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<document id="C3B9DE27DAA7A8B3D9AA4CFBC675A349" ID-CLB-Dataset="20416" ID-DOI="10.5852/ejt.2022.798.1671" ID-GBIF-Dataset="dc185c1a-2bc2-48f1-bbdd-e0ae42f852f1" ID-ISSN="2118-9773" ID-Zenodo-Dep="6323040" ID-ZooBank="F136E044-62C8-4FB3-8160-7DAE663D9600" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646225490206" checkinUser="felipe" docAuthor="Houben, Albrecht M., Monnens, Marlies, Proesmans, Willem &amp; Artois, Tom J." docDate="2022" docId="038A87DAA769FF8807C7FDC7FCBA0CD3" docLanguage="en" docName="ejt-798_70-102.pdf" docOrigin="European Journal of Taxonomy 798" docSource="http://dx.doi.org/10.5852/ejt.2022.798.1671" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hoplopera isis Houben, Proesmans &amp; Artois 2022, sp. nov." docType="treatment" docUuid="4DA3359A-D279-4286-B7B0-822382D258F7" docUuidSource="ZooBank" docVersion="11" lastPageNumber="88" masterDocId="FFB3FFA2A779FF9A0626FF8AFFA50F38" masterDocTitle="Limnoterrestrial ‘ Typhloplanidae’ (Rhabdocoela, Platyhelminthes), with the description of four new species and a new genus" masterLastPageNumber="102" masterPageNumber="70" pageNumber="86" updateTime="1699316596528" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1E9A04D2260FF899FAE1F14CE1AB327F">Limnoterrestrial ‘ Typhloplanidae’ (Rhabdocoela, Platyhelminthes), with the description of four new species and a new genus</mods:title>
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<mods:namePart id="659F35C23AC4660E57D9088689330EC5">Houben, Albrecht M.</mods:namePart>
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<mods:affiliation id="1FD1CFEB2077A96CAEB05409F96F2625">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
<mods:nameIdentifier id="904EBEDA89E2869D74DC5DA8BDC0D064" type="email">albrecht.houben@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C32FFDEED7EBCD84BF8DBE7B67CC2607">Monnens, Marlies</mods:namePart>
<mods:nameIdentifier id="8786C9D9C751AE07E55AF5F6998C186F" type="LSID">782F71E0-EF84-48DA-BE72-8E205CB78EAC</mods:nameIdentifier>
<mods:affiliation id="40F5EE3B782A42B0A42A82E3F57F533A">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
<mods:nameIdentifier id="EA8462014BD5018264A4E5D4E2D2C16C" type="email">marlies.monnens@uhasselt.be</mods:nameIdentifier>
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<mods:namePart id="2FC5E3C5252C71D7C0C42DEF424C062A">Proesmans, Willem</mods:namePart>
<mods:nameIdentifier id="76F953888A68DAA24B089EC048C7D230" type="LSID">41B81434-06EE-4BD0-8DCE-4DE73F2A6B30</mods:nameIdentifier>
<mods:affiliation id="BB9EE854A12B923EFE804BC44E79AC7F">Agroécologie, INRAE, 17 Rue Sully, F- 21065 Dijon, France.</mods:affiliation>
<mods:nameIdentifier id="B7567793CA3920CB57431172A3AAC3A2" type="email">willem.proesmans@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FE5882C19F8A7CE4FBCFA551CA802DA2">Artois, Tom J.</mods:namePart>
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<mods:affiliation id="B0EFFB92CD1A5DA657A6F76AB3434546">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
<mods:nameIdentifier id="51A8C7F48E6FCFAE6451DFF234CAEAEB" type="email">tom.artois@uhasselt.be</mods:nameIdentifier>
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<mods:title id="4028C8580E3F8890DAB8B5A100C3FDAE">European Journal of Taxonomy</mods:title>
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<mods:date id="AE2B35F089BA11B3080E7F128D1D90BA">2022</mods:date>
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<mods:number id="A3E184BD9184B79763F7F899A9DAC0E3">2022-03-01</mods:number>
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<treatment id="038A87DAA769FF8807C7FDC7FCBA0CD3" ID-DOI="http://doi.org/10.5281/zenodo.6328790" ID-GBIF-Taxon="193858136" ID-Zenodo-Dep="6328790" LSID="urn:lsid:zoobank.org:act:4DA3359A-D279-4286-B7B0-822382D258F7" httpUri="http://treatment.plazi.org/id/038A87DAA769FF8807C7FDC7FCBA0CD3" lastPageId="18" lastPageNumber="88" pageId="16" pageNumber="86">
<subSubSection id="C3396547A769FF8A07C7FDC7FBF60D50" box="[481,1107,589,616]" pageId="16" pageNumber="86" type="nomenclature">
<paragraph id="8B9C36CCA769FF8A07C7FDC7FBF60D50" blockId="16.[371,1217,589,686]" box="[481,1107,589,616]" pageId="16" pageNumber="86">
<heading id="D0D481A0A769FF8A07C7FDC7FBF60D50" box="[481,1107,589,616]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="86" reason="2">
<taxonomicName id="4C234D4FA769FF8A07C7FDC7FC540D5F" authority="Houben, Proesmans &amp; Artois" authorityName="Houben, Proesmans &amp; Artois" authorityYear="2022" box="[481,1009,589,616]" family="Typhloplanidae" genus="Hoplopera" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="16" pageNumber="86" phylum="Platyhelminthes" rank="species" species="isis" status="sp. nov.">
<emphasis id="B957EADEA769FF8A07C7FDC7FD370D5F" bold="true" box="[481,658,589,616]" italics="true" pageId="16" pageNumber="86">Hoplopera isis</emphasis>
Houben, Proesmans &amp; Artois
</taxonomicName>
<taxonomicNameLabel id="A26457A5A769FF8A05DFFDC4FBF60D50" box="[1017,1107,590,616]" pageId="16" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3396547A769FF8A0755FDFAFCFD0D96" pageId="16" pageNumber="86" type="description">
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<uri id="FFB23ACEA769FF8A0755FDFAFB640DB3" box="[371,1217,624,651]" pageId="16" pageNumber="86">
urn:lsid:zoobank.org:act:
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</uri>
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<paragraph id="8B9C36CCA769FF8A04FAFD19FCFD0D96" blockId="16.[371,1217,589,686]" box="[732,856,659,686]" pageId="16" pageNumber="86">
<figureCitation id="13182A49A769FF8A04FAFD19FCFD0D96" box="[732,856,659,686]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7A–C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3396547A769FF8A069BFD5FFD150C92" pageId="16" pageNumber="86" type="diagnosis">
<paragraph id="8B9C36CCA769FF8A069BFD5FFE920DD7" blockId="16.[189,311,725,751]" box="[189,311,725,751]" pageId="16" pageNumber="86">
<heading id="D0D481A0A769FF8A069BFD5FFE920DD7" bold="true" box="[189,311,725,751]" fontSize="11" level="3" pageId="16" pageNumber="86" reason="3">
<emphasis id="B957EADEA769FF8A069BFD5FFE920DD7" bold="true" box="[189,311,725,751]" pageId="16" pageNumber="86">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA769FF8A069BFC89FD150C92" blockId="16.[189,1399,771,939]" pageId="16" pageNumber="86">
Dark species of
<taxonomicName id="4C234D4FA769FF8A075CFC89FE5F0C25" authorityName="Reisinger" authorityYear="1924" box="[378,506,771,797]" family="Typhloplanidae" genus="Hoplopera" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="16" pageNumber="86" phylum="Platyhelminthes" rank="genus">
<emphasis id="B957EADEA769FF8A075CFC89FE5F0C25" box="[378,506,771,797]" italics="true" pageId="16" pageNumber="86">Hoplopera</emphasis>
</taxonomicName>
, about
<quantity id="4CDB9B29A769FF8A0477FC8EFD0E0C26" box="[593,683,772,798]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="86" unit="mm" value="0.8">0.8 mm</quantity>
long with inconspicuous tail glands. Dermal rhabdites small, adenal rhabdites in two tracks. Atrial glands present. Spindle-shaped testes. Copulatory organ 20 µm long, without sclerotised structures. Bipartite bursa with sclerotised structure containing two groups of parallel, vertical rods, separated from each other by vertical bars, as a whole resembling a fish pot. Female duct forming a seminal receptacle.
</paragraph>
</subSubSection>
<subSubSection id="C3396547A769FF8A069BFC58FA810B23" box="[189,1316,978,1051]" pageId="16" pageNumber="86" type="etymology">
<paragraph id="8B9C36CCA769FF8A069BFC58FEE00CD4" blockId="16.[189,325,978,1004]" box="[189,325,978,1004]" pageId="16" pageNumber="86">
<heading id="D0D481A0A769FF8A069BFC58FEE00CD4" bold="true" box="[189,325,978,1004]" fontSize="11" level="3" pageId="16" pageNumber="86" reason="3">
<emphasis id="B957EADEA769FF8A069BFC58FEE00CD4" bold="true" box="[189,325,978,1004]" pageId="16" pageNumber="86">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA769FF8A069BFB8AFA810B23" blockId="16.[189,1316,1024,1051]" box="[189,1316,1024,1051]" pageId="16" pageNumber="86">
The species epithet is dedicated to
<taxonomicName id="4C234D4FA769FF8A0470FB8AFD430B22" authority="Houben" authorityName="Houben, Proesmans &amp; Artois" authorityYear="2022" box="[598,742,1024,1050]" family="Typhloplanidae" genus="Hoplopera" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="16" pageNumber="89" phylum="Platyhelminthes" rank="species" species="isis">Isis Houben</taxonomicName>
, first daughter of the first author (A.M. Houben).
</paragraph>
</subSubSection>
<subSubSection id="C3396547A769FF8A069BFBCBFE660A44" pageId="16" pageNumber="86" type="materials_examined">
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<heading id="D0D481A0A769FF8A069BFBCBFE080B63" bold="true" box="[189,429,1089,1115]" fontSize="11" level="3" pageId="16" pageNumber="86" reason="3">
<emphasis id="B957EADEA769FF8A069BFBCBFE080B63" bold="true" box="[189,429,1089,1115]" pageId="16" pageNumber="86">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA769FF8A06CCFBFAFCAA0BCC" blockId="16.[187,1399,1136,1268]" pageId="16" pageNumber="86">
<materialsCitation id="3B4B3C91A769FF8A06CCFBFAFCAA0BCC" ID-GBIF-Occurrence="3695141319" collectedFrom="moss growing in a small stream in a pine forest" collectingDate="2011-08-23" collectionCode="FMNH" collectorName="A. M. Houben &amp; W. Proesmans" country="Austria" latitude="46.829445" location="AUSTRIA" longLatPrecision="18" longitude="15.1025" pageId="16" pageNumber="86" specimenCode="KV.688" specimenCount="1" typeStatus="holotype">
<emphasis id="B957EADEA769FF8A06CCFBFAFEF80BB2" bold="true" box="[234,349,1136,1162]" pageId="16" pageNumber="86">
<typeStatus id="5498886EA769FF8A06CCFBFAFEF80BB2" box="[234,349,1136,1162]" pageId="16" pageNumber="86" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F334765CA769FF8A069BFB1EFE9F0B96" box="[189,314,1172,1198]" name="Austria" pageId="16" pageNumber="86">AUSTRIA</collectingCountry>
•
<specimenCount id="9D25FD45A769FF8A0771FB1EFE080B96" box="[343,429,1172,1198]" pageId="16" pageNumber="86" type="generic">1 spec.</specimenCount>
, studied alive and horizontally sectioned;
<locationDeviation id="9A2E2FEEA769FF8A0583FB19FE910BE9" pageId="16" pageNumber="86">south of Graz, between Glashütten and Trahütten</locationDeviation>
;
<geoCoordinate id="EE17500BA769FF8A071BFB3CFE6A0BE9" box="[317,463,1205,1233]" degrees="46" direction="north" minutes="49" orientation="latitude" pageId="16" pageNumber="86" precision="15" seconds="46" value="46.829445">46°49′46″ N</geoCoordinate>
,
<geoCoordinate id="EE17500BA769FF8A07FFFB3CFDCF0BE9" box="[473,618,1205,1233]" degrees="15" direction="east" minutes="06" orientation="longitude" pageId="16" pageNumber="86" precision="15" seconds="09" value="15.1025">15°06′09″ E</geoCoordinate>
;
<date id="FF9D100CA769FF8A0453FB3DFCB70BE9" box="[629,786,1207,1233]" pageId="16" pageNumber="86" value="2011-08-23">
<collectingDate id="EFD9E9E4A769FF8A0453FB3DFCB70BE9" box="[629,786,1207,1233]" pageId="16" pageNumber="86" value="2011-08-23">23 Aug. 2011</collectingDate>
</date>
;
<collectorName id="26D6531AA769FF8A053DFB3DFC1E0BE9" box="[795,955,1207,1233]" pageId="16" pageNumber="86">A.M. Houben</collectorName>
and
<collectorName id="26D6531AA769FF8A05D6FB3DFB300BE9" box="[1008,1173,1207,1233]" pageId="16" pageNumber="86">W. Proesmans</collectorName>
leg.;
<collectedFrom id="383F7531A769FF8A02F5FB3DFD980BCC" pageId="16" pageNumber="86">moss growing in a small stream in a pine forest</collectedFrom>
;
<specimenCode id="DB859EB7A769FF8A046FFB50FD030BCC" box="[585,678,1242,1268]" pageId="16" pageNumber="86">KV.688</specimenCode>
;
<collectionCode id="ED32AE09A769FF8A0494FB50FCAE0BCC" box="[690,779,1242,1268]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="86" type="Museum">FMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9C36CCA769FF8A06CCFA91FE660A44" blockId="16.[189,1398,1307,1404]" pageId="16" pageNumber="86">
<materialsCitation id="3B4B3C91A769FF8A06CCFA91FE660A44" ID-GBIF-Occurrence="3695141308" collectedFrom="moss growing in a small stream in a pine forest" collectingDate="2011-08-23" collectionCode="HU" collectorName="A. M. Houben &amp; W. Proesmans" country="Austria" latitude="46.829445" location="AUSTRIA" longLatPrecision="18" longitude="15.1025" pageId="16" pageNumber="86" specimenCode="822" specimenCount="5" typeStatus="paratype">
<emphasis id="B957EADEA769FF8A06CCFA91FECC0A0D" bold="true" box="[234,361,1307,1333]" pageId="16" pageNumber="86">
<typeStatus id="5498886EA769FF8A06CCFA91FECC0A0D" box="[234,361,1307,1333]" pageId="16" pageNumber="86" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F334765CA769FF8A069BFAB5FE9F0A61" box="[189,314,1343,1369]" name="Austria" pageId="16" pageNumber="86">AUSTRIA</collectingCountry>
•
<specimenCount id="9D25FD45A769FF8A0772FAB4FE080A61" box="[340,429,1342,1369]" pageId="16" pageNumber="86">5 specs</specimenCount>
, studied alive, one of which whole mounted; same collection data as for holotype; paratype no.
<specimenCode id="DB859EB7A769FF8A0770FAE8FE200A44" box="[342,389,1378,1404]" pageId="16" pageNumber="86">822</specimenCode>
;
<collectionCode id="ED32AE09A769FF8A07B7FAE8FE1A0A44" box="[401,447,1378,1404]" pageId="16" pageNumber="86">HU</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3396547A769FF88069BFA29FEB80DD0" lastPageId="18" lastPageNumber="88" pageId="16" pageNumber="86" type="description">
<paragraph id="8B9C36CCA769FF8A069BFA29FEEA0A85" blockId="16.[189,335,1443,1469]" box="[189,335,1443,1469]" pageId="16" pageNumber="86">
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<emphasis id="B957EADEA769FF8A069BFA29FEEA0A85" bold="true" box="[189,335,1443,1469]" pageId="16" pageNumber="86">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA769FF8A069BFA58FD5D0941" blockId="16.[188,1399,1490,1657]" pageId="16" pageNumber="86">
Specimens are about
<quantity id="4CDB9B29A769FF8A07E0FA58FDFC0AD4" box="[454,601,1490,1516]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="16" pageNumber="86" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.7–0.9 mm</quantity>
long. The body is rounded anteriorly and possesses a small tail provided with small, inconspicuous tail glands (
<figureCitation id="13182A49A769FF8A04CBFA7FFCEC0937" box="[749,841,1525,1551]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7C</figureCitation>
: tg). Rostrally, sharply pointed adenal rhabdites (
<figureCitation id="13182A49A769FF8A06E3F992FE84090B" box="[197,289,1560,1587]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7B</figureCitation>
: ar) are arranged in two groups. Very small dermal rhabdites occur all over the body. The paired protonephridiopores (
<figureCitation id="13182A49A769FF8A079FF9B1FDBD096D" box="[441,536,1595,1621]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7A</figureCitation>
: pp) lie posterior and lateral to the mouth. The rosulate pharynx is located just behind the middle of the body (
<figureCitation id="13182A49A769FF8A0479F9D4FD1B0941" box="[607,702,1630,1657]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7A</figureCitation>
: ph).
</paragraph>
<paragraph id="8B9C36CCA769FF8A069BF92AFAF80839" blockId="16.[188,1399,1696,1793]" pageId="16" pageNumber="86">
The gonopore (
<figureCitation id="13182A49A769FF8A075CF92AFE5A0982" box="[378,511,1696,1722]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7B–C</figureCitation>
: gp) is situated at ±80% of the body and connected to a genital atrium (
<figureCitation id="13182A49A769FF8A06E0F949FEE309E5" box="[198,326,1731,1757]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7A–C</figureCitation>
: ga) that is lined with a high epithelium and surrounded by muscles of uncertain orientation. At its posterior side, the genital atrium receives a large group of eosinophilic glands (
<figureCitation id="13182A49A769FF8A0282F96CFA800838" box="[1188,1317,1766,1792]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7A–C</figureCitation>
: ag).
</paragraph>
<paragraph id="8B9C36CCA769FF88069BF8A2FC600D21" blockId="16.[189,1400,1832,1999]" lastBlockId="18.[188,1400,265,538]" lastPageId="18" lastPageNumber="88" pageId="16" pageNumber="86">
Two small, spindle-shaped testes (
<figureCitation id="13182A49A769FF8A0469F8A2FD0B087A" box="[591,686,1832,1858]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7A</figureCitation>
: t) lie in front of the pharynx (
<figureCitation id="13182A49A769FF8A0237F8A2FBD5087A" box="[1041,1136,1832,1858]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7A</figureCitation>
: ph) and ventral to the paired vitellaria (
<figureCitation id="13182A49A769FF8A07ACF8C1FDAB085D" box="[394,526,1867,1893]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7A, C</figureCitation>
: vi). In some specimens, one testis seems bigger than the other one. The paired vasa deferentia (
<figureCitation id="13182A49A769FF8A07F5F8E4FDF108B0" box="[467,596,1902,1928]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7B–C</figureCitation>
: vde) enter the egg-shaped copulatory organ laterally. Two layers of spiral muscles surround the 24µm long copulatory organ (
<figureCitation id="13182A49A769FF8A055DF81BFC780893" box="[891,989,1937,1963]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7A</figureCitation>
: co), which bears an intracapsular seminal vesicle (
<figureCitation id="13182A49A769FF8A0759F83EFE5808F7" box="[383,509,1972,1999]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7B–C</figureCitation>
: vs) and an ejaculatory duct (
<figureCitation id="13182A49A769FF8A056BF83EFC6908F7" box="[845,972,1972,1999]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="16" pageNumber="86">Fig. 7B–C</figureCitation>
: de). Coarse-grained intra- (
<figureCitation id="13182A49A769FF880336F83EFF710E1B" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" lastPageId="18" lastPageNumber="88" pageId="16" pageNumber="86">Fig. 7B– C</figureCitation>
: gg1) and extracapsular (
<figureCitation id="13182A49A76BFF8807D8FE83FDF90E1B" box="[510,604,265,291]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="18" pageNumber="88">Fig. 7C</figureCitation>
: gg2) eosinophilic glands are associated with the copulatory organ. The relatively long male duct (
<figureCitation id="13182A49A76BFF88041EFEA6FD3D0E7E" box="[568,664,300,326]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="18" pageNumber="88">Fig. 7C</figureCitation>
: md) receives the bursa (
<figureCitation id="13182A49A76BFF8805E8FEA6FBF70E7E" box="[974,1106,300,326]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="18" pageNumber="88">Fig. 7A–C</figureCitation>
: bu) before entering the genital atrium (
<figureCitation id="13182A49A76BFF880749FEC5FE550E51" box="[367,496,335,361]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="18" pageNumber="88">Fig. 7A–C</figureCitation>
: ga). This bipartite bursa consists of an 18 µm long, thin-walled proximal part containing sperm and a 22 µm long distal part, which could be considered the bursal stalk (
<figureCitation id="13182A49A76BFF8802D4FEF8FAD60EB5" box="[1266,1395,370,397]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="18" pageNumber="88">Fig. 7B–C</figureCitation>
: bs). This stalk is surrounded by circular muscles and contains a sclerotised structure (
<figureCitation id="13182A49A76BFF880290FE1CFAB00E88" box="[1206,1301,406,432]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="18" pageNumber="88">Fig. 7B</figureCitation>
: ss) that resembles the structure of a fish pot. It consists of vertical rods that are divided into two groups by a horizontal bar. The proximal and distal part are also delimited by a horizontal bar. As a whole, this bursal stalk is wrapped in a loose matrix surrounded by circular muscles.
</paragraph>
<caption id="DF5C6644A768FF8B069BF8CBFB1808D0" ID-DOI="http://doi.org/10.5281/zenodo.6323058" ID-Zenodo-Dep="6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="17" pageNumber="87" startId="17.[189,232,1857,1883]" targetBox="[318,1331,280,1797]" targetPageId="17">
<paragraph id="8B9C36CCA768FF8B069BF8CBFB1808D0" blockId="17.[188,1398,1857,2025]" pageId="17" pageNumber="87">
<emphasis id="B957EADEA768FF8B069BF8CBFEA90864" bold="true" box="[189,268,1857,1884]" pageId="17" pageNumber="87">Fig. 7.</emphasis>
Two species of
<taxonomicName id="4C234D4FA768FF8B07EAF8CBFCB50864" authority="Reisinger, 1924" authorityName="Reisinger" authorityYear="1924" box="[460,784,1857,1884]" family="Typhloplanidae" genus="Hoplopera" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="17" pageNumber="87" phylum="Platyhelminthes" rank="genus">
<emphasis id="B957EADEA768FF8B07EAF8CBFDE90863" box="[460,588,1857,1883]" italics="true" pageId="17" pageNumber="87">Hoplopera</emphasis>
<bibRefCitation id="EFB24B3DA768FF8B0472F8CBFCB50864" author="Reisinger E." box="[596,784,1857,1884]" pageId="17" pageNumber="87" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger, 1924</bibRefCitation>
</taxonomicName>
.
<emphasis id="B957EADEA768FF8B053BF8CBFCF30863" bold="true" box="[797,854,1857,1883]" pageId="17" pageNumber="87">A–C</emphasis>
.
<taxonomicName id="4C234D4FA768FF8B0543F8CBFAD30863" authority="Houben, Proesmans &amp; Artois" authorityName="Houben, Proesmans &amp; Artois" authorityYear="2022" box="[869,1398,1857,1884]" family="Typhloplanidae" genus="Hoplopera" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="17" pageNumber="87" phylum="Platyhelminthes" rank="species" species="isis" status="sp. nov.">
<emphasis id="B957EADEA768FF8B0543F8CBFBB00863" box="[869,1045,1857,1883]" italics="true" pageId="17" pageNumber="87">Hoplopera isis</emphasis>
Houben, Proesmans &amp; Artois
</taxonomicName>
<taxonomicNameLabel id="A26457A5A768FF8B069BF8EFFEBE0847" box="[189,283,1893,1919]" pageId="17" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B957EADEA768FF8B0703F8EEFE9F0846" bold="true" box="[293,314,1892,1918]" pageId="17" pageNumber="87">A</emphasis>
. Internal organisation of a live specimen.
<emphasis id="B957EADEA768FF8B0566F8EFFCF10847" bold="true" box="[832,852,1893,1919]" pageId="17" pageNumber="87">B</emphasis>
. Organisation of the genital system of a live specimen.
<emphasis id="B957EADEA768FF8B0719F802FEF1089A" bold="true" box="[319,340,1928,1954]" pageId="17" pageNumber="87">C</emphasis>
. Reconstruction of reproductive organs based on horizontal sections.
<emphasis id="B957EADEA768FF8B028BF802FB46089A" bold="true" box="[1197,1251,1928,1954]" pageId="17" pageNumber="87">D–F</emphasis>
.
<taxonomicName id="4C234D4FA768FF8B02D0F802FE6308FD" authority="Reisinger, 1924" authorityName="Reisinger" authorityYear="1924" family="Typhloplanidae" genus="Hoplopera" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="17" pageNumber="87" phylum="Platyhelminthes" rank="species" species="opaca">
<emphasis id="B957EADEA768FF8B02D0F802FEA008FD" italics="true" pageId="17" pageNumber="87">Hoplopera opaca</emphasis>
<bibRefCitation id="EFB24B3DA768FF8B072DF821FE6308FD" author="Reisinger E." box="[267,454,1963,1989]" pageId="17" pageNumber="87" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger, 1924</bibRefCitation>
</taxonomicName>
.
<emphasis id="B957EADEA768FF8B07E8F821FE4608FD" bold="true" box="[462,483,1963,1989]" pageId="17" pageNumber="87">D</emphasis>
. Internal organisation of a live specimen.
<emphasis id="B957EADEA768FF8B05EEF821FC7908FD" bold="true" box="[968,988,1963,1989]" pageId="17" pageNumber="87">E</emphasis>
. Organisation of the genital system of a live specimen.
<emphasis id="B957EADEA768FF8B0784F844FE1108D0" bold="true" box="[418,436,1998,2024]" pageId="17" pageNumber="87">F</emphasis>
. Reconstruction of reproductive organs based on sagittal sections.
</paragraph>
</caption>
<paragraph id="8B9C36CCA76BFF88069BFDCBFEB80DD0" blockId="18.[189,1400,577,744]" pageId="18" pageNumber="88">
The vitellaria (
<figureCitation id="13182A49A76BFF88074CFDCBFE4F0D63" box="[362,490,577,603]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="18" pageNumber="88">Fig. 7A, C</figureCitation>
: vi) extend from the anterior third of the body to the posterior end where they meet. The oviduct broadens and becomes a seminal receptacle (
<figureCitation id="13182A49A76BFF88058AFDEEFB8B0D46" box="[940,1070,612,638]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="18" pageNumber="88">Fig. 7A–C</figureCitation>
: rs), which further becomes a long female duct (
<figureCitation id="13182A49A76BFF880789FD0DFDA80D99" box="[431,525,647,673]" captionStart="Fig" captionStartId="17.[189,232,1857,1883]" captionTargetBox="[318,1331,280,1797]" captionTargetId="figure-13@17.[251,1336,265,1818]" captionTargetPageId="17" captionText="Fig. 7. Two species of Hoplopera Reisinger, 1924. A–C. Hoplopera isis Houben, Proesmans &amp; Artois sp. nov. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on horizontal sections. D–F. Hoplopera opaca Reisinger, 1924. D. Internal organisation of a live specimen. E. Organisation of the genital system of a live specimen. F. Reconstruction of reproductive organs based on sagittal sections." figureDoi="http://doi.org/10.5281/zenodo.6323058" httpUri="https://zenodo.org/record/6323058/files/figure.png" pageId="18" pageNumber="88">Fig. 7C</figureCitation>
: fd), lined with a high, nucleated epithelium and surrounded by muscles. The vitelloduct was not observed with certainty, although it is probably an unpaired one that meets the oviduct.
</paragraph>
</subSubSection>
<subSubSection id="C3396547A76BFF88069BFC85FCBA0CD3" pageId="18" pageNumber="88" type="discussion">
<paragraph id="8B9C36CCA76BFF88069BFC85FEE70C11" blockId="18.[189,322,783,809]" box="[189,322,783,809]" pageId="18" pageNumber="88">
<heading id="D0D481A0A76BFF88069BFC85FEE70C11" bold="true" box="[189,322,783,809]" fontSize="11" level="3" pageId="18" pageNumber="88" reason="3">
<emphasis id="B957EADEA76BFF88069BFC85FEE70C11" bold="true" box="[189,322,783,809]" pageId="18" pageNumber="88">Discussion</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA76BFF88069BFCB4FCB90C60" blockId="18.[189,796,829,856]" box="[189,796,829,856]" pageId="18" pageNumber="88">
See the general discussion on the genus
<taxonomicName id="4C234D4FA76BFF8804B3FCB7FCB00C6F" authorityName="Reisinger" authorityYear="1924" box="[661,789,829,855]" family="Typhloplanidae" genus="Hoplopera" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="18" pageNumber="88" phylum="Platyhelminthes" rank="genus">
<emphasis id="B957EADEA76BFF8804B3FCB7FCB00C6F" box="[661,789,829,855]" italics="true" pageId="18" pageNumber="88">Hoplopera</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C36CCA76BFF88069BFCF5FE950CA1" blockId="18.[189,304,895,921]" box="[189,304,895,921]" pageId="18" pageNumber="88">
<heading id="D0D481A0A76BFF88069BFCF5FE950CA1" bold="true" box="[189,304,895,921]" fontSize="11" level="3" pageId="18" pageNumber="88" reason="3">
<emphasis id="B957EADEA76BFF88069BFCF5FE950CA1" bold="true" box="[189,304,895,921]" pageId="18" pageNumber="88">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA76BFF88069BFC27FCBA0CD3" blockId="18.[189,1398,941,1003]" pageId="18" pageNumber="88">
The hard parts and copulatory structures are not visible on the whole-mounted specimen. Therefore, we designate a serially sectioned specimen as
<typeStatus id="5498886EA76BFF880497FC5BFCB90CD3" box="[689,796,977,1003]" pageId="18" pageNumber="88" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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