260 lines
28 KiB
XML
260 lines
28 KiB
XML
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<mods:title id="EA2B055447FA6824A9B90E2E672FD052">New insights into the identities of the elasmobranch fauna of Sri Lanka</mods:title>
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<mods:title id="2517C07447771E9EF7DCF9B8F7847549">Zootaxa</mods:title>
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<treatment id="362D2832DA343E4C0AC1FA48FB42FEE4" ID-DOI="http://doi.org/10.5281/zenodo.5945427" ID-GBIF-Taxon="156191941" ID-Zenodo-Dep="5945427" LSID="urn:lsid:plazi:treatment:362D2832DA343E4C0AC1FA48FB42FEE4" httpUri="http://treatment.plazi.org/id/362D2832DA343E4C0AC1FA48FB42FEE4" lastPageId="16" lastPageNumber="217" pageId="14" pageNumber="215">
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<subSubSection id="F69ECAAFDA343E520AC1FA48FDBAFA62" box="[151,564,1481,1507]" pageId="14" pageNumber="215" type="nomenclature">
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<paragraph id="BE3B9924DA343E520AC1FA48FDBAFA62" blockId="14.[151,564,1481,1541]" box="[151,564,1481,1507]" pageId="14" pageNumber="215">
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<heading id="E5732E48DA343E520AC1FA48FDBAFA62" bold="true" box="[151,564,1481,1507]" fontSize="11" level="1" pageId="14" pageNumber="215" reason="1">
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<taxonomicName id="7984E2A7DA343E520AC1FA48FDBAFA62" authority="(Macleay)" authorityName="Macleay" baseAuthorityName="Macleay" baseAuthorityYear="1883" box="[151,564,1481,1507]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="14" pageNumber="215" phylum="Chordata" rank="species" species="granulatus">
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<emphasis id="8CF04536DA343E520AC1FA48FDBAFA62" bold="true" box="[151,564,1481,1507]" pageId="14" pageNumber="215">
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<emphasis id="8CF04536DA343E520AC1FA48FE3EFA62" bold="true" box="[151,432,1481,1507]" italics="true" pageId="14" pageNumber="215">Urogymnus granulatus</emphasis>
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(Macleay)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="F69ECAAFDA343E4C0AC1FA6DFB42FEE4" lastPageId="16" lastPageNumber="217" pageId="14" pageNumber="215" type="description">
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<paragraph id="BE3B9924DA343E520AC1FA6DFEE5F985" blockId="14.[151,564,1481,1541]" box="[151,363,1516,1541]" pageId="14" pageNumber="215">
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(
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<figureCitation id="26BF85A1DA343E520AC9FA6DFE8DF985" box="[159,259,1516,1541]" captionStart="FIGURE 2" captionStartId="5.[151,250,1856,1878]" captionTargetBox="[174,1416,202,1827]" captionTargetId="figure@5.[163,1428,193,1835]" captionTargetPageId="5" captionText="FIGURE 2. Neighbor-Joining trees of p-distances based on 1,044 bp of aligned NADH2 sequence data for individual batoid genera or groups of genera. Taxon names are followed by country, host specimen number (in parentheses), and GenBank number. Outgroup taxa used in each analysis were omitted from the individual trees. Scale bar values indicate substitutions per site. Specimens deposited in the Blue Resources Trust Ichthyology Collection are indicted by stars (see text for details). A. Aetobatus (Aetobatidae). B. Brevitrygon (Dasyatidae). C. Himantura (Dasyatidae). D. Maculabatis (Dasyatidae). E. Neotrygon (Dasyatidae). F. Pastinachus (Dasyatidae). G. Pateobatis (Dasyatidae). H. Urogymnus (Dasyatidae)." httpUri="https://zenodo.org/record/2637240/files/figure.png" pageId="14" pageNumber="215">Figs. 2H</figureCitation>
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,
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<figureCitation id="26BF85A1DA343E520B58FA6DFEA6F985" box="[270,296,1516,1540]" captionStart="FIGURE 10" captionStartId="15.[151,250,1868,1890]" captionTargetBox="[151,1436,218,1819]" captionTargetId="figure@15.[151,1436,218,1819]" captionTargetPageId="15" captionText="FIGURE 10. Pateobatis jenkinsii (A–C) and Urogymnus granulatus (D–H) (Dasyatidae). A. Dorsal surface of mature female (SL-98). B. Wedge-shaped thorns and denticles on base of tail of mature female (SL-43). C. Dorsal surface of immature male (SL-101; BRT-I 0028). D. Dorsal surface of portion of disc of large specimen (partial; SL-13). E. Thorns and denticles on base of tail of mature specimen (SL-13). F. Ventral surface of portion of disc of large specimen (partial; SL-13). G. Dorsal surface of immature female (SL-116; BRT-I 0030). H. Scapular denticles of immature female (SL-116; BRT-I 0030)." httpUri="https://zenodo.org/record/2637256/files/figure.png" pageId="14" pageNumber="215">10</figureCitation>
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D–H)
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</paragraph>
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<paragraph id="BE3B9924DA343E520AC1F9B5FCA9F834" blockId="14.[151,1436,1588,1973]" pageId="14" pageNumber="215">
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The left portion of the disc of a specimen (SL-13) found in a fish market in Puttalam in the North Western Province was preliminarily identifiable as
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<taxonomicName id="7984E2A7DA343E520859F9D8FC9DF9F1" baseAuthorityName="Macleay" baseAuthorityYear="1883" box="[527,787,1623,1648]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="14" pageNumber="215" phylum="Chordata" rank="species" species="granulatus">
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<emphasis id="8CF04536DA343E520859F9D8FC9DF9F1" box="[527,787,1623,1648]" italics="true" pageId="14" pageNumber="215">Urogymnus granulatus</emphasis>
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</taxonomicName>
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based on the coloration of the dorsal surface of the disc which was yellowish brown with small white spots; it also had several irregular rows of enlarged, thorn-like denticles more or less restricted to the midline of the body. While this denticle pattern is inconsistent with that described for this species by
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<bibRefCitation id="DA15E4D5DA343E520BBCF944FD34F95C" author="Last" box="[490,698,1732,1757]" etAl="et al." firstAuthor="Last" pageId="14" pageNumber="215" pagination="345 - 368" refId="ref18604" refString="Last, P. R., Naylor, G. J. P. & Manjaji-Matsumoto, B. M. (2016 a) A revised classification of the family Dasyatidae (Chondrichthyes: Myliobatiformes) based on new morphological and molecular insights. Zootaxa, 4139, 345 - 368. https: // doi. org / 10.11646 / zootaxa. 4139.3.2" type="journal article" year="2016" yearSuffix="a">
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Last
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<emphasis id="8CF04536DA343E520872F944FDD1F95C" box="[548,607,1732,1757]" italics="true" pageId="14" pageNumber="215">et al.</emphasis>
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(2016a
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</bibRefCitation>
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,
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<bibRefCitation id="DA15E4D5DA343E52089EF945FD56F95C" author="Last, P. R. & White, W. T. & de Carvalho, M. R. & Seret, B. & Stehmann, M. F. W. & Naylor, G. J. P." box="[712,728,1732,1757]" pageId="14" pageNumber="215" refId="ref18781" refString="Last, P. R., White, W. T., de Carvalho, M. R., Seret, B., Stehmann, M. F. W. & Naylor, G. J. P. (2016 b) Rays of the World. Cornell University Press, Ithaca, NY, USA, pp. 790." type="book" year="2016">b</bibRefCitation>
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), it is consistent with that reported in the redescription of
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<taxonomicName id="7984E2A7DA343E520FD7F944FE9EF881" baseAuthorityName="Macleay" baseAuthorityYear="1883" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="14" pageNumber="215" phylum="Chordata" rank="species" species="granulatus">
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<emphasis id="8CF04536DA343E520FD7F944FE9EF881" italics="true" pageId="14" pageNumber="215">U. granulatus</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="DA15E4D5DA343E520B17F966FDB7F881" author="Ishihara" box="[321,569,1767,1792]" etAl="et al." firstAuthor="Ishihara" pageId="14" pageNumber="215" pagination="23 - 28" refId="ref18501" refString="Ishihara, H., Homma, K., Takeda, Y. & Randall, J. E. (1993) Redescription, distribution and food habits of the Indo-Pacific dasyatidid stingray Himantura granulata. Japanese Journal of Ichthyology, 40, 23 - 28." type="journal article" year="1993">
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Ishihara
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<emphasis id="8CF04536DA343E520BF3F968FE6EF881" box="[421,480,1767,1792]" italics="true" pageId="14" pageNumber="215">et al.</emphasis>
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(1993)
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</bibRefCitation>
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. This identification was supported by our molecular analysis in which this specimen was found to differ from our reference specimen of
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<taxonomicName id="7984E2A7DA343E52091AF88CFC66F8A4" baseAuthorityName="Macleay" baseAuthorityYear="1883" box="[844,1000,1804,1829]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="14" pageNumber="215" phylum="Chordata" rank="species" species="granulatus">
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<emphasis id="8CF04536DA343E52091AF88CFC66F8A4" box="[844,1000,1804,1829]" italics="true" pageId="14" pageNumber="215">U. granulatus</emphasis>
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</taxonomicName>
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(CM03-74;
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<accessionNumber id="A1D704C7DA343E520E22F88DFB67F8A4" box="[1140,1257,1804,1829]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ518808" pageId="14" pageNumber="215">JQ518808</accessionNumber>
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) from
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<collectingCountry id="C693D9B4DA343E520F63F88DFA12F8A4" box="[1333,1436,1804,1829]" name="Australia" pageId="14" pageNumber="215">Australia</collectingCountry>
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by only a single bp. Our molecular work also confirmed the identity of a specimen, which has been deposited in the BRT Ichthyology Collection (SL-116; BRT-I 0030), collected from the Valaichchenai Fisheries Harbor in the Eastern Province earlier in the year, as
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<taxonomicName id="7984E2A7DA343E52083FF8F8FC87F811" baseAuthorityName="Macleay" baseAuthorityYear="1883" box="[617,777,1911,1936]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="14" pageNumber="215" phylum="Chordata" rank="species" species="granulatus">
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<emphasis id="8CF04536DA343E52083FF8F8FC87F811" box="[617,777,1911,1936]" italics="true" pageId="14" pageNumber="215">U. granulatus</emphasis>
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</taxonomicName>
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. This immature specimen was identical to the reference specimen from
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<collectingCountry id="C693D9B4DA343E520B13F81DFE22F834" box="[325,428,1948,1973]" name="Australia" pageId="14" pageNumber="215">Australia</collectingCountry>
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and differed from SL-13 by 1 bp.
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</paragraph>
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<caption id="EAFBC9ACDA353E530AC1F8CDFB45F85D" httpUri="https://zenodo.org/record/2637256/files/figure.png" pageId="15" pageNumber="216" startId="15.[151,250,1868,1890]" targetBox="[151,1436,218,1819]" targetPageId="15">
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<paragraph id="BE3B9924DA353E530AC1F8CDFB45F85D" blockId="15.[151,1436,1868,2012]" pageId="15" pageNumber="216">
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<emphasis id="8CF04536DA353E530AC1F8CDFEAEF8E0" bold="true" box="[151,288,1868,1890]" pageId="15" pageNumber="216">FIGURE 10.</emphasis>
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<taxonomicName id="7984E2A7DA353E530B70F8CDFE61F8E3" baseAuthorityName="Annandale" baseAuthorityYear="1909" box="[294,495,1868,1890]" class="Chondrichthyes" family="Dasyatidae" genus="Pateobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="216" phylum="Chordata" rank="species" species="jenkinsii">
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<emphasis id="8CF04536DA353E530B70F8CDFE61F8E3" box="[294,495,1868,1890]" italics="true" pageId="15" pageNumber="216">Pateobatis jenkinsii</emphasis>
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</taxonomicName>
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(
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<emphasis id="8CF04536DA353E530BA9F8CDFDBEF8E3" bold="true" box="[511,560,1868,1890]" pageId="15" pageNumber="216">A–C</emphasis>
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) and
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<taxonomicName id="7984E2A7DA353E53083DF8CCFCDBF8E3" baseAuthorityName="Macleay" baseAuthorityYear="1883" box="[619,853,1868,1890]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="216" phylum="Chordata" rank="species" species="granulatus">
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<emphasis id="8CF04536DA353E53083DF8CCFCDBF8E3" box="[619,853,1868,1890]" italics="true" pageId="15" pageNumber="216">Urogymnus granulatus</emphasis>
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||
</taxonomicName>
|
||
(
|
||
<emphasis id="8CF04536DA353E530932F8CDFC19F8E0" bold="true" box="[868,919,1868,1890]" pageId="15" pageNumber="216">D–H</emphasis>
|
||
) (
|
||
<taxonomicName id="7984E2A7DA353E5309FBF8CDFBAAF8E3" authorityName="Jordan & Gilbert" authorityYear="1879" box="[941,1060,1868,1890]" class="Chondrichthyes" family="Dasyatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="216" phylum="Chordata" rank="family">Dasyatidae</taxonomicName>
|
||
).
|
||
<emphasis id="8CF04536DA353E530E62F8CDFBC3F8E0" bold="true" box="[1076,1101,1868,1889]" pageId="15" pageNumber="216">A.</emphasis>
|
||
Dorsal surface of mature female (SL-98).
|
||
<emphasis id="8CF04536DA353E530AA2F8EAFE85F801" bold="true" box="[244,267,1899,1920]" pageId="15" pageNumber="216">B.</emphasis>
|
||
Wedge-shaped thorns and denticles on base of tail of mature female (SL-43).
|
||
<emphasis id="8CF04536DA353E530E7AF8EBFBCBF801" bold="true" box="[1068,1093,1898,1920]" pageId="15" pageNumber="216">C.</emphasis>
|
||
Dorsal surface of immature male (SL-101; BRT-I 0028).
|
||
<emphasis id="8CF04536DA353E530BD3F808FE10F81F" bold="true" box="[389,414,1929,1950]" pageId="15" pageNumber="216">D.</emphasis>
|
||
Dorsal surface of portion of disc of large specimen (partial; SL-13).
|
||
<emphasis id="8CF04536DA353E530E0EF808FBE1F81F" bold="true" box="[1112,1135,1929,1950]" pageId="15" pageNumber="216">E.</emphasis>
|
||
Thorns and denticles on base of tail of mature specimen (SL-13).
|
||
<emphasis id="8CF04536DA353E530854F829FD9DF83C" bold="true" box="[514,531,1960,1981]" pageId="15" pageNumber="216">F.</emphasis>
|
||
Ventral surface of portion of disc of large specimen (partial; SL-13).
|
||
<emphasis id="8CF04536DA353E530E87F826FB6BF83C" bold="true" box="[1233,1253,1959,1981]" pageId="15" pageNumber="216">G.</emphasis>
|
||
Dorsal surface of immature female (SL-116; BRT-I 0030).
|
||
<emphasis id="8CF04536DA353E530860F846FDDEF85D" bold="true" box="[566,592,1991,2012]" pageId="15" pageNumber="216">H.</emphasis>
|
||
Scapular denticles of immature female (SL-116; BRT-I 0030).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BE3B9924DA2A3E4C0A91FF16FB42FEE4" blockId="16.[151,1436,151,357]" pageId="16" pageNumber="217">
|
||
This is the first report of
|
||
<taxonomicName id="7984E2A7DA2A3E4C0BB0FF18FD0CFF31" baseAuthorityName="Macleay" baseAuthorityYear="1883" box="[486,642,151,176]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="granulatus">
|
||
<emphasis id="8CF04536DA2A3E4C0BB0FF18FD0CFF31" box="[486,642,151,176]" italics="true" pageId="16" pageNumber="217">U. granulatus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="C693D9B4DA2A3E4C089CFF16FCB2FF31" box="[714,828,151,176]" name="Sri Lanka" pageId="16" pageNumber="217">Sri Lanka</collectingCountry>
|
||
. This may be the species referred to previously in
|
||
<collectingCountry id="C693D9B4DA2A3E4C0F2DFF16FF53FF54" name="Sri Lanka" pageId="16" pageNumber="217">Sri Lanka</collectingCountry>
|
||
as
|
||
<taxonomicName id="7984E2A7DA2A3E4C0B5EFF3CFD78FF54" authority="(Bloch & Schneider)" authorityName="Bloch & Schneider" baseAuthorityName="Bloch & Schneider" baseAuthorityYear="1801" box="[264,758,188,213]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="africanus">
|
||
<emphasis id="8CF04536DA2A3E4C0B5EFF3CFE70FF55" box="[264,510,188,212]" italics="true" pageId="16" pageNumber="217">Urogymnus africanus</emphasis>
|
||
(Bloch & Schneider)
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="DA15E4D5DA2A3E4C0970FF3CFC51FF55" author="De Silva" box="[806,991,188,213]" firstAuthor="De Silva" pageId="16" pageNumber="217" pagination="301 - 306" refId="ref17494" refString="De Silva, R. I. (1978) Some skates and rays of Sri Lanka. Loris, 14, 301 - 306." type="journal article" year="1978">De Silva (1978)</bibRefCitation>
|
||
or possibly as
|
||
<taxonomicName id="7984E2A7DA2A3E4C0EC4FF3CFE0CFF79" authority="(Bloch & Schneider)" authorityName="Bloch & Schneider" baseAuthorityName="Bloch & Schneider" baseAuthorityYear="1801" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="8CF04536DA2A3E4C0EC4FF3CFA12FF55" box="[1170,1436,189,212]" italics="true" pageId="16" pageNumber="217">Urogymnus asperrimus</emphasis>
|
||
(Bloch & Schneider)
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="DA15E4D5DA2A3E4C0B96FF61FD23FF79" author="De Bruin" box="[448,685,223,248]" etAl="et al." firstAuthor="De Bruin" pageId="16" pageNumber="217" refId="ref17417" refString="De Bruin, G. H. P., Russell, B. C. & Bogusch, A. (1995) FAO Species Identification Field Guide for Fisheries Purposes. The Marine Fishery Resources of Sri Lanka. FAO, Rome, Italy, 400 pp." type="book" year="1995">
|
||
De Bruin
|
||
<emphasis id="8CF04536DA2A3E4C0867FF60FDE4FF79" box="[561,618,223,248]" italics="true" pageId="16" pageNumber="217">et al.</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DA15E4D5DA2A3E4C08ECFF61FC05FF79" author="Moron" box="[698,907,223,248]" etAl="et al." firstAuthor="Moron" pageId="16" pageNumber="217" pagination="142 - 157" refId="ref19181" refString="Moron, J., Bertrand, B. & Last, P. (1998) A check-list of sharks and rays of western Sri Lanka. Journal of the Marine Biological Association of India, 40, 142 - 157." type="journal article" year="1998">
|
||
Morón
|
||
<emphasis id="8CF04536DA2A3E4C0958FF60FCC6FF79" box="[782,840,223,248]" italics="true" pageId="16" pageNumber="217">et al.</emphasis>
|
||
1998
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DA15E4D5DA2A3E4C09CEFF61FBB1FF79" author="De Silva" box="[920,1087,223,248]" firstAuthor="De Silva" pageId="16" pageNumber="217" pagination="294 - 301" refId="ref17594" refString="De Silva, R. I. (2006) Taxonomy and status of the sharks and rays of Sri Lanka. In: Bambaradeniya, C. N. B. (Ed.), The Fauna of Sri Lanka: Status of Taxonomy, Research and Conservation. The World Conservation Union (IUCN), Sri Lanka and Government of Sri Lanka, Colombo, Sri Lanka, pp. 294 - 301." type="book chapter" year="2006">De Silva 2006</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="DA15E4D5DA2A3E4C0E05FF61FAABFF79" author="Last" box="[1107,1317,223,248]" etAl="et al." firstAuthor="Last" pageId="16" pageNumber="217" refId="ref18781" refString="Last, P. R., White, W. T., de Carvalho, M. R., Seret, B., Stehmann, M. F. W. & Naylor, G. J. P. (2016 b) Rays of the World. Cornell University Press, Ithaca, NY, USA, pp. 790." type="book" year="2016" yearSuffix="b">
|
||
Last
|
||
<emphasis id="8CF04536DA2A3E4C0EDDFF60FB30FF79" box="[1163,1214,223,248]" italics="true" pageId="16" pageNumber="217">et al</emphasis>
|
||
. (2016b)
|
||
</bibRefCitation>
|
||
suggested that both
|
||
<taxonomicName id="7984E2A7DA2A3E4C0B52FE84FE2FFE9C" baseAuthorityName="Macleay" baseAuthorityYear="1883" box="[260,417,260,285]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="granulatus">
|
||
<emphasis id="8CF04536DA2A3E4C0B52FE84FE2FFE9C" box="[260,417,260,285]" italics="true" pageId="16" pageNumber="217">U. granulatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7984E2A7DA2A3E4C0B8DFE84FDF3FE9D" baseAuthorityName="Bloch & Schneider" baseAuthorityYear="1801" box="[475,637,261,284]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="8CF04536DA2A3E4C0B8DFE84FDF3FE9D" box="[475,637,261,284]" italics="true" pageId="16" pageNumber="217">U. asperrimus</emphasis>
|
||
</taxonomicName>
|
||
co-occur in
|
||
<collectingCountry id="C693D9B4DA2A3E4C0959FE85FC0FFE9C" box="[783,897,260,285]" name="Sri Lanka" pageId="16" pageNumber="217">Sri Lanka</collectingCountry>
|
||
, and a specimen consistent with
|
||
<taxonomicName id="7984E2A7DA2A3E4C0EACFE84FA12FE9D" baseAuthorityName="Bloch & Schneider" baseAuthorityYear="1801" box="[1274,1436,261,284]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="8CF04536DA2A3E4C0EACFE84FA12FE9D" box="[1274,1436,261,284]" italics="true" pageId="16" pageNumber="217">U. asperrimus</emphasis>
|
||
</taxonomicName>
|
||
collected since our survey confirms this. However, whether the form of this species in
|
||
<collectingCountry id="C693D9B4DA2A3E4C0ED2FEA6FB7BFEC1" box="[1156,1269,295,320]" name="Sri Lanka" pageId="16" pageNumber="217">Sri Lanka</collectingCountry>
|
||
is
|
||
<taxonomicName id="7984E2A7DA2A3E4C0F4CFEA8FE9BFEE5" baseAuthorityName="Bloch & Schneider" baseAuthorityYear="1801" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="8CF04536DA2A3E4C0F4CFEA8FE9BFEE5" italics="true" pageId="16" pageNumber="217">Urogymnus asperrimus</emphasis>
|
||
</taxonomicName>
|
||
1 or
|
||
<taxonomicName id="7984E2A7DA2A3E4C0B19FECCFDD8FEE5" baseAuthorityName="Bloch & Schneider" baseAuthorityYear="1801" box="[335,598,333,356]" class="Chondrichthyes" family="Dasyatidae" genus="Urogymnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="8CF04536DA2A3E4C0B19FECCFDD8FEE5" box="[335,598,333,356]" italics="true" pageId="16" pageNumber="217">Urogymnus asperrimus</emphasis>
|
||
</taxonomicName>
|
||
2 (sensu
|
||
<bibRefCitation id="DA15E4D5DA2A3E4C0896FECDFC14FEE5" author="Naylor" box="[704,922,332,357]" etAl="et al." firstAuthor="Naylor" pageId="16" pageNumber="217" pagination="1 - 262" refId="ref19577" refString="Naylor, G. J. P., Caira, J. N., Jensen, K., Rosana, K. A. M., White, W. T. & Last, P. R. (2012 a) A DNA sequence-based approach to the identification of shark and ray species and its implications for global elasmobranch diversity and parasitology. Bulletin of the American Museum of Natural History, 367, 1 - 262." type="journal article" year="2012" yearSuffix="a">
|
||
Naylor
|
||
<emphasis id="8CF04536DA2A3E4C0943FECCFCC3FEE4" box="[789,845,332,357]" italics="true" pageId="16" pageNumber="217">et al.</emphasis>
|
||
2012a
|
||
</bibRefCitation>
|
||
) remains to be determined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |