210 lines
20 KiB
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210 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.417.7733" ID-GBIF-Dataset="6ed372e2-700c-4945-b3a0-fe5872ef38db" ID-PMC="PMC4109464" ID-Pensoft-Pub="1313-2970-417-71" ID-PubMed="25061356" ID-ZBK="4675ED7211FA4D42836CBD36B77FC296" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-417-71" ModsDocOrigin="ZooKeys 417" ModsDocTitle="Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution" checkinTime="1451245732228" checkinUser="pensoft" docAuthor="Sokolov, Igor M., Reddell, James R. & Kavanaugh, David H." docDate="2014" docId="8C36FD7890A09A6984913B24F87724B3" docLanguage="en" docName="ZooKeys 417: 71-101" docOrigin="ZooKeys 417" docSource="http://dx.doi.org/10.3897/zookeys.417.7733" docTitle="Anillinus affabilis" docType="treatment" docVersion="3" lastPageNumber="80" masterDocId="FF97FFE46B074A506D13FF9B6152FFF2" masterDocTitle="Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution" masterLastPageNumber="101" masterPageNumber="71" pageNumber="77" updateTime="1668158702436" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reddell, James R.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>417</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>71</mods:start>
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<mods:end>101</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.417.7733</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.417.7733</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-417-71</mods:identifier>
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<mods:identifier type="ZBK">4675ED7211FA4D42836CBD36B77FC296</mods:identifier>
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<mods:identifier type="ZooBank">4675ED7211FA4D42836CBD36B77FC296</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152053694" LSID="urn:lsid:plazi:treatment:8C36FD7890A09A6984913B24F87724B3" httpUri="http://treatment.plazi.org/id/8C36FD7890A09A6984913B24F87724B3" lastPageId="9" lastPageNumber="80" pageId="6" pageNumber="77">
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<subSubSection pageId="6" pageNumber="77" type="multiple">
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<paragraph pageId="6" pageNumber="77">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="77" type="nomenclature">
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<paragraph pageId="6" pageNumber="77">
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<taxonomicName authority="Brues" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis (Brues)</taxonomicName>
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Figs 1A, 3D, 4D, 5A, 6
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 7
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<normalizedToken originalValue="B–C">B-C</normalizedToken>
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, 8
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="77" type="reference_group">
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<paragraph pageId="6" pageNumber="77">
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillus affabilis" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="affabilis">Anillus affabilis</taxonomicName>
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Brues, 1902: 366.
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<bibRefCitation author="Leng, CW" journalOrPublisher="John D. Sherman Jr., Mount Vernon (NY)" pageId="26" pageNumber="97" title="Catalogue of the Coleoptera of America, north of Mexico" year="1920">Leng 1920</bibRefCitation>
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</paragraph>
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<paragraph pageId="6" pageNumber="77">
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
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(Brues),
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<bibRefCitation author="Jeannel, R" journalOrPublisher="Revue Francaise d'Entomologie" pageId="26" pageNumber="97" pagination="241 - 339" title="Les Bembidiides endoges (Col. Carabidae). Monographie d'une lignee gondwanienne." volume="3" year="1937">Jeannel 1937</bibRefCitation>
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;
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<bibRefCitation author="Erwin, TL" journalOrPublisher="The Coleopoterists Bulletin" pageId="25" pageNumber="96" pagination="231 - 256" title="A catalogue of the primary types of Carabidae (incl. Cicindelinae) in the collections of the United States National Museum of Natural History (USNM) (Coleoptera)." volume="32" year="1978">Erwin and House 1978</bibRefCitation>
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;
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="Zookeys" pageId="25" pageNumber="96" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, North of Mexico." url="10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet 2012</bibRefCitation>
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</paragraph>
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<paragraph pageId="6" pageNumber="77">
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillodes affabilis" order="Coleoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="affabilis">Anillodes affabilis</taxonomicName>
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(Brues),
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<bibRefCitation author="Jeannel, R" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle (Serie A) Zoologie" pageId="26" pageNumber="97" pagination="33 - 204" title="Monographie des " Anillini ", bembidiides endoges [Coleoptera Trechidae]." volume="28" year="1963 a">Jeannel 1963a</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="78" pageId="6" pageNumber="77" type="type material">
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<paragraph pageId="6" pageNumber="77">Type material.</paragraph>
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<paragraph pageId="7" pageNumber="78">
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillus affabilis" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="affabilis">
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<pageBreakToken pageId="7" pageNumber="78" start="start">Anillus</pageBreakToken>
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affabilis
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</taxonomicName>
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Brues, 1902: 366. Lectotype (Fig. 1A), designated by
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<bibRefCitation author="Erwin, TL" journalOrPublisher="The Coleopoterists Bulletin" pageId="25" pageNumber="96" pagination="231 - 256" title="A catalogue of the primary types of Carabidae (incl. Cicindelinae) in the collections of the United States National Museum of Natural History (USNM) (Coleoptera)." volume="32" year="1978">Erwin and House (1978</bibRefCitation>
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: 233), a male, in NMNH, glued on a hair, dissected, and labeled: \ schmitti [handwritten] \ Type [typed] \ COTYPE [typed red label] \ Austin Texas [handwritten] \ HSB dissect of int. 1932 [handwritten] [these first five labels mounted on a single card] \
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillus affabilis" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="affabilis">Anillus affabilis</taxonomicName>
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Brues 1902 Mann Coll. 1932 [red handwritten label with male symbol] \ LECTOTYPE [male symbol]
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
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(Brues) By Erwin 78 \. Two paralectotypes: one female, also in NMNH, labeled: \ Austin Texas [printed] \ E. caecum [handwritten] \ type [typed with blue ink] \ TYPE [typed red label] \ ADP 115719 \
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillodes affabilis" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="affabilis">Anillodes affabilis</taxonomicName>
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Brues R, Jeannel det. 19 \ USNM TM #2067637 \ Paralectotype
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillus affabilis" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="affabilis">Anillus affabilis</taxonomicName>
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Brues recognized by Sokolov & Kavanaugh 2014 [printed yellow label] \; one male, in MCZ, labeled: \ Austin Tex \ Type [white label] \ H. C. Fall Collection \ Anillus affabilis Brues \ Paralectotype
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillus affabilis" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="affabilis">Anillus affabilis</taxonomicName>
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Brues recognized by Sokolov & Kavanaugh 2014 [printed yellow label]
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</paragraph>
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<caption pageId="7" pageNumber="78">
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<paragraph pageId="7" pageNumber="78">
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Figure 1. Images of type specimens of previously described Texas
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<taxonomicName genus="Anillina" lsidName="Anillina" pageId="7" pageNumber="78" rank="genus">Anillina</taxonomicName>
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species. A
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillus affabilis" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="affabilis">Anillus affabilis</taxonomicName>
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Brues, lectotype; B
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillodes sinuatus" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="sinuatus">Anillodes sinuatus</taxonomicName>
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Jeannel, holotype C
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<taxonomicName class="Insecta" family="Carabidae" genus="Micranillodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micranillodes depressus" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="depressus">Micranillodes depressus</taxonomicName>
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Jeannel, holotype. Scale = 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="78" type="notes on nomenclature and types">
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<paragraph pageId="7" pageNumber="78">Notes on nomenclature and types.</paragraph>
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<paragraph pageId="7" pageNumber="78">
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In his original description,
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<bibRefCitation author="Brues, CT" journalOrPublisher="The American Naturalist, an Illustrated Magazine of Natural History" pageId="25" pageNumber="96" pagination="365 - 378" title="New and little-known guests of the Texan legionary ants." volume="36" year="1902">Brues (1902)</bibRefCitation>
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noted that he had examined three specimens of this species, all from Austin, Texas. In 1978, the paralectotype specimens noted above were unknown to
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<bibRefCitation author="Erwin, TL" journalOrPublisher="The Coleopoterists Bulletin" pageId="25" pageNumber="96" pagination="231 - 256" title="A catalogue of the primary types of Carabidae (incl. Cicindelinae) in the collections of the United States National Museum of Natural History (USNM) (Coleoptera)." volume="32" year="1978">Erwin and House (1978</bibRefCitation>
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: 233) but they have been located in the MCZ and NMNH since then.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="78" type="recognition">
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<paragraph pageId="7" pageNumber="78">Recognition.</paragraph>
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<paragraph pageId="7" pageNumber="78">
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Females of
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
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are practically indistinguishable from females of
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus wisemanensis" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="wisemanensis">Anillinus wisemanensis</taxonomicName>
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, described below. From adults of other Texan species of the genus, those of
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
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can be distinguished by the following combination of external characters: large size, comparatively narrow and transverse pronotum with rectilinearly constricted lateral margins, rather wide and long elytra with rounded apices; and males can be further distinguished by the minute spine on the posterior edge of metafemora and shape of the median lobe.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="9" lastPageNumber="80" pageId="7" pageNumber="78" type="redescription">
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<paragraph pageId="7" pageNumber="78">Redescription.</paragraph>
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<paragraph pageId="7" pageNumber="78">
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Medium-sized for genus (SBL range 1.73-1.96 mm, mean 1.87
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.120 mm, n=3).
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</paragraph>
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<paragraph pageId="7" pageNumber="78">
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Habitus. Body form (Figs 1A, 5A) subdepressed, subparallel, slightly elongate (WE/SBL 0.38
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.010), head normally proportioned for genus (WH/WPm 0.76
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.026), pronotum moderately narrow in comparison to elytra (WPm/WE 0.78
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.007).
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</paragraph>
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<paragraph pageId="7" pageNumber="78">Color. Body rufotestaceous or brunneorufous, appendages testaceous.</paragraph>
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<paragraph pageId="7" pageNumber="78">Microsculpture. Distinct over all dorsal surfaces of head, pronotum and elytra, with slightly transverse polygonal meshes of more or less scaly appearance on elytra.</paragraph>
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<paragraph pageId="7" pageNumber="78">
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Prothorax. Pronotum of normal length (LP/LE 0.39
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.012) and slightly transverse for genus (WPm/LP 1.31
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.016), lateral margins almost rectilinear and moderately constricted posteriorly (WPm/WPp 1.32
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.008). Anterior angles indistinct, posterior angles slightly obtuse (100-110°). Width between anterior and posterior angles of equal length (WPa/WPp 1.00
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.026). Basal margin almost straight.
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</paragraph>
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<paragraph pageId="7" pageNumber="78">
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Elytra. Widely depressed along suture, of normal length (LE/SBL 0.58
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.008) and typical width for genus (WE/LE 0.66
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.029), with traces of 4-5 striae. Humeri distinct, rounded, in outline forming right angle with longitudinal axis of body. Lateral margins subparallel, slightly divergent at basal fourth, evenly rounded to apex in apical fourth, without subapical sinuation. Vestiture of elytra short (less than one-third length of discal setae). Apex of elytron rounded.
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</paragraph>
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<paragraph pageId="7" pageNumber="78">Legs. Male protarsomere 1 markedly dilated apico-laterally with rows of adhesive setae ventrally. Male hind legs modified: metafemora with minute tooth at middle along posteroventral margin (Fig. 4D).</paragraph>
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<paragraph pageId="8" pageNumber="79">
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<pageBreakToken pageId="8" pageNumber="79" start="start">Abdomen</pageBreakToken>
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. Abdominal ventrites 3 and 4 of males each with a pair of protuberances at the places of setal attachments (Fig. 3D). Last visible abdominal ventrite of male slightly depressed (Fig. 3D).
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</paragraph>
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<paragraph pageId="8" pageNumber="79">Male genitalia. Median lobe of aedeagus (Fig. 6A) with short basal lobe, long arcuate shaft, and strongly enlarged apex, broadly rounded at tip. Dorsal margin strongly sclerotized along almost all its length. Ventral margin enlarged along entire length from apex to basal orifice, bearing numerous poriferous canals. Dorsal sclerite in form of a semicircular blade-like structure with characteristic basal prolongations. Without distinct ventral sclerites, but with few sclerotized fields in apicoventral area. Dorsal membraneous field with small spines located dorsally from dorsal sclerite. Enlarged apical area of median lobe with a dark spine-like structure. Right paramere slightly enlarged, long and narrow, with numerous (>8) long setae (Fig. 6C), approximately equal in length to length of the paramere. Left paramere slightly enlarged apically and greatly so basally, with translucent keel and large lateral process comparable in size to paramere itself (Fig. 6B), without long setae.</paragraph>
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<paragraph lastPageId="9" lastPageNumber="80" pageId="8" pageNumber="79">
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Female genitalia. Gonocoxite 2 unguiform (Fig. 7B), rather long, with moderately curved blade and narrowly rounded apex, with nematiform and two ensiform setae, the lateral of which is thicker than the medial one. Laterotergite with 9-10 setae. Sper
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<pageBreakToken pageId="9" pageNumber="80" start="start">matheca</pageBreakToken>
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(Fig. 7C, sp) with distal part of cornu (dpc) markedly dilated and abruptly narrowed to the proximal part (ppc). Nodulus (n) short, ramus undifferentiated.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="80" type="geographical distribution">
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<paragraph pageId="9" pageNumber="80">Geographical distribution.</paragraph>
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<paragraph pageId="9" pageNumber="80">This species is known only from Travis County, Texas (Fig. 8, black triangle), in the vicinity of the Balcones Fault Zone. In addition to the lectotype and paralectotype, we have examined a total of five specimens (3 males and 2 females), all dissected: one male and one female labeled: \ TX: Travis Co., Tooth Cave, 24.V.1992, J. Reddell \ Texas Memorial Museum Invertebrate Zool Coll #27.145 \; one male and one female, in CNC, labeled: \ TX: Travis Co., Tooth Cave, 6.VI.1992, J. Reddell \ Texas Memorial Museum Invertebrate Zool Coll #27.146 \; one male, in TMM, labeled: \ TX: Travis Co., Three-Holer Cave, 18.VIII.1990, J. Reddell, M. Reyes \ Texas Memorial Museum Invertebrate Zool Coll #27.144 \.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="80" type="way of life">
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<paragraph pageId="9" pageNumber="80">Way of life.</paragraph>
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<paragraph pageId="9" pageNumber="80">
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All newly collected beetles were found in caves on the underside of rocks shallowly embedded in soil. Specimens of the type series were sifted from ant nests (
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<bibRefCitation author="Brues, CT" journalOrPublisher="The American Naturalist, an Illustrated Magazine of Natural History" pageId="25" pageNumber="96" pagination="365 - 378" title="New and little-known guests of the Texan legionary ants." volume="36" year="1902">Brues 1902</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="80" type="relationships">
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<paragraph pageId="9" pageNumber="80">Relationships.</paragraph>
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<paragraph pageId="9" pageNumber="80">
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The shape of the spermatheca in females and the armature of the internal sac and type of ventral enlargement of the median lobe in males suggest a likely relationship with
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |