C2A44FFA9FFAC70E05457FE34FABB" blockId="8.[160,1156,1286,1727]" box="[160,281,1286,1311]" pageId="8" pageNumber="524">
<emphasis id="51C7F656FFA9FFAC70E05457FE34FABB" bold="true" box="[160,281,1286,1311]" italics="true" pageId="8" pageNumber="524">Variations</emphasis>
</paragraph>
<paragraph id="630C2A44FFA9FFAC70E0547BFE72F91B" blockId="8.[160,1156,1286,1727]" pageId="8" pageNumber="524">
Material examined with L10 =
<quantity id="A44B87A1FFA9FFAC7257547BFDB7FAE6" box="[535,666,1322,1346]" metricMagnitude="-3" metricUnit="m" metricValue="8.15" metricValueMax="11.0" metricValueMin="5.3" pageId="8" pageNumber="524" unit="mm" value="8.15" valueMax="11.0" valueMin="5.3">
5.3
<emphasis id="51C7F656FFA9FFAC7278547BFD68FAE6" box="[568,581,1322,1346]" italics="true" pageId="8" pageNumber="524"></emphasis>
11 mm
</quantity>
and W10 =
<quantity id="A44B87A1FFA9FFAC737E547BFCE7FAE6" box="[830,970,1322,1346]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="6.7" metricValueMin="3.1" pageId="8" pageNumber="524" unit="mm" value="4.9" valueMax="6.7" valueMin="3.1">
3.1
<emphasis id="51C7F656FFA9FFAC731F547BFC41FAE6" box="[863,876,1322,1346]" italics="true" pageId="8" pageNumber="524"></emphasis>
6.7 mm
</quantity>
. One specimen complete with 158 chaetigers and pygidium in regeneration. Palps reaching middle of the first peristomial ring or first chaetiger. Lateral and median antennae reaching first or third chaetiger. The maxillary formula variations are MII 3
<emphasis id="51C7F656FFA9FFAC736454C3FC1CFA0E" box="[804,817,1426,1450]" italics="true" pageId="8" pageNumber="524"></emphasis>
4+3
<emphasis id="51C7F656FFA9FFAC731D54C3FC47FA0E" box="[861,874,1426,1450]" italics="true" pageId="8" pageNumber="524"></emphasis>
4,
<collectionCode id="05A2B281FFA9FFAC73C554C3FC83FA0E" box="[901,942,1426,1450]" pageId="8" pageNumber="524">MIII</collectionCode>
4
<emphasis id="51C7F656FFA9FFAC738254C3FCE2FA0E" box="[962,975,1426,1450]" italics="true" pageId="8" pageNumber="524"></emphasis>
5,
<collectionCode id="05A2B281FFA9FFAC73AA54C3FB38FA0E" box="[1002,1045,1426,1450]" country="Italy" httpUri="http://grbio.org/cool/mev6-t33n" name="Universita' degli Studi di Milano, Medicina Veterinaria, Sez. Parassitologia" pageId="8" pageNumber="524" type="University or college">MIV</collectionCode>
4
<emphasis id="51C7F656FFA9FFAC746A54C3FB1AFA0E" box="[1066,1079,1426,1450]" italics="true" pageId="8" pageNumber="524"></emphasis>
6+6
<emphasis id="51C7F656FFA9FFAC742354C3FB5DFA0E" box="[1123,1136,1426,1450]" italics="true" pageId="8" pageNumber="524"></emphasis>
7. The proportions of the maxillary apparatus vary as follows: the maxillary carriers are 2.8
<emphasis id="51C7F656FFA9FFAC743654E5FBAEFA68" box="[1142,1155,1460,1484]" italics="true" pageId="8" pageNumber="524"></emphasis>
4.1 times shorter than the length of the MI; the closing system is 5
<emphasis id="51C7F656FFA9FFAC73F35486FCEDFA4B" box="[947,960,1495,1519]" italics="true" pageId="8" pageNumber="524"></emphasis>
5.8 times shorter than the length of the MI; the cavity opening is 2.8
<emphasis id="51C7F656FFA9FFAC72A254ABFDC2F9B6" box="[738,751,1530,1554]" italics="true" pageId="8" pageNumber="524"></emphasis>
3.8 times shorter than the length of the MII. Branchiae from chaetigers 30
<emphasis id="51C7F656FFA9FFAC720F574DFD71F990" box="[591,604,1564,1588]" italics="true" pageId="8" pageNumber="524"></emphasis>
44 to last chaetiger. Maximum number of branchial filaments varied from four to five. Poorly developed postchaetal lobe present in the first 23
<emphasis id="51C7F656FFA9FFAC70AE5733FFD6F9DE" box="[238,251,1634,1658]" italics="true" pageId="8" pageNumber="524"></emphasis>
40 chaetigers. Ventral cirri with a swollen base from chaetigers 5 to 91. Start of subacicular hooks in chaetigers 30
<emphasis id="51C7F656FFA9FFAC726757D5FD19F938" box="[551,564,1668,1692]" italics="true" pageId="8" pageNumber="524"></emphasis>
39. Compound falcigers start from chaetigers 30
<emphasis id="51C7F656FFA9FFAC741A57D5FB4AF938" box="[1114,1127,1668,1692]" italics="true" pageId="8" pageNumber="524"></emphasis>
54 to last chaetiger.
</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFA8FFAD70E051DFFDBCFF6E" box="[160,657,142,202]" pageId="9" pageNumber="525" type="distribution">
<paragraph id="630C2A44FFA8FFAD70E051DFFE01FF03" blockId="9.[160,657,142,202]" box="[160,300,142,167]" pageId="9" pageNumber="525">
<emphasis id="51C7F656FFA8FFAD70E051DFFE01FF03" bold="true" box="[160,300,142,167]" italics="true" pageId="9" pageNumber="525">Distribution</emphasis>
</paragraph>
<paragraph id="630C2A44FFA8FFAD70E051E3FDBCFF6E" blockId="9.[160,657,142,202]" box="[160,657,178,202]" pageId="9" pageNumber="525">
Angola, Ghana, and the
<collectingCountry id="1BA46AD4FFA8FFAD71F651E3FDA0FF6E" box="[438,653,178,202]" name="Republic of the Congo" pageId="9" pageNumber="525">Republic of Congo</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="2BA979CFFFA8FFAD70E051A7FC93FDE3" pageId="9" pageNumber="525" type="discussion">
<paragraph id="630C2A44FFA8FFAD70E051A7FE2BFEAB" blockId="9.[160,1156,246,583]" box="[160,262,246,271]" pageId="9" pageNumber="525">
<emphasis id="51C7F656FFA8FFAD70E051A7FE2BFEAB" bold="true" box="[160,262,246,271]" italics="true" pageId="9" pageNumber="525">Remarks</emphasis>
</paragraph>
<paragraph id="630C2A44FFA8FFAD70E0504BFC93FDE3" blockId="9.[160,1156,246,583]" pageId="9" pageNumber="525">
<taxonomicName id="A4B351C7FFA8FFAD70E0504BFEECFE96" authorityName="Molina-Acevedo" authorityYear="2019" baseAuthorityName="Monroi" baseAuthorityYear="1936" box="[160,449,282,306]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="525" phylum="Annelida" rank="species" species="dartevellei" status="comb. nov.">
<emphasis id="51C7F656FFA8FFAD70E0504BFEECFE96" box="[160,449,282,306]" italics="true" pageId="9" pageNumber="525">Treadwellphysa dartevellei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFA8FFAD718A504BFD67FE96" box="[458,586,282,306]" pageId="9" pageNumber="525" rank="species">comb. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="A4B351C7FFA8FFAD728E504BFC6FFE96" baseAuthorityName="Fauchaldi" baseAuthorityYear="1977" box="[718,834,282,306]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="525" phylum="Annelida" rank="species" species="amadae">
<emphasis id="51C7F656FFA8FFAD728E504BFC6FFE96" box="[718,834,282,306]" italics="true" pageId="9" pageNumber="525">T. amadae</emphasis>
</taxonomicName>
,
<taxonomicName id="A4B351C7FFA8FFAD7312504BFC86FE96" authorityName="Molina-Acevedo" authorityYear="2019" baseAuthorityName="Fauchaldi" baseAuthorityYear="1970" box="[850,939,282,306]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="525" phylum="Annelida" rank="species" species="mixta" status="comb. nov.">
<emphasis id="51C7F656FFA8FFAD7312504BFC86FE96" box="[850,939,282,306]" italics="true" pageId="9" pageNumber="525">T. mixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFA8FFAD73F5504BFB19FE96" box="[949,1076,282,306]" pageId="9" pageNumber="525" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName id="A4B351C7FFA8FFAD7431504BFE01FEF0" authorityName="Molina-Acevedo and Carrera-Parrai" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="525" phylum="Annelida" rank="species" species="yucatanensis">
<emphasis id="51C7F656FFA8FFAD7431504BFE01FEF0" italics="true" pageId="9" pageNumber="525">T. yucatanensis</emphasis>
</taxonomicName>
in having compound spinigiers and falcigers in anterior chaetigers. However,
<taxonomicName id="A4B351C7FFA8FFAD7154500EFEB2FED3" authorityName="Molina-Acevedo" authorityYear="2019" baseAuthorityName="Monroi" baseAuthorityYear="1936" box="[276,415,351,375]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="525" phylum="Annelida" rank="species" species="dartevellei" status="comb. nov.">
<emphasis id="51C7F656FFA8FFAD7154500EFEB2FED3" box="[276,415,351,375]" italics="true" pageId="9" pageNumber="525">T. dartevellei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFA8FFAD71EA500EFD07FED3" box="[426,554,351,375]" pageId="9" pageNumber="525" rank="species">comb. nov.</taxonomicNameLabel>
lacks compound spinifalcigers and the subaciacular hooks are reddish with a translucent distal end; while
<taxonomicName id="A4B351C7FFA8FFAD736250D3FCB4FE3E" baseAuthorityName="Fauchaldi" baseAuthorityYear="1977" box="[802,921,386,410]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="525" phylum="Annelida" rank="species" species="amadae">
<emphasis id="51C7F656FFA8FFAD736250D3FCB4FE3E" box="[802,921,386,410]" italics="true" pageId="9" pageNumber="525">T. amadae</emphasis>
</taxonomicName>
and
<taxonomicName id="A4B351C7FFA8FFAD739A50D3FBAEFE3E" authorityName="Molina-Acevedo and Carrera-Parrai" authorityYear="2017" box="[986,1155,386,410]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="525" phylum="Annelida" rank="species" species="yucatanensis">
<emphasis id="51C7F656FFA8FFAD739A50D3FBAEFE3E" box="[986,1155,386,410]" italics="true" pageId="9" pageNumber="525">T. yucatanensis</emphasis>
</taxonomicName>
have spinifalcigers and translucent subacicular hooks. Moreover,
<taxonomicName id="A4B351C7FFA8FFAD733850F5FCE2FE18" authorityName="Molina-Acevedo" authorityYear="2019" baseAuthorityName="Fauchaldi" baseAuthorityYear="1970" box="[888,975,420,444]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="525" phylum="Annelida" rank="species" species="mixta" status="comb. nov.">
<emphasis id="51C7F656FFA8FFAD733850F5FCE2FE18" box="[888,975,420,444]" italics="true" pageId="9" pageNumber="525">T. mixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFA8FFAD739550F5FB7FFE18" box="[981,1106,420,444]" pageId="9" pageNumber="525" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName id="A4B351C7FFA8FFAD70E05096FE03FE7B" authorityName="Molina-Acevedo" authorityYear="2019" baseAuthorityName="Monroi" baseAuthorityYear="1936" box="[160,302,455,479]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="525" phylum="Annelida" rank="species" species="dartevellei" status="comb. nov.">
<emphasis id="51C7F656FFA8FFAD70E05096FE03FE7B" box="[160,302,455,479]" italics="true" pageId="9" pageNumber="525">T. dartevellei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFA8FFAD717A5096FE90FE7B" box="[314,445,455,479]" pageId="9" pageNumber="525" rank="species">comb. nov.</taxonomicNameLabel>
differ by the distribution of the compound chaetae; in the former, the compound spinigers are only in the anterior chaetigers and the compound falcigers are in all chaetigers; while
<taxonomicName id="A4B351C7FFA8FFAD7276535DFDEDFD80" authorityName="Molina-Acevedo" authorityYear="2019" baseAuthorityName="Monroi" baseAuthorityYear="1936" box="[566,704,524,548]" class="Polychaeta" family="Eunicidae" genus="Treadwellphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="525" phylum="Annelida" rank="species" species="dartevellei" status="comb. nov.">
<emphasis id="51C7F656FFA8FFAD7276535DFDEDFD80" box="[566,704,524,548]" italics="true" pageId="9" pageNumber="525">T. dartevellei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AF44B2DFFA8FFAD7289535DFC6AFD80" box="[713,839,524,548]" pageId="9" pageNumber="525" rank="species">comb. nov.</taxonomicNameLabel>
has compound spinigers in all chaetigers and compound falcigers present from chaetigers 30
<emphasis id="51C7F656FFA8FFAD73CF537EFCB1FDE3" box="[911,924,559,583]" italics="true" pageId="9" pageNumber="525"></emphasis>
54.
</paragraph>
</subSubSection>
</treatment>
</document>