treatments-xml/data/8C/38/1B/8C381B2A3AEE5E7D880458FDDE54FCD9.xml

<document ID-DOI="http://dx.doi.org/10.3897/herpetozoa.36.e106373" ID-Pensoft-Pub="2682-955X-36-177" ID-Pensoft-UUID="28C510E372505DCD94A4467DA672ED94" ID-ZooBank="FAB9FAE431F24084ABB4A38447A72783" ModsDocID="2682-955X-36-177" checkinTime="1689838388098" checkinUser="pensoft" docAuthor="Liu, Shuo, Hou, Mian, Wang, Qiaoyan, Rao, Dingqi &amp; Li, Song" docDate="2023" docId="8C381B2A3AEE5E7D880458FDDE54FCD9" docLanguage="en" docName="Herpetozoa 36: 177-188" docOrigin="Herpetozoa 36" docPubDate="2023-07-19" docSource="http://dx.doi.org/10.3897/herpetozoa.36.e106373" docTitle="Odorrana heatwolei" docType="treatment" docVersion="2" id="28C510E372505DCD94A4467DA672ED94" lastPageNumber="177" masterDocId="28C510E372505DCD94A4467DA672ED94" masterDocTitle="Resurrection and distribution extension of Odorrana heatwolei (Stuart &amp; Bain, 2005) (Anura, Ranidae)" masterLastPageNumber="188" masterPageNumber="177" pageNumber="177" updateTime="1689838689063" updateUser="ExternalLinkService">
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<mods:title id="C412E8D4E569792DC48F8D261DECD0C5">Resurrection and distribution extension of Odorrana heatwolei (Stuart &amp; Bain, 2005) (Anura, Ranidae)</mods:title>
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<mods:namePart id="2C96194DD9FF8AEDE00FD20C9BDC0A97">Liu, Shuo</mods:namePart>
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<mods:affiliation id="C74A65293957830994C2FE6E5F345F55">Kunming Natural History Museum of Zoology, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China</mods:affiliation>
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<mods:namePart id="C5DDF47B341771A81510E12931000126">Hou, Mian</mods:namePart>
<mods:affiliation id="714DE7B239AAC1500EADAB8607399493">Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
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<mods:namePart id="00CDDE1EDA04C4D3D2EFB90C0B44339F">Wang, Qiaoyan</mods:namePart>
<mods:affiliation id="B094F84947806AE43CFC4854518DC3DA">College of Continuing (Online) Education, Sichuan Normal University, Chengdu, Sichuan 610066, China</mods:affiliation>
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<mods:namePart id="35DD9E31BF3572D3A91423FD4D73185F">Rao, Dingqi</mods:namePart>
<mods:affiliation id="080F92386C7640EC0F11EF564C5FC8F7">Kunming Natural History Museum of Zoology, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China</mods:affiliation>
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<mods:namePart id="E5FD98AD4B8310B196011FE620E821ED">Li, Song</mods:namePart>
<mods:affiliation id="A251DD8B6BD8DE326E180B0B7F71BC32">Kunming Natural History Museum of Zoology, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China</mods:affiliation>
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<mods:number id="BBFF6D792E447A45C68773877DEC6C0D">36</mods:number>
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<treatment id="8C381B2A3AEE5E7D880458FDDE54FCD9" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9" httpUri="http://treatment.plazi.org/id/8C381B2A3AEE5E7D880458FDDE54FCD9" lastPageNumber="177" pageId="0" pageNumber="177">
<subSubSection id="B722595F3DAD9473BB3F439D566B8F43" pageId="0" pageNumber="177" type="nomenclature">
<paragraph id="BE2E870989033B9F687D0EFE365273CE" pageId="0" pageNumber="177">
<taxonomicName id="E6B0A70A794FC61E0D755D61D32C0EFD" ID-CoL="4RDS4" LSID="8C381B2A-3AEE-5E7D-8804-58FDDE54FCD9" authority="(Stuart &amp; Bain, 2005)" baseAuthorityName="Stuart &amp; Bain" baseAuthorityYear="2005" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana heatwolei" order="Anura" pageId="0" pageNumber="177" phylum="Chordata" rank="species" species="heatwolei">Odorrana heatwolei (Stuart &amp; Bain, 2005)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A29B4200D0ECB1A3D832F313226EFFB2" pageId="0" pageNumber="177" type="description">
<paragraph id="63D613558403A2C0D446A406D2F19B98" pageId="0" pageNumber="177">
<figureCitation id="CF17A030262479BBF195AA3FC3FD7D4C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Comparison between Odorrana heatwolei and O. tiannanensis in life. A. Adult male O. heatwolei from Mengla, Yunnan, China; B. Adult male O. tiannanensis from Hekou, Yunnan, China; C. Adult female O. heatwolei from Mengla, Yunnan, China; D. Adult female O. tiannanensis from Hekou, Yunnan, China." figureDoi="10.3897/herpetozoa.36.e106373.figure3" httpUri="https://binary.pensoft.net/fig/880822" pageId="0" pageNumber="177">Figs 3A, C</figureCitation>
<figureCitation id="D906FE28D7B26840188D469F43346504" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal view (A), lateral view (B), and ventral view (C) of the male (KIZ 20194255) and dorsal view (D), lateral view (E), and ventral view (F) of an adult female (escaped) of Odorrana heatwolei from China in life." figureDoi="10.3897/herpetozoa.36.e106373.figure4" httpUri="https://binary.pensoft.net/fig/880823" pageId="0" pageNumber="177">, 4</figureCitation>
<figureCitation id="F4C178EF1B2DC03B30E6ED325520A6A1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsal view (A) and ventral view (B) of the specimens of Odorrana heatwolei from China in preservative." figureDoi="10.3897/herpetozoa.36.e106373.figure6" httpUri="https://binary.pensoft.net/fig/880825" pageId="0" pageNumber="177">, 6</figureCitation>
<figureCitation id="433644CD5FBA9DE529F800DEAB7BBAFA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Close-up views of the head (A), vola (B), and planta (C) of the male (KIZ 2019541) and close-up views of the head (D), vola (E), and planta (F) of the female (KIZ 2019542) of Odorrana heatwolei from China in preservative." figureDoi="10.3897/herpetozoa.36.e106373.figure7" httpUri="https://binary.pensoft.net/fig/880826" pageId="0" pageNumber="177">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5E060226283693E0FA50FE9AE32DF4D9" pageId="0" pageNumber="177" type="reference_group">
<paragraph id="C5B2DB2A4E423121E2F073C4DACC686B" pageId="0" pageNumber="177">
<taxonomicName id="2C83AD5CCE80AC12BCEC4743E6BCA463" authorityName="Stuart &amp; Bain" authorityYear="2005" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana heatwolei" order="Anura" pageId="0" pageNumber="177" phylum="Chordata" rank="species" species="heatwolei">Rana heatwolei</taxonomicName>
Stuart &amp; Bain, 2005
</paragraph>
<paragraph id="A9975E5C3518F3973399DF777D817678" pageId="0" pageNumber="177">
<taxonomicName id="9ED4BA5539B06E1C4DDE7C22CCD05F34" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana tiannanensis" order="Anura" pageId="0" pageNumber="177" phylum="Chordata" rank="species" species="tiannanensis">Rana tiannanensis</taxonomicName>
- Ohler, 2007
</paragraph>
<paragraph id="41F6CED7216A0049DA3929B7CCA3A76B" pageId="0" pageNumber="177">
<taxonomicName id="E5C90AF4399FF36649BC2658CD3F25A1" baseAuthorityName="Stuart &amp; Bain" baseAuthorityYear="2005" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana heatwolei" order="Anura" pageId="0" pageNumber="177" phylum="Chordata" rank="species" species="heatwolei">Odorrana heatwolei</taxonomicName>
- 
<bibRefCitation id="D2CBCA18AAF772AEACAFCBBB521005F0" DOI="https://doi.org/10.30906/1026-2296-2021-28-3A-1-110" author="Poyarkov, NA" journalOrPublisher="Alytes" pageId="0" pageNumber="177" refId="B14" refString="Poyarkov, NA, Nguyen, TV, Popov, ES, Geissler, P, Pawangkhanant, P, Neang, T, Suwannapoom, C, Orlov, NL, 2021. Recent progress in taxonomic studies, biogeographic analysis, and revised checklist of amphibians of Indochina. Russian Journal of Herpetology 28(3A): 1-110. https://doi.org/10.30906/1026-2296-2021-28-3A-1-110" title="Recent progress in taxonomic studies, biogeographic analysis, and revised checklist of amphibians of Indochina. Russian Journal of Herpetology 28 (3 A): 1 - 110." url="https://doi.org/10.30906/1026-2296-2021-28-3A-1-110" year="2021">Poyarkov et al., 2021</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="FA4432BAF1815978954D159D02083E77" pageId="0" pageNumber="177" type="type material">
<paragraph id="B4021EFE3141A0BEC4C903B4ECAC070E" pageId="0" pageNumber="177">Type material.</paragraph>
<paragraph id="F366251AA60252D8573C670B77F2F51B" pageId="0" pageNumber="177">
<emphasis id="D86C71F79B7CD09C70CDEB93F4C25D4D" bold="true" italics="true" pageId="0" pageNumber="177">Holotype</emphasis>
: FMNH 258134, adult male.
</paragraph>
<paragraph id="ED0054691181F90E3E676FE6EA880B61" pageId="0" pageNumber="177">
<emphasis id="40C01BA2D0867CACC3E5B23442BD4366" bold="true" pageId="0" pageNumber="177">
<emphasis id="322E1FA6BF2836CACF1804BF1E666484" italics="true" pageId="0" pageNumber="177">Paratypes</emphasis>
.
</emphasis>
FMNH 258279-83, 258598, six females; FMNH 258135-39, five males.
</paragraph>
</subSubSection>
<subSubSection id="F98E7830BCA6063B11A20E1C23015148" pageId="0" pageNumber="177" type="type locality">
<paragraph id="AED2C6C838938A8A2D4CA5A35ED12FE3" pageId="0" pageNumber="177">Type locality.</paragraph>
<paragraph id="9807E79DC5D2780459559C11433F6313" pageId="0" pageNumber="177">
Phou Dendin National Biodiversity Conservation Area, Phongsaly District, Phongsaly Province, Laos (
<geoCoordinate id="91E8D87913351BBA623CFB1B74A12F34" degrees="22" direction="north" minutes="05" orientation="latitude" precision="15" seconds="38" value="22.09389">22°05'38&quot;N</geoCoordinate>
, 
<geoCoordinate id="AA07B509C73EF51452B0B9D42E65D48E" degrees="102" direction="east" minutes="12" orientation="longitude" precision="15" seconds="50" value="102.21388">102°12'50&quot;E</geoCoordinate>
; alt. 600 m).
</paragraph>
<caption id="30E873D2F0537334D3E6CDA333DFB096" doi="10.3897/herpetozoa.36.e106373.figure6" httpUri="https://binary.pensoft.net/fig/880825" pageId="0" pageNumber="177" start="Figure 6" startId="F6">
<paragraph id="D0058CFC0423588072F2DD41D80A17CF" pageId="0" pageNumber="177">
<emphasis id="8385EF3994DDA7E19C3356740852193B" bold="true" pageId="0" pageNumber="177">Figure 6.</emphasis>
Dorsal view (
<emphasis id="9D2BDABF268B677D595E82DAF312874B" bold="true" pageId="0" pageNumber="177">A</emphasis>
) and ventral view (
<emphasis id="5014869F086D114992119FFFD1C30DAE" bold="true" pageId="0" pageNumber="177">B</emphasis>
) of the specimens of 
<taxonomicName id="9AA5F2EF8F7008AA2AD4938B187A0DF6" baseAuthorityName="Stuart &amp; Bain" baseAuthorityYear="2005" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana heatwolei" order="Anura" pageId="0" pageNumber="177" phylum="Chordata" rank="species" species="heatwolei">
<emphasis id="71A63DE5168BA82CE59EF770E3052008" italics="true" pageId="0" pageNumber="177">Odorrana heatwolei</emphasis>
</taxonomicName>
from China in preservative.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7885A08390E4BFBC89165B7EF5A787F5" pageId="0" pageNumber="177" type="etymology">
<paragraph id="265917418A58717805C4313FC3389841" pageId="0" pageNumber="177">Etymology.</paragraph>
<paragraph id="E54363C8034303EC858640A375EBDCA3" pageId="0" pageNumber="177">
The specific epithet is a patronym for Dr. Harold F. Heatwole. We propose &quot;Dense-spotted Odorous Frog&quot; for the English name and 
<normalizedToken id="CB8588F36EBC7A0B81219B577EE5F4F0" originalValue="“麻点臭蛙”">&quot;麻点臭蛙&quot;</normalizedToken>
(
<normalizedToken id="90261AB9715B3701EF3D9543DE56F6C1" originalValue="má">ma</normalizedToken>
<normalizedToken id="E3CEFBE85CCF37E7A74F911CE4BFFA51" originalValue="diǎn">diǎn</normalizedToken>
<normalizedToken id="D47E183476E9D1529D4EA12C5D031FF1" originalValue="chòu">chou</normalizedToken>
<normalizedToken id="6B6EADADC2A65CE7F643C152B9644B9C" originalValue="wā">wa</normalizedToken>
) for the Chinese name.
</paragraph>
<caption id="513229C8A1D8885EA36FB246774A26FC" doi="10.3897/herpetozoa.36.e106373.figure7" httpUri="https://binary.pensoft.net/fig/880826" pageId="0" pageNumber="177" start="Figure 7" startId="F7">
<paragraph id="BE57B705FBB519BE7AD516D20E66901F" pageId="0" pageNumber="177">
<emphasis id="9E7753741E9742634140998E035BC6E6" bold="true" pageId="0" pageNumber="177">Figure 7.</emphasis>
Close-up views of the head (
<emphasis id="B89E5F8FAFED7734942D1353B56BDB16" bold="true" pageId="0" pageNumber="177">A</emphasis>
), vola (
<emphasis id="8FD39741FB44266A31998A3631C121C2" bold="true" pageId="0" pageNumber="177">B</emphasis>
), and planta (
<emphasis id="3FB738E79EA4ECC8EC882060D8A74B3F" bold="true" pageId="0" pageNumber="177">C</emphasis>
) of the male (KIZ2019541) and close-up views of the head (
<emphasis id="8021C5486B97FFC79FC057E1F24F266D" bold="true" pageId="0" pageNumber="177">D</emphasis>
), vola (
<emphasis id="1C549F1462F8240751557E340450616C" bold="true" pageId="0" pageNumber="177">E</emphasis>
), and planta (
<emphasis id="9E3CF6B57D25EA2DD56F3A87615218B7" bold="true" pageId="0" pageNumber="177">F</emphasis>
) of the female (KIZ2019542) of 
<taxonomicName id="FD2269B6DA5C5A929574636A4C77C3FE" baseAuthorityName="Stuart &amp; Bain" baseAuthorityYear="2005" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana heatwolei" order="Anura" pageId="0" pageNumber="177" phylum="Chordata" rank="species" species="heatwolei">
<emphasis id="AD9481A1863BA4F74C3F379FC3BCCA13" italics="true" pageId="0" pageNumber="177">Odorrana heatwolei</emphasis>
</taxonomicName>
from China in preservative.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4FE9D4C4F938343700B78D2660DE94A8" pageId="0" pageNumber="177" type="updated diagnosis">
<paragraph id="0455BAA461B5258C34C79E7E8676ED51" pageId="0" pageNumber="177">Updated diagnosis.</paragraph>
<paragraph id="18D5BC23FD1679D1068F44671568F648" pageId="0" pageNumber="177">Habitus robust; body sizes of females approximately two to three times that of males, SVL 37.1-57.3 mm in adult males, 97.3-106.7 mm in adult females; head length slightly greater than head width; dorsal skin shagreened, skin on flanks granular; supratympanic fold present; dorsolateral folds distinct in males and indistinct in females; tympanum large in males (TD/ED 0.58-0.73) and relatively small in females (TD/ED 0.45-0.58); vomerine teeth distinct; relative lengths of fingers III&gt; IV&gt; I&gt; II; inner metacarpal tubercle and outer metacarpal tubercle present, outer metatarsal tubercle absent; all finger and toe tips slightly expanded; toes with entire webbing to disks; uniform light brown or dark brown on whole body, dense tiny black dots scattered on dorsum; paired external vocal sacs and nuptial pad on the base of finger I present in adult males.</paragraph>
</subSubSection>
<subSubSection id="AF4A3E977972E1DFA0E6FABC4580C881" pageId="0" pageNumber="177" type="materials_examined">
<paragraph id="0693AD9E12CDB42F0D9864888CCDE9B2" pageId="0" pageNumber="177">Specimens examined.</paragraph>
<paragraph id="2302B1A4CB70B7A183E30AEAF3862D95" pageId="0" pageNumber="177">
<materialsCitation id="FC658EE1E65ED66E55AC4441EC428E59" collectingDate="2019-04-24" collectingDateMax="2019-05-04" collectingDateMin="2019-04-24" country="China" county="Mengla County" elevation="780" latitude="21.916388" location="Mandan village" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" specimenCount="three" specimenCount-adult="2" stateProvince="Yunnan Province">
KIZ20194255, adult male, and KIZ20194256-KIZ20194258, 
<specimenCount id="05D688F688DA183245EC3DD5C70D98CD" type="adult">three adult</specimenCount>
females, collected on 
<collectingDate id="5F5626187A4271197D42182B4FE9BE78" value="2019-04-24">24 April 2019</collectingDate>
from 
<location id="8AE76251853B3C1777973E6A8E28747A" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:8AE76251853B3C1777973E6A8E28747A" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Mandan village" stateProvince="Yunnan Province">Mandan village</location>
, 
<location id="FCECA42B2B0D40532DE455B71760640C" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:FCECA42B2B0D40532DE455B71760640C" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Mengla Town" stateProvince="Yunnan Province">Mengla Town</location>
, 
<collectingCounty id="7D6D7722BEAADBF86BC0108E9159C447">Mengla County</collectingCounty>
, 
<location id="B6604DC2C84448BAB723B9D873D7EECA" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:B6604DC2C84448BAB723B9D873D7EECA" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Xishuangbanna Prefecture" stateProvince="Yunnan Province">Xishuangbanna Prefecture</location>
, 
<collectingRegion id="FFF8865973B6654000C703FEDCABCA41" country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCountry id="1CD60C2A04FB7DDDE48981BD05316FEF" name="China">China</collectingCountry>
(
<geoCoordinate id="591054B689179C4A4A5FF7FE77350349" degrees="21" direction="north" minutes="28" orientation="latitude" precision="15" seconds="59" value="21.483055">21°28'59&quot;N</geoCoordinate>
, 
<geoCoordinate id="264401396F6968DB5F8ED0E2D8EBE4AC" degrees="101" direction="east" minutes="40" orientation="longitude" precision="15" seconds="29" value="101.67472">101°40'29&quot;E</geoCoordinate>
, 
<quantity id="D63B859F00D4061C6A4F564E6601DAB4" metricMagnitude="2" metricUnit="m" metricValue="7.8" unit="m" value="780.0">
<elevation id="D58B5EDCD37B5B0837ADA1E142362678" metricMagnitude="2" metricUnit="m" metricValue="7.8" unit="m" value="780.0">780 m</elevation>
</quantity>
elevation) by 
<location id="5DFC0DA64BAF432C6015818BFE7D5AD7" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:5DFC0DA64BAF432C6015818BFE7D5AD7" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Shuo Liu" stateProvince="Yunnan Province">Shuo Liu</location>
; KIZ20194279, adult female, collected on 
<collectingDate id="9B52771097B13B20C54BEFF689434668" value="2019-04-27">27 April 2019</collectingDate>
from 
<location id="1B479F2EFBCAFA2DE1953B9F5850CBA8" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:1B479F2EFBCAFA2DE1953B9F5850CBA8" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Mengyuan village" stateProvince="Yunnan Province">Mengyuan village</location>
, 
<location id="6AC4947555CED2A69911F5F43E86BF32" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:6AC4947555CED2A69911F5F43E86BF32" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Guanlei Town" stateProvince="Yunnan Province">Guanlei Town</location>
, 
<location id="49A8F09F872A85C456175E2E923FED09" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:49A8F09F872A85C456175E2E923FED09" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Mengla County" stateProvince="Yunnan Province">Mengla County</location>
, 
<location id="E3D0457F51AD0396AFBAA4B6B749B2C2" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:E3D0457F51AD0396AFBAA4B6B749B2C2" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Xishuangbanna Prefecture" stateProvince="Yunnan Province">Xishuangbanna Prefecture</location>
, 
<collectingRegion id="C266F615435C840951D64E1C68F055FE" country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCountry id="625F0E31F6BF7A48332A0BDABDB4716F" name="China">China</collectingCountry>
(
<geoCoordinate id="3113F43B5C80352F705AA466F57CEC08" degrees="21" direction="north" minutes="38" orientation="latitude" precision="15" seconds="35" value="21.643055">21°38'35&quot;N</geoCoordinate>
, 
<geoCoordinate id="8DCFF310F91DECF508DAE543998F2F6F" degrees="101" direction="east" minutes="26" orientation="longitude" precision="15" seconds="54" value="101.44833">101°26'54&quot;E</geoCoordinate>
, 
<quantity id="4C867C26C4FFB010228D698640C8D161" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">
<elevation id="41B96F1A0F2AF604F498F2F765162CBC" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">950 m</elevation>
</quantity>
elevation) by 
<location id="E0C2ACC2073AF225AAE083D8CFCFB3EC" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:E0C2ACC2073AF225AAE083D8CFCFB3EC" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Shuo Liu" stateProvince="Yunnan Province">Shuo Liu</location>
; KIZ20194301-KIZ20194302, 
<specimenCount id="4F0B7A8C7B22C013C28D95552942714D" type="adult">two adult</specimenCount>
males, collected on 
<collectingDate id="85C17C32CF39D4C488D7318DB34FB5B8" value="2019-04-30">30 April 2019</collectingDate>
from 
<location id="66EC022B6DDA2C12D8AC6E218306F748" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:66EC022B6DDA2C12D8AC6E218306F748" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Bubeng village" stateProvince="Yunnan Province">Bubeng village</location>
, 
<location id="EE7B398BC974390EFA837051ED1F963C" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:EE7B398BC974390EFA837051ED1F963C" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Mengla Town" stateProvince="Yunnan Province">Mengla Town</location>
, 
<location id="07698879FC85F9ABA785C4890F240696" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:07698879FC85F9ABA785C4890F240696" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Mengla County" stateProvince="Yunnan Province">Mengla County</location>
, 
<location id="48B160FBADAAA751655F6D74957B221A" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:48B160FBADAAA751655F6D74957B221A" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Xishuangbanna Prefecture" stateProvince="Yunnan Province">Xishuangbanna Prefecture</location>
, 
<collectingRegion id="DABEB47A410EE8633910AEB2369F7793" country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCountry id="41B19F53DF690879F25B90751A724174" name="China">China</collectingCountry>
(
<geoCoordinate id="67488CAEE607EC80C1C210D702F3654C" degrees="21" direction="north" minutes="36" orientation="latitude" precision="15" seconds="21" value="21.605833">21°36'21&quot;N</geoCoordinate>
, 
<geoCoordinate id="5BFE08D320B119C5164195146C942E0B" degrees="101" direction="east" minutes="33" orientation="longitude" precision="15" seconds="27" value="101.5575">101°33'27&quot;E</geoCoordinate>
, 
<quantity id="85D1CA2C9BBD86FC8E24E16F356E6BF8" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<elevation id="BCCEF2E62110D353A4CE66EF16752DF6" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
</quantity>
elevation) by 
<location id="B8BCE7A7A3CCD86924CB71A414305279" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:B8BCE7A7A3CCD86924CB71A414305279" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Shuo Liu" stateProvince="Yunnan Province">Shuo Liu</location>
; KIZ2019541, adult male, and KIZ2019542, adult female, collected on 
<collectingDate id="6BB455610A694BA75F6B7BB45F098632" value="2019-05-04">4 May 2019</collectingDate>
from 
<location id="2172A4EA173B7000183C7864E85BB3A2" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:2172A4EA173B7000183C7864E85BB3A2" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Xinshan village" stateProvince="Yunnan Province">Xinshan village</location>
, 
<location id="EC8FAE6F8A183F6F69739AACC61D9195" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:EC8FAE6F8A183F6F69739AACC61D9195" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Yaoqu Township" stateProvince="Yunnan Province">Yaoqu Township</location>
, 
<location id="57186AF0D6CDA7D1690534009B296221" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:57186AF0D6CDA7D1690534009B296221" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Mengla County" stateProvince="Yunnan Province">Mengla County</location>
, 
<location id="05255FD38777B85671116061D5C73E38" LSID="urn:lsid:plazi:treatment:8C381B2A3AEE5E7D880458FDDE54FCD9:05255FD38777B85671116061D5C73E38" country="China" county="Mengla County" latitude="21.916388" longLatPrecision="21" longitude="101.53778" municipality="Shuo Liu" name="Xishuangbanna Prefecture" stateProvince="Yunnan Province">Xishuangbanna Prefecture</location>
, 
<collectingRegion id="2666A8C9609396DE6095A1AC5A89447E" country="China" name="Yunnan">Yunnan Province</collectingRegion>
, 
<collectingCountry id="A35DC45F7183D26630783FA4A0161C59" name="China">China</collectingCountry>
(
<geoCoordinate id="FFCB3C99F3C866B0061BA277D1B34C43" degrees="21" direction="north" minutes="54" orientation="latitude" precision="15" seconds="59" value="21.916388">21°54'59&quot;N</geoCoordinate>
, 
<geoCoordinate id="E52305F97082DBE6B7470920827C9101" degrees="101" direction="east" minutes="32" orientation="longitude" precision="15" seconds="16" value="101.53778">101°32'16&quot;E</geoCoordinate>
, 
<quantity id="72EB0277E4ABA7A9008E8392B04CA82B" metricMagnitude="2" metricUnit="m" metricValue="8.4" unit="m" value="840.0">
<elevation id="B17FB9DA8321D7B1D0A2ABAEA35FCC67" metricMagnitude="2" metricUnit="m" metricValue="8.4" unit="m" value="840.0">840 m</elevation>
</quantity>
elevation) by 
<collectingMunicipality id="41442D6135871DB3DAA89DA650C2E9C2">Shuo Liu</collectingMunicipality>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3A1744E2310D129A03342817DC63F4DD" pageId="0" pageNumber="177" type="description">
<paragraph id="22EB4A67699D9E14620859F7D2BA9EB2" pageId="0" pageNumber="177">Description of the specimens from China.</paragraph>
<paragraph id="4FE1E628483739DCB557A4E0658A2692" pageId="0" pageNumber="177">
Morphometric and meristic data are presented in Table 
<tableCitation id="2B61C9D432636A41ECD7498F699EEB7C" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphological measurements (mm) of the specimens of Odorrana heatwolei from China." httpUri="http://table.plazi.org/id/C33B2203AACD2D4E813D4D42FA6186B4" pageId="0" pageNumber="177" tableUuid="C33B2203AACD2D4E813D4D42FA6186B4">3</tableCitation>
. SVL 46.8-51.7 mm in males, 97.4-106.7 mm in females; head width slightly shorter than length (HW/HL 0.87-0.91 in males, 0.93-0.98 in females); snout obtuse, pointed in dorsal view and rounded in profile, obviously projecting beyond lower jaw; nostril closer to snout than to eye; canthus rostralis distinct; loreal region concave; internarial distance greater than interorbital distance; snout length greater than eye diameter; pineal body distinct; supratympanic fold horizontal and straight; tympanum large in males (TD/ED 0.66-0.73) and relatively small in females (TD/ED 0.48-0.58), round and transparent; vomerine teeth developed into mass on two oblique ridges between the two choanae; tongue cordiform, posterior notch enlarged and formed as U-shaped; vocal sac openings on floor of mouth in each corner and an external vocal sac behind each angle of mouth in males.
</paragraph>
<caption id="C33B2203AACD2D4E813D4D42FA6186B4" ID-Table-UUID="C33B2203AACD2D4E813D4D42FA6186B4" httpUri="http://table.plazi.org/id/C33B2203AACD2D4E813D4D42FA6186B4" pageId="0" pageNumber="177" start="Table 3" startId="T3">
<paragraph id="72F1FE678D7FF57E84701B0844DCD053" pageId="0" pageNumber="177">
<emphasis id="6E2B5055EDF3BBFA1A6E270466876466" bold="true" pageId="0" pageNumber="177">Table 3.</emphasis>
Morphological measurements (mm) of the specimens of 
<taxonomicName id="6C866CEA44915842C1C54D622AB80515" baseAuthorityName="Stuart &amp; Bain" baseAuthorityYear="2005" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana heatwolei" order="Anura" pageId="0" pageNumber="177" phylum="Chordata" rank="species" species="heatwolei">
<emphasis id="1242FB055EEC37F4954679FAB8EBED0E" italics="true" pageId="0" pageNumber="177">Odorrana heatwolei</emphasis>
</taxonomicName>
from China.
</paragraph>
</caption>
<paragraph id="BCBEBABF5DDEE6A994AFF3D68B77B241" pageId="0" pageNumber="177">
<table id="43A1BAE181A8714D30CF13138C7FE5D9" pageId="0" pageNumber="177">
<tr id="A90477A8ECAA0609A2D794CA9851B0A4" pageId="0" pageNumber="177">
<th id="B00678CD48E90B58118D26A11F392251" colspan="1" isEmpty="true" pageId="0" pageNumber="177" rowspan="1">-</th>
<th id="6DE8A0C4E1DA5278217EAAE59267325E" colspan="1" pageId="0" pageNumber="177" rowspan="1">KIZ2019541 Male</th>
<th id="3A757675AC91C66696E2A1A71505C176" colspan="1" pageId="0" pageNumber="177" rowspan="1">KIZ20194301 Male</th>
<th id="7F476B403DA8E38CE2770CE788AC5C1A" colspan="1" pageId="0" pageNumber="177" rowspan="1">KIZ20194302 Male</th>
<th id="25097E03C0E8FDDAC421B472DBA9506C" colspan="1" pageId="0" pageNumber="177" rowspan="1">KIZ20194255 Male</th>
<th id="FB7CE30CB359589537D1621912E4B59B" colspan="1" pageId="0" pageNumber="177" rowspan="1">KIZ20194256 Female</th>
<th id="8AC4580C144750902357F2C35900E424" colspan="1" pageId="0" pageNumber="177" rowspan="1">KIZ20194257 Female</th>
<th id="2EEEF0331DD2A072924352503C81D953" colspan="1" pageId="0" pageNumber="177" rowspan="1">KIZ20194258 Female</th>
<th id="D8D0B1F23A1036322AFC9952EF1423A0" colspan="1" pageId="0" pageNumber="177" rowspan="1">KIZ20194279 Female</th>
<th id="519D5525D5E727FBEE9C8D00C83940E6" colspan="1" pageId="0" pageNumber="177" rowspan="1">KIZ2019542 Female</th>
</tr>
<tr id="68DCCD97E71916174354B2F685FACA8B" pageId="0" pageNumber="177">
<td id="06961316772BAE99BF5403A8C6626715" colspan="1" pageId="0" pageNumber="177" rowspan="1">SVL</td>
<td id="B5E5C5A6492416D928790794FCEB3BC1" colspan="1" pageId="0" pageNumber="177" rowspan="1">51.7</td>
<td id="DE49E16EBC48B612A349D493B70E8421" colspan="1" pageId="0" pageNumber="177" rowspan="1">48.3</td>
<td id="8C5BB37262543B9A19FDF1EB5BA808CC" colspan="1" pageId="0" pageNumber="177" rowspan="1">48.8</td>
<td id="5CC8A54E5BD5C2917A96A2E9B0F069A6" colspan="1" pageId="0" pageNumber="177" rowspan="1">46.8</td>
<td id="D6B2436171A8A405894379315D1B9356" colspan="1" pageId="0" pageNumber="177" rowspan="1">97.4</td>
<td id="44E0162F0EB7DFD9AA99591345FE42E1" colspan="1" pageId="0" pageNumber="177" rowspan="1">102.7</td>
<td id="E75184F1B17E03FA37664B802C402D8D" colspan="1" pageId="0" pageNumber="177" rowspan="1">103.4</td>
<td id="2923FC8828FE20ADA211927C1EE53E75" colspan="1" pageId="0" pageNumber="177" rowspan="1">106.7</td>
<td id="60E60351F512A7F5F3D876216C565E80" colspan="1" pageId="0" pageNumber="177" rowspan="1">104.2</td>
</tr>
<tr id="83C7B7966E520D72EE68299B862B669A" pageId="0" pageNumber="177">
<td id="1E22738EEE089B266C95F8A0E54A2F66" colspan="1" pageId="0" pageNumber="177" rowspan="1">HL</td>
<td id="D679F7300AD337027324A7575C2E2CC5" colspan="1" pageId="0" pageNumber="177" rowspan="1">21.8</td>
<td id="B9D4854F22F9C43BC1078FB86D6EA6E6" colspan="1" pageId="0" pageNumber="177" rowspan="1">20.3</td>
<td id="9306D6B0D99062AD946E90C02CCF6DC6" colspan="1" pageId="0" pageNumber="177" rowspan="1">19.6</td>
<td id="9869A32BDFFF0DE3C884551AE973D749" colspan="1" pageId="0" pageNumber="177" rowspan="1">19.7</td>
<td id="CDF3C5F26172047B65CED0165FF8F355" colspan="1" pageId="0" pageNumber="177" rowspan="1">37.5</td>
<td id="93D4AF93AA4CF0033D7610FC7BA124A1" colspan="1" pageId="0" pageNumber="177" rowspan="1">37.7</td>
<td id="FBEFED6331B4FC922DFFB629BDCFB815" colspan="1" pageId="0" pageNumber="177" rowspan="1">37.4</td>
<td id="AAD70B49FFBFEE9B3E255A823B5FA7EC" colspan="1" pageId="0" pageNumber="177" rowspan="1">39.2</td>
<td id="3B33FA2C86DDCCDED0ED5111BFC4730C" colspan="1" pageId="0" pageNumber="177" rowspan="1">39.0</td>
</tr>
<tr id="54AC72746B82E363D423E3FF0DEFD421" pageId="0" pageNumber="177">
<td id="E9BE7F66613CBBE6F0D2152A674D2F8F" colspan="1" pageId="0" pageNumber="177" rowspan="1">HW</td>
<td id="5A1A7D4238A6272073629002797CC237" colspan="1" pageId="0" pageNumber="177" rowspan="1">19.1</td>
<td id="8976596E75EB99D6D77D157B9378F6A1" colspan="1" pageId="0" pageNumber="177" rowspan="1">17.7</td>
<td id="54E9D1204BCBEFA0A1E42C3D8B66C91B" colspan="1" pageId="0" pageNumber="177" rowspan="1">17.9</td>
<td id="3337C040A3541CBC78BC485B0E38CEC5" colspan="1" pageId="0" pageNumber="177" rowspan="1">17.2</td>
<td id="36BA4F84ECC3777A391DEF3C2AD246A6" colspan="1" pageId="0" pageNumber="177" rowspan="1">34.9</td>
<td id="19B5F5F2D23A08FF1EE0957BCA749747" colspan="1" pageId="0" pageNumber="177" rowspan="1">36.1</td>
<td id="7846CF518942C477F1EA15F3707DBE72" colspan="1" pageId="0" pageNumber="177" rowspan="1">35.7</td>
<td id="DD50BE3B6C489AFD9278AA360BE183F8" colspan="1" pageId="0" pageNumber="177" rowspan="1">36.9</td>
<td id="D1FE2DC8B98B21B32C383E1EDCD6B3BB" colspan="1" pageId="0" pageNumber="177" rowspan="1">38.4</td>
</tr>
<tr id="B1C60C14D122E6EECE1CD143719C3E58" pageId="0" pageNumber="177">
<td id="9F0FC4101022A26E23455A78855DE31F" colspan="1" pageId="0" pageNumber="177" rowspan="1">SL</td>
<td id="96CC926572F923ED8F694AAD1C8AF141" colspan="1" pageId="0" pageNumber="177" rowspan="1">8.9</td>
<td id="4BCC8E527D88429BFF905DC1923EC163" colspan="1" pageId="0" pageNumber="177" rowspan="1">8.8</td>
<td id="248424D083A6173D561DE12D6A76C3FA" colspan="1" pageId="0" pageNumber="177" rowspan="1">8.3</td>
<td id="707E916950F37A167763F99CA53A641F" colspan="1" pageId="0" pageNumber="177" rowspan="1">8.5</td>
<td id="ACE015855B4814BDA256B9C9FEFC89C1" colspan="1" pageId="0" pageNumber="177" rowspan="1">16.4</td>
<td id="798F4C241F105797EF060E3F02CE3964" colspan="1" pageId="0" pageNumber="177" rowspan="1">15.7</td>
<td id="B2E7FD380165499482F89AD41331AACE" colspan="1" pageId="0" pageNumber="177" rowspan="1">16.3</td>
<td id="FE24412417C278594995A7FA969F3358" colspan="1" pageId="0" pageNumber="177" rowspan="1">16.4</td>
<td id="F8D79A18818F460FC532273E45148BC4" colspan="1" pageId="0" pageNumber="177" rowspan="1">16.6</td>
</tr>
<tr id="2613E63E93F0716CDB7AA78F2C2E4CC2" pageId="0" pageNumber="177">
<td id="A631B2F1F48EFBB2562ECB2F5FBDF898" colspan="1" pageId="0" pageNumber="177" rowspan="1">IND</td>
<td id="C0448684E44C7FC52F8D725B74C78B89" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.4</td>
<td id="46AD2B4A6710A482A7DC5709D55957EF" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.8</td>
<td id="EA6CE3D3AA2494018FD2475EB3BDE62D" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.3</td>
<td id="A307B5AB06D4487929EB85A9E3DCABE9" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.5</td>
<td id="6B39A4FE43370ED382E4160929E6B12E" colspan="1" pageId="0" pageNumber="177" rowspan="1">10.3</td>
<td id="B4BAD0ED3F67D63B27E0EB0C9D8BA569" colspan="1" pageId="0" pageNumber="177" rowspan="1">10.0</td>
<td id="1FD5DD871187C332AEFBF3EB995E796A" colspan="1" pageId="0" pageNumber="177" rowspan="1">9.8</td>
<td id="3599FD9DAF5803F59D4D083C4E334EC9" colspan="1" pageId="0" pageNumber="177" rowspan="1">10.4</td>
<td id="1DD73BF2A83E1333D39B8E2C7A85706E" colspan="1" pageId="0" pageNumber="177" rowspan="1">11.0</td>
</tr>
<tr id="8C4E58791EFB6D72D13A81A0F6381307" pageId="0" pageNumber="177">
<td id="6A94ABA626843C98CAE833DEBB4FCFBB" colspan="1" pageId="0" pageNumber="177" rowspan="1">IOD</td>
<td id="16BF06FDE1A5A46AC9A0B48CD45C2663" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.2</td>
<td id="68CF776749692A16C39B562321CCD803" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.1</td>
<td id="135323034A7527FAE8182938374D9572" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.0</td>
<td id="5DDA3961FDC95E4D249A9B25AAF9514E" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.6</td>
<td id="AA20127B1D9B785B40A49DFE0FFA7FA9" colspan="1" pageId="0" pageNumber="177" rowspan="1">8.7</td>
<td id="AF0D25C67B39CD75CB39E0E17ED7B125" colspan="1" pageId="0" pageNumber="177" rowspan="1">9.4</td>
<td id="D6F57834272ABBC87B9DA8F5FDA64254" colspan="1" pageId="0" pageNumber="177" rowspan="1">8.1</td>
<td id="B8A97D0530F8BABD951BCCCB52C04BA0" colspan="1" pageId="0" pageNumber="177" rowspan="1">9.6</td>
<td id="35512ABA6063A99653CD2CD8CCC79908" colspan="1" pageId="0" pageNumber="177" rowspan="1">9.5</td>
</tr>
<tr id="5F1D328F0DDFEE6166B7C54153093474" pageId="0" pageNumber="177">
<td id="EF3ADE474C93B08401A11E44A1A1F882" colspan="1" pageId="0" pageNumber="177" rowspan="1">ED</td>
<td id="BBAF1536EC5AD96F1F8571D5ECDBB4ED" colspan="1" pageId="0" pageNumber="177" rowspan="1">7.2</td>
<td id="4FF76ECC76638D3C2488565247ABA906" colspan="1" pageId="0" pageNumber="177" rowspan="1">7.6</td>
<td id="858B06192B687C472CC81F67115D7BE2" colspan="1" pageId="0" pageNumber="177" rowspan="1">7.1</td>
<td id="BBAFF70436BDC60BFB746F755EB36524" colspan="1" pageId="0" pageNumber="177" rowspan="1">7.0</td>
<td id="182F0B2BBD9324CAAC02A51ACB1DEC97" colspan="1" pageId="0" pageNumber="177" rowspan="1">10.5</td>
<td id="0CEB21AA2E2CC0CBFD79E9B95ECB81F8" colspan="1" pageId="0" pageNumber="177" rowspan="1">11.0</td>
<td id="38474A4836F8593D4E0DA7153EB8319B" colspan="1" pageId="0" pageNumber="177" rowspan="1">10.9</td>
<td id="990061B519BD6800239E1354ABF6E5E0" colspan="1" pageId="0" pageNumber="177" rowspan="1">12.2</td>
<td id="F7E039E5A34E84E56BD0541D6FD0F891" colspan="1" pageId="0" pageNumber="177" rowspan="1">11.1</td>
</tr>
<tr id="5C58FEB1EC70942B74C07285329A52A2" pageId="0" pageNumber="177">
<td id="5B08944A2CF23B4DFD9EFB05633BFBF4" colspan="1" pageId="0" pageNumber="177" rowspan="1">UEW</td>
<td id="0FA22225561EAE52A4A7ADCA45BBC806" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.2</td>
<td id="9DCEDA6ADDF6274C663E14E97781BFC9" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.1</td>
<td id="26E3531620F140B5D7382F6FB7EF03E7" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.8</td>
<td id="3CA8734C0432FABA4287A1480C94264D" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.8</td>
<td id="FB700C950E566AC5D8D0016C682164B7" colspan="1" pageId="0" pageNumber="177" rowspan="1">7.5</td>
<td id="203DB5010162F3D1AEB4F0E9D3D4C18F" colspan="1" pageId="0" pageNumber="177" rowspan="1">7.7</td>
<td id="A1C85C671C766D3E5CBD9E95BB13B372" colspan="1" pageId="0" pageNumber="177" rowspan="1">7.6</td>
<td id="8D8F643B903BCD50B24A3C9A61520704" colspan="1" pageId="0" pageNumber="177" rowspan="1">7.7</td>
<td id="2C50B6BC5D37C38212DFB621BAA19A0F" colspan="1" pageId="0" pageNumber="177" rowspan="1">8.4</td>
</tr>
<tr id="BB8D7769C78D2E4CC3B2A4C1D758CA06" pageId="0" pageNumber="177">
<td id="5BFC6C00C7BA7F21A56C1E89D3D3CB5C" colspan="1" pageId="0" pageNumber="177" rowspan="1">TD</td>
<td id="2F0B2A786468EEAD946576C0C537A258" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.2</td>
<td id="FFD26DBCAD63DA99EFD874B26CA32152" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.3</td>
<td id="3E18B0152442C56D96D1D52C92552387" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.7</td>
<td id="84544F89BB603848FFE22473CA9736DD" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.1</td>
<td id="5BBC0D9538223585EED01FDF0E860A57" colspan="1" pageId="0" pageNumber="177" rowspan="1">6.1</td>
<td id="73D4B11DB1B90292B1C4D76A03104215" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.9</td>
<td id="0D18EE6519D1CE85BDCAB8D200A1C50F" colspan="1" pageId="0" pageNumber="177" rowspan="1">6.0</td>
<td id="E8A17E6A38E22239F8858727EE6D33F0" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.8</td>
<td id="A8EE7C104C0E15F4FB7F02D9CC5CFBC7" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.7</td>
</tr>
<tr id="3224B2CB7ECDFFF12373AFAC56E36264" pageId="0" pageNumber="177">
<td id="428587DFCEC4AE4C4C6FB724886FFDA5" colspan="1" pageId="0" pageNumber="177" rowspan="1">TED</td>
<td id="2DD78C4E8488461CB1BF5F40DC224200" colspan="1" pageId="0" pageNumber="177" rowspan="1">1.7</td>
<td id="60FE9307504390C386E8EEA51DFA380E" colspan="1" pageId="0" pageNumber="177" rowspan="1">1.5</td>
<td id="15235C0A4557D7C5853B209344BCF3F3" colspan="1" pageId="0" pageNumber="177" rowspan="1">1.5</td>
<td id="63FC826D64886EAFD7B77C374A271CCA" colspan="1" pageId="0" pageNumber="177" rowspan="1">1.6</td>
<td id="C12960932057B1C88543FC3B364EAD90" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.5</td>
<td id="B6BF0AFC4B788FB4B78174AA9B1E1C20" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.4</td>
<td id="A56357742DA4AB3002BE0E838D703E40" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.7</td>
<td id="329B23930F2BC19621B90199C13097DF" colspan="1" pageId="0" pageNumber="177" rowspan="1">5.0</td>
<td id="0232B2CB680249CC6E506E8ECE6838E8" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.8</td>
</tr>
<tr id="51D6098403382ECBBE43DC18CD50F465" pageId="0" pageNumber="177">
<td id="104D9FF1D40A1FF131A863CD679F85C6" colspan="1" pageId="0" pageNumber="177" rowspan="1">FHL</td>
<td id="13E9A4A9EE0330B4EEB20C3235C82C8E" colspan="1" pageId="0" pageNumber="177" rowspan="1">28.2</td>
<td id="26134C350DDD6878BCF226C8BEDC61BD" colspan="1" pageId="0" pageNumber="177" rowspan="1">25.7</td>
<td id="B422580A65147299F02FCA42BA388DA5" colspan="1" pageId="0" pageNumber="177" rowspan="1">26.4</td>
<td id="17C314B2E659D5EC1973AA404242101E" colspan="1" pageId="0" pageNumber="177" rowspan="1">26.3</td>
<td id="93B28C192568FA4B83039F197FD48038" colspan="1" pageId="0" pageNumber="177" rowspan="1">49.1</td>
<td id="53958E34F8D2C9E9924329A2F616C3D9" colspan="1" pageId="0" pageNumber="177" rowspan="1">51.2</td>
<td id="F8187F8470D4EB85C8C91FFBFEEC82F1" colspan="1" pageId="0" pageNumber="177" rowspan="1">50.3</td>
<td id="7ABD922D624E69B7442B79CA383F852F" colspan="1" pageId="0" pageNumber="177" rowspan="1">53.0</td>
<td id="7EF02B95B9601AF94A38E3719AB2BC5A" colspan="1" pageId="0" pageNumber="177" rowspan="1">55.1</td>
</tr>
<tr id="784BBDCA034C4512A5AE5922B29818CA" pageId="0" pageNumber="177">
<td id="682A492E23ABC61655A28BECFC46C891" colspan="1" pageId="0" pageNumber="177" rowspan="1">HAL</td>
<td id="56F21011F5048425582C3C0EB4FA2D56" colspan="1" pageId="0" pageNumber="177" rowspan="1">16.0</td>
<td id="17A5C351A95504812706053F8A09A7FC" colspan="1" pageId="0" pageNumber="177" rowspan="1">15.1</td>
<td id="56B594161B32D1C4D33152B0700E5487" colspan="1" pageId="0" pageNumber="177" rowspan="1">15.3</td>
<td id="6F282444E468C83F8AD551EE9432F3E7" colspan="1" pageId="0" pageNumber="177" rowspan="1">15.6</td>
<td id="03764447FB6576CCBAE6C4FFC9E78C92" colspan="1" pageId="0" pageNumber="177" rowspan="1">28.6</td>
<td id="E1C5CA41CCC78198EC4E854A0FA1BE18" colspan="1" pageId="0" pageNumber="177" rowspan="1">30.1</td>
<td id="4D6493C942D05BBDF2A06230C55B5F1E" colspan="1" pageId="0" pageNumber="177" rowspan="1">29.6</td>
<td id="619FF0BFF0A6BBA7008AC768F804C05D" colspan="1" pageId="0" pageNumber="177" rowspan="1">31.4</td>
<td id="E45B4AE8A39FE58FBC9F4079B51FDAA8" colspan="1" pageId="0" pageNumber="177" rowspan="1">31.9</td>
</tr>
<tr id="8912DB4EF0FC46A87F217FEDB22EC105" pageId="0" pageNumber="177">
<td id="5716A3A0F2D0E2AD77E40A1EB3AEEB5A" colspan="1" pageId="0" pageNumber="177" rowspan="1">TL</td>
<td id="A03EAF084B01C4F2997A04A607C65151" colspan="1" pageId="0" pageNumber="177" rowspan="1">34.3</td>
<td id="B756D336AF93FD2FCD36F4DE18EDD856" colspan="1" pageId="0" pageNumber="177" rowspan="1">31.5</td>
<td id="5FFFC794B7B1AFA321BF68FB95264ADF" colspan="1" pageId="0" pageNumber="177" rowspan="1">32.9</td>
<td id="C3DF6DE35D2967435C41A0A8E695EA72" colspan="1" pageId="0" pageNumber="177" rowspan="1">31.2</td>
<td id="933994BBAA48280D926E4B33C28DB0FC" colspan="1" pageId="0" pageNumber="177" rowspan="1">65.6</td>
<td id="D960519A5849B6CB39E25AAE64638174" colspan="1" pageId="0" pageNumber="177" rowspan="1">66.5</td>
<td id="F7964D99B6CF2AC6679ABC8FA190DAF5" colspan="1" pageId="0" pageNumber="177" rowspan="1">65.9</td>
<td id="2B4404DB110F2533D09381C5650A1CAF" colspan="1" pageId="0" pageNumber="177" rowspan="1">70.4</td>
<td id="FEA50658E9AAF9787AC3A13447CB0C5B" colspan="1" pageId="0" pageNumber="177" rowspan="1">71.8</td>
</tr>
<tr id="0557B69CD262A872F3F8091FECEC1C25" pageId="0" pageNumber="177">
<td id="5056F589591ED064525A24EE7FDCE030" colspan="1" pageId="0" pageNumber="177" rowspan="1">TFL</td>
<td id="17634C4B7FEA9A4838955652338E3F16" colspan="1" pageId="0" pageNumber="177" rowspan="1">44.1</td>
<td id="2FE3F60579EF3C9195EA21C2E99F2C1B" colspan="1" pageId="0" pageNumber="177" rowspan="1">40.3</td>
<td id="0BA12ADCF387E2A9E0A9F7BB38DD24FE" colspan="1" pageId="0" pageNumber="177" rowspan="1">42.0</td>
<td id="2416926FFFF1A7B617DD955AE2AE407D" colspan="1" pageId="0" pageNumber="177" rowspan="1">40.6</td>
<td id="E914A7EEE345DA65EF22F02C67305614" colspan="1" pageId="0" pageNumber="177" rowspan="1">83.2</td>
<td id="83F9274253A6EED925ED8242D027AA86" colspan="1" pageId="0" pageNumber="177" rowspan="1">86.0</td>
<td id="02CC500B78DF5AF631C694F7FB0A982F" colspan="1" pageId="0" pageNumber="177" rowspan="1">82.6</td>
<td id="C8E6E871E5DE189C1138D9F3A2B4DD38" colspan="1" pageId="0" pageNumber="177" rowspan="1">90.6</td>
<td id="75066EEE03BEB09453733B4729A89528" colspan="1" pageId="0" pageNumber="177" rowspan="1">94.1</td>
</tr>
<tr id="C6FAD7A5EA5880441EF04CC1ADD66665" pageId="0" pageNumber="177">
<td id="713486EA6E7F636A478531D03F5017AE" colspan="1" pageId="0" pageNumber="177" rowspan="1">FTL</td>
<td id="2CF3F805426D0A6E52BB1C8C2B779497" colspan="1" pageId="0" pageNumber="177" rowspan="1">29.5</td>
<td id="D9E8A45F60FD0D0F9F65C3CD2E64638B" colspan="1" pageId="0" pageNumber="177" rowspan="1">27.3</td>
<td id="EC674C97A0B8387FC3B04F5104BFB2F2" colspan="1" pageId="0" pageNumber="177" rowspan="1">28.4</td>
<td id="EFA2BA4037DCAF2A9B3B426048336E62" colspan="1" pageId="0" pageNumber="177" rowspan="1">26.9</td>
<td id="6B6A6E322ECB2FC26CF5DE87B92C400F" colspan="1" pageId="0" pageNumber="177" rowspan="1">55.7</td>
<td id="0F8574B73AA1404B06BC5133189CFDDD" colspan="1" pageId="0" pageNumber="177" rowspan="1">59.0</td>
<td id="A08FC6093DDA093CAB09C05A7CEEC4BE" colspan="1" pageId="0" pageNumber="177" rowspan="1">56.5</td>
<td id="C4ED2ED3B93E63FB3326F37E9D65F04E" colspan="1" pageId="0" pageNumber="177" rowspan="1">60.7</td>
<td id="7A55B9C4CB7CFF38D6AEE7AEE7882FC7" colspan="1" pageId="0" pageNumber="177" rowspan="1">65.0</td>
</tr>
<tr id="AEEAF84EFA3582FF9F5F1655526334CA" pageId="0" pageNumber="177">
<td id="32F25631E35F4D7DAE7697FECBB6DB9A" colspan="1" pageId="0" pageNumber="177" rowspan="1">FDW3</td>
<td id="FAA0D8662230B8CC1F2C9553CB5E37B5" colspan="1" pageId="0" pageNumber="177" rowspan="1">1.9</td>
<td id="71834588270CD9457FF4AA0D1192A2F8" colspan="1" pageId="0" pageNumber="177" rowspan="1">1.9</td>
<td id="7387BACF7E234B3292A0F691D4BCC8BB" colspan="1" pageId="0" pageNumber="177" rowspan="1">1.8</td>
<td id="59A31E4204AAF5DB41959E6BAC05A986" colspan="1" pageId="0" pageNumber="177" rowspan="1">1.8</td>
<td id="6A97B3B0D8F4A13C7E9C5AF753EDBF4C" colspan="1" pageId="0" pageNumber="177" rowspan="1">3.7</td>
<td id="EA64578865C4DAF80E159ACE0273BD5F" colspan="1" pageId="0" pageNumber="177" rowspan="1">3.5</td>
<td id="F55087BBC5A2EE41177E5FDF60A5E27A" colspan="1" pageId="0" pageNumber="177" rowspan="1">2.6</td>
<td id="4DAB702BDF14A1CFB2A3AF3D67B37694" colspan="1" pageId="0" pageNumber="177" rowspan="1">3.6</td>
<td id="45A8A55606F4062A8EF0EF0B4FC9D95E" colspan="1" pageId="0" pageNumber="177" rowspan="1">4.0</td>
</tr>
</table>
</paragraph>
<paragraph id="F8F11542AA56DDCB40A10012774779A1" pageId="0" pageNumber="177">Forelimbs robust; relative lengths of fingers III&gt; IV&gt; I&gt; II; all finger tips slightly expanded; lateral fringes and webbing on fingers absent; subarticular tubercle round and prominent; one oval thenar tubercle prominent on the ventral of finger I; inner metacarpal tubercle and outer metacarpal tubercle distinct; supernumerary tubercles at the base of fingers II-IV; grayish-yellow nuptial pad on finger I in adult males.</paragraph>
<paragraph id="1BE0FBDC7DAB0FEDB5E039EEA93128DE" pageId="0" pageNumber="177">Hind limbs long, tibia longer than thigh; toes long, relative lengths IV&gt; V&gt; III&gt; II&gt; I; all toe tips slightly expanded; toes with entire webbing to disks; subarticular tuber-cles prominent and longitudinally ovoid; inner metatarsal tubercle prominent and oval; outer metatarsal and supernumerary tubercles absent.</paragraph>
<paragraph id="C06A554AF3192604D408B1D071D881D8" pageId="0" pageNumber="177">Dorsal skin shagreened with some very indistinct tubercles, skin on flanks granular, ventral skin smooth, and dorsolateral folds distinct in males. Skin slightly smoother and dorsolateral folds more indistinct in females.</paragraph>
</subSubSection>
<subSubSection id="933B5DE846350B3A43925D4F412283F9" pageId="0" pageNumber="177" type="coloration in life">
<paragraph id="3D8A2CE53F962468FF73B3C9F1F518BA" pageId="0" pageNumber="177">Coloration in life.</paragraph>
<paragraph id="CA995BC2EEEA5E6052E06A3AEE95F376" pageId="0" pageNumber="177">Dorsum brown scattered with dense tiny brownish black dots; upper lip yellowish white with indistinct dark spots; lower lip creamy white with distinct dark spots; lateral side of head and body brown with many irregular dark brown spots; dorsal surfaces of limbs brown with some indistinct dark brown bands; ventral surface of head and forelimbs, chest, and belly creamy white; ventral surface of hindlimbs light yellow; buttocks slightly pink; tympanum dark brown; and iris yellow in males. Coloration in females very similar to but usually slightly lighter than that in males.</paragraph>
</subSubSection>
<subSubSection id="29F8FA29FE8EFF968ECFC69E921A1F4F" pageId="0" pageNumber="177" type="coloration in preservative">
<paragraph id="29F6C2D6EFC8E162E69AD14397389A17" pageId="0" pageNumber="177">Coloration in preservative.</paragraph>
<paragraph id="8FD9A3878327347391AB5B8E5CF98E64" pageId="0" pageNumber="177">Dorsum color turned darker to grayish black, the dense tiny black dots on dorsum still visible, dorsal color of limbs turned darker to brownish black, the bands on limbs became indistinct. Ventral color of head and forelimbs, chest, and belly faded to pale white, ventral color of hindlimbs turned to flesh color. Iris became black, pupil became white.</paragraph>
</subSubSection>
<subSubSection id="45CB2F3FCCB21B3E86086D288A3D1A33" pageId="0" pageNumber="177" type="natural history">
<paragraph id="F08A7F3CF7D23466DE2E299B210412CA" pageId="0" pageNumber="177">Natural history.</paragraph>
<paragraph id="A1585C6E8B3BA6724294A9BB62E7E1C9" pageId="0" pageNumber="177">
The specimens from China were all found on the banks of rivers at night. No eggs or tadpoles were found from April to May. There were many stones and forests with good vegetation on the banks of the river (Fig. 
<figureCitation id="C2B5486311305D587DB49581003E6A81" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. The habitat at the collection site of the specimens of Odorrana heatwolei in Mengla, Yunnan, China." figureDoi="10.3897/herpetozoa.36.e106373.figure8" httpUri="https://binary.pensoft.net/fig/880827" pageId="0" pageNumber="177">8</figureCitation>
). An 
<taxonomicName id="E8D1AE938A00205CB4084988D552A7D8" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops" order="Anura" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="D0CF21DD64EAF1A8B7948423EBDBE20E" italics="true" pageId="0" pageNumber="177">Amolops</emphasis>
</taxonomicName>
species of 
<taxonomicName id="01014D11B10C6C1192479A2046E9BF86" baseAuthorityName="Bain, Stuart &amp; Orlov" baseAuthorityYear="2006" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops vitreus" order="Anura" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="vitreus">Amolops cf. vitreus</taxonomicName>
(Bain, Stuart &amp; Orlov, 2006) is distributed sympatric with this species.
</paragraph>
<caption id="62C6EB22C7F2C660FED2F48C3495E144" doi="10.3897/herpetozoa.36.e106373.figure8" httpUri="https://binary.pensoft.net/fig/880827" pageId="0" pageNumber="177" start="Figure 8" startId="F8">
<paragraph id="439382C182FC0BF1986AD323D63085E5" pageId="0" pageNumber="177">
<emphasis id="5AFE09D51D59864F4613787C27014617" bold="true" pageId="0" pageNumber="177">Figure 8.</emphasis>
The habitat at the collection site of the specimens of 
<taxonomicName id="6011BC72EEDC54702E73E7527527FC77" baseAuthorityName="Stuart &amp; Bain" baseAuthorityYear="2005" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana heatwolei" order="Anura" pageId="0" pageNumber="177" phylum="Chordata" rank="species" species="heatwolei">
<emphasis id="9198D4BACE2C5C5F85282C2DD468BAB9" italics="true" pageId="0" pageNumber="177">Odorrana heatwolei</emphasis>
</taxonomicName>
in Mengla, Yunnan, China.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="08E96E7C932AF02E64BD93C4276F2AD7" pageId="0" pageNumber="177" type="distribution">
<paragraph id="30D6D630D3D103F34C0CBB64459885E5" pageId="0" pageNumber="177">Distribution.</paragraph>
<paragraph id="085B2B8698052C14AD45ADD6639AE20E" pageId="0" pageNumber="177">
Currently known in Phongsaly Province, Laos, and Xishuangbanna Prefecture, Yunnan Province, China (Fig. 
<figureCitation id="BD5DF1A15E8395895BAC4454C24C010E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map showing the type locality (black dot) of O. tiannanensis, the type locality (black triangle) of O. heatwolei, the type locality (black square) of O. macrotympana, the new collection site (black star) of O. heatwolei, and the new collection sites (black diamonds) of O. tiannanensis." figureDoi="10.3897/herpetozoa.36.e106373.figure1" httpUri="https://binary.pensoft.net/fig/880820" pageId="0" pageNumber="177">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>