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<mods:title id="B415F095380C37B86963030D699734C8">Notes on Hexapodidae (Crustacea, Brachyura) from Indonesia and Malaysia, with description of a new species of Mariaplax Rahayu &amp; Ng, 2014, from Lombok, Indonesia</mods:title>
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<mods:namePart id="5FA6E7D4D3705AAE635263E6B9B7CA58">Ng, Peter K. L.</mods:namePart>
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<treatment id="038DFD0AFF98E267FF0DFC25FE29CE2D" ID-DOI="http://doi.org/10.5281/zenodo.5457336" ID-GBIF-Taxon="119634730" ID-Zenodo-Dep="5457336" LSID="urn:lsid:plazi:treatment:038DFD0AFF98E267FF0DFC25FE29CE2D" httpUri="http://treatment.plazi.org/id/038DFD0AFF98E267FF0DFC25FE29CE2D" lastPageId="9" lastPageNumber="134" pageId="1" pageNumber="126">
<subSubSection id="C33E1F97FF98E26FFF0DFC25FEA9CF48" pageId="1" pageNumber="126" type="nomenclature">
<paragraph id="8B9B4C1CFF98E26FFF0DFC25FDF9C8AE" blockId="1.[151,578,992,1053]" box="[151,578,992,1019]" pageId="1" pageNumber="126">
<heading id="D0D3FB70FF98E26FFF0DFC25FDF9C8AE" bold="true" box="[151,578,992,1019]" fontSize="11" level="1" pageId="1" pageNumber="126" reason="1">
<taxonomicName id="4C24379FFF98E26FFF0DFC25FDF9C8AE" ID-CoL="3LGMV" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[151,578,992,1019]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF98E26FFF0DFC25FDF9C8AE" bold="true" box="[151,578,992,1019]" pageId="1" pageNumber="126">
<emphasis id="B950900EFF98E26FFF0DFC25FECBC8AF" bold="true" box="[151,368,992,1018]" italics="true" pageId="1" pageNumber="126">Hexapinus latipes</emphasis>
(De
<bibRefCitation id="EFB531EDFF98E26FFE31FC24FD81C8AE" author="Haan" box="[427,570,993,1019]" pageId="1" pageNumber="126" refString="Haan, W. de (1835) Crustacea. Fauna Japonica, sive Descriptio animalium, quae in itinere per Japoniam, jussu et auspiciis superiorum, qui summum in India Batava imperium tenent, suscepto, annis 1823 - 1830 collegit, notis, observationibus et adumbrationibus illustravit P. F. de Siebold. Conjunctis studiis C. J. Temminck et H. Schlegel pro Vertebratis atque W. de Haan pro Invertebratis elaborata Regis aupicus edita. P. F. v. Siebold. Leiden, Lugundi-Batavorum. Decas II, 25 - 64, pls. 9 - 15, 17, C, D." type="book chapter" year="1835">Haan, 1835</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9B4C1CFF98E26FFF0DFBC1FEA9CF48" blockId="1.[151,578,992,1053]" box="[151,274,1028,1053]" pageId="1" pageNumber="126">
(
<figureCitation id="131F5099FF98E26FFF05FBC1FEB1CF48" box="[159,266,1028,1053]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6. A F" captionStartId-0="3.[151,250,1855,1877]" captionStartId-1="4.[151,250,1921,1943]" captionStartId-2="5.[151,250,1917,1939]" captionStartId-3="6.[151,250,1875,1897]" captionStartId-4="7.[151,250,1246,1268]" captionTargetBox-0="[151,1436,193,1832]" captionTargetBox-1="[151,1436,193,1898]" captionTargetBox-2="[210,1369,193,1875]" captionTargetBox-3="[151,1436,193,1853]" captionTargetBox-4="[163,1419,187,1223]" captionTargetId-0="figure@3.[151,1436,193,1833]" captionTargetId-1="figure@4.[151,1436,193,1900]" captionTargetId-2="figure@5.[210,1371,193,1876]" captionTargetId-3="figure@6.[151,1436,193,1853]" captionTargetId-4="figure@7.[151,1435,175,1233]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." captionText-1="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." captionText-2="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." captionText-3="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." captionText-4="FIGURE 6. A F, Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; G, H, Mariaplax aspera n. sp., holotype female (6.4 × 4.4 mm) (MZB), Lombok, Indonesia. A, ventral view of left G 1; B, dorsal view of left G 1; C, ventral view of distal part of left G 1; D, dorsal view of distal part of left G 1 (setae not drawn); E, left G 2 (ventral view); F, distal half of left G 2 (dorsal view); G, left third maxilliped; H, female abdomen. Scales: A, B, E, G, H = 1.0 mm; C, D, F = 0.5 mm." httpUri-0="https://zenodo.org/record/241817/files/figure.png" httpUri-1="https://zenodo.org/record/241818/files/figure.png" httpUri-2="https://zenodo.org/record/241819/files/figure.png" httpUri-3="https://zenodo.org/record/241820/files/figure.png" httpUri-4="https://zenodo.org/record/241821/files/figure.png" pageId="1" pageNumber="126">Figs. 26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E1F97FF98E26FFF0DFB88FD62CF89" pageId="1" pageNumber="126" type="reference_group">
<paragraph id="8B9B4C1CFF98E26FFF0DFB88FDA4CFD5" blockId="1.[151,1114,1100,1183]" pageId="1" pageNumber="126">
<treatmentCitationGroup id="AB346B32FF98E26FFF0DFB88FDA4CFD5" pageId="1" pageNumber="126">
<taxonomicName id="4C24379FFF98E26FFF0DFB88FE83CF37" ID-CoL="3LGRL" box="[151,312,1101,1122]" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="sexpes">
<emphasis id="B950900EFF98E26FFF0DFB88FE83CF37" box="[151,312,1101,1122]" italics="true" pageId="1" pageNumber="126">Hexapus sexpes</emphasis>
</taxonomicName>
—De
<treatmentCitation id="0A856A0DFF98E26FFEEFFB88FDA9CF37" author="Man" box="[373,530,1100,1122]" page="322" pageId="1" pageNumber="126" year="1888">
<bibRefCitation id="EFB531EDFF98E26FFEEFFB88FE5BCF37" author="Man" box="[373,480,1100,1122]" pageId="1" pageNumber="126" refString="Man, J. G. de (1888) Bericht uber die von Herrn Dr. J. Brock im indischen Archipel gesammelten Decapoden und Stomatopoden. Archiv fur Naturgeschichte, Berlin, 53 (1), 289 - 600, pls. 11 - 22." type="journal article" year="1888">Man 1888</bibRefCitation>
: 322
</treatmentCitation>
, pl. 13 figs. 3, 3a, 3b (not
<taxonomicName id="4C24379FFF98E26FFC86FB89FBE8CF37" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[796,1107,1100,1122]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="sexpes">
<emphasis id="B950900EFF98E26FFC86FB89FC15CF37" box="[796,942,1100,1122]" italics="true" pageId="1" pageNumber="126">Cancer sexpes</emphasis>
<bibRefCitation id="EFB531EDFF98E26FFC2FFB89FBE8CF37" author="Fabricius" box="[949,1107,1100,1122]" pageId="1" pageNumber="126" refString="Fabricius, J. C. (1798) Supplementum Entomologiae systematicae. Proft et Storch, Hafniae, 572 pp." type="book" year="1798">Fabricius, 1798</bibRefCitation>
</taxonomicName>
)
<taxonomicName id="4C24379FFF98E26FFF0DFBAEFE60CFD5" ID-CoL="3LGRK" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[151,475,1130,1152]" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF98E26FFF0DFBAEFE81CFD5" box="[151,314,1130,1152]" italics="true" pageId="1" pageNumber="126">Hexapus latipes</emphasis>
De
<bibRefCitation id="EFB531EDFF98E26FFEF9FBAEFE60CFD5" author="Haan" box="[355,475,1130,1152]" pageId="1" pageNumber="126" refString="Haan, W. de (1835) Crustacea. Fauna Japonica, sive Descriptio animalium, quae in itinere per Japoniam, jussu et auspiciis superiorum, qui summum in India Batava imperium tenent, suscepto, annis 1823 - 1830 collegit, notis, observationibus et adumbrationibus illustravit P. F. de Siebold. Conjunctis studiis C. J. Temminck et H. Schlegel pro Vertebratis atque W. de Haan pro Invertebratis elaborata Regis aupicus edita. P. F. v. Siebold. Leiden, Lugundi-Batavorum. Decas II, 25 - 64, pls. 9 - 15, 17, C, D." type="book chapter" year="1835">Haan, 1835</bibRefCitation>
</taxonomicName>
: pl. D.
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</paragraph>
<paragraph id="8B9B4C1CFF98E26FFF0DFB4FFD5ECFCA" blockId="1.[151,1114,1100,1183]" box="[151,741,1161,1183]" pageId="1" pageNumber="126">
<treatmentCitationGroup id="AB346B32FF98E26FFF0DFB4FFD5ECFCA" box="[151,741,1161,1183]" pageId="1" pageNumber="126">
<taxonomicName id="4C24379FFF98E26FFF0DFB4FFEF6CFCA" ID-CoL="3LGMV" box="[151,333,1161,1183]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF98E26FFF0DFB4FFEF6CFCA" box="[151,333,1161,1183]" italics="true" pageId="1" pageNumber="126">Hexapinus latipes</emphasis>
</taxonomicName>
<treatmentCitation id="0A856A0DFF98E26FFEFDFB4CFDD9CFCA" author="Rahayu" box="[359,610,1161,1183]" httpUri="http://treatment.plazi.org/id/03EB87DA3B36E344FC7AFA15FB1F9A70" page="409" pageId="1" pageNumber="126" year="2014">
<bibRefCitation id="EFB531EDFF98E26FFEFDFB4CFD8BCFCA" author="Rahayu" box="[359,560,1161,1183]" pageId="1" pageNumber="126" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng 2014</bibRefCitation>
: 409
</treatmentCitation>
, figs. 1013.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9B4C1CFF98E26FFF0DFB03FD62CF89" blockId="1.[151,729,1222,1244]" box="[151,729,1222,1244]" pageId="1" pageNumber="126">
See
<bibRefCitation id="EFB531EDFF98E26FFF58FB03FE70CF89" author="Rahayu" box="[194,459,1222,1244]" pageId="1" pageNumber="126" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng (2014: 409)</bibRefCitation>
for a complete synonymy.
</paragraph>
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<emphasis id="B950900EFF98E26FFF0DFACCFEC0CE77" bold="true" box="[151,379,1289,1314]" pageId="1" pageNumber="126">Material examined</emphasis>
.
<typeStatus id="549FF2BEFF98E26FFE11FACFFE40CE77" box="[395,507,1290,1314]" pageId="1" pageNumber="126" type="lectotype">Lectotype</typeStatus>
female (10.0 ×
<quantity id="4CDCE1F9FF98E26FFD2EFACCFCB5CE77" box="[692,782,1289,1314]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="1" pageNumber="126" unit="mm" value="6.7">6.7 mm</quantity>
) (
<collectionCode id="ED35D4D9FF98E26FFCBFFACFFCC0CE77" LSID="urn:lsid:biocol.org:col:34992" box="[805,891,1290,1314]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="126">RMNH</collectionCode>
31783d),
<collectingCountry id="F3330C8CFF98E26FFC6BFACFFB8FCE74" box="[1009,1076,1290,1313]" name="Japan" pageId="1" pageNumber="126">Japan</collectingCountry>
. Others:
<emphasis id="B950900EFF98E26FFB06FACCFB5ECE77" bold="true" box="[1180,1253,1289,1314]" pageId="1" pageNumber="126">
<collectingCountry id="F3330C8CFF98E26FFB06FACCFB5ECE77" box="[1180,1253,1289,1314]" name="Japan" pageId="1" pageNumber="126">Japan</collectingCountry>
</emphasis>
:
<specimenCount id="9D228795FF98E26FFB6CFACFFAE3CE77" box="[1270,1368,1289,1314]" pageId="1" pageNumber="126" type="female">1 female</specimenCount>
(10.2 ×
<quantity id="4CDCE1F9FF98E26FFF3FFAE8FEBBCE13" box="[165,256,1325,1350]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="1" pageNumber="126" unit="mm" value="7.6">7.6 mm</quantity>
) (NSMT-Cr 5694), Kushimoto, Kii Peninsula, Wakayama Prefecture,
<geoCoordinate id="EE102ADBFF98E26FFBBCFAEBFB2CCE13" box="[1062,1175,1325,1350]" direction="north" orientation="latitude" pageId="1" pageNumber="126" precision="92" value="33.453335">
33
<emphasis id="B950900EFF98E26FFBDBFAE8FBF7CE13" bold="true" box="[1089,1100,1325,1350]" pageId="1" pageNumber="126">°</emphasis>
27.2'N
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135
<emphasis id="B950900EFF98E26FFB50FAE8FB6ECE13" bold="true" box="[1226,1237,1325,1350]" pageId="1" pageNumber="126">°</emphasis>
45.4'E
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33
<emphasis id="B950900EFF98E26FFADCFAE8FAEACE13" bold="true" box="[1350,1361,1325,1350]" pageId="1" pageNumber="126">°</emphasis>
27.2'N
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135
<emphasis id="B950900EFF98E26FFF5BFA94FF77CE3F" bold="true" box="[193,204,1361,1386]" pageId="1" pageNumber="126">°</emphasis>
45.6'E
</geoCoordinate>
,
<quantity id="4CDCE1F9FF98E26FFEBCFA97FE2BCE3C" box="[294,400,1362,1386]" metricMagnitude="1" metricUnit="m" metricValue="2.3" metricValueMax="2.7" metricValueMin="1.9" pageId="1" pageNumber="126" unit="m" value="23.0" valueMax="27.0" valueMin="19.0">1927 m</quantity>
, dredge, coll. M. Takeda,
<date id="FF9A6ADCFF98E26FFD57FA97FCD1CE3F" box="[717,874,1361,1386]" pageId="1" pageNumber="126" value="1978-07-17">17 July 1978</date>
;
<specimenCount id="9D228795FF98E26FFCE1FA97FC5ACE3F" box="[891,993,1361,1386]" pageId="1" pageNumber="126" type="female">1 female</specimenCount>
(30.4 ×
<quantity id="4CDCE1F9FF98E26FFBCAFA97FB02CE3F" box="[1104,1209,1362,1386]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="1" pageNumber="126" unit="mm" value="17.9">17.9 mm</quantity>
) (KPM NH 7123), Enoshima Island, T. Sakai Collection;
<specimenCount id="9D228795FF98E26FFDDFFAB3FD1DCEDB" box="[581,678,1397,1422]" pageId="1" pageNumber="126" type="female">1 female</specimenCount>
(28.7 ×
<quantity id="4CDCE1F9FF98E26FFC9FFAB0FCD2CEDB" box="[773,873,1397,1422]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="1" pageNumber="126" unit="mm" value="16.7">16.7 mm</quantity>
) (KPM NH 7460) Minabe, Wakayama Prefecture, T. Sakai Collection;
<specimenCount id="9D228795FF98E26FFE11FA5FFE4BCEE7" box="[395,496,1433,1458]" pageId="1" pageNumber="126" type="female">1 female</specimenCount>
(30.4 ×
<quantity id="4CDCE1F9FF98E26FFDC4FA5CFD73CEE7" box="[606,712,1433,1458]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="126" unit="mm" value="17.5">17.5 mm</quantity>
) (KPM NH 6331), Mikawa-Ishiki, Aichi Prefecture, T. Sakai Collection.
<collectingCountry id="F3330C8CFF98E26FFE83FA78FE35CE83" box="[281,398,1469,1494]" name="Indonesia" pageId="1" pageNumber="126">
<emphasis id="B950900EFF98E26FFE83FA78FE35CE83" bold="true" box="[281,398,1469,1494]" pageId="1" pageNumber="126">Indonesia</emphasis>
</collectingCountry>
:
<specimenCount id="9D228795FF98E26FFE07FA7BFE53CE83" box="[413,488,1469,1494]" pageId="1" pageNumber="126" type="male">1 male</specimenCount>
(20.9 ×
<quantity id="4CDCE1F9FF98E26FFDD2FA78FD0BCE83" box="[584,688,1469,1494]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="126" unit="mm" value="12.5">12.5 mm</quantity>
),
<specimenCount id="9D228795FF98E26FFD59FA7BFC9FCE83" box="[707,804,1469,1494]" pageId="1" pageNumber="126" type="female">1 female</specimenCount>
(18.9 ×
<quantity id="4CDCE1F9FF98E26FFC19FA7BFC52CE83" box="[899,1001,1470,1494]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="126" unit="mm" value="12.1">12.1 mm</quantity>
) (
<collectionCode id="ED35D4D9FF98E26FFC65FA7BFB82CE83" LSID="urn:lsid:biocol.org:col:34838" box="[1023,1081,1470,1494]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="126">SMF</collectionCode>
<collectionCode id="ED35D4D9FF98E26FFBDAFA7BFBC5CE83" box="[1088,1150,1470,1494]" httpUri="http://grbio.org/cool/gs4h-1fee" name="Zoologischen Museums Greifswald" pageId="1" pageNumber="126">ZMG</collectionCode>
160), South
<collectingCountry id="F3330C8CFF98E26FFA95FA78FAE9CE83" box="[1295,1362,1469,1494]" name="China" pageId="1" pageNumber="126">China</collectingCountry>
Sea at
<collectingCountry id="F3330C8CFF98E26FFF0DFA24FEBFCEAF" box="[151,260,1505,1530]" name="Indonesia" pageId="1" pageNumber="126">Moluccas</collectingCountry>
Seas,
<collectingCountry id="F3330C8CFF98E26FFED6FA24FE18CEAF" box="[332,419,1505,1530]" name="Indonesia" pageId="1" pageNumber="126">Ambon</collectingCountry>
,
<geoCoordinate id="EE102ADBFF98E26FFE34FA27FD96CEAF" box="[430,557,1506,1530]" direction="south" orientation="latitude" pageId="1" pageNumber="126" precision="9" value="-13.7005">13°42.03S</geoCoordinate>
<geoCoordinate id="EE102ADBFF98E26FFDAEFA24FD7ECEAF" box="[564,709,1505,1530]" direction="east" orientation="longitude" pageId="1" pageNumber="126" precision="1" value="128.16444">128°9.867E</geoCoordinate>
, coll. J.G.F. Brock,
<date id="FF9A6ADCFF98E26FFC3AFA27FBD4CEAF" box="[928,1135,1505,1530]" pageId="1" pageNumber="126" value="1885-09-07">7 September 1885</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E1F97FF98E26CFF5DF9C0FE6ECB80" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126" type="diagnosis">
<paragraph id="8B9B4C1CFF98E26CFF5DF9C0FE6ECB80" blockId="1.[151,1437,1289,2034]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126">
<emphasis id="B950900EFF98E26FFF5DF9C0FEFACD4B" bold="true" box="[199,321,1541,1566]" pageId="1" pageNumber="126">Diagnosis.</emphasis>
Carapace subquadrate, much wider than long, width 1.31.7 times length, dorsal surfaces almost entirely covered with distinct pits (
<figureCitation id="131F5099FF98E26FFDAEF9ECFDC5CD17" box="[564,638,1577,1602]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="1" pageNumber="126">Fig. 2</figureCitation>
); anterolateral margin lined with distinct rounded granules (
<figureCitation id="131F5099FF98E26FFADEF9ECFA36CD17" box="[1348,1421,1577,1602]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="1" pageNumber="126">Fig. 2</figureCitation>
); suborbital, subhepatic, and upper part of pterygostomial regions covered with flattened granules, and punctate (
<figureCitation id="131F5099FF98E26FFAF5F988FF1FCDDF" captionStart="FIGURE 3" captionStartId="4.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1898]" captionTargetId="figure@4.[151,1436,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="1" pageNumber="126">Fig. 3</figureCitation>
A, B); pterygostomial region with oblique striae, dense setae under striae, adjacent to Milne Edwards opening (
<figureCitation id="131F5099FF98E26FFF05F950FF53CDFB" box="[159,232,1685,1710]" captionStart="FIGURE 3" captionStartId="4.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1898]" captionTargetId="figure@4.[151,1436,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="1" pageNumber="126">Fig. 3</figureCitation>
B, C); outer surfaces of third maxillipeds covered with numerous small granules; ischium expanded proximolaterally, distomesially, subequal in length to merus medially, with gently convex mesial margin (
<figureCitation id="131F5099FF98E26FFAAFF97CFA38CD87" box="[1333,1411,1721,1746]" captionStart="FIGURE 3" captionStartId="4.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1898]" captionTargetId="figure@4.[151,1436,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="1" pageNumber="126">Figs. 3</figureCitation>
B, C, 4AC); outer surfaces of chelipeds granulated, adult major male chela with cluster of granules expanded inward forming triangular structure in upper view, followed downward by median tubercles (
<figureCitation id="131F5099FF98E26FFBEDF8C4FB05CC4F" box="[1143,1214,1793,1818]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="1" pageNumber="126">Fig. 4</figureCitation>
E, F); P2P4 short, stout, outer surfaces with numerous distinct small granules, short setae (
<figureCitation id="131F5099FF98E26FFC27F8E0FBB6CC6B" box="[957,1037,1829,1854]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="1" pageNumber="126">Figs. 2</figureCitation>
,
<figureCitation id="131F5099FF98E26FFB83F8E3FB93CC6B" box="[1049,1064,1830,1854]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="1" pageNumber="126">4</figureCitation>
GI); male thoracic sternum with sternites 1, 2 fused, forming subtriangular structure, each with deep transverse groove that does not extend to lateral margins, posteromedian part of sternite 3 with 2 granulated projections that bracket telson (
<figureCitation id="131F5099FF98E26FFA96F8A8FAEACCD3" box="[1292,1361,1901,1926]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="1" pageNumber="126">Fig. 5</figureCitation>
A, D); sternite 3 delimited from sternite 4 by deep tranverse groove that forms thoracic sternal groove, relatively shallower along margins (
<figureCitation id="131F5099FF98E26FFE24F870FDB9CC9B" box="[446,514,1973,1998]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="1" pageNumber="126">Fig. 5</figureCitation>
A, D); male abdomen (
<figureCitation id="131F5099FF98E26FFC96F870FCF4CC9B" box="[780,847,1973,1998]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="1" pageNumber="126">Fig. 5</figureCitation>
AC) with somites 1, 2 free; somites 35 fused but median parts of sutures still visible, somite 6 longer than broad, shorter than fused somites 35, longer than telson, telson pentagonal (
<figureCitation id="131F5099FF9BE26CFEE2FF52FE79CBE5" box="[376,450,151,176]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="2" pageNumber="127">Fig. 5</figureCitation>
A, B); G1 slender, curved along distal third, distalmost part gradually tapering to subtruncate tip (
<figureCitation id="131F5099FF9BE26CFED6FF79FE72CB80" box="[332,457,188,213]" captionStart="FIGURE 6. A F" captionStartId="7.[151,250,1246,1268]" captionTargetBox="[163,1419,187,1223]" captionTargetId="figure@7.[151,1435,175,1233]" captionTargetPageId="7" captionText="FIGURE 6. A F, Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; G, H, Mariaplax aspera n. sp., holotype female (6.4 × 4.4 mm) (MZB), Lombok, Indonesia. A, ventral view of left G 1; B, dorsal view of left G 1; C, ventral view of distal part of left G 1; D, dorsal view of distal part of left G 1 (setae not drawn); E, left G 2 (ventral view); F, distal half of left G 2 (dorsal view); G, left third maxilliped; H, female abdomen. Scales: A, B, E, G, H = 1.0 mm; C, D, F = 0.5 mm." httpUri="https://zenodo.org/record/241821/files/figure.png" pageId="2" pageNumber="127">Fig. 6AD</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5B1C94FF9BE26CFF0DF842FA9FCC8F" httpUri="https://zenodo.org/record/241816/files/figure.png" pageId="2" pageNumber="127" targetBox="[151,1436,265,1904]" targetPageId="2">
<paragraph id="8B9B4C1CFF9BE26CFF0DF842FA9FCC8F" blockId="2.[151,1436,1927,2010]" pageId="2" pageNumber="127">
<emphasis id="B950900EFF9BE26CFF0DF842FEA3CCC9" bold="true" box="[151,280,1927,1949]" pageId="2" pageNumber="127">FIGURE 1.</emphasis>
Overall habitus. A,
<taxonomicName id="4C24379FFF9BE26CFE62F84DFCE5CCC8" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[504,862,1927,1949]" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="sexpes">
<emphasis id="B950900EFF9BE26CFE62F84DFD25CCC8" box="[504,670,1928,1949]" italics="true" pageId="2" pageNumber="127">Hexapus sexpes</emphasis>
(Fabricius, 1798)
</taxonomicName>
, male (7.3 × 4.5 mm, 6.0 × 3.6 mm) (ZRC), Jeram, Peninsular Malaysia; B,
<taxonomicName id="4C24379FFF9BE26CFE00F862FD47CCEE" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[410,764,1957,1980]" class="Malacostraca" family="Hexapodidae" genus="Hexapus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="sexpes">
<emphasis id="B950900EFF9BE26CFE00F862FD85CCE9" box="[410,574,1959,1980]" italics="true" pageId="2" pageNumber="127">Hexapus sexpes</emphasis>
(Fabricius, 1798)
</taxonomicName>
, female (7.6 × 4.7 mm) (ZRC), Jeram, Peninsular Malaysia; C,
<taxonomicName id="4C24379FFF9BE26CFF0DF800FDB4CC8F" authority="Rahayu &amp; Ng, 2014" authorityName="Rahayu &amp; Ng" authorityYear="2014" box="[151,527,1988,2010]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B950900EFF9BE26CFF0DF800FE81CC8F" box="[151,314,1988,2010]" italics="true" pageId="2" pageNumber="127">Hexapinus latus</emphasis>
Rahayu &amp; Ng, 2014
</taxonomicName>
, female (7.3 × 4.2 mm) (NNM-ZMA Crust De 241762b), Paternoster Islands.
</paragraph>
</caption>
<caption id="DF5B1C94FF9AE26DFF0DF8FAFF51CCC7" httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="3" pageNumber="128" targetBox="[151,1436,193,1832]" targetPageId="3">
<paragraph id="8B9B4C1CFF9AE26DFF0DF8FAFF51CCC7" blockId="3.[151,1435,1855,1938]" pageId="3" pageNumber="128">
<emphasis id="B950900EFF9AE26DFF0DF8FAFEAECC01" bold="true" box="[151,277,1855,1877]" pageId="3" pageNumber="128">FIGURE 2.</emphasis>
<taxonomicName id="4C24379FFF9AE26DFE84F885FD2BCC00" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[286,656,1855,1877]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF9AE26DFE84F885FE6CCC00" box="[286,471,1855,1877]" italics="true" pageId="3" pageNumber="128">Hexapinus latipes</emphasis>
(De Haan, 1835)
</taxonomicName>
, overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160b), Ambon.
</paragraph>
</caption>
<caption id="DF5B1C94FF9DE26AFF0DF844FE49CC81" httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="4" pageNumber="129" targetBox="[151,1436,193,1898]" targetPageId="4">
<paragraph id="8B9B4C1CFF9DE26AFF0DF844FE49CC81" blockId="4.[151,1436,1921,2004]" pageId="4" pageNumber="129">
<emphasis id="B950900EFF9DE26AFF0DF844FEA9CCC3" bold="true" box="[151,274,1921,1943]" pageId="4" pageNumber="129">FIGURE 3.</emphasis>
<taxonomicName id="4C24379FFF9DE26AFE82F847FD3BCCC2" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[280,640,1921,1943]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF9DE26AFE82F847FE74CCC2" box="[280,463,1921,1943]" italics="true" pageId="4" pageNumber="129">Hexapinus latipes</emphasis>
(De Haan, 1835)
</taxonomicName>
, male (20.9 × 12.5 mm) (SMF ZMG 160a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 14.
</paragraph>
</caption>
<caption id="DF5B1C94FF9CE26BFF0DF8B8FC18CCBA" httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="5" pageNumber="130" targetBox="[210,1369,193,1875]" targetPageId="5">
<paragraph id="8B9B4C1CFF9CE26BFF0DF8B8FC18CCBA" blockId="5.[151,1436,1917,2031]" pageId="5" pageNumber="130">
<emphasis id="B950900EFF9CE26BFF0DF8B8FEA9CCC7" bold="true" box="[151,274,1917,1939]" pageId="5" pageNumber="130">FIGURE 4.</emphasis>
<taxonomicName id="4C24379FFF9CE26BFE82F8BBFD3ACCC6" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[280,641,1917,1939]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF9CE26BFE82F8BBFE74CCC6" box="[280,463,1917,1939]" italics="true" pageId="5" pageNumber="130">Hexapinus latipes</emphasis>
(De Haan, 1835)
</taxonomicName>
. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160b), Ambon. AC, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P4; H, right P3; I, right P2.
</paragraph>
</caption>
<caption id="DF5B1C94FF9FE268FF0DF896FEBECCB6" httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="6" pageNumber="131" targetBox="[151,1436,193,1853]" targetPageId="6">
<paragraph id="8B9B4C1CFF9FE268FF0DF896FEBECCB6" blockId="6.[151,1436,1875,2019]" pageId="6" pageNumber="131">
<emphasis id="B950900EFF9FE268FF0DF896FEAFCC3D" bold="true" box="[151,276,1875,1897]" pageId="6" pageNumber="131">FIGURE 5.</emphasis>
<taxonomicName id="4C24379FFF9FE268FE86F891FD37CC3C" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[284,652,1875,1897]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF9FE268FE86F891FE6FCC3C" box="[284,468,1875,1897]" italics="true" pageId="6" pageNumber="131">Hexapinus latipes</emphasis>
(De Haan, 1835)
</taxonomicName>
. AD, male (20.9 × 12.5 mm) (SMF ZMG 160a), Ambon, Indonesia; EH, female (18.9 × 12.1 mm) (SMF ZMG 160b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 36; B, F, thoracic sternum and abdominal somites 16; C, G, posterior thoracic sternum and abdominal somites 15 showing sternite 8 (st8); D, sternoabdominal cavity, G1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v).
</paragraph>
</caption>
<caption id="DF5B1C94FF9EE269FF0DFB1BFE3DCE3A" httpUri="https://zenodo.org/record/241821/files/figure.png" pageId="7" pageNumber="132" targetBox="[163,1419,187,1223]" targetPageId="7">
<paragraph id="8B9B4C1CFF9EE269FF0DFB1BFE3DCE3A" blockId="7.[151,1436,1246,1391]" pageId="7" pageNumber="132">
<emphasis id="B950900EFF9EE269FF0DFB1BFEAFCFA6" bold="true" box="[151,276,1246,1268]" pageId="7" pageNumber="132">FIGURE 6.</emphasis>
AF,
<taxonomicName id="4C24379FFF9EE269FECCFB1AFD7CCFA1" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[342,711,1246,1268]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF9EE269FECCFB1AFDB5CFA1" box="[342,526,1246,1268]" italics="true" pageId="7" pageNumber="132">Hexapinus latipes</emphasis>
(De Haan, 1835)
</taxonomicName>
, male (20.9 × 12.5 mm) (SMF ZMG 160a), Ambon, Indonesia; G, H,
<taxonomicName id="4C24379FFF9EE269FF0DFB39FEF4CE46" box="[151,335,1276,1299]" class="Malacostraca" family="Hexapodidae" genus="Mariaplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
<emphasis id="B950900EFF9EE269FF0DFB39FEF4CE46" box="[151,335,1276,1299]" italics="true" pageId="7" pageNumber="132">Mariaplax aspera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2632D75FF9EE269FECDFB38FE29CE47" box="[343,402,1277,1298]" pageId="7" pageNumber="132" rank="species">
<emphasis id="B950900EFF9EE269FECDFB38FE29CE47" bold="true" box="[343,402,1277,1298]" pageId="7" pageNumber="132">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype female (6.4 × 4.4 mm) (MZB), Lombok, Indonesia. A, ventral view of left G1; B, dorsal view of left G1; C, ventral view of distal part of left G1; D, dorsal view of distal part of left G1 (setae not drawn); E, left G2 (ventral view); F, distal half of left G2 (dorsal view); G, left third maxilliped; H, female abdomen. Scales: A, B, E, G, H = 1.0 mm; C, D, F = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C33E1F97FF9EE266FF5DFA53FEBDCC89" lastPageId="8" lastPageNumber="133" pageId="7" pageNumber="132" type="description">
<paragraph id="8B9B4C1CFF9EE269FF5DFA53FE40CCF2" blockId="7.[151,1437,1430,2031]" pageId="7" pageNumber="132">
<emphasis id="B950900EFF9EE269FF5DFA53FEE2CEFA" bold="true" box="[199,345,1430,1455]" pageId="7" pageNumber="132">Description.</emphasis>
Carapace subquadrate, much wider than long, width 1.31.7 times length, prominently pitted on almost entire dorsal surface, regions indistinct, median H-shaped depression distinct (
<figureCitation id="131F5099FF9EE269FBC2FA7FFB25CE86" box="[1112,1182,1466,1491]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="7" pageNumber="132">Fig. 2</figureCitation>
). Anterolateral margin arcuate, lined with distinct rounded granules; merging with distinctly diverging sinuous posterolateral margin sinuous, surface with numerous granules, widest point convex; posterolateral corner with round prominence over base of coxa of P4; lateral branchial surfaces produced, visible from dorsal view (
<figureCitation id="131F5099FF9EE269FBA2F9E3FBC5CD6A" box="[1080,1150,1574,1599]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="7" pageNumber="132">Fig. 2</figureCitation>
). Front narrow, deflexed, truncate, divided into 2 lobes with almost straight margins, projecting just beyond lower edge of orbits (
<figureCitation id="131F5099FF9EE269FA83F98FFAD3CD36" box="[1305,1384,1610,1635]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="7" pageNumber="132">Figs. 2</figureCitation>
,
<figureCitation id="131F5099FF9EE269FAE9F98EFA3BCD36" box="[1395,1408,1611,1635]" captionStart="FIGURE 3" captionStartId="4.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1898]" captionTargetId="figure@4.[151,1436,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="7" pageNumber="132">3</figureCitation>
A, B). Orbit distinct, transversely ovate; eye small, movable, cornea small, as wide as peduncle (
<figureCitation id="131F5099FF9EE269FB3CF9ABFB5CCDD2" box="[1190,1255,1646,1671]" captionStart="FIGURE 3" captionStartId="4.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1898]" captionTargetId="figure@4.[151,1436,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="7" pageNumber="132">Fig. 3</figureCitation>
A, B). Epistome longitudinally narrow; posterior margin concave with broadly triangular median projection (
<figureCitation id="131F5099FF9EE269FA9BF957FAF7CDFE" box="[1281,1356,1682,1707]" captionStart="FIGURE 3" captionStartId="4.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1898]" captionTargetId="figure@4.[151,1436,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="7" pageNumber="132">Fig. 3</figureCitation>
A, B). Suborbital, pterygostomial, subhepatic regions covered with flattened granules, punctate (
<figureCitation id="131F5099FF9EE269FB60F973FAF2CD9A" box="[1274,1353,1718,1743]" captionStart="FIGURE 3" captionStartId="4.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1898]" captionTargetId="figure@4.[151,1436,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="7" pageNumber="132">Fig. 3</figureCitation>
A, B). Pterygostomial region with oblique striae, dense setae under striae adjacent to Milne Edwards opening (
<figureCitation id="131F5099FF9EE269FAA4F91FFA39CDA6" box="[1342,1410,1754,1779]" captionStart="FIGURE 3" captionStartId="4.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1898]" captionTargetId="figure@4.[151,1436,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="7" pageNumber="132">Fig. 3</figureCitation>
B, C). Third maxillipeds broad, outer surfaces covered with numerous small granules; ischium laterally expanded, subequal in length to merus, expanded distally, with gently convex mesial margin, merus wider than long, carpus, propodus, dactylus cylindrical; dactylus longer than propodus, combined length of dactylus, propodus, carpus subequal to length of merus, ischium; exopod relatively broad, about 0.4 times width of ischium, flagellum well developed (
<figureCitation id="131F5099FF9EE269FE83F84BFEDDCCF2" box="[281,358,1934,1959]" captionStart="FIGURE 3" captionStartId="4.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1898]" captionTargetId="figure@4.[151,1436,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="7" pageNumber="132">Figs. 3</figureCitation>
B, C, 4AC).
</paragraph>
<paragraph id="8B9B4C1CFF9EE266FF5DF877FF71CADD" blockId="7.[151,1437,1430,2031]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="133" pageId="7" pageNumber="132">
Chelipeds stout, unequal; outer surfaces covered with numerous small rounded granules (
<figureCitation id="131F5099FF9EE269FB42F877FA9BCC9E" box="[1240,1312,1970,1995]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="7" pageNumber="132">Fig. 4</figureCitation>
D). Carpus ovate, without setae on upper outer surface, inner angle low, granular, outer face uneven or with small granules; merus short, upper outer surface covered with granules, fringe of setae dorsomesially (
<figureCitation id="131F5099FF91E266FBE4FF52FB7FCBE5" box="[1150,1220,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="8" pageNumber="133">Fig. 2</figureCitation>
B, C). Major chela with gaping fingers when closed; palm slightly longer than wide, inner surface with subdorsal expanded triangular granular structure, median tubercle (
<figureCitation id="131F5099FF91E266FDA8FF1AFDCDCBAD" box="[562,630,223,248]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="8" pageNumber="133">Fig. 4</figureCitation>
E, F); dactylus relatively short, dorsal margin with small granules, tufts of setae proximally; cutting edge of dactylus with 2 large proximal teeth, rest of edge crenulated; propodal finger smooth, cutting edges with low teeth (
<figureCitation id="131F5099FF91E266FDD5FEE2FD2ECA15" box="[591,661,295,320]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="8" pageNumber="133">Fig. 4</figureCitation>
D). Minor chela similar but palm more slender, ventral margin with median tufts of short setae; fingers strongly gaping when closed; dorsal margin with rounded granules, setae (
<figureCitation id="131F5099FF91E266FAF5FE89FF1CCADD" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="8" pageNumber="133">Fig. 4</figureCitation>
D).
</paragraph>
<paragraph id="8B9B4C1CFF91E266FF5DFE51FBAAC86D" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="133">
P2P4 short, stout; P3 longest (
<figureCitation id="131F5099FF91E266FDA9FE51FD3ECAF9" box="[563,645,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="8" pageNumber="133">Figs. 2</figureCitation>
,
<figureCitation id="131F5099FF91E266FD09FE51FD19CAF9" box="[659,674,404,428]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="8" pageNumber="133">4</figureCitation>
GI). P2 shortest, stoutest; merus almost 3 times length of carpus, outer surface with numerous distinct small granules, short setae, upper, lower margins with short setae, unarmed otherwise; dactylus slender, gently upcurved, fringe of setae on upper, lower margins, slightly longer than propodus (
<figureCitation id="131F5099FF91E266FF05FE3AFF5FC94D" box="[159,228,511,536]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="8" pageNumber="133">Fig. 4</figureCitation>
G). P3 relatively stouter, longer than P2; surfaces relatively more granular; merus almost 3 times length of carpus, outer surface with numerous distinct, small granules, setae, upper, lower margins with setae, unarmed otherwise; carpus as long as propodus, outer surface with granules, denser on upper half, surface with short setae; outer surface of propodus granular, lower margin with setae, crenulations, spines present distally; relatively strongest among pereopods (
<figureCitation id="131F5099FF91E266FE43FD4AFDA7C9FD" box="[473,540,655,680]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="8" pageNumber="133">Fig. 4</figureCitation>
H). P4 most slender; merus more than twice length of carpus, outer surface with small granules, short setae, upper, lower margins with tufts of dense short setae, unarmed otherwise; carpus as long as propodus, outer surface with granules, denser on upper half, surface with short setae; outer surface of propodus with low granules, lower margin with setae, crenulations, spines present distally; dactylus slender, gently upcurved, fringe of setae on upper, lower margins, slightly longer than propodus (
<figureCitation id="131F5099FF91E266FC2EFCDAFC42C86D" box="[948,1017,799,824]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="8" pageNumber="133">
Fig.
<date id="FF9A6ADCFF91E266FC73FCE5FC42C86D" box="[1001,1017,800,824]" pageId="8" pageNumber="133">4</date>
</figureCitation>
I).
</paragraph>
<paragraph id="8B9B4C1CFF91E266FF5DFC81FDEACE58" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="133">
Male thoracic sternum broad; sternites 1, 2 fused, forming subtriangular structure, each with deep transverse groove that does not extend to lateral margins, separated from sternite 3 by distinct ridge, adjacent deep transverse groove; sternite 3 longitudinally narrow, posteromedian part with 2 granular projections that bracket telson; sternite 3 delimited from sternite 4 by deep tranverse groove that forms thoracic sternal groove, relatively shallower along margins; sternites 47 well developed, separated by distinct sutures, surfaces distinctly punctate; sternite 8 just exposed when abdomen closed, visible as small triangular plate between abdomen, sternite 7, about half length as male abdominal somite 1 (
<figureCitation id="131F5099FF91E266FE4FFBD9FD9CCF60" box="[469,551,1052,1077]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="8" pageNumber="133">Figs. 5</figureCitation>
C). Sternoabdominal cavity elongated, reaching to just below base of third maxilliped, posterior margin of thoracic sternite 3 (
<figureCitation id="131F5099FF91E266FD47FBFAFC90CF0D" box="[733,811,1087,1112]" captionStart="FIGURE 3" captionStartId="4.[151,250,1921,1943]" captionTargetBox="[151,1436,193,1898]" captionTargetId="figure@4.[151,1436,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; frontal view of cephalothorax. A, dorsal surface of carapace, front, epistome and orbit; B, C, buccal field, third maxillipeds, pterygostomial regions and thoracic sternites 1 4." httpUri="https://zenodo.org/record/241818/files/figure.png" pageId="8" pageNumber="133">Figs. 3</figureCitation>
C, 5A). Abdominal press button locking mechanism of male distinct, present as large rounded tubercle on anterior edge of sternite 5, overlapping posterior part of sternite 4 (
<figureCitation id="131F5099FF91E266FF2DFB42FF47CFF5" box="[183,252,1159,1184]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="8" pageNumber="133">Fig. 5</figureCitation>
D). Male abdomen relatively long, slender, surfaces granular; somites 1, 2 free; somites 35 fused but median parts of sutures still visible as shallow grooves, lateral margins sinuous; somite 6 longer than broad, shorter than fused somites 35, longer than telson, lateral margins convex; telson pentagonal, with rounded distal margin, angular anterolateral edge (
<figureCitation id="131F5099FF91E266FE53FB31FDB1CE58" box="[457,522,1268,1293]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="8" pageNumber="133">Fig. 5</figureCitation>
AC).
</paragraph>
<paragraph id="8B9B4C1CFF91E266FF5DFADDFDCFCEC9" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="133">
G1 simple, slender, curved along distal third, lying in oblique groove on anterior part of sternum, distal onefifth not concealed under abdomen; distalmost part gradually tapering to subtruncate tip; lateral margins, distal surfaces lined with long plumose setae which partially obscure surfaces (
<figureCitation id="131F5099FF91E266FC5EFA9AFBFBCE2D" box="[964,1088,1375,1400]" captionStart="FIGURE 6. A F" captionStartId="7.[151,250,1246,1268]" captionTargetBox="[163,1419,187,1223]" captionTargetId="figure@7.[151,1435,175,1233]" captionTargetPageId="7" captionText="FIGURE 6. A F, Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; G, H, Mariaplax aspera n. sp., holotype female (6.4 × 4.4 mm) (MZB), Lombok, Indonesia. A, ventral view of left G 1; B, dorsal view of left G 1; C, ventral view of distal part of left G 1; D, dorsal view of distal part of left G 1 (setae not drawn); E, left G 2 (ventral view); F, distal half of left G 2 (dorsal view); G, left third maxilliped; H, female abdomen. Scales: A, B, E, G, H = 1.0 mm; C, D, F = 0.5 mm." httpUri="https://zenodo.org/record/241821/files/figure.png" pageId="8" pageNumber="133">Fig. 6AD</figureCitation>
). G2 short, about quarter of G1 length; distal part spatuliform (
<figureCitation id="131F5099FF91E266FE68FA41FDDCCEC9" box="[498,615,1412,1437]" captionStart="FIGURE 6. A F" captionStartId="7.[151,250,1246,1268]" captionTargetBox="[163,1419,187,1223]" captionTargetId="figure@7.[151,1435,175,1233]" captionTargetPageId="7" captionText="FIGURE 6. A F, Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; G, H, Mariaplax aspera n. sp., holotype female (6.4 × 4.4 mm) (MZB), Lombok, Indonesia. A, ventral view of left G 1; B, dorsal view of left G 1; C, ventral view of distal part of left G 1; D, dorsal view of distal part of left G 1 (setae not drawn); E, left G 2 (ventral view); F, distal half of left G 2 (dorsal view); G, left third maxilliped; H, female abdomen. Scales: A, B, E, G, H = 1.0 mm; C, D, F = 0.5 mm." httpUri="https://zenodo.org/record/241821/files/figure.png" pageId="8" pageNumber="133">Fig. 6E, F</figureCitation>
).
</paragraph>
<paragraph id="8B9B4C1CFF91E266FF5DFA62FE92CDB5" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="133">
<emphasis id="B950900EFF91E266FF5DFA62FE92CE95" bold="true" box="[199,297,1447,1472]" pageId="8" pageNumber="133">Females</emphasis>
. Chelipeds approximately symmetrical, inner surface subdorsal margin has raised patch of granules but no obvious projection. Female thoracic sternum relatively broad (
<figureCitation id="131F5099FF91E266FC5DFA09FBB6CEB0" box="[967,1037,1484,1509]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="8" pageNumber="133">Fig. 5</figureCitation>
E), sternites 1, 2 fused, each with transverse groove, separated from sternite 3 by distinct ridge, transverse groove; sternite 3 separated from sternite 4 by ridge laterally, thoracic sternal groove short, shallow, extends obliquely from sternoabdominal cavity (
<figureCitation id="131F5099FF91E266FADBF9D1FA3FCD78" box="[1345,1412,1556,1581]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="8" pageNumber="133">Fig. 5</figureCitation>
E, H); sternites 47 distinct, separated by distinct sutures; sternite 8 just visible as small triangular plate when abdomen closed, about half length of abdominal somite 1 (
<figureCitation id="131F5099FF91E266FCDDF999FC36CD20" box="[839,909,1628,1653]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="8" pageNumber="133">Fig. 5</figureCitation>
G). Vulvae large, without opercular cover, on anterior half on sternite 6, opening subovate (
<figureCitation id="131F5099FF91E266FD3DF9BAFD57CDCD" box="[679,748,1663,1688]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="8" pageNumber="133">Fig. 5</figureCitation>
H). Sternoabdominal cavity reaches distal part of sternite 4. Female abdomen subovate, with 6 free somites; somite 6 longest, trapezoidal; telson subtriangular with rounded tip (
<figureCitation id="131F5099FF91E266FF05F902FF5ACDB5" box="[159,225,1735,1760]" captionStart="FIGURE 5" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[151,1436,193,1853]" captionTargetId="figure@6.[151,1436,193,1853]" captionTargetPageId="6" captionText="FIGURE 5. Hexapinus latipes (De Haan, 1835). A D, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; E H, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A, E, anterior thoracic sternum; telson and abdominal somites 3 6; B, F, thoracic sternum and abdominal somites 1 6; C, G, posterior thoracic sternum and abdominal somites 1 5 showing sternite 8 (st 8); D, sternoabdominal cavity, G 1 and tubercle of abdominal locking mechanism (bp); H, female sternoabdominal cavity and vulvae (v)." httpUri="https://zenodo.org/record/241820/files/figure.png" pageId="8" pageNumber="133">Fig. 5</figureCitation>
EG,).
</paragraph>
<paragraph id="8B9B4C1CFF91E266FF5DF929FEBDCC89" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="133">
<emphasis id="B950900EFF91E266FF5DF929FE8ECC50" bold="true" box="[199,309,1772,1797]" pageId="8" pageNumber="133">Variation</emphasis>
. As in
<taxonomicName id="4C24379FFF91E266FE0EF928FDD7CC51" box="[404,620,1772,1796]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="133" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B950900EFF91E266FE0EF928FDD7CC51" box="[404,620,1772,1796]" italics="true" pageId="8" pageNumber="133">Hexapinus simplex</emphasis>
</taxonomicName>
, the ambulatory meri of the male (
<figureCitation id="131F5099FF91E266FB94F929FBDBCC51" box="[1038,1120,1772,1797]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="8" pageNumber="133">Figs. 2</figureCitation>
B, 4GI) is proportionately longer than those of females (
<figureCitation id="131F5099FF91E266FD8EF8CAFDDBCC7D" box="[532,608,1807,1832]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="8" pageNumber="133">Fig. 2</figureCitation>
A, C). There is also a distinct tendency for specimens to become proportionately broader as they increase in size. The largest females from
<collectingCountry id="F3330C8CFF91E266FC6DF8F0FB8CCC19" box="[1015,1079,1845,1868]" name="Japan" pageId="8" pageNumber="133">Japan</collectingCountry>
have carapace width to length ratios of 1.711.73, with the intermediate sized ones from
<collectingCountry id="F3330C8CFF91E266FCADF892FC30CC25" box="[823,907,1879,1904]" name="Indonesia" pageId="8" pageNumber="133">Ambon</collectingCountry>
and
<collectingCountry id="F3330C8CFF91E266FC5CF89CFBBDCC25" box="[966,1030,1881,1904]" name="Japan" pageId="8" pageNumber="133">Japan</collectingCountry>
1.561.67. The smallest specimens (including the
<typeStatus id="549FF2BEFF91E266FEDAF8B9FE13CCC0" box="[320,424,1916,1941]" pageId="8" pageNumber="133" type="lectotype">lectotype</typeStatus>
) are 1.341.49. The form of the merus of the third maxilliped also varies, with smaller specimens having a slightly more quadrate structure (
<figureCitation id="131F5099FF91E266FD6AF865FC8ECCED" box="[752,821,1952,1977]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="8" pageNumber="133">Fig. 4</figureCitation>
B, C) while in the largest ones, they are more subovate (
<figureCitation id="131F5099FF91E266FF05F801FF58CC89" box="[159,227,1988,2013]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="8" pageNumber="133">Fig. 4</figureCitation>
A).
</paragraph>
</subSubSection>
<subSubSection id="C33E1F97FF90E267FF5DFF52FE29CE2D" pageId="9" pageNumber="134" type="discussion">
<paragraph id="8B9B4C1CFF90E267FF5DFF52FD53CA30" blockId="9.[151,1437,151,1401]" pageId="9" pageNumber="134">
<emphasis id="B950900EFF90E267FF5DFF52FE8FCBE5" bold="true" box="[199,308,151,176]" pageId="9" pageNumber="134">Remarks</emphasis>
. This is a poorly known species and hitherto known only from females.
<bibRefCitation id="EFB531EDFF90E267FBEBFF52FA27CBE5" author="Rahayu" box="[1137,1436,151,176]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng (2014: 409)</bibRefCitation>
commented that: “De Mans (1888: 322, pl. 13, fig. 3) female specimen from
<collectingCountry id="F3330C8CFF90E267FB9FFF79FBE2CB80" box="[1029,1113,188,213]" name="Indonesia" pageId="9" pageNumber="134">Ambon</collectingCountry>
is also likely to be a species of
<taxonomicName id="4C24379FFF90E267FF2FFF25FE96CBA2" box="[181,301,224,247]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B950900EFF90E267FF2FFF25FE96CBA2" box="[181,301,224,247]" italics="true" pageId="9" pageNumber="134">Hexapinus</emphasis>
</taxonomicName>
although he does not show the maxillipeds; it resembles
<taxonomicName id="4C24379FFF90E267FC30FF24FB9ACBAD" box="[938,1057,224,248]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B950900EFF90E267FC30FF24FB9ACBAD" box="[938,1057,224,248]" italics="true" pageId="9" pageNumber="134">H. simplex</emphasis>
</taxonomicName>
described from nearby Lombok”. De
<bibRefCitation id="EFB531EDFF90E267FF24FEC0FEC7CA49" author="Man" box="[190,380,260,284]" pageId="9" pageNumber="134" refString="Man, J. G. de (1888) Bericht uber die von Herrn Dr. J. Brock im indischen Archipel gesammelten Decapoden und Stomatopoden. Archiv fur Naturgeschichte, Berlin, 53 (1), 289 - 600, pls. 11 - 22." type="journal article" year="1888">Man (1888: 322)</bibRefCitation>
actually had two specimens (a male and a female) and figured the male. Examination of these two specimens, which originally belonged to the Museum of Zoology, Göttingen University but are now conserved in SMF, shows that they belong to
<taxonomicName id="4C24379FFF90E267FD86FE88FD31CA31" box="[540,650,332,356]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF90E267FD86FE88FD31CA31" box="[540,650,332,356]" italics="true" pageId="9" pageNumber="134">H. latipes</emphasis>
</taxonomicName>
instead.
</paragraph>
<paragraph id="8B9B4C1CFF90E267FF5DFEAAFA8DC9D1" blockId="9.[151,1437,151,1401]" pageId="9" pageNumber="134">
<bibRefCitation id="EFB531EDFF90E267FF5DFEAAFDBBCADD" author="Rahayu" box="[199,512,367,392]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng (2014: 423)</bibRefCitation>
noted that
<taxonomicName id="4C24379FFF90E267FD11FEB5FCD8CADD" box="[651,867,368,392]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B950900EFF90E267FD11FEB5FCD8CADD" box="[651,867,368,392]" italics="true" pageId="9" pageNumber="134">Hexapinus simplex</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C24379FFF90E267FB1AFEB5FAF5CADD" box="[1152,1358,368,392]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF90E267FB1AFEB5FAF5CADD" box="[1152,1358,368,392]" italics="true" pageId="9" pageNumber="134">Hexapinus latipes</emphasis>
</taxonomicName>
by its relatively wider carapace, the dorsal surface of the carapace is less prominently pitted, and the ischium of the third maxilliped is relatively shorter, with the mesial margin more strongly convex. The form of the carapace surface and proportions of the third maxilliped are obviously different in the present specimens of both species, although the proportions of the carapace is not an easy character to use as it can vary. The pits on the carapace of
<taxonomicName id="4C24379FFF90E267FABEFDC5FF59C969" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF90E267FABEFDC5FF59C969" italics="true" pageId="9" pageNumber="134">Hexapinus latipes</emphasis>
</taxonomicName>
are very pronounced and all over the dorsal surface. This is evident even on the small
<typeStatus id="549FF2BEFF90E267FB79FDE1FAFEC968" box="[1251,1349,548,573]" pageId="9" pageNumber="134" type="holotype">holotype</typeStatus>
female (
<bibRefCitation id="EFB531EDFF90E267FF04FD82FE42C935" author="Rahayu" box="[158,505,583,608]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng, 2014: fig. 10A</bibRefCitation>
). In
<taxonomicName id="4C24379FFF90E267FDB7FD8DFCB9C935" box="[557,770,584,608]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B950900EFF90E267FDB7FD8DFCB9C935" box="[557,770,584,608]" italics="true" pageId="9" pageNumber="134">Hexapinus simplex</emphasis>
</taxonomicName>
, while there are also pits, they are less distinct, shallower, more scattered even when more prominent and not over the entire surface (
<bibRefCitation id="EFB531EDFF90E267FC44FDA9FA92C9D1" author="Rahayu" box="[990,1321,620,645]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng 2014: fig. 18C</bibRefCitation>
).
</paragraph>
<paragraph id="8B9B4C1CFF90E267FF5DFD4AFC28C89D" blockId="9.[151,1437,151,1401]" pageId="9" pageNumber="134">
Another useful character is the form of the chela. In adult male
<taxonomicName id="4C24379FFF90E267FC2BFD55FBC6C9FD" box="[945,1149,656,680]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF90E267FC2BFD55FBC6C9FD" box="[945,1149,656,680]" italics="true" pageId="9" pageNumber="134">Hexapinus latipes</emphasis>
</taxonomicName>
, the inner surface of the major palm has a well-developed dorsal, subdorsal granular projection (
<figureCitation id="131F5099FF90E267FC56FD71FBA6C999" box="[972,1053,692,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="9" pageNumber="134">Figs. 2</figureCitation>
B, 4E, F). It is also visible on the smaller chela; and in female chelae, but is relatively shorter (
<figureCitation id="131F5099FF90E267FCF9FD12FC13C9A5" box="[867,936,727,752]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="9" pageNumber="134">Fig. 2</figureCitation>
A, C). This character is visible even in the
<typeStatus id="549FF2BEFF90E267FF0DFD39FF42C840" box="[151,249,764,789]" pageId="9" pageNumber="134" type="holotype">holotype</typeStatus>
female and Japanese female specimens (
<bibRefCitation id="EFB531EDFF90E267FD5BFD39FB2BC841" author="Rahayu" box="[705,1168,764,789]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng 2014: figs. 10A, 11A, 12A</bibRefCitation>
). In
<taxonomicName id="4C24379FFF90E267FB5BFD38FA2FC841" box="[1217,1428,764,788]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B950900EFF90E267FB5BFD38FA2FC841" box="[1217,1428,764,788]" italics="true" pageId="9" pageNumber="134">Hexapinus simplex</emphasis>
</taxonomicName>
, the inner surface of the palm is almost smooth and not raised (
<bibRefCitation id="EFB531EDFF90E267FCCEFCDAFB2FC86D" author="Rahayu" box="[852,1172,799,824]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng, 2014: fig. 18</bibRefCitation>
). The G1 of
<taxonomicName id="4C24379FFF90E267FABEFCE5FF59C809" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF90E267FABEFCE5FF59C809" italics="true" pageId="9" pageNumber="134">Hexapinus latipes</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C24379FFF90E267FE59FC80FD2CC809" box="[451,663,836,860]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B950900EFF90E267FE59FC80FD2CC809" box="[451,663,836,860]" italics="true" pageId="9" pageNumber="134">Hexapinus simplex</emphasis>
</taxonomicName>
, but the median part is relatively stouter and towards the distal third, it bends outwards, with the distal part gradually tapering to a subtruncate tip (
<figureCitation id="131F5099FF90E267FB97FCA2FB37C8D5" box="[1037,1164,871,896]" captionStart="FIGURE 6. A F" captionStartId="7.[151,250,1246,1268]" captionTargetBox="[163,1419,187,1223]" captionTargetId="figure@7.[151,1435,175,1233]" captionTargetPageId="7" captionText="FIGURE 6. A F, Hexapinus latipes (De Haan, 1835), male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; G, H, Mariaplax aspera n. sp., holotype female (6.4 × 4.4 mm) (MZB), Lombok, Indonesia. A, ventral view of left G 1; B, dorsal view of left G 1; C, ventral view of distal part of left G 1; D, dorsal view of distal part of left G 1 (setae not drawn); E, left G 2 (ventral view); F, distal half of left G 2 (dorsal view); G, left third maxilliped; H, female abdomen. Scales: A, B, E, G, H = 1.0 mm; C, D, F = 0.5 mm." httpUri="https://zenodo.org/record/241821/files/figure.png" pageId="9" pageNumber="134">Fig. 6AD</figureCitation>
). In
<taxonomicName id="4C24379FFF90E267FB25FCADFA2FC8D5" box="[1215,1428,872,896]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B950900EFF90E267FB25FCADFA2FC8D5" box="[1215,1428,872,896]" italics="true" pageId="9" pageNumber="134">Hexapinus simplex</emphasis>
</taxonomicName>
, the median part of the G1 is more slender, the entire G1 gently curves outwards and the distal part tapers down suddenly to a slender rounded tip (
<bibRefCitation id="EFB531EDFF90E267FD80FC6AFC3DC89D" author="Rahayu" box="[538,902,943,968]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng 2014: fig. 21A, B</bibRefCitation>
).
</paragraph>
<paragraph id="8B9B4C1CFF90E267FF5DFC11FEC9CFBD" blockId="9.[151,1437,151,1401]" pageId="9" pageNumber="134">
Also noteworthy, as discussed by
<bibRefCitation id="EFB531EDFF90E267FDD9FC11FC89C8B9" author="Rahayu" box="[579,818,980,1005]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng (2014)</bibRefCitation>
,
<taxonomicName id="4C24379FFF90E267FCA4FC10FBB3C8B9" box="[830,1032,980,1004]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF90E267FCA4FC10FBB3C8B9" box="[830,1032,980,1004]" italics="true" pageId="9" pageNumber="134">Hexapinus latipes</emphasis>
</taxonomicName>
grows to much larger adult sizes. A female measuring 10.2 by
<quantity id="4CDCE1F9FF90E267FE58FC32FDA2CF5A" box="[450,537,1015,1040]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="134" unit="mm" value="7.6">7.6 mm</quantity>
from
<collectingCountry id="F3330C8CFF90E267FDC4FC3DFD25CF5A" box="[606,670,1016,1039]" name="Japan" pageId="9" pageNumber="134">Japan</collectingCountry>
was still immature, with the abdomen narrow and the gonopods not evident (
<bibRefCitation id="EFB531EDFF90E267FF63FBD9FDD0CF61" author="Rahayu" box="[249,619,1052,1077]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng 2014: 414, fig. 11</bibRefCitation>
). The largest specimen of
<taxonomicName id="4C24379FFF90E267FC0CFBD8FBDACF61" box="[918,1121,1052,1076]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF90E267FC0CFBD8FBDACF61" box="[918,1121,1052,1076]" italics="true" pageId="9" pageNumber="134">Hexapinus latipes</emphasis>
</taxonomicName>
, a mature female, measures
<quantity id="4CDCE1F9FF90E267FF0DFB85FF40CF02" box="[151,251,1088,1112]" metricMagnitude="-2" metricUnit="m" metricValue="3.04" pageId="9" pageNumber="134" unit="mm" value="30.4">30.4 mm</quantity>
in carapace width (
<figureCitation id="131F5099FF90E267FE4DFBFAFDA1CF0D" box="[471,538,1087,1112]" captionStart="FIGURE 2" captionStartId="3.[151,250,1855,1877]" captionTargetBox="[151,1436,193,1832]" captionTargetId="figure@3.[151,1436,193,1833]" captionTargetPageId="3" captionText="FIGURE 2. Hexapinus latipes (De Haan, 1835), overall habitus. A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon." httpUri="https://zenodo.org/record/241817/files/figure.png" pageId="9" pageNumber="134">Fig. 2</figureCitation>
A;
<bibRefCitation id="EFB531EDFF90E267FDA6FBFAFCA7CF0D" author="Rahayu" box="[572,796,1087,1112]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng 2014</bibRefCitation>
: 412). Specimens of
<taxonomicName id="4C24379FFF90E267FB9CFB85FB61CF0D" box="[1030,1242,1088,1112]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B950900EFF90E267FB9CFB85FB61CF0D" box="[1030,1242,1088,1112]" italics="true" pageId="9" pageNumber="134">Hexapinus simplex</emphasis>
</taxonomicName>
measuring about 6.0 to 7.0 mm in carapace width are already mature, with the males having well-developed gonopods and the females having an ovate abdomen with setose pleopods (
<bibRefCitation id="EFB531EDFF90E267FC89FB42FB97CFF5" author="Rahayu" box="[787,1068,1159,1185]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng 2014: 418</bibRefCitation>
). The present adult specimens of
<taxonomicName id="4C24379FFF90E267FF0DFB68FEDBCF91" box="[151,352,1196,1220]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B950900EFF90E267FF0DFB68FEDBCF91" box="[151,352,1196,1220]" italics="true" pageId="9" pageNumber="134">Hexapinus latipes</emphasis>
</taxonomicName>
from
<collectingCountry id="F3330C8CFF90E267FE3EFB69FE43CF90" box="[420,504,1196,1221]" name="Indonesia" pageId="9" pageNumber="134">Ambon</collectingCountry>
are mature, with the male measuring
<quantity id="4CDCE1F9FF90E267FC04FB69FBB9CF91" box="[926,1026,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="9" pageNumber="134" unit="mm" value="20.9">20.9 mm</quantity>
and the female
<quantity id="4CDCE1F9FF90E267FB29FB69FAACCF91" box="[1203,1303,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="9" pageNumber="134" unit="mm" value="18.9">18.9 mm</quantity>
in carapace width, respectively.
</paragraph>
<paragraph id="8B9B4C1CFF90E267FF5DFB31FE29CE2D" blockId="9.[151,1437,151,1401]" pageId="9" pageNumber="134">
The form of the merus of the third maxilliped appears to vary, being more quadrate in smaller specimens (
<figureCitation id="131F5099FF90E267FAF5FB31FF1CCE65" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="9" pageNumber="134">Fig. 4</figureCitation>
B, C) but more ovate in large ones (
<figureCitation id="131F5099FF90E267FDA2FAD2FDC7CE65" box="[568,636,1303,1328]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="9" pageNumber="134">Fig. 4</figureCitation>
A); as has been noted by
<bibRefCitation id="EFB531EDFF90E267FC03FAD2FAAFCE65" author="Rahayu" box="[921,1300,1303,1329]" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng (2014: fig. 13CE)</bibRefCitation>
. The mesial margin of the ischium, however, is much less convex (
<figureCitation id="131F5099FF90E267FD63FAF9FC80CE01" box="[761,827,1340,1365]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1939]" captionTargetBox="[210,1369,193,1875]" captionTargetId="figure@5.[210,1371,193,1876]" captionTargetPageId="5" captionText="FIGURE 4. Hexapinus latipes (De Haan, 1835). A, female (30.4 × 17.9 mm) (KPM NH 7123), Enoshima Island, Japan; B, D I, male (20.9 × 12.5 mm) (SMF ZMG 160 a), Ambon, Indonesia; C, female (18.9 × 12.1 mm) (SMF ZMG 160 b), Ambon. A C, left third maxillipeds and pterygostomial regions; D, outer views of chelae; E, inner surface of chela of major right chela; F, dorsal view of chela of major right chela; G, right P 4; H, right P 3; I, right P 2." httpUri="https://zenodo.org/record/241819/files/figure.png" pageId="9" pageNumber="134">Fig. 4</figureCitation>
AC) compared to that of
<taxonomicName id="4C24379FFF90E267FBF8FAF8FA8DCE01" box="[1122,1334,1340,1364]" class="Malacostraca" family="Hexapodidae" genus="Hexapinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B950900EFF90E267FBF8FAF8FA8DCE01" box="[1122,1334,1340,1364]" italics="true" pageId="9" pageNumber="134">Hexapinus simplex</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB531EDFF90E267FADFFAF9FE3ECE2D" author="Rahayu" pageId="9" pageNumber="134" refString="Rahayu, D. L. &amp; Ng, P. K. L. (2014) New genera and new species of Hexapodidae (Crustacea, Brachyura) from the Indo-West Pacific and east Atlantic. Raffles Bulletin of Zoology, 62, 396 - 483." type="journal article" year="2014">Rahayu &amp; Ng 2014: fig. 20G</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>