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<document id="60397EB5AC67B04C2419E1F3AD858801" ID-CLB-Dataset="31530" ID-DOI="10.5281/zenodo.1050060" ID-GBIF-Dataset="2d28bebf-e3c5-457b-9152-8a6877ded281" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510742923475" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="C063786AFF84FFD7FF0D10D45CF492CA" docLanguage="en" docName="zt03074p198.pdf.imf" docOrigin="Zootaxa 3074" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena beniensis Perkins, 2011, new species" docType="treatment" docVersion="7" lastPageNumber="43" masterDocId="3C5A0012FFAEFFFDFF9A11335F219540" masterDocTitle="New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)" masterLastPageNumber="198" masterPageNumber="1" pageNumber="43" updateTime="1698485333395" updateUser="ExternalLinkService">
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<mods:title id="233E1F14584DDCCA9B71AFF33F2C0FC6">New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="7277149067D9D500F3CC886CEA2F36B0">Perkins, Philip D.</mods:namePart>
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<mods:date id="5FFEF677BB7724CA8D674D370D3EF7EA">2011</mods:date>
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<mods:number id="23EB027A1E324145DC6C56013CFBA7A8">2011-10-28</mods:number>
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<treatment id="C063786AFF84FFD7FF0D10D45CF492CA" ID-DOI="http://doi.org/10.5281/zenodo.6184494" ID-GBIF-Taxon="137306990" ID-Zenodo-Dep="6184494" LSID="urn:lsid:plazi:treatment:C063786AFF84FFD7FF0D10D45CF492CA" httpUri="http://treatment.plazi.org/id/C063786AFF84FFD7FF0D10D45CF492CA" lastPageNumber="43" pageId="42" pageNumber="43">
<subSubSection id="00D09AF7FF84FFD7FF0D10D45DA99765" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="4875C97CFF84FFD7FF0D10D45D049741" blockId="42.[151,648,487,549]" box="[151,549,487,513]" pageId="42" pageNumber="43">
<heading id="133D7E10FF84FFD7FF0D10D45D049741" bold="true" box="[151,549,487,513]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<emphasis id="7ABE156EFF84FFD7FF0D10D45D049741" bold="true" box="[151,549,487,513]" pageId="42" pageNumber="43">
<taxonomicName id="8FCAB2FFFF84FFD7FF0D10D45EA79741" ID-CoL="6MHF9" box="[151,390,487,513]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="beniensis" status="sp. nov.">
<emphasis id="7ABE156EFF84FFD7FF0D10D45EA79741" bold="true" box="[151,390,487,513]" italics="true" pageId="42" pageNumber="43">Hydraena beniensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="618DA815FF84FFD7FE0F10D45D049741" box="[405,549,487,513]" pageId="42" pageNumber="43" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4875C97CFF84FFD7FF0D133E5DA99765" blockId="42.[151,648,487,549]" box="[151,648,525,549]" pageId="42" pageNumber="43">
<figureCitation id="D0F1D5F9FF84FFD7FF0D133E5FD09765" box="[151,241,525,549]" captionStart="FIGURE 75" captionStartId="125.[151,250,1612,1635]" captionTargetBox="[226,1357,193,1591]" captionTargetId="figure@125.[226,1360,193,1591]" captionTargetPageId="125" captionText="FIGURE 75. Hydraena beniensis, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050144/files/figure.png" pageId="42" pageNumber="43">Figs. 75</figureCitation>
(habitus), 79 (aedeagus), 199 (map)
</paragraph>
</subSubSection>
<subSubSection id="00D09AF7FF84FFD7FF0D136B5E1D96FF" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="4875C97CFF84FFD7FF0D136B5D4E97FA" blockId="42.[151,1437,599,1931]" pageId="42" pageNumber="43">
<emphasis id="7ABE156EFF84FFD7FF0D136B5E699730" bold="true" box="[151,328,599,624]" pageId="42" pageNumber="43">
<typeStatus id="977177DEFF84FFD7FF0D136B5FF39730" box="[151,210,600,624]" pageId="42" pageNumber="43">Type</typeStatus>
Material.
</emphasis>
<typeStatus id="977177DEFF84FFD7FED413645E97972F" box="[334,438,599,623]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="30DD89ECFF84FFD7FD8613645D519730" box="[540,624,599,624]" name="Bolivia" pageId="42" pageNumber="43">
<emphasis id="7ABE156EFF84FFD7FD8613645D519730" bold="true" box="[540,624,599,624]" pageId="42" pageNumber="43">Bolivia</emphasis>
</collectingCountry>
: Beni,
<quantity id="8F326499FF84FFD7FD5213645C2E972F" box="[712,783,599,623]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="42" pageNumber="43" unit="km" value="40.0">40 km</quantity>
E. San Borja, Estacion Biologica Beni, Palm Camp at Rio Curiraba, ex. small pool on sand bar, edge of sunlit river,
<date id="3C74EFBCFF84FFD7FC82134E5CB297D5" box="[792,915,637,661]" pageId="42" pageNumber="43" value="1987-09-14">14 ix 1987</date>
, W. E. Steiner. Deposited in the
<collectionCode id="2EDB51B9FF84FFD7FB60134E5A6E97D5" LSID="urn:lsid:biocol.org:col:34871" box="[1274,1359,637,661]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="42" pageNumber="43">USNM</collectionCode>
.
<emphasis id="7ABE156EFF84FFD7FAC3134E5FFF97FB" bold="true" pageId="42" pageNumber="43">
<typeStatus id="977177DEFF84FFD7FAC3134E5FF897FB" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same data as
<typeStatus id="977177DEFF84FFD7FE1F13915EC697FA" box="[389,487,674,698]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
(5
<collectionCode id="2EDB51B9FF84FFD7FD9713915D4297FA" LSID="urn:lsid:biocol.org:col:34871" box="[525,611,674,698]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="42" pageNumber="43">USNM</collectionCode>
).
</paragraph>
<paragraph id="4875C97CFF84FFD7FF5D13F45E1D96FF" blockId="42.[151,1437,599,1931]" pageId="42" pageNumber="43">
<emphasis id="7ABE156EFF84FFD7FF5D13F45EF397A0" bold="true" box="[199,466,711,736]" pageId="42" pageNumber="43">Differential Diagnosis.</emphasis>
A small species (ca.
<quantity id="8F326499FF84FFD7FD5B13F45C07979F" box="[705,806,711,735]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="42" pageNumber="43" unit="mm" value="1.25">1.25 mm</quantity>
) with a distinctive pronotal fascia. The dorsal punctures are relatively small, but not widely separated, interstices ca. 1xpd on both the pronotal disc and the basal 1/3 of the elytra. Similar in size and dorsal habitus to
<taxonomicName id="8FCAB2FFFF84FFD7FD1112275DDE966B" box="[651,767,787,811]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="delvasi">
<emphasis id="7ABE156EFF84FFD7FD1112275DDE966B" box="[651,767,787,811]" italics="true" pageId="42" pageNumber="43">H. delvasi</emphasis>
</taxonomicName>
(
<figureCitation id="D0F1D5F9FF84FFD7FC9512215C4E966A" box="[783,879,786,810]" captionStart="FIGURE 75" captionStartId="125.[151,250,1612,1635]" captionTargetBox="[226,1357,193,1591]" captionTargetId="figure@125.[226,1360,193,1591]" captionTargetPageId="125" captionText="FIGURE 75. Hydraena beniensis, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050144/files/figure.png" pageId="42" pageNumber="43">Figs. 75</figureCitation>
,
<figureCitation id="D0F1D5F9FF84FFD7FCE612215CBD966A" box="[892,924,786,810]" captionStart="FIGURE 96" captionStartId="138.[151,250,1650,1673]" captionTargetBox="[249,1335,193,1626]" captionTargetId="figure@138.[249,1338,193,1629]" captionTargetPageId="138" captionText="FIGURE 96. Hydraena delvasi, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050164/files/figure.png" pageId="42" pageNumber="43">96</figureCitation>
); differing therefrom by the smaller pronotal fascia, the shining (non-microreticulate) disc of the frons, and the more elongate elytra; the aedeagi of the two species differ markedly (
<figureCitation id="D0F1D5F9FF84FFD7FE11126E5ECB9635" box="[395,490,861,885]" captionStart="FIGURES 78 81" captionStartId="128.[151,264,1650,1673]" captionText="FIGURES 78 81. Aedeagi of holotypes." pageId="42" pageNumber="43">Figs. 79</figureCitation>
,
<figureCitation id="D0F1D5F9FF84FFD7FE6D126E5D369635" box="[503,535,861,885]" captionStart="FIGURES 99 101" captionStartId="141.[151,264,1613,1636]" captionTargetBox="[246,1337,236,1561]" captionTargetId="figure@141.[226,1360,193,1592]" captionTargetPageId="141" captionText="FIGURES 99 101. Aedeagi of holotypes, and aedeagus of H. delvasi non-type from Venezuela." httpUri="https://zenodo.org/record/1050172/files/figure.png" pageId="42" pageNumber="43">99</figureCitation>
). The aedeagus of
<taxonomicName id="8FCAB2FFFF84FFD7FD74126D5C5B9635" box="[750,890,861,885]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="beniensis">
<emphasis id="7ABE156EFF84FFD7FD74126D5C5B9635" box="[750,890,861,885]" italics="true" pageId="42" pageNumber="43">H. beniensis</emphasis>
</taxonomicName>
is more similar to that of
<taxonomicName id="8FCAB2FFFF84FFD7FB3C126D5ABD9635" authority="Perkins" authorityName="Perkins" box="[1190,1436,861,885]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trinidensis">
<emphasis id="7ABE156EFF84FFD7FB3C126D5A1E9635" box="[1190,1343,861,885]" italics="true" pageId="42" pageNumber="43">H. trinidensis</emphasis>
Perkins
</taxonomicName>
(
<figureCitation id="D0F1D5F9FF84FFD7FF0512B15FD596DA" box="[159,244,898,922]" captionStart="FIGURES 71 73" captionStartId="123.[151,264,1582,1605]" captionText="FIGURES 71 73. Aedeagi of holotypes." pageId="42" pageNumber="43">Fig. 73</figureCitation>
), but that species is larger (ca. 1.34 vs.
<quantity id="8F326499FF84FFD7FD2512B15C0896DA" box="[703,809,898,922]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="42" pageNumber="43" unit="mm" value="1.25">1.25 mm</quantity>
). Reliable determinations will require dissection of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="00D09AF7FF84FFD7FF5D12FE5AAE927F" pageId="42" pageNumber="43" type="description">
<paragraph id="4875C97CFF84FFD7FF5D12FE5B4F9115" blockId="42.[151,1437,599,1931]" pageId="42" pageNumber="43">
<emphasis id="7ABE156EFF84FFD7FF5D12FE5E7096A6" bold="true" box="[199,337,973,998]" pageId="42" pageNumber="43">Description</emphasis>
. Size:
<typeStatus id="977177DEFF84FFD7FE3712FE5D2E96A5" box="[429,527,973,997]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.25/0.51; head 0.18/0.32; pronotum 0.31/0.41, PA 0.35, PB 0.38; elytra 0.76/0.51. Dorsum of head piceous; pronotum testaceous in front of and behind piceous fascia, laterally fascia becoming smaller and brown, ratios of color bands, as measured in midline, ca.
<date id="3C74EFBCFF84FFD7FF6C150E5E619115" box="[246,320,1085,1109]" pageId="42" pageNumber="43" value="2011-05-05">5/11/5</date>
; elytra brown; legs testaceous; maxillary palpi testaceous, tip not darker.
</paragraph>
<paragraph id="4875C97CFF84FFD7FF5D15505D21904F" blockId="42.[151,1437,599,1931]" pageId="42" pageNumber="43">Frons punctures ca. 1xef, slightly larger and denser near eyes than medially; interstices laterally effacedly microreticulate, weakly shining, ca. 0.51xpd, medially shining, ca. 12xpd. Clypeus microreticulate laterally, very finely sparsely punctulate medially. Mentum moderately sparsely moderately finely punctulate, shining; postmentum anteriorly shining, sparsely punctulate, posteriorly very finely densely micropunctulate, dull. Genae raised, shining, without posterior ridge.</paragraph>
<paragraph id="4875C97CFF84FFD7FF5D142D5BBE903F" blockId="42.[151,1437,599,1931]" pageId="42" pageNumber="43">Pronotum weakly arcuate laterally; anterior margin straight behind eyes, emarginate behind frons, scintilla absent or obsolete; punctures on disc ca. 1xpd those on frons, interstices shining, 14xpd on disc, punctures slightly larger and denser at anterior and posterior; PF1, PF2, and PF4 absent; PF3 moderately deep.</paragraph>
<paragraph id="4875C97CFF84FFD7FF5D14BE5CB0937A" blockId="42.[151,1437,599,1931]" pageId="42" pageNumber="43">Elytra weakly arcuate laterally; summit of posterior declivity slightly before midlength; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming slightly smaller and more widely spaced toward posterior. Intervals not raised, shining, on basal 1/3 ca. 12xpd, as are interstices between punctures of a row, a few punctures subserial. Apices in dorsal aspect conjointly, rather sharply rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="4875C97CFF84FFD7FF5D17745AAE927F" blockId="42.[151,1437,599,1931]" pageId="42" pageNumber="43">Ratios of P2 width and plaque shape (P2/w/l/s) ca. 2/1.5/5/4. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel or nearly so, apex blunt, raised slightly above mesoventral intercoxal process. Plaques moderately narrow, straight, very slightly converging toward one another anteriorly, weakly transversely rounded, located at sides of moderately deep median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at straight posterior margin ca. 1.5x P2. Protibia weakly arcuate in proximal 1/2, medial margin slightly widened in distal 1/2. Meso- and metatibia slender, straight. Abdominal apex symmetrical; last tergite with deep apicomedian notch. Females not yet known.</paragraph>
</subSubSection>
<subSubSection id="00D09AF7FF84FFD7FF5D167E5C2E9225" box="[199,783,1869,1894]" pageId="42" pageNumber="43" type="etymology">
<paragraph id="4875C97CFF84FFD7FF5D167E5C2E9225" blockId="42.[151,1437,599,1931]" box="[199,783,1869,1894]" pageId="42" pageNumber="43">
<emphasis id="7ABE156EFF84FFD7FF5D167E5E6F9225" bold="true" box="[199,334,1869,1893]" pageId="42" pageNumber="43">Etymology.</emphasis>
Named in reference to the
<typeStatus id="977177DEFF84FFD7FDE4167D5D8F9226" box="[638,686,1870,1894]" pageId="42" pageNumber="43">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="00D09AF7FF84FFD7FF5D16415CF492CA" box="[199,981,1906,1931]" pageId="42" pageNumber="43" type="distribution">
<paragraph id="4875C97CFF84FFD7FF5D16415CF492CA" blockId="42.[151,1437,599,1931]" box="[199,981,1906,1931]" pageId="42" pageNumber="43">
<emphasis id="7ABE156EFF84FFD7FF5D16415E4192CB" bold="true" box="[199,352,1906,1931]" pageId="42" pageNumber="43">Distribution.</emphasis>
Currently known only from the
<typeStatus id="977177DEFF84FFD7FD5616405DDD92CB" box="[716,764,1907,1931]" pageId="42" pageNumber="43">type</typeStatus>
locality (
<figureCitation id="D0F1D5F9FF84FFD7FCFC16415CE892CA" box="[870,969,1906,1930]" captionStart="FIGURE 199" captionStartId="192.[151,250,1650,1673]" captionText="FIGURE 199. Geographical distributions of Hydraena species." pageId="42" pageNumber="43">Fig. 199</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>