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<document id="3E4AB11971B4837CA062C39AE577774E" ID-CLB-Dataset="55750" ID-DOI="10.5281/zenodo.5385198" ID-GBIF-Dataset="c7f95b94-53f8-420d-97c5-4e27d7c211bf" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385198" ID-ZooBank="AC7F1491-CD89-4F10-9D61-55354B55E368" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630300948061" checkinUser="valdenar" docAuthor="Hájek, Jiří &amp; Brancucci, Michel" docDate="2015" docId="038A87C4FFAFFF96F13E93C5FA7CB809" docLanguage="en" docName="RafflesBZool.63.309-326.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Laccophilus jaechi Hájek &amp; Brancucci 2015, new species" docType="treatment" docUuid="A77D6585-7D1D-4449-94C5-9FC4467845DC" docUuidSource="ZooBank" docVersion="10" lastPageNumber="319" masterDocId="FFB3FFBCFFA5FF9CF02A9279FFD5BB5C" masterDocTitle="A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)" masterLastPageNumber="326" masterPageNumber="309" pageNumber="319" updateTime="1699240741915" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="800DA4BCD922F8DE77410071233F4DD4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="474BDDB5DE3D0A211BF770C0192D21A9">A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="EF3640DD107B0E2997C96BE434632711" type="personal">
<mods:role id="861F7A640D5F63DDBC76130A8C3499B6">
<mods:roleTerm id="015AF032D2DD18899FEC0DDD247004C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E52C85F98675D5954A8D038A3AAE0C8">Hájek, Jiří</mods:namePart>
<mods:affiliation id="B37BB68F36C581249AA2054089D7CFF9">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 – Horní Počernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="AE9C86722BCC709A3562976C47AEDA73" type="email">jiri_hajek@nm.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="828B3A5D90C66027B74E8007E8BCDAD5" type="personal">
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</mods:role>
<mods:namePart id="940C6B07F2B54EBA99797684217FB2F1">Brancucci, Michel</mods:namePart>
<mods:affiliation id="09B078C4833770A8CC2ECAC7FC851953">Naturhistorisches Museum, Basel, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B5A1585EEA58CC66056B2F89C5E2CB5A">text</mods:typeOfResource>
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<mods:title id="F55EA971081CD2E11AA48BE20AE3688C">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="B34E933E74DE9422E37D5A967A4391C4">
<mods:date id="DDDB92AB2367AD064C9C9C10D17225BA">2015</mods:date>
<mods:detail id="17C2954DAD5EBFCB34FDDF1B24F056F6" type="pubDate">
<mods:number id="7CBA29D68E868E13F329EB1C3C8F8E3A">2015-08-06</mods:number>
</mods:detail>
<mods:detail id="29D5837A1C83A733988A166A1F5157D5" type="volume">
<mods:number id="31A7EC8A4B18DD47503345F06B7D62EE">63</mods:number>
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<mods:classification id="A044322844434AFC061526ECA40E4E26">journal article</mods:classification>
<mods:identifier id="43DE0D59D2F70B286FC6383B71758D55" type="CLB-Dataset">55750</mods:identifier>
<mods:identifier id="4A7FB40F57651B75FB557A5F30DC71AC" type="DOI">10.5281/zenodo.5385198</mods:identifier>
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<mods:identifier id="76B9C3EFF699B4274860C2A750CBCD19" type="ISSN">2345-7600</mods:identifier>
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</mods:mods>
<treatment id="038A87C4FFAFFF96F13E93C5FA7CB809" ID-DOI="http://doi.org/10.5281/zenodo.5466887" ID-GBIF-Taxon="194955223" ID-Zenodo-Dep="5466887" LSID="urn:lsid:zoobank.org:act:A77D6585-7D1D-4449-94C5-9FC4467845DC" httpUri="http://treatment.plazi.org/id/038A87C4FFAFFF96F13E93C5FA7CB809" lastPageNumber="319" pageId="10" pageNumber="319">
<subSubSection id="C3396559FFAFFF96F13E93C5FDAEBA89" box="[276,635,444,469]" pageId="10" pageNumber="319" type="nomenclature">
<paragraph id="8B9C36D2FFAFFF96F13E93C5FDAEBA89" blockId="10.[276,635,444,501]" box="[276,635,444,469]" pageId="10" pageNumber="319">
<heading id="D0D481BEFFAFFF96F13E93C5FDAEBA89" bold="true" box="[276,635,444,469]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="319" reason="2">
<emphasis id="B957EAC0FFAFFF96F13E93C5FDAEBA89" bold="true" box="[276,635,444,469]" pageId="10" pageNumber="319">
<taxonomicName id="4C234D51FFAFFF96F13E93C5FE32BA88" ID-CoL="8LXXF" authority="Hájek &amp; Brancucci, 2015" authorityName="Hájek &amp; Brancucci" authorityYear="2015" box="[276,487,444,468]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="319" phylum="Arthropoda" rank="species" species="jaechi" status="sp. nov.">
<emphasis id="B957EAC0FFAFFF96F13E93C5FE32BA88" bold="true" box="[276,487,444,468]" italics="true" pageId="10" pageNumber="319">Laccophilus jaechi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26457BBFFAFFF96F1DC93C4FDAEBA89" box="[502,635,445,469]" pageId="10" pageNumber="319" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAFFF96F1AE93A4FDDEBAA9" box="[388,523,477,501]" pageId="10" pageNumber="319" type="description">
<paragraph id="8B9C36D2FFAFFF96F1AE93A4FDDEBAA9" blockId="10.[276,635,444,501]" box="[388,523,477,501]" pageId="10" pageNumber="319">
(
<figureCitation id="13182A57FFAFFF96F1A693A4FE0CBAA8" box="[396,473,477,501]" captionStart="Figs" captionStartId="3.[113,157,2051,2073]" captionTargetBox="[115,1452,190,2035]" captionTargetId="figure-10@3.[97,1463,180,2039]" captionTargetPageId="3" captionText="Figs. 1–9. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385200" httpUri="https://zenodo.org/record/5385200/files/figure.png" pageId="10" pageNumber="319">Figs. 7</figureCitation>
,
<figureCitation id="13182A57FFAFFF96F1CD93A4FDD6BAA9" box="[487,515,477,501]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="10" pageNumber="319">24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAFFF96F0A29064FD39B809" pageId="10" pageNumber="319" type="materials_examined">
<paragraph id="8B9C36D2FFAFFF96F0A29064FD2BB969" blockId="10.[136,766,541,565]" box="[136,766,541,565]" pageId="10" pageNumber="319">
<emphasis id="B957EAC0FFAFFF96F0A29064FEF1B969" bold="true" box="[136,292,541,565]" pageId="10" pageNumber="319">
<typeStatus id="54988870FFAFFF96F0A29064FF17B969" box="[136,194,541,565]" pageId="10" pageNumber="319">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3347642FFAFFF96F1079064FE5AB969" box="[301,399,541,565]" name="Thailand" pageId="10" pageNumber="319">Thailand</collectingCountry>
,
<collectingRegion id="49E7F830FFAFFF96F1B19064FDE7B969" box="[411,562,541,565]" country="Thailand" name="Trat" pageId="10" pageNumber="319">Trat Province</collectingRegion>
,
<location id="8EFC6009FFAFFF96F2149064FD2EB969" LSID="urn:lsid:plazi:treatment:038A87C4FFAFFF96F13E93C5FA7CB809:8EFC6009FFAFFF96F2149064FD2EB969" box="[574,763,541,565]" country="Thailand" name="Ko Chang Island" pageId="10" pageNumber="319" stateProvince="Trat">Ko Chang Island</location>
.
</paragraph>
<paragraph id="8B9C36D2FFAFFF96F0A29024FD39B809" blockId="10.[136,775,605,855]" pageId="10" pageNumber="319">
<emphasis id="B957EAC0FFAFFF96F0A29024FEB2B929" bold="true" box="[136,359,605,629]" pageId="10" pageNumber="319">Material examined.</emphasis>
<materialsCitation id="3B4B3C8FFFAFFF96F1479024FDC6B9E9" ID-GBIF-Occurrence="3760488401" collectionCode="NHMW" collectorName="Jach" country="Thailand" location="THAILAND" pageId="10" pageNumber="319" specimenCount="1" typeStatus="holotype">
<typeStatus id="54988870FFAFFF96F1479024FE05B929" box="[365,464,605,629]" pageId="10" pageNumber="319" type="holotype">Holotype</typeStatus>
male, labelled: O [eastern] –
<collectingCountry id="F3347642FFAFFF96F0A29004FEC7B9C9" box="[136,274,637,661]" name="Thailand" pageId="10" pageNumber="319">THAILAND</collectingCountry>
1990 / Ko Chang [Island] (12) / White Sands Beach / leg.
<collectorName id="26D65304FFAFFF96F13190E4FE9EB9E9" box="[283,331,669,693]" pageId="10" pageNumber="319">Jäch</collectorName>
10.12. (
<collectionCode id="ED32AE17FFAFFF96F18190E4FDDFB9E9" box="[427,522,669,693]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="319" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFAFFF96F20A90E4FD39B989" ID-GBIF-Occurrence="3760488407" collectionCode="NMPC, NHMW" collectorName="Jach" country="Thailand" location="THAILAND" pageId="10" pageNumber="319" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54988870FFAFFF96F20A90E4FD5BB9E9" box="[544,654,669,693]" pageId="10" pageNumber="319" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D25FD5BFFAFFF96F2B690E4FD3EB9E9" box="[668,747,669,693]" pageId="10" pageNumber="319" type="male">1 male</specimenCount>
,
<specimenCount id="9D25FD5BFFAFFF96F2D090E4FF0EB989" pageId="10" pageNumber="319" type="female">2 females</specimenCount>
: same label data as holotype (
<collectionCode id="ED32AE17FFAFFF96F20190C4FDAEB989" box="[555,635,701,725]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="319">NMPC</collectionCode>
,
<collectionCode id="ED32AE17FFAFFF96F2A390C4FD30B989" box="[649,741,701,725]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="319" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFAFFF96F2D390C4FF24B849" ID-GBIF-Occurrence="3760488378" collectingDate="1981-11" collectionCode="NHMB" collectorName="Coll. T. Lander" country="Thailand" location="Khao Sebap" pageId="10" pageNumber="319" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D25FD5BFFAFFF96F2D390C4FF6AB9A9" pageId="10" pageNumber="319" type="male">1 male</specimenCount>
:
<location id="8EFC6009FFAFFF96F0ED90A4FE91B9A9" LSID="urn:lsid:plazi:treatment:038A87C4FFAFFF96F13E93C5FA7CB809:8EFC6009FFAFFF96F0ED90A4FE91B9A9" box="[199,324,733,757]" country="Thailand" name="Khao Sebap" pageId="10" pageNumber="319">Khao Sebap</location>
/
<collectingCountry id="F3347642FFAFFF96F17F90A4FE39B9A9" box="[341,492,733,757]" name="Thailand" pageId="10" pageNumber="319">THAILANDE</collectingCountry>
/
<date id="FF9D1012FFAFFF96F1D790A4FD8DB9A9" box="[509,600,733,757]" pageId="10" pageNumber="319" value="1981-11">
<collectingDate id="EFD9E9FAFFAFFF96F1D790A4FD8DB9A9" box="[509,600,733,757]" pageId="10" pageNumber="319" value="1981-11">XI. 1981</collectingDate>
</date>
/
<collectorName id="26D65304FFAFFF96F24390A4FCD2B9A9" box="[617,775,733,757]" pageId="10" pageNumber="319">Coll. T. Lander</collectorName>
(
<collectionCode id="ED32AE17FFAFFF96F0A59084FF30B849" box="[143,229,765,789]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="10" pageNumber="319">NHMB</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4B3C8FFFAFFF96F0D29084FD39B809" ID-GBIF-Occurrence="3760488443" collectingDate="2005-05-03" collectingDateMax="2005-05-19" collectingDateMin="2005-05-03" collectionCode="NHMB, NMPC" collectorName="E. Jendek &amp; O. Sausa" country="Cambodia" elevation="0" latitude="11.566667" location="Tatai riv." longitude="103.116667" pageId="10" pageNumber="319" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="KOH KONG">
<specimenCount id="9D25FD5BFFAFFF96F0D29084FE98B849" box="[248,333,765,789]" pageId="10" pageNumber="319" type="male">4 males</specimenCount>
,
<specimenCount id="9D25FD5BFFAFFF96F1729084FE11B849" box="[344,452,765,789]" pageId="10" pageNumber="319" type="female">4 females</specimenCount>
:
<collectingCountry id="F3347642FFAFFF96F1E49084FD8AB849" box="[462,607,765,789]" name="Cambodia" pageId="10" pageNumber="319">CAMBODIA</collectingCountry>
SW,
<locationDeviation id="9A2E2FF0FFAFFF96F2B69084FD2DB849" box="[668,760,765,789]" pageId="10" pageNumber="319">20kmSE</locationDeviation>
/
<collectingRegion id="49E7F830FFAFFF96F0A29164FECCB869" box="[136,281,797,821]" country="Cambodia" name="Koh Kong" pageId="10" pageNumber="319">KOH KONG</collectingRegion>
:
<location id="8EFC6009FFAFFF96F10E9164FE53B869" LSID="urn:lsid:plazi:treatment:038A87C4FFAFFF96F13E93C5FA7CB809:8EFC6009FFAFFF96F10E9164FE53B869" box="[292,390,797,821]" country="Cambodia" name="Tatai riv." pageId="10" pageNumber="319" stateProvince="KOH KONG">Tatai riv.</location>
/ 11°34′, N103°07′E /
<date id="FF9D1012FFAFFF96F2539164FCD7B869" box="[633,770,797,821]" pageId="10" pageNumber="319" value="2005-05-03" valueMax="2005-05-19" valueMin="2005-05-03">
<collectingDate id="EFD9E9FAFFAFFF96F2539164FCD7B869" box="[633,770,797,821]" pageId="10" pageNumber="319" value="2005-05-03" valueMax="2005-05-19" valueMin="2005-05-03">3.-19.v.2005</collectingDate>
</date>
,
<elevation id="000ED1E1FFAFFF96F0A29144FF26B808" box="[136,243,829,853]" pageId="10" pageNumber="319">
50-
<quantity id="4CDB9B37FFAFFF96F0809144FF26B808" box="[170,243,829,853]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="319" unit="m" value="300.0">300 m</quantity>
</elevation>
, /
<collectorName id="26D65304FFAFFF96F13A9144FEA4B809" box="[272,369,829,853]" pageId="10" pageNumber="319">E.Jendek</collectorName>
&amp;
<collectorName id="26D65304FFAFFF96F1BD9144FE25B80B" box="[407,496,829,855]" pageId="10" pageNumber="319">O.Šauša</collectorName>
leg. (
<collectionCode id="ED32AE17FFAFFF96F21A9144FD51B809" box="[560,644,829,853]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="10" pageNumber="319">NHMB</collectionCode>
,
<collectionCode id="ED32AE17FFAFFF96F2BA9144FD34B809" box="[656,737,829,853]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="319">NMPC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAFFF96F0A29104FEE0BFE9" pageId="10" pageNumber="319" type="diagnosis">
<paragraph id="8B9C36D2FFAFFF96F0A29104FEE0BFE9" blockId="10.[136,777,893,1205]" pageId="10" pageNumber="319">
<emphasis id="B957EAC0FFAFFF96F0A29104FF28B8C9" bold="true" box="[136,253,893,917]" pageId="10" pageNumber="319">Diagnosis.</emphasis>
Medium sized, oval oblong species (body length
<quantity id="4CDB9B37FFAFFF96F0A291E4FEDAB8E9" box="[136,271,925,949]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="10" pageNumber="319" unit="mm" value="4.25" valueMax="4.4" valueMin="4.1">4.1–4.4 mm</quantity>
). It is very similar to
<taxonomicName id="4C234D51FFAFFF96F1D491E7FD41B8E9" authorityName="Brancucci" authorityYear="1983" box="[510,660,925,949]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="319" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="B957EAC0FFAFFF96F1D491E7FD41B8E9" box="[510,660,925,949]" italics="true" pageId="10" pageNumber="319">L. tonkinensis</emphasis>
</taxonomicName>
in habitus and elytral colouration, and to
<taxonomicName id="4C234D51FFAFFF96F1E591C7FD84B889" authorityName="Hájek &amp; Brancucci" authorityYear="2015" box="[463,593,957,981]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="319" phylum="Arthropoda" rank="species" species="hendrichi">
<emphasis id="B957EAC0FFAFFF96F1E591C7FD84B889" box="[463,593,957,981]" italics="true" pageId="10" pageNumber="319">L. hendrichi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C234D51FFAFFF96F2AF91C7FCD2B889" authorityName="Regimbart" authorityYear="1899" box="[645,775,958,981]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="319" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="B957EAC0FFAFFF96F2AF91C7FCD2B889" box="[645,775,958,981]" italics="true" pageId="10" pageNumber="319">L. javanicus</emphasis>
</taxonomicName>
in general shape of penis. The new species differs from all three mentioned species by the distinct shape of the median lobe distal part, which is ventrally very weakly sinuate, apically with small inflexed point.
<taxonomicName id="4C234D51FFAFFF96F1D99644FD69BF09" authorityName="Hájek &amp; Brancucci" authorityYear="2015" box="[499,700,1085,1109]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="319" phylum="Arthropoda" rank="species" species="jaechi">
<emphasis id="B957EAC0FFAFFF96F1D99644FD69BF09" box="[499,700,1085,1109]" italics="true" pageId="10" pageNumber="319">Laccophilus jaechi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C234D51FFAFFF96F0E99627FE90BF29" authorityName="Hájek &amp; Brancucci" authorityYear="2015" box="[195,325,1117,1141]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="319" phylum="Arthropoda" rank="species" species="hendrichi">
<emphasis id="B957EAC0FFAFFF96F0E99627FE90BF29" box="[195,325,1117,1141]" italics="true" pageId="10" pageNumber="319">L. hendrichi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C234D51FFAFFF96F1539627FE29BF29" authorityName="Regimbart" authorityYear="1899" box="[377,508,1118,1141]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="319" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="B957EAC0FFAFFF96F1539627FE29BF29" box="[377,508,1118,1141]" italics="true" pageId="10" pageNumber="319">L. javanicus</emphasis>
</taxonomicName>
also in the constant very broad testaceous elytral subbasal band not separated from the elytral base.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAFFF96F0A296A4FAC3B929" pageId="10" pageNumber="319" type="description">
<paragraph id="8B9C36D2FFAFFF96F0A296A4FCDDBEC8" blockId="10.[136,776,1245,1429]" pageId="10" pageNumber="319">
<emphasis id="B957EAC0FFAFFF96F0A296A4FEC6BFA9" bold="true" box="[136,275,1245,1269]" pageId="10" pageNumber="319">Description.</emphasis>
Colouration: Head and pronotum testaceous, elytra brownish-black with testaceous pattern consisting of broad sinuous subbasal transverse band not separated from base of elytra, indistinct latero-medial spot, narrow preapical transverse band interrupted on suture, and a small indistinct apical spot (
<figureCitation id="13182A57FFAFFF96F1229704FE9CBEC9" box="[264,329,1405,1429]" captionStart="Figs" captionStartId="3.[113,157,2051,2073]" captionTargetBox="[115,1452,190,2035]" captionTargetId="figure-10@3.[97,1463,180,2039]" captionTargetPageId="3" captionText="Figs. 1–9. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385200" httpUri="https://zenodo.org/record/5385200/files/figure.png" pageId="10" pageNumber="319">Fig. 7</figureCitation>
). Appendages and ventral part testaceous.
</paragraph>
<paragraph id="8B9C36D2FFAFFF96F0A297C4FE9DB369" blockId="10.[136,776,1469,2101]" pageId="10" pageNumber="319">Surface structures and sculpture: Head reticulation simple, composed of large irregular polygonal meshes, which are moderately deeply impressed. Frons between eyes, and clypeus with several large punctures. Reticulation of pronotum same as that of head. Pronotum along fore and basal margins and in posterior angles with numerous large setigerous punctures. Reticulation of elytra simple, composed of large irregular polygonal meshes similar to that of head or pronotum, but slightly more impressed. Meshes mostly complete, inner space often with several (1–3) small punctures. Each elytron with indistinct, sutural, discal and lateral row of large serial setigerous punctures. Ventral part reticulated with fine elongated meshes. Prosternal apophysis lanceolate. Abdominal ventrites with numerous oblique grooves and several large setigerous punctures, lateral margin bordered. Apical ventrite rounded posteriorly, with large setigerous punctures and transverse grooves on disc. Male pro- and mesotarsomeres 1–3 weakly dilated with bundle of adhesive setae on ventral side. Claws of pro- and mesotarsi simple and equal.</paragraph>
<paragraph id="8B9C36D2FFAFFF96F36992C4FC60BBA9" blockId="10.[835,1474,189,245]" pageId="10" pageNumber="319">
Male genitalia: Median lobe of aedeagus in lateral view as in
<figureCitation id="13182A57FFAFFF96F34B92A4FC67BBA9" box="[865,946,221,245]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="10" pageNumber="319">Fig. 24</figureCitation>
.
</paragraph>
<paragraph id="8B9C36D2FFAFFF96F3699364FC6BBA09" blockId="10.[835,1474,285,341]" pageId="10" pageNumber="319">Female genitalia: Similar to male; pro- and mesotarsomeres not dilated.</paragraph>
<paragraph id="8B9C36D2FFAFFF96F3699304FBF8BAA9" blockId="10.[835,1475,381,501]" pageId="10" pageNumber="319">
<emphasis id="B957EAC0FFAFFF96F3699304FC12BAC9" bold="true" box="[835,967,381,405]" pageId="10" pageNumber="319">Variability.</emphasis>
Specimens of the
<typeStatus id="54988870FFAFFF96F4B89307FB15BACA" box="[1170,1216,382,406]" pageId="10" pageNumber="319">type</typeStatus>
series vary in extent of elytral colour pattern, especially of the subbasal testaceous band, the major part of which is separated in some specimens from the elytral base.
</paragraph>
<paragraph id="8B9C36D2FFAFFF96F3699067FAC3B929" blockId="10.[835,1473,541,629]" pageId="10" pageNumber="319">
Measurements (N= 13). Body length
<quantity id="4CDB9B37FFAFFF96F4C79064FAA1B969" box="[1261,1396,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="10" pageNumber="319" unit="mm" value="4.25" valueMax="4.4" valueMin="4.1">4.1–4.4 mm</quantity>
(mean value =
<quantity id="4CDB9B37FFAFFF96F38F9044FC2AB909" box="[933,1023,573,597]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="10" pageNumber="319" unit="mm" value="4.3">4.3 mm</quantity>
,
<typeStatus id="54988870FFAFFF96F4259044FBA5B909" box="[1039,1136,573,597]" pageId="10" pageNumber="319" type="holotype">holotype</typeStatus>
<quantity id="4CDB9B37FFAFFF96F4519044FB0DB909" box="[1147,1240,573,597]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="10" pageNumber="319" unit="mm" value="4.1">4.1 mm</quantity>
); width
<quantity id="4CDB9B37FFAFFF96F5109044FA14B909" box="[1338,1473,573,597]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="10" pageNumber="319" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3–2.5 mm</quantity>
(mean value =
<quantity id="4CDB9B37FFAFFF96F3C39024FBEAB929" box="[1001,1087,605,629]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="319" unit="mm" value="2.4">2.4 mm</quantity>
,
<typeStatus id="54988870FFAFFF96F4669024FB7CB929" box="[1100,1193,605,629]" pageId="10" pageNumber="319" type="holotype">holotype</typeStatus>
<quantity id="4CDB9B37FFAFFF96F4989024FADFB929" box="[1202,1290,605,629]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="319" unit="mm" value="2.3">2.3 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAFFF96F36990E4FB5CB9A9" pageId="10" pageNumber="319" type="etymology">
<paragraph id="8B9C36D2FFAFFF96F36990E4FB5CB9A9" blockId="10.[835,1474,669,757]" pageId="10" pageNumber="319">
<emphasis id="B957EAC0FFAFFF96F36990E4FC10B9E9" bold="true" box="[835,965,669,693]" pageId="10" pageNumber="319">Etymology.</emphasis>
The new species is dedicated to its collector, our colleague Manfred A. Jäch (NHMW), a specialist on
<taxonomicName id="4C234D51FFAFFF96F36990A4FC1EB9A9" authorityName="Mulsant" authorityYear="1844" box="[835,971,733,757]" class="Insecta" family="Hydraenidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="319" phylum="Arthropoda" rank="family">Hydraenidae</taxonomicName>
and Dryopoidea.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAFFF96F3699164FA7CB809" pageId="10" pageNumber="319" type="distribution">
<paragraph id="8B9C36D2FFAFFF96F3699164FA7CB809" blockId="10.[835,1474,797,853]" pageId="10" pageNumber="319">
<emphasis id="B957EAC0FFAFFF96F3699164FC03B869" bold="true" box="[835,982,797,821]" pageId="10" pageNumber="319">Distribution.</emphasis>
So far known only from a small area of southeastern
<collectingCountry id="F3347642FFAFFF96F3BD9144FC23B809" box="[919,1014,829,853]" name="Thailand" pageId="10" pageNumber="319">Thailand</collectingCountry>
and south-western
<collectingCountry id="F3347642FFAFFF96F4E69144FAEFB809" box="[1228,1338,829,853]" name="Cambodia" pageId="10" pageNumber="319">Cambodia</collectingCountry>
(
<figureCitation id="13182A57FFAFFF96F5619144FA48B809" box="[1355,1437,829,853]" captionStart="Figs" captionStartId="9.[113,157,2093,2115]" captionTargetBox="[141,1427,190,2077]" captionTargetPageId="9" captionText="Figs. 40–41. Distribution of the species of the Laccophilus javanicus group" figureDoi="http://doi.org/10.5281/zenodo.5385208" httpUri="https://zenodo.org/record/5385208/files/figure.png" pageId="10" pageNumber="319">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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