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<document id="83AB7EA37E5DBD83F82F720DA752F866" ID-CLB-Dataset="26923" ID-DOI="10.11646/zootaxa.4594.1.1" ID-GBIF-Dataset="3fc46d02-9b74-434b-a777-46d70967d61f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2667081" ID-ZooBank="4308D364-BCD7-473D-83BC-5B4130C32287" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556963471166" checkinUser="plazi" docAuthor="Leong, Chi-Man, Guénard, Benoit, Shiao, Shiuh-Feng &amp; Lin, Chung-Chi" docDate="2019" docId="6E6AB6166F65FF8CFF065BA6FBA9CBB5" docLanguage="en" docName="zootaxa.4594.1.1.pdf.imf" docOrigin="Zootaxa 4594 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ponera alisana Terayama 1986" docType="treatment" docVersion="11" lastPageNumber="27" masterDocId="9253CE6E6F71FF96FF915F65FFB6CD2E" masterDocTitle="Taxonomic revision of the genus Ponera Latreille, 1804 (Hymenoptera: Formicidae) of Taiwan and Japan, with a key to East Asian species" masterLastPageNumber="86" masterPageNumber="1" pageNumber="21" updateTime="1698725828403" updateUser="ExternalLinkService">
<mods:mods id="09AD9A49642F67943175EAA97D603E79" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="6610F2859494E87C75E6E718CD1AB38D">Taxonomic revision of the genus Ponera Latreille, 1804 (Hymenoptera: Formicidae) of Taiwan and Japan, with a key to East Asian species</mods:title>
</mods:titleInfo>
<mods:name id="9CE2AEA7E409E62D597BB52ABBBE8645" type="personal">
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<mods:namePart id="1319E77B6419246977E9C857CBDAC9C3">Leong, Chi-Man</mods:namePart>
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<mods:namePart id="3E10D7285BE9541FA75785DFEDFB5231">Guénard, Benoit</mods:namePart>
</mods:name>
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<mods:role id="392593D59B6B0CDB06F7A6E14E7D9502">
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<mods:namePart id="90CED73194E02FA8FEC29193992967B0">Shiao, Shiuh-Feng</mods:namePart>
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<mods:namePart id="D034C1CA4983158E4A6D2A99C7D17CD9">Lin, Chung-Chi</mods:namePart>
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<mods:title id="19206155F5F6444C09DE611CEEBA7A98">Zootaxa</mods:title>
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<mods:date id="1C8B89599A2AB0633B1CD9463F775337">2019</mods:date>
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<mods:number id="DA4F1506FCF9B4C1DE60230FB79E8FBB">2019-05-03</mods:number>
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<mods:number id="863258FECDC4878263804B0F4A5D83A4">4594</mods:number>
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<mods:identifier id="A640DC0C6315F517CAF50933FDAF4D28" type="DOI">10.11646/zootaxa.4594.1.1</mods:identifier>
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<treatment id="6E6AB6166F65FF8CFF065BA6FBA9CBB5" ID-DOI="http://doi.org/10.5281/zenodo.5669046" ID-GBIF-Taxon="157059607" ID-Zenodo-Dep="5669046" LSID="urn:lsid:plazi:treatment:6E6AB6166F65FF8CFF065BA6FBA9CBB5" httpUri="http://treatment.plazi.org/id/6E6AB6166F65FF8CFF065BA6FBA9CBB5" lastPageId="26" lastPageNumber="27" pageId="20" pageNumber="21">
<subSubSection id="AED9548B6F65FF82FF065BA6FDABC9F0" box="[151,541,1219,1246]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="E67C07006F65FF82FF065BA6FDABC9F0" blockId="20.[151,875,1219,1280]" box="[151,541,1219,1246]" pageId="20" pageNumber="21">
<heading id="BD34B06C6F65FF82FF065BA6FDABC9F0" bold="true" box="[151,541,1219,1246]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<taxonomicName id="21C37C836F65FF82FF065BA6FDABC9F0" ID-CoL="4LTMM" authority="Terayama, 1986" authorityName="Terayama" authorityYear="1986" box="[151,541,1219,1246]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="alisana">
<emphasis id="D4B7DB126F65FF82FF065BA6FDABC9F0" bold="true" box="[151,541,1219,1246]" pageId="20" pageNumber="21">
<emphasis id="D4B7DB126F65FF82FF065BA6FEF8C9F3" bold="true" box="[151,334,1219,1245]" italics="true" pageId="20" pageNumber="21">Ponera alisana</emphasis>
<bibRefCitation id="82527AF16F65FF82FEC45BA1FDABC9F0" author="Terayama, M." box="[341,541,1220,1246]" pageId="20" pageNumber="21" pagination="591 - 595" refId="ref36618" refString="Terayama, M. (1986) Two new ants of the genus Ponera (Hymenoptera, Formicidae) from Taiwan. Kontyu, 54, 591 - 595." type="journal article" year="1986">Terayama, 1986</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F65FF82FF065B82FCDDC82E" box="[151,875,1255,1280]" pageId="20" pageNumber="21" type="description">
<paragraph id="E67C07006F65FF82FF065B82FCDDC82E" blockId="20.[151,875,1219,1280]" box="[151,875,1255,1280]" pageId="20" pageNumber="21">
(
<figureCitation id="7EF81B856F65FF82FF0E5B82FF42C9D1" box="[159,244,1255,1280]" captionStart="FIGURE 31" captionStartId="21.[151,250,1990,2012]" captionTargetBox="[155,1408,193,1965]" captionTargetId="figure@21.[155,1408,193,1965]" captionTargetPageId="21" captionText="FIGURE 31. P. alisana worker (CMPon045), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667145/files/figure.png" pageId="20" pageNumber="21">Fig. 31</figureCitation>
: worker;
<figureCitation id="7EF81B856F65FF82FECD5B82FE0FC9D1" box="[348,441,1255,1280]" captionStart="FIGURE 32" captionStartId="22.[151,250,1723,1745]" captionTargetBox="[167,1420,193,1701]" captionTargetId="figure@22.[167,1420,193,1701]" captionTargetPageId="22" captionText="FIGURE 32. P. alisana alate queen (CMPon050), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667147/files/figure.png" pageId="20" pageNumber="21">Figs. 32</figureCitation>
,
<figureCitation id="7EF81B856F65FF82FE545B82FE53C9D1" box="[453,485,1255,1279]" captionStart="FIGURE 33" captionStartId="23.[151,250,1900,1922]" captionTargetBox="[167,1420,812,1878]" captionTargetId="figure@23.[167,1420,812,1878]" captionTargetPageId="23" captionText="FIGURE 33. P. alisana alate queen (CMPon050), A: left-forewing, B: left-hindwing." httpUri="https://zenodo.org/record/2667149/files/figure.png" pageId="20" pageNumber="21">33</figureCitation>
: dealate queen;
<figureCitation id="7EF81B856F65FF82FD095B82FD42C9D1" box="[664,756,1255,1280]" captionStart="FIGURE 34" captionStartId="24.[151,250,1953,1975]" captionTargetBox="[158,1428,193,1932]" captionTargetId="figure@24.[158,1428,193,1932]" captionTargetPageId="24" captionText="FIGURE 34. P. alisana male (CMPon049), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: antennae." httpUri="https://zenodo.org/record/2667151/files/figure.png" pageId="20" pageNumber="21">Figs. 34</figureCitation>
,
<figureCitation id="7EF81B856F65FF82FC915B82FC96C82E" box="[768,800,1255,1280]" captionStart="FIGURE 35" captionStartId="25.[151,250,1109,1131]" captionTargetBox="[167,1420,193,1086]" captionTargetId="figure@25.[167,1420,193,1088]" captionTargetPageId="25" captionText="FIGURE 35. P. alisana male (CMPon049), A: left-forewing, B: right-hindwing." httpUri="https://zenodo.org/record/2667153/files/figure.png" pageId="20" pageNumber="21">35</figureCitation>
: male)
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F65FF82FF065A55FF5FC84D" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="E67C07006F65FF82FF065A55FF5FC84D" blockId="20.[151,1436,1327,1379]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="C6D3202E6F65FF82FF065A55FF5FC84D" pageId="20" pageNumber="21">
<taxonomicName id="21C37C836F65FF82FF065A55FDA2C86B" ID-CoL="4LTMM" authority="Terayama, 1986: 591" authorityName="Terayama" authorityPageNumber="591" authorityYear="1986" box="[151,532,1327,1349]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="alisana">
<emphasis id="D4B7DB126F65FF82FF065A55FE82C86B" box="[151,308,1327,1349]" italics="true" pageId="20" pageNumber="21">Ponera alisana</emphasis>
<treatmentCitation id="676221116F65FF82FEAA5A55FDA2C86B" author="Terayama, M." box="[315,532,1327,1349]" page="591" pageId="20" pageNumber="21" year="1986">
<bibRefCitation id="82527AF16F65FF82FEAA5A55FE55C86B" author="Terayama, M." box="[315,483,1327,1349]" pageId="20" pageNumber="21" pagination="591 - 595" refId="ref36618" refString="Terayama, M. (1986) Two new ants of the genus Ponera (Hymenoptera, Formicidae) from Taiwan. Kontyu, 54, 591 - 595." type="journal article" year="1986">Terayama, 1986</bibRefCitation>
: 591
</treatmentCitation>
</taxonomicName>
(w. q.).
<typeStatus id="3978B9A26F65FF82FDFC5A55FD29C86B" box="[621,671,1328,1349]" pageId="20" pageNumber="21">Type</typeStatus>
locality:
<collectingCountry id="9ED447906F65FF82FC935A4AFCF9C86B" box="[770,847,1327,1349]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
,
<collectingRegion id="2407C9E26F65FF82FCCB5A4AFC55C86B" box="[858,995,1327,1349]" country="Taiwan" name="Chiayi" pageId="20" pageNumber="21">Chiayi Hsien</collectingRegion>
, Fenchihu.
<treatmentCitation id="676221116F65FF82FBCA5A4AFAC3C86B" author="Yoshimura, M. &amp; Hosoishi, S. &amp; Kuboki, Y. &amp; Onoyama, K. &amp; Ogata, K." box="[1115,1397,1327,1349]" page="198" pageId="20" pageNumber="21" year="2009">
<bibRefCitation id="82527AF16F65FF82FBCA5A4AFAF5C86B" author="Yoshimura, M. &amp; Hosoishi, S. &amp; Kuboki, Y. &amp; Onoyama, K. &amp; Ogata, K." box="[1115,1347,1327,1349]" pageId="20" pageNumber="21" pagination="194 - 201" refId="ref37048" refString="Yoshimura, M., Hosoishi, S., Kuboki, Y., Onoyama, K. &amp; Ogata, K. (2009) New synonym and new Japanese record of the ant genus Ponera (Hymenoptera: Formicidae). Entomological Science, 12, 194 - 201. https: // doi. org / 10.1111 / j. 1479 - 8298.2009.00325. x" type="journal article" year="2009">
Yoshimura
<emphasis id="D4B7DB126F65FF82FB405A55FB49C86B" box="[1233,1279,1327,1349]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
. 2009
</bibRefCitation>
: 198
</treatmentCitation>
(w. q.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F65FF82FF065AF5FCFCCA53" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="E67C07006F65FF82FF065AF5FB4FC8E3" blockId="20.[151,1436,1424,1917]" pageId="20" pageNumber="21">
<emphasis id="D4B7DB126F65FF82FF065AF5FD8FC887" bold="true" box="[151,569,1424,1449]" pageId="20" pageNumber="21">
Type material examined:
<materialsCitation id="56AB0D5D6F65FF82FE595AF5FD8FC887" ID-GBIF-Occurrence="2237845462" box="[456,569,1424,1449]" collectingDate="1980-03-03" collectionCode="TARI" collectorName="M Terayama" country="Taiwan" elevation="1400" location="Fenchihu" pageId="20" pageNumber="21" specimenCount="1" specimenCount-worker="1" stateProvince="Chiayi" typeStatus="paratype">
<collectingCountry id="9ED447906F65FF82FE595AF5FD8FC887" box="[456,569,1424,1449]" name="Taiwan" pageId="20" pageNumber="21">TAIWAN</collectingCountry>
</materialsCitation>
</emphasis>
.
<typeStatus id="3978B9A26F65FF82FDD65AF4FD01C887" box="[583,695,1425,1449]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="F0C5CC896F65FF82FD535AF4FC90C887" box="[706,806,1424,1449]" pageId="20" pageNumber="21" type="worker">1 worker</specimenCount>
and 1 alate queen,
<collectingRegion id="2407C9E26F65FF82FB915AF5FB28C887" box="[1024,1182,1424,1449]" country="Taiwan" name="Chiayi" pageId="20" pageNumber="21">Chiayi county</collectingRegion>
,
<location id="E31C51DB6F65FF82FB3B5AF5FAA3C887" LSID="urn:lsid:plazi:treatment:6E6AB6166F65FF8CFF065BA6FBA9CBB5:E31C51DB6F65FF82FB3B5AF5FAA3C887" box="[1194,1301,1424,1449]" country="Taiwan" name="Fenchihu" pageId="20" pageNumber="21" stateProvince="Chiayi">Fenchihu</location>
, ca.
<quantity id="213BAAE56F65FF82FAD65AF4FA2AC886" box="[1351,1436,1425,1449]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="20" pageNumber="21" unit="m" value="1400.0">
<elevation id="6DEEE0336F65FF82FAD65AF4FA2AC886" box="[1351,1436,1425,1449]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="20" pageNumber="21" unit="m" value="1400.0">1400 m</elevation>
</quantity>
alt.,
<collectingDate id="8239D8286F65FF82FF595AD0FE8DC8E3" box="[200,315,1461,1485]" pageId="20" pageNumber="21" value="1980-03-03">
<date id="927D21C06F65FF82FF595AD0FE8DC8E3" box="[200,315,1461,1485]" pageId="20" pageNumber="21" value="1980-03-03">3 III 1980</date>
</collectingDate>
,
<collectorName id="4B3662D66F65FF82FED75AD0FE62C8E3" box="[326,468,1461,1485]" pageId="20" pageNumber="21">M Terayama</collectorName>
leg (TARI),
<materialsCitation id="56AB0D5D6F65FF82FDF55AD0FB43C8E3" ID-GBIF-Occurrence="2237845420" box="[612,1269,1460,1485]" collectionCode="NIAES" country="Taiwan" pageId="20" pageNumber="21" specimenCount="1" specimenCount-worker="1" typeStatus="paratype">
<specimenCount id="F0C5CC896F65FF82FDF55AD0FD7EC8E3" box="[612,712,1460,1485]" pageId="20" pageNumber="21" type="worker">1 worker</specimenCount>
and 1 alate queen, same collecting data (
<collectionCode id="80D29FC56F65FF82FB055AD0FB5BC8E3" box="[1172,1261,1461,1485]" country="Japan" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="20" pageNumber="21">NIAES</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E67C07006F65FF82FF565ABDFCD0CB17" blockId="20.[151,1436,1424,1917]" pageId="20" pageNumber="21">
<emphasis id="D4B7DB126F65FF82FF565ABDFD28C8DF" bold="true" box="[199,670,1496,1521]" pageId="20" pageNumber="21">
Non-type material examined:
<materialsCitation id="56AB0D5D6F65FF82FDBC5ABDFD28C8DF" ID-GBIF-Occurrence="2237845401" box="[557,670,1496,1521]" country="Taiwan" location="TAIWAN" pageId="20" pageNumber="21" specimenCount="3" specimenCount-worker="3">
<collectingCountry id="9ED447906F65FF82FDBC5ABDFD28C8DF" box="[557,670,1496,1521]" name="Taiwan" pageId="20" pageNumber="21">TAIWAN</collectingCountry>
</materialsCitation>
</emphasis>
.
<specimenCount id="F0C5CC896F65FF82FD3D5ABCFCA9C8DF" box="[684,799,1496,1521]" pageId="20" pageNumber="21" type="worker">3 workers</specimenCount>
, 6 alate queens and
<materialsCitation id="56AB0D5D6F65FF82FB905ABCFCD4CB17" ID-GBIF-Occurrence="2237845426" collectingDate="1997-10-08" collectionCode="NTU" collectorName="WH Tsai" country="Taiwan" elevation="1400" location="Fenchihu" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Chiayi">
<specimenCount id="F0C5CC896F65FF82FB905ABCFBF9C8DF" box="[1025,1103,1496,1521]" pageId="20" pageNumber="21" type="male">1 male</specimenCount>
,
<collectingRegion id="2407C9E26F65FF82FBCA5ABDFB4CC8DF" box="[1115,1274,1496,1521]" country="Taiwan" name="Chiayi" pageId="20" pageNumber="21">Chiayi county</collectingRegion>
,
<location id="E31C51DB6F65FF82FA975ABDFAC7C8DF" LSID="urn:lsid:plazi:treatment:6E6AB6166F65FF8CFF065BA6FBA9CBB5:E31C51DB6F65FF82FA975ABDFAC7C8DF" box="[1286,1393,1496,1521]" country="Taiwan" name="Fenchihu" pageId="20" pageNumber="21" stateProvince="Chiayi">Fenchihu</location>
, ca.
<elevation id="6DEEE0336F65FF82FF065A98FF44CB3A" box="[151,242,1533,1557]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="20" pageNumber="21" unit="m" value="1400.0">
<quantity id="213BAAE56F65FF82FF065A98FF44CB3A" box="[151,242,1533,1557]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="20" pageNumber="21" unit="m" value="1400.0">1400 m</quantity>
</elevation>
alt.,
<collectingDate id="8239D8286F65FF82FEA75A98FE18CB3B" box="[310,430,1533,1557]" pageId="20" pageNumber="21" value="1997-10-08">
<date id="927D21C06F65FF82FEA75A98FE18CB3B" box="[310,430,1533,1557]" pageId="20" pageNumber="21" value="1997-10-08">8 X 1997</date>
</collectingDate>
,
<collectorName id="4B3662D66F65FF82FE2E5A98FD9FCB3B" box="[447,553,1532,1557]" pageId="20" pageNumber="21">WH Tsai</collectorName>
leg (
<collectionCode id="80D29FC56F65FF82FDFE5A98FD18CB3B" box="[623,686,1533,1557]" pageId="20" pageNumber="21">NTU</collectionCode>
: CMPon046, CMPon047, CMPon048, CMPon049, CMPon050, CMPon051, CMPon052, CMPon053, CMPon054, CMPon055)
</materialsCitation>
.
</paragraph>
<paragraph id="E67C07006F65FF82FF565920FEA1CBE7" blockId="20.[151,1436,1424,1917]" pageId="20" pageNumber="21">Measurements (mm) and indices. Workers (n=4): HL 0.760.81; HW 0.620.65; SL 0.610.66; A06L 0.05; A07L 0.06; A08L 0.09; A09L 0.10; A10L 0.11; PrW 0.450.53; WL 1.051.16; PeH 0.450.49; PeNL 0.240.27; PeW 0.350.38; ATL 0.530.55; ATW 0.610.65; CI 7883, SI 98105, PeI 7079, LPeI 5160, DPeI 141151, ATI 8589.</paragraph>
<paragraph id="E67C07006F65FF82FF5659B0FB3BCA1B" blockId="20.[151,1436,1424,1917]" pageId="20" pageNumber="21">Queens (n=6): HL 0.820.84; HW 0.650.70; SL 0.6472; A06L 0.06; A07L 0.07; A08L 0.09; A09L 0.10; A10L 0.12; PrW 0.600.65; WL 1.281.32; PeH 0.530.56; PeNL 0.250.29; PeW 0.420.43; ATL 0.620.64; ATW 0.7279; CI 7983, SI 103106, PeI 6672, LPeI 4552, DPeI 146171, ATI 8187.</paragraph>
<paragraph id="E67C07006F65FF82FF565825FCFCCA53" blockId="20.[151,1436,1424,1917]" pageId="20" pageNumber="21">Male (n=1): HL 0.62; HW 0.55; SL 0.06; PrW 0.77; WL 1.27; PeH 0.37; PeNL 0.20; PeW 0.30; ATL 0.62; ATW 0.60; CI 88, SI 11, PeI 39, LPeI 53, DPeI 151, ATI 104.</paragraph>
</subSubSection>
<caption id="B2BC57886F64FF83FF0658A3FAAFCAF2" box="[151,1305,1990,2012]" httpUri="https://zenodo.org/record/2667145/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1990,2012]" targetBox="[155,1408,193,1965]" targetPageId="21">
<paragraph id="E67C07006F64FF83FF0658A3FAAFCAF2" blockId="21.[151,1305,1990,2012]" box="[151,1305,1990,2012]" pageId="21" pageNumber="22">
<emphasis id="D4B7DB126F64FF83FF0658A3FEA9CAF2" bold="true" box="[151,287,1990,2012]" pageId="21" pageNumber="22">FIGURE 31.</emphasis>
<taxonomicName id="21C37C836F64FF83FEB458A2FE3CCAF2" authorityName="Terayama" authorityYear="1986" box="[293,394,1990,2012]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="alisana">
<emphasis id="D4B7DB126F64FF83FEB458A2FE84CAF2" box="[293,306,1991,2012]" italics="true" pageId="21" pageNumber="22">P</emphasis>
.
<emphasis id="D4B7DB126F64FF83FEAE58A3FE3CCAF2" box="[319,394,1990,2012]" italics="true" pageId="21" pageNumber="22">alisana</emphasis>
</taxonomicName>
worker (CMPon045), A: body, lateral view, B: body, dorsal view, C: head, full-face view.
</paragraph>
</caption>
<caption id="B2BC57886F67FF80FF0659DEFAF2CBFF" box="[151,1348,1723,1745]" httpUri="https://zenodo.org/record/2667147/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1723,1745]" targetBox="[167,1420,193,1701]" targetPageId="22">
<paragraph id="E67C07006F67FF80FF0659DEFAF2CBFF" blockId="22.[151,1348,1723,1745]" box="[151,1348,1723,1745]" pageId="22" pageNumber="23">
<emphasis id="D4B7DB126F67FF80FF0659DEFEA9CBFF" bold="true" box="[151,287,1723,1745]" pageId="22" pageNumber="23">FIGURE 32.</emphasis>
<taxonomicName id="21C37C836F67FF80FEB459D9FE3CCBFF" authorityName="Terayama" authorityYear="1986" box="[293,394,1723,1745]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="alisana">
<emphasis id="D4B7DB126F67FF80FEB459D9FE84CBFF" box="[293,306,1724,1745]" italics="true" pageId="22" pageNumber="23">P</emphasis>
.
<emphasis id="D4B7DB126F67FF80FEAE59DEFE3CCBFF" box="[319,394,1723,1745]" italics="true" pageId="22" pageNumber="23">alisana</emphasis>
</taxonomicName>
alate queen (CMPon050), A: body, lateral view, B: body, dorsal view, C: head, full-face view.
</paragraph>
</caption>
<subSubSection id="AED9548B6F67FF80FF56599DFBDBCA8E" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="E67C07006F67FF80FF56599DFBDBCA8E" blockId="22.[151,1436,1784,2025]" pageId="22" pageNumber="23">
<emphasis id="D4B7DB126F67FF80FF56599DFE0ECA3F" bold="true" box="[199,440,1784,1809]" pageId="22" pageNumber="23">Diagnosis (worker).</emphasis>
This large species (HW:
<quantity id="213BAAE56F67FF80FD60599DFC2DCA3E" box="[753,923,1784,1809]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="6.5" metricValueMin="5.8" pageId="22" pageNumber="23" unit="mm" value="0.615" valueMax="0.65" valueMin="0.58">0.580.65 mm</quantity>
) is characterized by a long antennal scape exceeding the posterior margin of the head; a clypeus with acute medial tooth; a subpetiolar process with large teeth; and a deeply incised metanotal groove.
<taxonomicName id="21C37C836F67FF80FD315824FCF8CA77" authorityName="Terayama" authorityYear="1986" box="[672,846,1856,1881]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="alisana">
<emphasis id="D4B7DB126F67FF80FD315824FCF8CA77" box="[672,846,1856,1881]" italics="true" pageId="22" pageNumber="23">Ponera alisana</emphasis>
</taxonomicName>
presents similarities with
<taxonomicName id="21C37C836F67FF80FBE95824FAA8CA77" box="[1144,1310,1856,1881]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nangonshan">
<emphasis id="D4B7DB126F67FF80FBE95824FB30CA76" box="[1144,1158,1857,1880]" italics="true" pageId="22" pageNumber="23">P</emphasis>
.
<emphasis id="D4B7DB126F67FF80FB045825FAA8CA77" box="[1173,1310,1856,1881]" italics="true" pageId="22" pageNumber="23">nangonshan</emphasis>
</taxonomicName>
but can be distinguished from the latter by the long antennal scape, which exceeds the posterior margin of head and the presence of an acute medial tooth on the clypeus (blunt medial tooth in
<taxonomicName id="21C37C836F67FF80FC2958ECFBEBCA8F" box="[952,1117,1928,1953]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nangonshan">
<emphasis id="D4B7DB126F67FF80FC2958ECFC70CA8E" box="[952,966,1929,1952]" italics="true" pageId="22" pageNumber="23">P</emphasis>
.
<emphasis id="D4B7DB126F67FF80FC4558EDFBEBCA8F" box="[980,1117,1928,1953]" italics="true" pageId="22" pageNumber="23">nangonshan</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F67FF8CFF5658C9FB26CB59" lastPageId="26" lastPageNumber="27" pageId="22" pageNumber="23" type="description">
<paragraph id="E67C07006F67FF81FF5658C9FF6ACC45" blockId="22.[151,1436,1784,2025]" lastBlockId="23.[151,1437,151,759]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<emphasis id="D4B7DB126F67FF80FF5658C9FD9CCAEB" bold="true" box="[199,554,1964,1989]" pageId="22" pageNumber="23">Description of worker. Head.</emphasis>
In full-face view, head subrectangular and distinctly longer than broad (CI: 7883), with concave posterior margin, convex lateral margins and strongly rounded posterolateral corners (
<figureCitation id="7EF81B856F67FF81FAFE58B5FF7FCD9E" captionStart="FIGURE 31" captionStartId="21.[151,250,1990,2012]" captionTargetBox="[155,1408,193,1965]" captionTargetId="figure@21.[155,1408,193,1965]" captionTargetPageId="21" captionText="FIGURE 31. P. alisana worker (CMPon045), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667145/files/figure.png" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">Fig. 31C</figureCitation>
). Eye small; composed of a total of 34 indistinct facets. Anterior clypeal margin with strong and acute medial tooth. Masticatory margin of mandible with a series of about 15 indistinct denticles, and three large teeth on the apical part. Antennal scape, when laid backward, exceeding by about 10% of the scape length past the posterodorsal corner; average ratio of the length of antennomeres = 1.19: 1.93: 2.17: 2.26 (n=4).
</paragraph>
<paragraph id="E67C07006F66FF81FF565E13FB51CF31" blockId="23.[151,1437,151,759]" pageId="23" pageNumber="24">
<emphasis id="D4B7DB126F66FF81FF565E13FEFACCA1" bold="true" box="[199,332,374,399]" pageId="23" pageNumber="24">Mesosoma.</emphasis>
Mesosomal dorsum in lateral view almost straight (
<figureCitation id="7EF81B856F66FF81FC265E13FB92CCA1" box="[951,1060,374,399]" captionStart="FIGURE 31" captionStartId="21.[151,250,1990,2012]" captionTargetBox="[155,1408,193,1965]" captionTargetId="figure@21.[155,1408,193,1965]" captionTargetPageId="21" captionText="FIGURE 31. P. alisana worker (CMPon045), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667145/files/figure.png" pageId="23" pageNumber="24">Fig. 31A</figureCitation>
). Pronotum in dorsal view with acutely convex anterior margin, and broadly convex lateral margins (
<figureCitation id="7EF81B856F66FF81FC3A5EFEFBA5CC9D" box="[939,1043,411,436]" captionStart="FIGURE 31" captionStartId="21.[151,250,1990,2012]" captionTargetBox="[155,1408,193,1965]" captionTargetId="figure@21.[155,1408,193,1965]" captionTargetPageId="21" captionText="FIGURE 31. P. alisana worker (CMPon045), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667145/files/figure.png" pageId="23" pageNumber="24">Fig. 31B</figureCitation>
). Metanotal groove in dorsal view and lateral view strongly and broadly incised. Lateral mesopleural suture in lateral view deeply incised. Propodeal dorsum in dorsal view broad with straight lateral margins, narrow anteriad. Posterodorsal corner of propodeum in lateral view angular, propodeal dorsum and declivity forming approximatively a 120 degree angle.
</paragraph>
<paragraph id="E67C07006F66FF81FF565D4EFAFACFD9" blockId="23.[151,1437,151,759]" pageId="23" pageNumber="24">
<emphasis id="D4B7DB126F66FF81FF565D4EFEFCCF6A" bold="true" box="[199,330,555,580]" pageId="23" pageNumber="24">Metasoma.</emphasis>
Petiolar node in dorsal view thick and arched, with slightly convex anterior margin, almost straight posterior margin, and broadly convex lateral margins (
<figureCitation id="7EF81B856F66FF81FD675D2BFCEACF49" box="[758,860,590,615]" captionStart="FIGURE 31" captionStartId="21.[151,250,1990,2012]" captionTargetBox="[155,1408,193,1965]" captionTargetId="figure@21.[155,1408,193,1965]" captionTargetPageId="21" captionText="FIGURE 31. P. alisana worker (CMPon045), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667145/files/figure.png" pageId="23" pageNumber="24">Fig. 31B</figureCitation>
). Petiolar node in lateral view moderately thick and rectangular, with straight anterior and posterior margins, and slightly convex dorsum, posterodorsal corner higher than anterodorsal corner (
<figureCitation id="7EF81B856F66FF81FE515DF3FD9DCF81" box="[448,555,662,687]" captionStart="FIGURE 31" captionStartId="21.[151,250,1990,2012]" captionTargetBox="[155,1408,193,1965]" captionTargetId="figure@21.[155,1408,193,1965]" captionTargetPageId="21" captionText="FIGURE 31. P. alisana worker (CMPon045), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667145/files/figure.png" pageId="23" pageNumber="24">Fig. 31A</figureCitation>
). Subpetiolar process with big and oval fenestra, anteroventral corner blunt, concave ventral margin, and posteroventral corner concave with a pair of developed teeth. Third abdominal tergum distinctly broader than long (ATI: 8589), with straight anterior margin and slightly convex lateral margins.
</paragraph>
<caption id="B2BC57886F66FF81FF065809FC4ACAAC" box="[151,1020,1900,1922]" httpUri="https://zenodo.org/record/2667149/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1900,1922]" targetBox="[167,1420,812,1878]" targetPageId="23">
<paragraph id="E67C07006F66FF81FF065809FC4ACAAC" blockId="23.[151,1020,1900,1922]" box="[151,1020,1900,1922]" pageId="23" pageNumber="24">
<emphasis id="D4B7DB126F66FF81FF065809FE96CAAC" bold="true" box="[151,288,1900,1922]" pageId="23" pageNumber="24">FIGURE 33.</emphasis>
<taxonomicName id="21C37C836F66FF81FEB45808FE3CCAAC" authorityName="Terayama" authorityYear="1986" box="[293,394,1900,1922]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="alisana">
<emphasis id="D4B7DB126F66FF81FEB45808FE84CAAC" box="[293,306,1901,1922]" italics="true" pageId="23" pageNumber="24">P</emphasis>
.
<emphasis id="D4B7DB126F66FF81FEAE5809FE3CCAAC" box="[319,394,1900,1922]" italics="true" pageId="23" pageNumber="24">alisana</emphasis>
</taxonomicName>
alate queen (CMPon050), A: left-forewing, B: left-hindwing.
</paragraph>
</caption>
<caption id="B2BC57886F69FF8EFF0658C4FA3ECA99" box="[151,1416,1953,1975]" httpUri="https://zenodo.org/record/2667151/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1953,1975]" targetBox="[158,1428,193,1932]" targetPageId="24">
<paragraph id="E67C07006F69FF8EFF0658C4FA3ECA99" blockId="24.[151,1416,1953,1975]" box="[151,1416,1953,1975]" pageId="24" pageNumber="25">
<emphasis id="D4B7DB126F69FF8EFF0658C4FEA9CA99" bold="true" box="[151,287,1953,1975]" pageId="24" pageNumber="25">FIGURE 34.</emphasis>
<taxonomicName id="21C37C836F69FF8EFEB458C7FE3CCA99" authorityName="Terayama" authorityYear="1986" box="[293,394,1953,1975]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="alisana">
<emphasis id="D4B7DB126F69FF8EFEB458C7FE84CA99" box="[293,306,1954,1975]" italics="true" pageId="24" pageNumber="25">P</emphasis>
.
<emphasis id="D4B7DB126F69FF8EFEAE58C4FE3CCA99" box="[319,394,1953,1975]" italics="true" pageId="24" pageNumber="25">alisana</emphasis>
</taxonomicName>
male (CMPon049), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: antennae.
</paragraph>
</caption>
<caption id="B2BC57886F68FF8FFF065B30FC7FC945" box="[151,969,1109,1131]" httpUri="https://zenodo.org/record/2667153/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1109,1131]" targetBox="[167,1420,193,1086]" targetPageId="25">
<paragraph id="E67C07006F68FF8FFF065B30FC7FC945" blockId="25.[151,969,1109,1131]" box="[151,969,1109,1131]" pageId="25" pageNumber="26">
<emphasis id="D4B7DB126F68FF8FFF065B30FE96C945" bold="true" box="[151,288,1109,1131]" pageId="25" pageNumber="26">FIGURE 35.</emphasis>
<taxonomicName id="21C37C836F68FF8FFEB45B33FE3CC945" authorityName="Terayama" authorityYear="1986" box="[293,394,1109,1131]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="alisana">
<emphasis id="D4B7DB126F68FF8FFEB45B33FE84C945" box="[293,306,1110,1131]" italics="true" pageId="25" pageNumber="26">P</emphasis>
.
<emphasis id="D4B7DB126F68FF8FFEAE5B30FE3CC945" box="[319,394,1109,1131]" italics="true" pageId="25" pageNumber="26">alisana</emphasis>
</taxonomicName>
male (CMPon049), A: left-forewing, B: right-hindwing.
</paragraph>
</caption>
<paragraph id="E67C07006F68FF8FFF565BFDFA9BC86E" blockId="25.[151,1437,1176,1633]" pageId="25" pageNumber="26">
<emphasis id="D4B7DB126F68FF8FFF565BFDFEF4C99F" bold="true" box="[199,322,1176,1201]" pageId="25" pageNumber="26">Sculpture.</emphasis>
Head densely punctate. Mandible sparsely punctate. Pronotum and mesonotum evenly punctate. Mesopleuron and metapleuron weakly striate. Propodeal dorsum sparsely punctate. Propodeum in lateral view with weakly striate lower portion and evenly punctate upper portion. Propodeal declivity smooth with few punctures. Lateral and anterior faces of petiole evenly punctate, smooth posterior face, dorsum with few punctures. The third and fourth abdominal segments evenly punctate, other segments smooth and shining with few punctures.
</paragraph>
<paragraph id="E67C07006F68FF8FFF565A29FD1DC8FF" blockId="25.[151,1437,1176,1633]" pageId="25" pageNumber="26">
<emphasis id="D4B7DB126F68FF8FFF565A29FEE1C84B" bold="true" box="[199,343,1356,1381]" pageId="25" pageNumber="26">Pubescence.</emphasis>
Head, antennae, mesosoma, petiole, and gaster with evenly distributed short hairs; mesopleuron, metapleuron, propodeum and lateral face of petiole with scattered short hairs. Dorsal and ventral faces of head, anterior margin of clypeus, sides of mandibles, dorsum of petiolar node and gaster with many long erect hairs. Subpetiolar process with a few long erect hairs.
</paragraph>
<paragraph id="E67C07006F68FF8FFF565AB9FB40C8DB" blockId="25.[151,1437,1176,1633]" box="[199,1270,1500,1525]" pageId="25" pageNumber="26">
<emphasis id="D4B7DB126F68FF8FFF565AB9FEA6C8DB" bold="true" box="[199,272,1500,1525]" pageId="25" pageNumber="26">Color.</emphasis>
Body color reddish brown. Mandible, clypeus, antennae, legs, and apex of gaster lighter.
</paragraph>
<paragraph id="E67C07006F68FF8FFF565965FD6BCB8E" blockId="25.[151,1437,1176,1633]" lastBlockId="25.[393,733,1671,1696]" pageId="25" pageNumber="26">
<emphasis id="D4B7DB126F68FF8FFF565965FD96CB37" bold="true" box="[199,544,1536,1561]" pageId="25" pageNumber="26">Description of queen. Head.</emphasis>
(
<figureCitation id="7EF81B856F68FF8FFDA25965FD2ACB36" box="[563,668,1536,1561]" captionStart="FIGURE 32" captionStartId="22.[151,250,1723,1745]" captionTargetBox="[167,1420,193,1701]" captionTargetId="figure@22.[167,1420,193,1701]" captionTargetPageId="22" captionText="FIGURE 32. P. alisana alate queen (CMPon050), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667147/files/figure.png" pageId="25" pageNumber="26">Fig. 32C</figureCitation>
) Similar to worker caste, but with large and oval eye, maximum diameter of each eye about
<quantity id="213BAAE56F68FF8FFE475940FD8DCB12" box="[470,571,1573,1597]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="mm" value="0.2">0.20 mm</quantity>
with 11 ommatidia along the maximum diameter. Three ocelli present, and forming an equilateral triangle. Antennal scape similar to worker; average ratio of the length of antennomeres = 1.20: 1.54: 1.82: 2.07 (n=6).
</paragraph>
<paragraph id="E67C07006F68FF8FFF5659DEFD53CAA6" blockId="25.[151,1437,1723,2036]" pageId="25" pageNumber="26">
<emphasis id="D4B7DB126F68FF8FFF5659DEFEFACBFA" bold="true" box="[199,332,1723,1748]" pageId="25" pageNumber="26">Mesosoma.</emphasis>
Mesosomal dorsum in lateral view broadly convex (
<figureCitation id="7EF81B856F68FF8FFC2A59DEFB91CBFA" box="[955,1063,1723,1748]" captionStart="FIGURE 32" captionStartId="22.[151,250,1723,1745]" captionTargetBox="[167,1420,193,1701]" captionTargetId="figure@22.[167,1420,193,1701]" captionTargetPageId="22" captionText="FIGURE 32. P. alisana alate queen (CMPon050), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667147/files/figure.png" pageId="25" pageNumber="26">Fig. 32A</figureCitation>
). Pronotum in dorsal view with acutely convex anterior margin and broadly convex lateral margins (
<figureCitation id="7EF81B856F68FF8FFC0859BAFBB7CBD9" box="[921,1025,1759,1784]" captionStart="FIGURE 32" captionStartId="22.[151,250,1723,1745]" captionTargetBox="[167,1420,193,1701]" captionTargetId="figure@22.[167,1420,193,1701]" captionTargetPageId="22" captionText="FIGURE 32. P. alisana alate queen (CMPon050), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667147/files/figure.png" pageId="25" pageNumber="26">Fig. 32B</figureCitation>
). Scutum subtrapezoidal and narrow posteriad, with moderately convex anterior margin and straight posterior margin. Transcutal suture distinct. Scutellum oval. Anapleural sulcus distinctly incised. Propodeal-metapleural suture indistinct. Propodeal dorsum in dorsal view broad with straight lateral margins. Propodeal corner in lateral view angular; propodeal dorsum and declivity forming approximatively 115 degree angle.
</paragraph>
<paragraph id="E67C07006F68FF8CFF5658F6FAF0CDFA" blockId="25.[151,1437,1723,2036]" lastBlockId="26.[151,1437,151,1691]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="D4B7DB126F68FF8FFF5658F6FEFCCA82" bold="true" box="[199,330,1939,1964]" pageId="25" pageNumber="26">Metasoma.</emphasis>
Petiolar node in dorsal view thin and broader than long, with broadly convex anterior margin and slightly convex posterior margin (
<figureCitation id="7EF81B856F68FF8FFD8158D2FDC0CAFE" box="[528,630,1975,2000]" captionStart="FIGURE 32" captionStartId="22.[151,250,1723,1745]" captionTargetBox="[167,1420,193,1701]" captionTargetId="figure@22.[167,1420,193,1701]" captionTargetPageId="22" captionText="FIGURE 32. P. alisana alate queen (CMPon050), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667147/files/figure.png" pageId="25" pageNumber="26">Fig. 32B</figureCitation>
). Petiolar node in lateral view thin and trapezoidal, with straight anterior and posterior margins; posterodorsal corner higher than anterodorsal corner (
<figureCitation id="7EF81B856F68FF8FFB9558BEFBD8CADA" box="[1028,1134,2011,2036]" captionStart="FIGURE 32" captionStartId="22.[151,250,1723,1745]" captionTargetBox="[167,1420,193,1701]" captionTargetId="figure@22.[167,1420,193,1701]" captionTargetPageId="22" captionText="FIGURE 32. P. alisana alate queen (CMPon050), A: body, lateral view, B: body, dorsal view, C: head, full-face view." httpUri="https://zenodo.org/record/2667147/files/figure.png" pageId="25" pageNumber="26">Fig. 32A</figureCitation>
). Subpetiolar process with big and oval fenestra, anteroventral corner blunt, distinctly concave ventral margin, and posteroventral corner concave with a pair of developed teeth. The third abdominal segment similar to worker caste (ATI: 8187).
</paragraph>
<paragraph id="E67C07006F6BFF8CFF565FBAFF4CCC33" blockId="26.[151,1437,151,1691]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF565FBAFEF4CDD6" bold="true" box="[199,322,223,248]" pageId="26" pageNumber="27">Sculpture.</emphasis>
Similar to worker caste; except metapleuron and lateral propodeum with increased puncture and striation.
</paragraph>
<paragraph id="E67C07006F6BFF8CFF565E42FEE1CC4B" blockId="26.[151,1437,151,1691]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF565E42FEE1CC6E" bold="true" box="[199,343,295,320]" pageId="26" pageNumber="27">Pubescence.</emphasis>
Similar to worker caste; except metapleuron and lateral propodeum with increased short decumbent hairs.
</paragraph>
<paragraph id="E67C07006F6BFF8CFF565E0AFAC1CCDB" blockId="26.[151,1437,151,1691]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF565E0AFEB9CCA6" bold="true" box="[199,271,367,392]" pageId="26" pageNumber="27">Wing.</emphasis>
Forewing (
<figureCitation id="7EF81B856F6BFF8CFE015E0AFE4FCCA6" box="[400,505,367,392]" captionStart="FIGURE 33" captionStartId="23.[151,250,1900,1922]" captionTargetBox="[167,1420,812,1878]" captionTargetId="figure@23.[167,1420,812,1878]" captionTargetPageId="23" captionText="FIGURE 33. P. alisana alate queen (CMPon050), A: left-forewing, B: left-hindwing." httpUri="https://zenodo.org/record/2667149/files/figure.png" pageId="26" pageNumber="27">Fig. 33A</figureCitation>
): Rsf1 and Mf1 forming a blunt angle; Rsf1 long, Mf1 distinctly straight. Mf2 long and straight, Rs+M juncture Mf2 with distad 1m-cu, cell dc1 subrectangular. 2rs-m juncture with Rsf4 distad 2r-rs; Rsf4 and Rsf23 with triangular gap. Mf1 diverging from M+Cu distad cu-a. Cell smc2 distinctly shorter than cell mc1. Hindwing (
<figureCitation id="7EF81B856F6BFF8CFEC55EB9FE0ACCDA" box="[340,444,476,501]" captionStart="FIGURE 33" captionStartId="23.[151,250,1900,1922]" captionTargetBox="[167,1420,812,1878]" captionTargetId="figure@23.[167,1420,812,1878]" captionTargetPageId="23" captionText="FIGURE 33. P. alisana alate queen (CMPon050), A: left-forewing, B: left-hindwing." httpUri="https://zenodo.org/record/2667149/files/figure.png" pageId="26" pageNumber="27">Fig. 33B</figureCitation>
): A vein reaching wing outer margin, but Rsf, Cuf, Mf almost reaching outer margin.
</paragraph>
<paragraph id="E67C07006F6BFF8CFF565E9AFD95CF39" blockId="26.[151,1437,151,1691]" box="[199,547,511,536]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF565E9AFEA6CF36" bold="true" box="[199,272,511,536]" pageId="26" pageNumber="27">Color.</emphasis>
Similar to worker caste.
</paragraph>
<paragraph id="E67C07006F6BFF8CFF565D41FBE2CE71" blockId="26.[151,1437,151,1691]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF565D41FDB5CF13" bold="true" box="[199,515,548,573]" pageId="26" pageNumber="27">Description of male. Head.</emphasis>
In full-face view, head circular and longer than broad (CI: 88), excluding eye (
<figureCitation id="7EF81B856F6BFF8CFAFF5D41FF7FCF4E" captionStart="FIGURE 34" captionStartId="24.[151,250,1953,1975]" captionTargetBox="[158,1428,193,1932]" captionTargetId="figure@24.[158,1428,193,1932]" captionTargetPageId="24" captionText="FIGURE 34. P. alisana male (CMPon049), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: antennae." httpUri="https://zenodo.org/record/2667151/files/figure.png" pageId="26" pageNumber="27">Fig. 34C</figureCitation>
). Three ocelli present, and forming an equilateral triangle. Compound eye in lateral view big and oval, with convex anterior margin and straight posterior margin, the maximum diameter of each eye about
<quantity id="213BAAE56F6BFF8CFB4B5D09FA89CFAA" box="[1242,1343,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="26" pageNumber="27" unit="mm" value="0.28">0.28 mm</quantity>
with 20 ommatidia along the maximum diameter. Labrum suboval, with convex anterior margin, and as long as the mandible. Palpal formula 4, 2; third maxillary palpomere slightly shorter than fourth maxillary palpomere; second labial palpomere broader than first labial palpomere, and 1/3 times as long as first labial palpomere. Mandible subtriangular with convex medial margin and straight lateral margins, masticatory margin edentate with sharp apex. A total of 13 antennal segments, scape short; the average ratio of the length of antennomeres approximately 1
<superScript id="11B6AA486F6BFF8CFF345C26FF19CE7F" attach="left" box="[165,175,835,849]" fontSize="6" pageId="26" pageNumber="27">st</superScript>
to 13
<superScript id="11B6AA486F6BFF8CFF7E5C26FF4DCE7F" attach="left" box="[239,251,835,849]" fontSize="6" pageId="26" pageNumber="27">th</superScript>
= 1.0: 1.3: 4.1: 3.4: 3.9: 3.7: 3.7: 3.7: 4.2: 3.6: 3.6: 3.6: 5.9 from base (n=1).
</paragraph>
<paragraph id="E67C07006F6BFF8CFF565C0FFAD3C919" blockId="26.[151,1437,151,1691]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF565C0FFEF3CEAD" bold="true" box="[199,325,874,899]" pageId="26" pageNumber="27">Mesosoma</emphasis>
. Mesosomal dorsum in lateral view uneven, with sub-right anterior corner (
<figureCitation id="7EF81B856F6BFF8CFB385C0FFAA5CEAD" box="[1193,1299,874,899]" captionStart="FIGURE 34" captionStartId="24.[151,250,1953,1975]" captionTargetBox="[158,1428,193,1932]" captionTargetId="figure@24.[158,1428,193,1932]" captionTargetPageId="24" captionText="FIGURE 34. P. alisana male (CMPon049), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: antennae." httpUri="https://zenodo.org/record/2667151/files/figure.png" pageId="26" pageNumber="27">Fig. 34A</figureCitation>
). Scutum in dorsal view well-developed and large, with broadly convex anterior margin, slightly convex lateral and posterior margins (
<figureCitation id="7EF81B856F6BFF8CFE975CD7FEC4CEE5" box="[262,370,946,971]" captionStart="FIGURE 34" captionStartId="24.[151,250,1953,1975]" captionTargetBox="[158,1428,193,1932]" captionTargetId="figure@24.[158,1428,193,1932]" captionTargetPageId="24" captionText="FIGURE 34. P. alisana male (CMPon049), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: antennae." httpUri="https://zenodo.org/record/2667151/files/figure.png" pageId="26" pageNumber="27">Fig. 34B</figureCitation>
). Scutum in lateral view broadly convex. Transcutal suture distinct and broad. Scutellum subcircular. Anapleural sulcus distinctly and clearly incised. Propodeal-metapleural suture weakly incised. Propodeal dorsum in dorsal view broad. Propodeal corner in lateral view angular; propodeal dorsum and declivity forming approximatively a 130 degree angle. Each fore, middle and hind leg bearing one pectinate tibial spur.
</paragraph>
<paragraph id="E67C07006F6BFF8CFF565B27FE87C9C5" blockId="26.[151,1437,151,1691]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF565B27FEFCC975" bold="true" box="[199,330,1090,1115]" pageId="26" pageNumber="27">Metasoma.</emphasis>
Petiolar node in dorsal view thin, and broader than long. Petiolar node in lateral view look subtriangular (broken), with straight and sloping anterior and posterior margins (
<figureCitation id="7EF81B856F6BFF8CFBAA5B03FB10C951" box="[1083,1190,1126,1151]" captionStart="FIGURE 34" captionStartId="24.[151,250,1953,1975]" captionTargetBox="[158,1428,193,1932]" captionTargetId="figure@24.[158,1428,193,1932]" captionTargetPageId="24" captionText="FIGURE 34. P. alisana male (CMPon049), A: body, lateral view, B: body, dorsal view, C: head, full-face view, D: antennae." httpUri="https://zenodo.org/record/2667151/files/figure.png" pageId="26" pageNumber="27">Fig. 34A</figureCitation>
). Subpetiolar process subtriangular without fenestra. Third abdominal tergum arched (ATI: 64), and third abdominal sternum prior with a small spine. Helcium broad and crossribbed. Pygidium triangular, apex forming a median spine. Hypopygium subtriangular.
</paragraph>
<paragraph id="E67C07006F6BFF8CFF565B93FDEEC8EC" blockId="26.[151,1437,151,1691]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF565B93FEF4C821" bold="true" box="[199,322,1270,1295]" pageId="26" pageNumber="27">Sculpture.</emphasis>
Head sparely punctate. Pronotum in dorsal view with abundance short striae forming two striped lines, with longitudinal striate on their posterior portion. Posterior portion of scutum with longitudinal striae. Anepisternum evenly punctate. Katepisternum and mesopleuron with scattered striae. Posterodorsal portion of propodeum evenly punctate. Propodeal declivity smooth with few punctures. Dorsal face of petiole smooth evenly punctate, smooth posterior face. The third and fourth abdominal segments with sparsely punctate, other segments smooth and shining with few punctures.
</paragraph>
<paragraph id="E67C07006F6BFF8CFF565AABFE7ECB01" blockId="26.[151,1437,151,1691]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF565AABFEE1C8C9" bold="true" box="[199,343,1486,1511]" pageId="26" pageNumber="27">Pubescence.</emphasis>
Body with evenly distributed short hairs. Dorsal and ventral faces of head, anterior margin of clypeus, sides of mandibles, dorsum of petiolar node and gaster with many long erect hairs. Subpetiolar process with a few long erect hairs.
</paragraph>
<paragraph id="E67C07006F6BFF8CFF56595FFB1BCB7D" blockId="26.[151,1437,151,1691]" box="[199,1197,1594,1619]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF56595FFEB9CB7D" bold="true" box="[199,271,1594,1619]" pageId="26" pageNumber="27">Wing.</emphasis>
Similar to gyne caste, except for Cu1f1 slightly convex in forewing (
<figureCitation id="7EF81B856F6BFF8CFB89595FFBC3CB7D" box="[1048,1141,1594,1619]" captionStart="FIGURE 33" captionStartId="23.[151,250,1900,1922]" captionTargetBox="[167,1420,812,1878]" captionTargetId="figure@23.[167,1420,812,1878]" captionTargetPageId="23" captionText="FIGURE 33. P. alisana alate queen (CMPon050), A: left-forewing, B: left-hindwing." httpUri="https://zenodo.org/record/2667149/files/figure.png" pageId="26" pageNumber="27">Figs. 33</figureCitation>
,
<figureCitation id="7EF81B856F6BFF8CFB10595FFB16CB7D" box="[1153,1184,1594,1619]" captionStart="FIGURE 35" captionStartId="25.[151,250,1109,1131]" captionTargetBox="[167,1420,193,1086]" captionTargetId="figure@25.[167,1420,193,1088]" captionTargetPageId="25" captionText="FIGURE 35. P. alisana male (CMPon049), A: left-forewing, B: right-hindwing." httpUri="https://zenodo.org/record/2667153/files/figure.png" pageId="26" pageNumber="27">35</figureCitation>
).
</paragraph>
<paragraph id="E67C07006F6BFF8CFF56593BFB26CB59" blockId="26.[151,1437,151,1691]" box="[199,1168,1630,1656]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF56593BFEA6CB59" bold="true" box="[199,272,1630,1655]" pageId="26" pageNumber="27">Color.</emphasis>
Body color reddish brown; mandible, antennae, legs, and apex of gaster lighter.
</paragraph>
</subSubSection>
<subSubSection id="AED9548B6F6BFF8CFF5659E7FBA9CBB5" box="[199,1055,1666,1691]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="E67C07006F6BFF8CFF5659E7FBA9CBB5" blockId="26.[151,1437,151,1691]" box="[199,1055,1666,1691]" pageId="26" pageNumber="27">
<emphasis id="D4B7DB126F6BFF8CFF5659E7FED6CBB5" bold="true" box="[199,352,1666,1691]" pageId="26" pageNumber="27">Distribution.</emphasis>
<collectingCountry id="9ED447906F6BFF8CFEF659E6FE11CBB4" box="[359,423,1667,1690]" name="Japan" pageId="26" pageNumber="27">Japan</collectingCountry>
(Yakushima Island),
<collectingCountry id="9ED447906F6BFF8CFD0759E7FD5ECBB5" box="[662,744,1666,1691]" name="Taiwan" pageId="26" pageNumber="27">Taiwan</collectingCountry>
(Chiyai county: Fenchihu).
</paragraph>
</subSubSection>
</treatment>
</document>