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<document id="F45CA1F9F4CB1A351B73BEA6A88C1792" ID-CLB-Dataset="5151" ID-DOI="10.11646/zootaxa.3856.2.3" ID-GBIF-Dataset="2d7003e5-498f-4fa8-92df-6c407fd18953" ID-ISSN="1175-5326" ID-Zenodo-Dep="4929871" ID-ZooBank="919C3E5E-B97E-40D8-A72A-A74A974147A5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623440692681" checkinUser="felipe" docAuthor="Miranda, Vinícius Da Rocha &amp; Brasil, Ana Claudia Dos Santos" docDate="2014" docId="EF572A28422CFF99FF607B90F261F850" docLanguage="en" docName="zt03856p226.pdf" docOrigin="Zootaxa 3856 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pholoides sinepapillatus Miranda &amp; Brasil 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="215" masterDocId="136E5250422EFF9DFFF77D31F328FFC5" masterDocTitle="Two new species and a new record of Scale-worms (Polychaeta) from Southwest Atlantic deep-sea coral mounds" masterLastPageNumber="226" masterPageNumber="211" pageNumber="213" updateTime="1699012918814" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3F89529B02DDB81EC17E684222A85BAF">Two new species and a new record of Scale-worms (Polychaeta) from Southwest Atlantic deep-sea coral mounds</mods:title>
</mods:titleInfo>
<mods:name id="B645063B3A8A73238CE7FDD4C7D1DA76" type="personal">
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<mods:namePart id="6A72AF81AE430830E744B58C488B475F">Miranda, Vinícius Da Rocha</mods:namePart>
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<mods:namePart id="58FEA1B3E32E28368C12ED92FA58BA5B">Brasil, Ana Claudia Dos Santos</mods:namePart>
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<mods:typeOfResource id="85DB5DEBA063B8167286C9CF9961895D">text</mods:typeOfResource>
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<mods:title id="05ED62C74B66F8AC2C0739A9AB6E950C">Zootaxa</mods:title>
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<mods:part id="6A889F08DAEA815235D9C77A34D5FEB6">
<mods:date id="800E57434969830C0E26EA60B22E81C4">2014</mods:date>
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<mods:number id="6406368F2B1E7D902DF0C8644F0BE5C5">2014-08-22</mods:number>
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<mods:number id="8144DA7B34B0729AEECDEF2AC7C3CEF0">3856</mods:number>
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<mods:classification id="66C373CBB200F5FAD7A9C22B2C022ADE">journal article</mods:classification>
<mods:identifier id="029C09BE8DEFA25691154A786F71D72E" type="CLB-Dataset">5151</mods:identifier>
<mods:identifier id="F5F5B4043AEFC17F419605E29110373B" type="DOI">10.11646/zootaxa.3856.2.3</mods:identifier>
<mods:identifier id="B20AE389EC48FF3EFA5EB39652E6F289" type="GBIF-Dataset">2d7003e5-498f-4fa8-92df-6c407fd18953</mods:identifier>
<mods:identifier id="FE409030B14316580F2CCBD667752900" type="ISSN">1175-5326</mods:identifier>
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<treatment id="EF572A28422CFF99FF607B90F261F850" ID-DOI="http://doi.org/10.5281/zenodo.5167232" ID-GBIF-Taxon="183923435" ID-Zenodo-Dep="5167232" LSID="urn:lsid:plazi:treatment:EF572A28422CFF99FF607B90F261F850" httpUri="http://treatment.plazi.org/id/EF572A28422CFF99FF607B90F261F850" lastPageId="4" lastPageNumber="215" pageId="2" pageNumber="213">
<subSubSection id="2FE4C8B5422CFF9FFF607B90F2B4F918" pageId="2" pageNumber="213" type="nomenclature">
<paragraph id="67419B3E422CFF9FFF607B90F109F97E" blockId="2.[151,545,1697,1757]" box="[151,545,1697,1723]" pageId="2" pageNumber="213">
<heading id="3C092C52422CFF9FFF607B90F109F97E" bold="true" box="[151,545,1697,1723]" fontSize="11" level="1" pageId="2" pageNumber="213" reason="1">
<emphasis id="558A472C422CFF9FFF607B90F109F97E" bold="true" box="[151,545,1697,1723]" pageId="2" pageNumber="213">
<taxonomicName id="A0FEE0BD422CFF9FFF607B90F295F97E" ID-CoL="4GMLY" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[151,445,1697,1723]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="213" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422CFF9FFF607B90F295F97E" bold="true" box="[151,445,1697,1723]" italics="true" pageId="2" pageNumber="213">Pholoides sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422CFF9FFE337B90F109F97E" box="[452,545,1697,1723]" pageId="2" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="67419B3E422CFF9FFF607BF5F2B4F918" blockId="2.[151,545,1697,1757]" box="[151,412,1732,1757]" pageId="2" pageNumber="213">Figures: 2 AE; 3 AK</paragraph>
</subSubSection>
<subSubSection id="2FE4C8B5422CFF9FFF607A3DF72FF8AE" pageId="2" pageNumber="213" type="materials_examined">
<paragraph id="67419B3E422CFF9FFF607A3DF72FF8AE" blockId="2.[151,1437,1804,2009]" pageId="2" pageNumber="213">
<materialsCitation id="D7969163422CFF9FFF607A3DF0EEF88D" ID-GBIF-Occurrence="3331515301" collectingDate="2008-12-22" collectionCode="IBUFRJ" elevation="10" latitude="-22.37819" longLatPrecision="1" longitude="-40.095848" pageId="2" pageNumber="213" specimenCode="IBUFRJ-2131" specimenCount="2" typeStatus="holotype">
<emphasis id="558A472C422CFF9FFF607A3DF288F8E0" bold="true" box="[151,416,1804,1829]" pageId="2" pageNumber="213">Material Examined:</emphasis>
4 spms.;
<typeStatus id="B845259C422CFF9FFDA37A3DF1C4F8E0" box="[596,748,1804,1829]" pageId="2" pageNumber="213" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="71F850B7422CFF9FFCF97A3DF056F8E6" box="[782,894,1804,1829]" pageId="2" pageNumber="213">2 spms.</specimenCount>
,
<specimenCode id="37583345422CFF9FFC567A3DF76BF8E0" box="[929,1091,1804,1829]" pageId="2" pageNumber="213">IBUFRJ-2131</specimenCode>
, ECOPROF
<quantity id="A00636DB422CFF9FFAF57A3DF6BFF8E6" box="[1282,1431,1804,1829]" metricMagnitude="0" metricUnit="m" metricValue="9.639" pageId="2" pageNumber="213" unit="m" value="9.639">
<elevation id="ECD37C0D422CFF9FFAF57A3DF6BFF8E6" box="[1282,1431,1804,1829]" metricMagnitude="0" metricUnit="m" metricValue="9.639" pageId="2" pageNumber="213" unit="m" value="9.639">9, 639 m</elevation>
</quantity>
,
<geoCoordinate id="02CAFDF9422CFF9FFF607A01F21DF88C" box="[151,309,1840,1865]" degrees="40" direction="west" minutes="5" orientation="longitude" pageId="2" pageNumber="213" precision="1" seconds="45,06" value="-40.095848">40º5'45,06&quot;W</geoCoordinate>
<geoCoordinate id="02CAFDF9422CFF9FFEB27A01F2CCF88C" box="[325,484,1840,1865]" degrees="22" direction="south" minutes="22" orientation="latitude" pageId="2" pageNumber="213" precision="1" seconds="41,48" value="-22.37819">22º22'41,48&quot;S</geoCoordinate>
,
<date id="1340BDFE422CFF9FFE017A01F151F88C" box="[502,633,1840,1865]" pageId="2" pageNumber="213" value="2008-12-22">
<collectingDate id="03044416422CFF9FFE017A01F151F88C" box="[502,633,1840,1865]" pageId="2" pageNumber="213" value="2008-12-22">22/12/2008</collectingDate>
</date>
, on
<taxonomicName id="A0FEE0BD422CFF9FFD427A01F097F88D" baseAuthorityName="Duncan" baseAuthorityYear="1873" box="[693,959,1840,1864]" class="Anthozoa" family="Caryophylliidae" genus="Solenosmilia" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="213" phylum="Cnidaria" rank="species" species="variabilis">
<emphasis id="558A472C422CFF9FFD427A01F097F88D" box="[693,959,1840,1864]" italics="true" pageId="2" pageNumber="213">Solenosmilia variabilis</emphasis>
</taxonomicName>
.
</materialsCitation>
<typeStatus id="B845259C422CFF9FFC237A01F75BF88C" box="[980,1139,1840,1865]" pageId="2" pageNumber="213" type="paratype">PARATYPES</typeStatus>
: 2 spms., IBUFRJ-2130, ECOPROF
<quantity id="A00636DB422CFF9FFEEB7A65F2A7F8AE" box="[284,399,1876,1901]" metricMagnitude="1" metricUnit="m" metricValue="1.0622" pageId="2" pageNumber="213" unit="m" value="10.622">10, 622 m</quantity>
,
<geoCoordinate id="02CAFDF9422CFF9FFE6C7A65F113F8A8" box="[411,571,1876,1901]" degrees="40" direction="west" minutes="5" orientation="longitude" pageId="2" pageNumber="213" precision="1" seconds="39,24" value="-40.09423">40º5'39,24&quot;W</geoCoordinate>
<geoCoordinate id="02CAFDF9422CFF9FFDBD7A65F1F2F8A8" box="[586,730,1876,1901]" degrees="22" direction="south" minutes="22" orientation="latitude" pageId="2" pageNumber="213" precision="1" seconds="3,21" value="-22.367558">22º22'3,21&quot;S</geoCoordinate>
,
<date id="1340BDFE422CFF9FFD127A65F04FF8A8" box="[741,871,1876,1901]" pageId="2" pageNumber="213" value="2009-01-17">17/01/2009</date>
, on
<taxonomicName id="A0FEE0BD422CFF9FFC627A64F72FF8AE" box="[917,1031,1876,1900]" class="Hydrozoa" family="Stylasteridae" genus="Errina" kingdom="Animalia" order="Anthoathecata" pageId="2" pageNumber="213" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="558A472C422CFF9FFC627A64F0C8F8A9" box="[917,992,1877,1900]" italics="true" pageId="2" pageNumber="213">Errina</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="2FE4C8B5422CFF9FFF307A49F1D0F81C" pageId="2" pageNumber="213" type="diagnosis">
<paragraph id="67419B3E422CFF9FFF307A49F1D0F81C" blockId="2.[151,1437,1804,2009]" pageId="2" pageNumber="213">
<emphasis id="558A472C422CFF9FFF307A49F26BF854" bold="true" box="[199,323,1912,1937]" pageId="2" pageNumber="213">Diagnosis:</emphasis>
Prostomium globular, with two pair of eyes. Lips of the mouth, the surface of the elytra and the ventral surface of the body smooth. Biramous parapodia with noto- and neuropodium of the same size. Elytra with faint concentric rings easily observed by transparency.
</paragraph>
</subSubSection>
<caption id="3381CBB6422DFF9EFF6078B5F23CFA33" ID-DOI="http://doi.org/10.5281/zenodo.4929877" ID-Zenodo-Dep="4929877" httpUri="https://zenodo.org/record/4929877/files/figure.png" pageId="3" pageNumber="214" startId="3.[151,250,1412,1434]" targetBox="[151,1436,239,1375]" targetPageId="3">
<paragraph id="67419B3E422DFF9EFF6078B5F23CFA33" blockId="3.[151,1436,1412,1526]" pageId="3" pageNumber="214">
<emphasis id="558A472C422DFF9EFF6078B5F23CFA5C" bold="true" box="[151,276,1412,1434]" pageId="3" pageNumber="214">FIGURE 2.</emphasis>
<taxonomicName id="A0FEE0BD422DFF9EFEEB78B5F131FA5F" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[284,537,1412,1434]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="214" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422DFF9EFEEB78B5F131FA5F" box="[284,537,1412,1434]" italics="true" pageId="3" pageNumber="214">Pholoides sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422DFF9EFDD778B4F159FA5F" box="[544,625,1413,1434]" pageId="3" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
(A) View of the Holotype in a hole on dead
<taxonomicName id="A0FEE0BD422DFF9EFBB578B5F607FA5F" baseAuthorityName="Duncan" baseAuthorityYear="1873" box="[1090,1327,1412,1434]" class="Anthozoa" family="Caryophylliidae" genus="Solenosmilia" kingdom="Animalia" order="Scleractinia" pageId="3" pageNumber="214" phylum="Cnidaria" rank="species" species="variabilis">
<emphasis id="558A472C422DFF9EFBB578B5F607FA5F" box="[1090,1327,1412,1434]" italics="true" pageId="3" pageNumber="214">Solenosmilia variabilis</emphasis>
</taxonomicName>
; (B) SEM of the dorsal surface of middle right elytra; (C) Detail of the tip of a notochaeta from a middle parapodium; (D) Blade and distal shaft of upper neurochaeta; (E) Blade and distal shaft of lower neurochaeta. (Scales bars: A=1 mm; B=200 µm; C, D, E= 10 µm).
</paragraph>
</caption>
<subSubSection id="2FE4C8B5422DFF99FF307B2EF7E8FDDD" lastPageId="4" lastPageNumber="215" pageId="3" pageNumber="214" type="description">
<paragraph id="67419B3E422DFF9EFF307B2EF60CF999" blockId="3.[151,1437,1567,1988]" pageId="3" pageNumber="214">
<emphasis id="558A472C422DFF9EFF307B2EF273F9FD" bold="true" box="[199,347,1567,1592]" pageId="3" pageNumber="214">Description:</emphasis>
<typeStatus id="B845259C422DFF9EFE9F7B2EF2F8F9FD" box="[360,464,1567,1592]" pageId="3" pageNumber="214" type="holotype">Holotype</typeStatus>
with 36 segments (length: 5.07 mm; width 0.7 mm);
<typeStatus id="B845259C422DFF9EFB9D7B11F7FDF9FD" box="[1130,1237,1568,1592]" pageId="3" pageNumber="214" type="paratype">paratypes</typeStatus>
with number of chaetigers varying from 27 (length: 2.6 mm; width: 0.6 mm) up to 35 (length: 4.62 mm; width 0.7 mm).
</paragraph>
<paragraph id="67419B3E422DFF9EFF307B56F653F8F1" blockId="3.[151,1437,1567,1988]" pageId="3" pageNumber="214">
Body dorsoventrally flattened, dorsal and ventral surfaces without papillae or tubercles (
<figureCitation id="FFC587BB422DFF9EFB527B56F7D7F945" box="[1189,1279,1639,1664]" captionStart="FIGURE 2" captionStartId="3.[151,250,1412,1434]" captionTargetBox="[151,1436,239,1375]" captionTargetId="figure-284@3.[151,1436,239,1375]" captionTargetPageId="3" captionText="FIGURE 2. Pholoides sinepapillatus sp. nov. (A) View of the Holotype in a hole on dead Solenosmilia variabilis; (B) SEM of the dorsal surface of middle right elytra; (C) Detail of the tip of a notochaeta from a middle parapodium; (D) Blade and distal shaft of upper neurochaeta; (E) Blade and distal shaft of lower neurochaeta. (Scales bars: A=1 mm; B=200 µm; C, D, E= 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929877" httpUri="https://zenodo.org/record/4929877/files/figure.png" pageId="3" pageNumber="214">Fig. 2A</figureCitation>
). Prostomium and tentacular segment fused (
<figureCitation id="FFC587BB422DFF9EFE0D7BBAF170F961" box="[506,600,1675,1700]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="3" pageNumber="214">Fig. 3A</figureCitation>
). Prostomium globular, as wide as long, one pair of eyes; antenna with ceratophore at anterior margin of prostomium, style papillate and inflated subdistally, 1.5 times longer than prostomium. First segment anterior and lateral to prostomium (
<figureCitation id="FFC587BB422DFF9EFCAE7BE2F098F929" box="[857,944,1747,1772]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="3" pageNumber="214">Fig. 3B</figureCitation>
), parapodia uniramous with notoacicula not extending beyond epidermis and without papillae on the ventral side; each parapodia with bundle of long, serrate capillary chaetae (
<figureCitation id="FFC587BB422DFF9EFE947A2AF2F6F8F1" box="[355,478,1819,1844]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="3" pageNumber="214">Fig. 3CD</figureCitation>
); dorsal cirrus (
<figureCitation id="FFC587BB422DFF9EFD7A7A2AF1C9F8F1" box="[653,737,1819,1844]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="3" pageNumber="214">Fig. 3E</figureCitation>
) length similar to median antenna, and a stout ventral palp.
</paragraph>
<paragraph id="67419B3E422DFF9EFF607A0EF079F801" blockId="3.[151,1437,1567,1988]" pageId="3" pageNumber="214">
Elytra on bulbous elytrophores on segments 2, 4, 5, 7, continuing on alternate segments to end of body; last two segments without elytra or dorsal tubercles. Elytra large, thick, subtriangular (
<figureCitation id="FFC587BB422DFF9EFBD97A52F7AEF8B9" box="[1070,1158,1891,1916]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="3" pageNumber="214">Fig. 3F</figureCitation>
) to reniform (
<figureCitation id="FFC587BB422DFF9EFAC77A52F6A6F8B9" box="[1328,1422,1891,1916]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="3" pageNumber="214">Fig. 3G</figureCitation>
), papillae on margins not covered by adjacent elytra; smooth surfaces and faint concentric rings visible (
<figureCitation id="FFC587BB422DFF9EFADD7AB6F6A7F865" box="[1322,1423,1927,1952]" captionStart="FIGURE 2" captionStartId="3.[151,250,1412,1434]" captionTargetBox="[151,1436,239,1375]" captionTargetId="figure-284@3.[151,1436,239,1375]" captionTargetPageId="3" captionText="FIGURE 2. Pholoides sinepapillatus sp. nov. (A) View of the Holotype in a hole on dead Solenosmilia variabilis; (B) SEM of the dorsal surface of middle right elytra; (C) Detail of the tip of a notochaeta from a middle parapodium; (D) Blade and distal shaft of upper neurochaeta; (E) Blade and distal shaft of lower neurochaeta. (Scales bars: A=1 mm; B=200 µm; C, D, E= 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929877" httpUri="https://zenodo.org/record/4929877/files/figure.png" pageId="3" pageNumber="214">Figs. 2B</figureCitation>
). Segments without elytra with small dorsal tubercles (
<figureCitation id="FFC587BB422DFF9EFD1D7A9AF06CF801" box="[746,836,1963,1988]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="3" pageNumber="214">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="67419B3E422AFF99FF307DA6F14EFE85" blockId="4.[151,1440,151,1941]" pageId="4" pageNumber="215">
Second segment with biramous parapodia (
<figureCitation id="FFC587BB422AFF99FD477DA6F023FF75" box="[688,779,151,176]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="4" pageNumber="215">Fig. 3H</figureCitation>
), one acicula per ramus, not extending beyond epidermis; ventral cirri with short, papillate cirrostyles. Notochaetae (
<figureCitation id="FFC587BB422AFF99FCA37D8DF096FF10" box="[852,958,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1412,1434]" captionTargetBox="[151,1436,239,1375]" captionTargetId="figure-284@3.[151,1436,239,1375]" captionTargetPageId="3" captionText="FIGURE 2. Pholoides sinepapillatus sp. nov. (A) View of the Holotype in a hole on dead Solenosmilia variabilis; (B) SEM of the dorsal surface of middle right elytra; (C) Detail of the tip of a notochaeta from a middle parapodium; (D) Blade and distal shaft of upper neurochaeta; (E) Blade and distal shaft of lower neurochaeta. (Scales bars: A=1 mm; B=200 µm; C, D, E= 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929877" httpUri="https://zenodo.org/record/4929877/files/figure.png" pageId="4" pageNumber="215">Figs. 2C</figureCitation>
;
<figureCitation id="FFC587BB422AFF99FC277D8DF73CFF10" box="[976,1044,188,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="4" pageNumber="215">3CD</figureCitation>
) capilliform, curved and serrate. Neurochaetae compound (
<figureCitation id="FFC587BB422AFF99FE247DEEF149FF3D" box="[467,609,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1412,1434]" captionTargetBox="[151,1436,239,1375]" captionTargetId="figure-284@3.[151,1436,239,1375]" captionTargetPageId="3" captionText="FIGURE 2. Pholoides sinepapillatus sp. nov. (A) View of the Holotype in a hole on dead Solenosmilia variabilis; (B) SEM of the dorsal surface of middle right elytra; (C) Detail of the tip of a notochaeta from a middle parapodium; (D) Blade and distal shaft of upper neurochaeta; (E) Blade and distal shaft of lower neurochaeta. (Scales bars: A=1 mm; B=200 µm; C, D, E= 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929877" httpUri="https://zenodo.org/record/4929877/files/figure.png" pageId="4" pageNumber="215">Figs. 2DE</figureCitation>
;
<figureCitation id="FFC587BB422AFF99FD807DEEF183FF3D" box="[631,683,223,248]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="4" pageNumber="215">3IJ</figureCitation>
), supracicular shorter than subacicular; shafts and blades of supracicular neurochaetae smooth; shafts of subacicular neurochaetae subdistally spinose, blades serrate and longer than blades of supracicular neurochaetae.
</paragraph>
<paragraph id="67419B3E422AFF99FF307C7DF7E8FDDD" blockId="4.[151,1440,151,1941]" pageId="4" pageNumber="215">
Remaining parapodia biramous (
<figureCitation id="FFC587BB422AFF99FDC87C7DF1B4FEA0" box="[575,668,332,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="4" pageNumber="215">Fig. 3K</figureCitation>
); notopodia as long as neuropodia, distally rounded to subconical, with serrate capilliform notochaetae (
<figureCitation id="FFC587BB422AFF99FDB97C5EF19FFE4D" box="[590,695,367,392]" captionStart="FIGURE 2" captionStartId="3.[151,250,1412,1434]" captionTargetBox="[151,1436,239,1375]" captionTargetId="figure-284@3.[151,1436,239,1375]" captionTargetPageId="3" captionText="FIGURE 2. Pholoides sinepapillatus sp. nov. (A) View of the Holotype in a hole on dead Solenosmilia variabilis; (B) SEM of the dorsal surface of middle right elytra; (C) Detail of the tip of a notochaeta from a middle parapodium; (D) Blade and distal shaft of upper neurochaeta; (E) Blade and distal shaft of lower neurochaeta. (Scales bars: A=1 mm; B=200 µm; C, D, E= 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929877" httpUri="https://zenodo.org/record/4929877/files/figure.png" pageId="4" pageNumber="215">Figs. 2C</figureCitation>
;
<figureCitation id="FFC587BB422AFF99FD307C5EF023FE4D" box="[711,779,367,392]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="4" pageNumber="215">3CD</figureCitation>
) and notoaciculae slightly extending beyond epidermis. Neuropodia with compound neurochaetae of two kinds: supracicular smaller than subacicular, shafts and blades of supracicular smooth; shafts of subacicular subdistally spinose, blades serrate and longer than supracicular (
<figureCitation id="FFC587BB422AFF99FA937C86F3F3FE30" captionStart="FIGURE 2" captionStartId="3.[151,250,1412,1434]" captionTargetBox="[151,1436,239,1375]" captionTargetId="figure-284@3.[151,1436,239,1375]" captionTargetPageId="3" captionText="FIGURE 2. Pholoides sinepapillatus sp. nov. (A) View of the Holotype in a hole on dead Solenosmilia variabilis; (B) SEM of the dorsal surface of middle right elytra; (C) Detail of the tip of a notochaeta from a middle parapodium; (D) Blade and distal shaft of upper neurochaeta; (E) Blade and distal shaft of lower neurochaeta. (Scales bars: A=1 mm; B=200 µm; C, D, E= 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929877" httpUri="https://zenodo.org/record/4929877/files/figure.png" pageId="4" pageNumber="215">Figs. 2DE</figureCitation>
;
<figureCitation id="FFC587BB422AFF99FF117CEDF232FE30" box="[230,282,476,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="4" pageNumber="215">3IJ</figureCitation>
); neuroaciculae not extending beyond epidermis, each neuroacicular lobe with small digitiform process near distal end of acicula. Ventral cirri present, with few short papillae on cirrostyles (
<figureCitation id="FFC587BB422AFF99FBAF7CCEF79BFDDD" box="[1112,1203,511,536]" captionStart="FIGURE 3" captionStartId="5.[151,250,1755,1777]" captionTargetBox="[184,1369,217,1631]" captionTargetId="figure-115@5.[151,1436,197,1696]" captionTargetPageId="5" captionText="FIGURE 3. Pholoides sinepapillatus sp. nov. (Holotype, IBUFRJ-3121) (A) Dorsal view of the anterior end; (B) Tentaculophore from right side (without dorsal cirrus); (C) Notochaeta; (D): Tip of same; (E) Style of dorsal cirrus of tentaculophore; (F) First elytra, right side; (G) Middle elytra, right side; (H) Parapodium from second segment; (I) Lower compound neurochaeta; (J): Upper compound neurochaeta; (K) Left middle elytrigerous parapodium. (Scales bars: A= 500 µm; B= 250 µm C= 50 µm, D, I and J= 150 µm and E= 75 µm; F, G, H and K = 300 µm)." figureDoi="http://doi.org/10.5281/zenodo.4929881" httpUri="https://zenodo.org/record/4929881/files/figure.png" pageId="4" pageNumber="215">Fig. 3K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2FE4C8B5422AFF99FF307F15F3F6FD6D" pageId="4" pageNumber="215" type="etymology">
<paragraph id="67419B3E422AFF99FF307F15F604FDA5" blockId="4.[151,1440,151,1941]" pageId="4" pageNumber="215">
<emphasis id="558A472C422AFF99FF307F15F279FDF8" bold="true" box="[199,337,548,573]" pageId="4" pageNumber="215">Etymology:</emphasis>
The name of the species refers to the fact that it doesn´t have papillae over the dorsal and ventral surfaces of the body, neither over the surface of the elytra, as observed in the other species of the genera.
</paragraph>
<paragraph id="67419B3E422AFF99FF307F5DF3F6FD6D" blockId="4.[151,1440,151,1941]" pageId="4" pageNumber="215">
<emphasis id="558A472C422AFF99FF307F5DF21FFD40" bold="true" box="[199,311,620,645]" pageId="4" pageNumber="215">Habitats:</emphasis>
Associated with
<taxonomicName id="A0FEE0BD422AFF99FE0D7F5DF1D5FD41" baseAuthorityName="Duncan" baseAuthorityYear="1873" box="[506,765,620,644]" class="Anthozoa" family="Caryophylliidae" genus="Solenosmilia" kingdom="Animalia" order="Scleractinia" pageId="4" pageNumber="215" phylum="Cnidaria" rank="species" species="variabilis">
<emphasis id="558A472C422AFF99FE0D7F5DF1D5FD41" box="[506,765,620,644]" italics="true" pageId="4" pageNumber="215">Solenosmilia variabilis</emphasis>
</taxonomicName>
and Dendrophiliidae corals, in a depth range from
<quantity id="A00636DB422AFF99FAC87F5DF657FD40" box="[1343,1407,620,645]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="215" unit="m" value="600.0">600m</quantity>
to
<specimenCount id="71F850B7422AFF99FF607FBEF3F6FD6D" box="[151,222,655,680]" pageId="4" pageNumber="215" type="generic">
<quantity id="A00636DB422AFF99FF607FBEF3F2FD6D" box="[151,218,655,680]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="4" pageNumber="215" unit="m" value="640.0">640m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="2FE4C8B5422AFF99FF307F85F7F8F888" pageId="4" pageNumber="215" type="discussion">
<paragraph id="67419B3E422AFF99FF307F85F2DAFC0D" blockId="4.[151,1440,151,1941]" pageId="4" pageNumber="215">
<emphasis id="558A472C422AFF99FF307F85F215FD08" bold="true" box="[199,317,692,717]" pageId="4" pageNumber="215">Remarks:</emphasis>
<taxonomicName id="A0FEE0BD422AFF99FEB07F84F12CFD09" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[327,516,692,716]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422AFF99FEB07F84F12CFD09" box="[327,516,692,716]" italics="true" pageId="4" pageNumber="215">P. sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422AFF99FDF97F85F141FD0E" box="[526,617,692,715]" pageId="4" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
differs from the other two recognized species of
<taxonomicName id="A0FEE0BD422AFF99FB597F85F636FD09" authorityName="Pruvot" authorityYear="1895" box="[1198,1310,692,716]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="genus">
<emphasis id="558A472C422AFF99FB597F85F636FD09" box="[1198,1310,692,716]" italics="true" pageId="4" pageNumber="215">Pholoides</emphasis>
</taxonomicName>
by having smooth ventral and dorsal surfaces: papillae are present on the ventral surface of the body of the two other
<taxonomicName id="A0FEE0BD422AFF99FF607FCDF24BFCD0" box="[151,355,764,789]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="558A472C422AFF99FF607FCDF22FFCD1" box="[151,263,764,788]" italics="true" pageId="4" pageNumber="215">Pholoides</emphasis>
species
</taxonomicName>
(
<taxonomicName id="A0FEE0BD422AFF99FE807FCCF007FCD0" authority="(Marenzeller, 1893)" baseAuthorityName="Marenzeller" baseAuthorityYear="1893" box="[375,815,764,789]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="dorsipapillatus">
<emphasis id="558A472C422AFF99FE807FCCF169FCD1" box="[375,577,764,788]" italics="true" pageId="4" pageNumber="215">P. dorsipapillatus</emphasis>
(
<bibRefCitation id="036FE6CF422AFF99FDA37FCDF00FFCD0" author="Marenzeller, E. von" box="[596,807,764,789]" pageId="4" pageNumber="215" pagination="25 - 48" refId="ref9440" refString="Marenzeller, E. von. (1893) Berichte der Commission fur Oceanographische erforschung des Ostlichen mittelmeeres. Zweite Reihe. VI. Zoologische Ergebnisse II. Polychaten des Grundes, gesammelt 1890, 1891, 1892. Denkschriften der Akademie der Wissenschaften, 60, 25 - 48. [Wien]" type="journal article" year="1893">Marenzeller, 1893</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="A0FEE0BD422AFF99FC997FCCF79DFCD0" authority="(Johnson, 1897))" authorityName="" baseAuthorityName="Johnson" baseAuthorityYear="1897" box="[878,1205,764,789]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="asperus">
<emphasis id="558A472C422AFF99FC997FCCF0CCFCD1" box="[878,996,765,788]" italics="true" pageId="4" pageNumber="215">P. asperus</emphasis>
(
<bibRefCitation id="036FE6CF422AFF99FC0F7FCDF78AFCD0" author="Johnson, H. P." box="[1016,1186,764,789]" pageId="4" pageNumber="215" pagination="153 - 199" refId="ref9047" refString="Johnson, H. P. (1897) A preliminary account of the marine annelids of the Pacific coast, with descriptions of new species. Proceedings of the California Academy of Sciences, 1 (5), 153 - 199." type="journal article" year="1897">Johnson, 1897</bibRefCitation>
))
</taxonomicName>
. The surface of the buccal lips differs among the three species. While
<taxonomicName id="A0FEE0BD422AFF99FD3E7E10F0A6FCFD" baseAuthorityName="Marenzeller" baseAuthorityYear="1893" box="[713,910,800,824]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="dorsipapillatus">
<emphasis id="558A472C422AFF99FD3E7E10F0A6FCFD" box="[713,910,800,824]" italics="true" pageId="4" pageNumber="215">P. dorsipapillatus</emphasis>
</taxonomicName>
has papillae over the ventral lip and
<taxonomicName id="A0FEE0BD422AFF99FADD7E10F6B4FCFD" authorityName="" baseAuthorityName="Johnson" baseAuthorityYear="1897" box="[1322,1436,801,824]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="asperus">
<emphasis id="558A472C422AFF99FADD7E10F6B4FCFD" box="[1322,1436,801,824]" italics="true" pageId="4" pageNumber="215">P. asperus</emphasis>
</taxonomicName>
both lips papillated,
<taxonomicName id="A0FEE0BD422AFF99FE8A7E74F11FFC99" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[381,567,836,860]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422AFF99FE8A7E74F2A3FC99" box="[381,395,837,860]" italics="true" pageId="4" pageNumber="215">P</emphasis>
.
<emphasis id="558A472C422AFF99FE6D7E75F11FFC99" box="[410,567,836,860]" italics="true" pageId="4" pageNumber="215">sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422AFF99FDC87E75F1BEFC9E" box="[575,662,836,859]" pageId="4" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
has both lips smooth. Additionally specimen of
<taxonomicName id="A0FEE0BD422AFF99FB4C7E74F65CFC99" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[1211,1396,836,860]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422AFF99FB4C7E74F65CFC99" box="[1211,1396,836,860]" italics="true" pageId="4" pageNumber="215">P. sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422AFF99FA8B7E75F3EFFCBA" pageId="4" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
did not show papillae on the dorsal tubercle, as is observed in the other two species of
<taxonomicName id="A0FEE0BD422AFF99FB487E59F607FC45" authorityName="Pruvot" authorityYear="1895" box="[1215,1327,872,896]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="genus">
<emphasis id="558A472C422AFF99FB487E59F607FC45" box="[1215,1327,872,896]" italics="true" pageId="4" pageNumber="215">Pholoides</emphasis>
</taxonomicName>
. As in
<emphasis id="558A472C422AFF99FA707E58F122FC61" italics="true" pageId="4" pageNumber="215">
<taxonomicName id="A0FEE0BD422AFF99FA707E58F26BFC61" baseAuthorityName="Marenzeller" baseAuthorityYear="1893" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="dorsipapillatus">P. dorsipapillatus</taxonomicName>
,
<taxonomicName id="A0FEE0BD422AFF99FEA77EBCF122FC61" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[336,522,908,932]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">P. sinepapillatus</taxonomicName>
</emphasis>
<taxonomicNameLabel id="4EB9FA57422AFF99FDE57EBDF143FC66" box="[530,619,908,931]" pageId="4" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
did not show papilla on the ventral side of the first segment, while this papilla is present in
<emphasis id="558A472C422AFF99FE8F7E80F2DAFC0D" box="[376,498,945,968]" italics="true" pageId="4" pageNumber="215">
<taxonomicName id="A0FEE0BD422AFF99FE8F7E80F2C7FC0D" authorityName="" baseAuthorityName="Johnson" baseAuthorityYear="1897" box="[376,495,945,968]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="asperus">P. asperus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="67419B3E422AFF99FF307EE5F608FB9D" blockId="4.[151,1440,151,1941]" pageId="4" pageNumber="215">
In both species already described, the elytra show papillae on the dorsal surface, while
<taxonomicName id="A0FEE0BD422AFF99FB797EE4F66EFC29" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[1166,1350,980,1004]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422AFF99FB797EE4F7B4FC29" box="[1166,1180,981,1004]" italics="true" pageId="4" pageNumber="215">P</emphasis>
.
<emphasis id="558A472C422AFF99FB5E7EE5F66EFC29" box="[1193,1350,980,1004]" italics="true" pageId="4" pageNumber="215">sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422AFF99FABA7EE5F6B4FC2E" box="[1357,1436,980,1003]" pageId="4" pageNumber="215" rank="species">sp.nov.</taxonomicNameLabel>
has smooth dorsal and ventral surface of elytra.
<bibRefCitation id="036FE6CF422AFF99FD3B7EC6F0A3FBD5" author="Pettibone, M. H." box="[716,907,1015,1040]" pageId="4" pageNumber="215" pagination="1 - 24" refId="ref9968" refString="Pettibone, M. H. (1992) Contribution to the polychaete family Pholoidae Kinberg. Smithsonian Contributions to Zoology, 532, 1 - 24." type="journal article" year="1992">Pettibone (1992)</bibRefCitation>
already emphasized the presence of knobbed papillae near the margins and middle region of elytra in
<taxonomicName id="A0FEE0BD422AFF99FCDE792CF0B6FBF1" authorityName="" baseAuthorityName="Johnson" baseAuthorityYear="1897" box="[809,926,1053,1076]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="asperus">
<emphasis id="558A472C422AFF99FCDE792CF0B6FBF1" box="[809,926,1053,1076]" italics="true" pageId="4" pageNumber="215">P. asperus</emphasis>
</taxonomicName>
. While
<taxonomicName id="A0FEE0BD422AFF99FC0A792CF75BFBF1" authorityName="" baseAuthorityName="Johnson" baseAuthorityYear="1897" box="[1021,1139,1053,1076]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="asperus">
<emphasis id="558A472C422AFF99FC0A792CF75BFBF1" box="[1021,1139,1053,1076]" italics="true" pageId="4" pageNumber="215">P. asperus</emphasis>
</taxonomicName>
presents strongly marked concentric rings,
<taxonomicName id="A0FEE0BD422AFF99FEAF7970F139FB9D" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[344,529,1088,1112]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422AFF99FEAF7970F139FB9D" box="[344,529,1088,1112]" italics="true" pageId="4" pageNumber="215">P. sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422AFF99FDEF7971F147FB92" box="[536,623,1088,1111]" pageId="4" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="A0FEE0BD422AFF99FD537970F042FB9D" baseAuthorityName="Marenzeller" baseAuthorityYear="1893" box="[676,874,1088,1112]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="dorsipapillatus">
<emphasis id="558A472C422AFF99FD537970F042FB9D" box="[676,874,1088,1112]" italics="true" pageId="4" pageNumber="215">P. dorsipapillatus</emphasis>
</taxonomicName>
, show faintly marked concentric rings.
</paragraph>
<paragraph id="67419B3E422AFF99FF307955F218FABD" blockId="4.[151,1440,151,1941]" pageId="4" pageNumber="215">
According to the description provided by
<bibRefCitation id="036FE6CF422AFF99FD507955F04CFBB8" author="Pettibone, M. H." box="[679,868,1124,1149]" pageId="4" pageNumber="215" pagination="1 - 24" refId="ref9968" refString="Pettibone, M. H. (1992) Contribution to the polychaete family Pholoidae Kinberg. Smithsonian Contributions to Zoology, 532, 1 - 24." type="journal article" year="1992">Pettibone (1992)</bibRefCitation>
for
<emphasis id="558A472C422AFF99FC617954F74CFBB9" box="[918,1124,1124,1148]" italics="true" pageId="4" pageNumber="215">
<taxonomicName id="A0FEE0BD422AFF99FC617954F748FBB9" baseAuthorityName="Marenzeller" baseAuthorityYear="1893" box="[918,1120,1124,1148]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="dorsipapillatus">P. dorsipapillatus</taxonomicName>
,
</emphasis>
<taxonomicName id="A0FEE0BD422AFF99FB9A7954F600FBB9" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[1133,1320,1124,1148]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422AFF99FB9A7954F753FBB9" box="[1133,1147,1125,1148]" italics="true" pageId="4" pageNumber="215">P</emphasis>
.
<emphasis id="558A472C422AFF99FB7C7955F600FBB9" box="[1163,1320,1124,1148]" italics="true" pageId="4" pageNumber="215">sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422AFF99FAC67955F6A9FBBE" box="[1329,1409,1124,1147]" pageId="4" pageNumber="215" rank="species">sp.nov.</taxonomicNameLabel>
is not significantly different regarding the parapodium and the compound chaetae. Both species present noto- and neuropodium almost with the same size; upper neurochaetae with blades longer than lower neurochaetae, presenting faintly spinose or smooth, and the shafts can be smooth or subdistally spinose. But when compared with
<taxonomicName id="A0FEE0BD422AFF99FF6079C4F238FAC9" authorityName="" baseAuthorityName="Johnson" baseAuthorityYear="1897" box="[151,272,1269,1292]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="asperus">
<emphasis id="558A472C422AFF99FF6079C4F238FAC9" box="[151,272,1269,1292]" italics="true" pageId="4" pageNumber="215">P. asperus</emphasis>
</taxonomicName>
some differences can be noticed such as the notopodium in
<emphasis id="558A472C422AFF99FC0079C4F75FFAC9" box="[1015,1143,1269,1292]" italics="true" pageId="4" pageNumber="215">
<taxonomicName id="A0FEE0BD422AFF99FC0079C4F75BFAC9" authorityName="" baseAuthorityName="Johnson" baseAuthorityYear="1897" box="[1015,1139,1269,1292]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="asperus">P. asperus</taxonomicName>
,
</emphasis>
that is smaller than the neuropodium and presents a pair of papillae in the presetal lobe and postsetal lobe covered with papillae; the neurochaetae of
<taxonomicName id="A0FEE0BD422AFF99FEA5780CF2EEFA91" authorityName="" baseAuthorityName="Johnson" baseAuthorityYear="1897" box="[338,454,1341,1364]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="asperus">
<emphasis id="558A472C422AFF99FEA5780CF2EEFA91" box="[338,454,1341,1364]" italics="true" pageId="4" pageNumber="215">P. asperus</emphasis>
</taxonomicName>
differ from
<taxonomicName id="A0FEE0BD422AFF99FDA4780CF025FA91" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[595,781,1340,1364]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422AFF99FDA4780CF149FA91" box="[595,609,1341,1364]" italics="true" pageId="4" pageNumber="215">P</emphasis>
.
<emphasis id="558A472C422AFF99FD87780DF025FA91" box="[624,781,1340,1364]" italics="true" pageId="4" pageNumber="215">sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422AFF99FCE2780DF04DFA96" box="[789,869,1340,1363]" pageId="4" pageNumber="215" rank="species">sp.nov.</taxonomicNameLabel>
by having larger and serrated blades and a longer spinose shaft.
</paragraph>
<paragraph id="67419B3E422AFF99FF3078B5F2B6F98A" blockId="4.[151,1440,151,1941]" pageId="4" pageNumber="215">
Another outstanding distinctive character is the number of eyes, while
<taxonomicName id="A0FEE0BD422AFF99FC1078B4F786FA59" baseAuthorityName="Marenzeller" baseAuthorityYear="1893" box="[999,1198,1412,1436]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="dorsipapillatus">
<emphasis id="558A472C422AFF99FC1078B4F786FA59" box="[999,1198,1412,1436]" italics="true" pageId="4" pageNumber="215">P. dorsipapillatus</emphasis>
</taxonomicName>
and
<taxonomicName id="A0FEE0BD422AFF99FB1278B4F671FA59" authorityName="" baseAuthorityName="Johnson" baseAuthorityYear="1897" box="[1253,1369,1413,1436]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="asperus">
<emphasis id="558A472C422AFF99FB1278B4F671FA59" box="[1253,1369,1413,1436]" italics="true" pageId="4" pageNumber="215">P. asperus</emphasis>
</taxonomicName>
show two pairs of eyes,
<taxonomicName id="A0FEE0BD422AFF99FE937898F135FA05" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[356,541,1448,1472]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422AFF99FE937898F25AFA05" box="[356,370,1449,1472]" italics="true" pageId="4" pageNumber="215">P</emphasis>
.
<emphasis id="558A472C422AFF99FE777899F135FA05" box="[384,541,1448,1472]" italics="true" pageId="4" pageNumber="215">sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422AFF99FDD37899F15BFA7A" box="[548,627,1448,1471]" pageId="4" pageNumber="215" rank="species">sp.nov.</taxonomicNameLabel>
has only one pair of eyes situated on the widest part of the prostomium.
<taxonomicName id="A0FEE0BD422AFF99FF6078FDF173FA20" authority="(Averincev 1978)" baseAuthorityName="Averincev" baseAuthorityYear="1978" box="[151,603,1484,1509]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="mendeleevi">
<emphasis id="558A472C422AFF99FF6078FDF2A5FA21" box="[151,397,1484,1508]" italics="true" pageId="4" pageNumber="215">Pholoides mendeleevi</emphasis>
(
<bibRefCitation id="036FE6CF422AFF99FE6B78FDF17AFA20" author="Averincev, V. G." box="[412,594,1484,1509]" pageId="4" pageNumber="215" pagination="51 - 72" refId="ref7724" refString="Averincev, V. G. (1978) The Polychaetous Annelids of the Aphroditiformia of the Shelf and Upper Bathyal of Australian and New Zealand Region and of Macquarie Island (on the Base Data of 16 th Cruise of R / V DM. Mendeleev). Transactions of the P. P. Shirov Institute of Oceanology Academy of Sciences of the USSR, Moscow, 113, 51 - 72." type="journal article" year="1978">Averincev 1978</bibRefCitation>
)
</taxonomicName>
was illustrated with one pair of eyes (Fig. 8: 76
<specimenCount id="71F850B7422AFF99FB7978FDF7EFFA26" box="[1166,1223,1484,1509]" pageId="4" pageNumber="215" type="generic">80 p.</specimenCount>
69), so this would not be the first register of
<taxonomicName id="A0FEE0BD422AFF99FE3378C1F11CF9CD" authorityName="Pruvot" authorityYear="1895" box="[452,564,1520,1544]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="genus">
<emphasis id="558A472C422AFF99FE3378C1F11CF9CD" box="[452,564,1520,1544]" italics="true" pageId="4" pageNumber="215">Pholoides</emphasis>
</taxonomicName>
presenting this character, which is not mentioned by
<bibRefCitation id="036FE6CF422AFF99FB6278DEF67AF9CD" author="Pettibone, M. H." box="[1173,1362,1519,1544]" pageId="4" pageNumber="215" pagination="1 - 24" refId="ref9968" refString="Pettibone, M. H. (1992) Contribution to the polychaete family Pholoidae Kinberg. Smithsonian Contributions to Zoology, 532, 1 - 24." type="journal article" year="1992">Pettibone (1992)</bibRefCitation>
in her revision of the genus. We believe that the presence of at least one pair of eyes could be used as a basic characteristic of the genus
<taxonomicName id="A0FEE0BD422AFF99FED07B09F2BFF995" authorityName="Pruvot" authorityYear="1895" box="[295,407,1592,1616]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="genus">
<emphasis id="558A472C422AFF99FED07B09F2BFF995" box="[295,407,1592,1616]" italics="true" pageId="4" pageNumber="215">Pholoides</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="67419B3E422AFF99FF307B6DF7F8F888" blockId="4.[151,1440,151,1941]" pageId="4" pageNumber="215">
<taxonomicName id="A0FEE0BD422AFF99FF307B6DF2F3F9B1" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[199,475,1628,1652]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422AFF99FF307B6DF2F3F9B1" box="[199,475,1628,1652]" italics="true" pageId="4" pageNumber="215">Pholoides sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422AFF99FE147B6DF11AF9B6" box="[483,562,1628,1651]" pageId="4" pageNumber="215" rank="species">sp.nov.</taxonomicNameLabel>
is not the first species of
<taxonomicName id="A0FEE0BD422AFF99FCA37B6DF0ECF9B1" authorityName="Pruvot" authorityYear="1895" box="[852,964,1628,1652]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="genus">
<emphasis id="558A472C422AFF99FCA37B6DF0ECF9B1" box="[852,964,1628,1652]" italics="true" pageId="4" pageNumber="215">Pholoides</emphasis>
</taxonomicName>
recorded for Brazilian waters. Sumida
<emphasis id="558A472C422AFF99FA767B6CF385F95D" italics="true" pageId="4" pageNumber="215">et. al</emphasis>
(2004) reported
<taxonomicName id="A0FEE0BD422AFF99FE867BB0F2CEF95D" authorityName="" baseAuthorityName="Johnson" baseAuthorityYear="1897" box="[369,486,1665,1688]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="asperus">
<emphasis id="558A472C422AFF99FE867BB0F2CEF95D" box="[369,486,1665,1688]" italics="true" pageId="4" pageNumber="215">P. asperus</emphasis>
</taxonomicName>
occurring in seabed pockmarks at Santos Basin, off the coast of
<collectingRegion id="A53A55DC422AFF99FB2E7B4EF665F95D" box="[1241,1357,1663,1688]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="215">São Paulo</collectingRegion>
(26ºS; 46ºW) at a depth of almost
<specimenCount id="71F850B7422AFF99FE397B95F13EF978" box="[462,534,1700,1725]" pageId="4" pageNumber="215" type="generic">
<quantity id="A00636DB422AFF99FE397B95F139F978" box="[462,529,1700,1725]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="215" unit="m" value="700.0">700m</quantity>
.
</specimenCount>
<bibRefCitation id="036FE6CF422AFF99FDEA7B95F182F978" author="Pinto, N. O. P." box="[541,682,1700,1725]" pageId="4" pageNumber="215" refId="ref9778" refString="Pinto, N. O. P. (2014) Poliquetas de escama da regiao Sudeste do Brasil. Universidade Estadual de Campinas, Campinas, 189 pp." type="book" year="2014">Pinto (2014)</bibRefCitation>
also reported another species of
<taxonomicName id="A0FEE0BD422AFF99FBE97B95F7A6F979" authorityName="Pruvot" authorityYear="1895" box="[1054,1166,1700,1724]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="genus">
<emphasis id="558A472C422AFF99FBE97B95F7A6F979" box="[1054,1166,1700,1724]" italics="true" pageId="4" pageNumber="215">Pholoides</emphasis>
</taxonomicName>
for
<collectingCountry id="1FE9DBAE422AFF99FB497B95F62DF978" box="[1214,1285,1700,1725]" name="Brazil" pageId="4" pageNumber="215">Brazil</collectingCountry>
, however the species described by her possess oval prostomium, first segment with prominent ventral papilla, body surface presenting papillae, elytra with faint concentric rings and notopodia smaller than neuropodia; instead of the global prostomium, first segment without ventral papilla, elytra with strongly marked concentric rings and noto- and neuropodia with the same size, as observed in
<taxonomicName id="A0FEE0BD422AFF99FD687A05F09BF889" authority="Miranda &amp; Brasil, 2014" authorityName="Miranda &amp; Brasil" authorityYear="2014" box="[671,947,1844,1868]" class="Polychaeta" family="Sigalionidae" genus="Pholoides" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="215" phylum="Annelida" rank="species" species="sinepapillatus" status="sp. nov.">
<emphasis id="558A472C422AFF99FD687A05F09BF889" box="[671,947,1844,1868]" italics="true" pageId="4" pageNumber="215">Pholoides sinepapillatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4EB9FA57422AFF99FC4D7A05F721F88E" box="[954,1033,1844,1867]" pageId="4" pageNumber="215" rank="species">sp.nov.</taxonomicNameLabel>
described herein.
</paragraph>
</subSubSection>
<subSubSection id="2FE4C8B5422AFF99FF307A66F261F850" pageId="4" pageNumber="215" type="distribution">
<paragraph id="67419B3E422AFF99FF307A66F261F850" blockId="4.[151,1440,151,1941]" pageId="4" pageNumber="215">
<emphasis id="558A472C422AFF99FF307A66F271F8B5" bold="true" box="[199,345,1879,1904]" pageId="4" pageNumber="215">Distribution</emphasis>
: Southwest Atlantic, currently known only from the
<typeStatus id="B845259C422AFF99FC5B7A69F0F4F8B5" box="[940,988,1880,1904]" pageId="4" pageNumber="215">type</typeStatus>
locality, Campos Basin, State of
<collectingRegion id="A53A55DC422AFF99FAA57A66F3C0F850" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="215">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="1FE9DBAE422AFF99FF0B7A4DF26CF850" box="[252,324,1916,1941]" name="Brazil" pageId="4" pageNumber="215">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>