226 lines
38 KiB
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226 lines
38 KiB
XML
<document id="D5AEB4490C9AA4CDCAB069619FD63D19" ID-DOI="10.11646/zootaxa.5027.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5448088" ID-ZooBank="D8FE51C2-C34F-47CE-87FC-80BAB16C54BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630464300981" checkinUser="plazi" docAuthor="Nazaré-Silva, Everton E. & Silva, Fernando A. B." docDate="2021" docId="038A87A7FFFFFFAD71F5FA5CB8C71980" docLanguage="en" docName="zootaxa.5027.1.3.pdf" docOrigin="Zootaxa 5027 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudocanthon Bates 1887" docType="key" docVersion="3" lastPageNumber="63" masterDocId="FFB3FFDFFFFDFFAF7162FFFBBD5A1E57" masterDocTitle="A taxonomic revision of the South American species of Pseudocanthon Bates, 1887 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)" masterLastPageNumber="86" masterPageNumber="61" pageNumber="63" updateTime="1699280357525" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C89FC2591B1CD7D390888CDBB1468D75">A taxonomic revision of the South American species of Pseudocanthon Bates, 1887 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:namePart id="BB42C7CA8E519311282FED455BBE6439">Nazaré-Silva, Everton E.</mods:namePart>
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<mods:affiliation id="2E61CD1067BD8F3270A70AD3EADC721A">Universidade Federal do Pará, Instituto de Ciências Biológicas. Programa de Pós-graduação em Zoologia da Universidade Federal do Pará e Museu Paraense Emílio Goeldi. Campus Belém. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil.</mods:affiliation>
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<mods:namePart id="A362F3F07D9EDA806D52107CCD5FF15B">Silva, Fernando A. B.</mods:namePart>
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<mods:affiliation id="4BAAE7183063DB92C0FAEE6D3995D874">Universidade Federal Rural de Pernambuco, Unidade Acadêmica de Serra Talhada-UAST. Av. Gregório Ferraz Nogueira. Mailbox 063. Serra Talhada, PE 56909 - 535, Brazil.</mods:affiliation>
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<mods:title id="39E8B0A2D0085878FCA6DBB695CAF4E4">Zootaxa</mods:title>
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<mods:date id="11448B596780914A6F97DF6186061DE8">2021</mods:date>
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<paragraph id="8B9C36B1FFFFFFAD71F5FA5CBE9A1B97" blockId="2.[151,960,1446,1473]" box="[151,960,1446,1473]" pageId="2" pageNumber="63">
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<heading id="D0D481DDFFFFFFAD71F5FA5CBE9A1B97" bold="true" box="[151,960,1446,1473]" fontSize="11" level="1" pageId="2" pageNumber="63" reason="1">
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<emphasis id="B957EAA3FFFFFFAD71F5FA5CBE9A1B97" bold="true" box="[151,960,1446,1473]" pageId="2" pageNumber="63">
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Key to the South American species of
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<taxonomicName id="4C234D32FFFFFFAD730DFA5DBE9A1B97" ID-CoL="642XK" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[623,960,1446,1473]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAA3FFFFFFAD730DFA5DBE701B96" bold="true" box="[623,810,1446,1473]" italics="true" pageId="2" pageNumber="63">Pseudocanthon</emphasis>
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<bibRefCitation id="EFB24B40FFFFFFAD7250FA5CBE9A1B97" author="Bates, H. W." box="[818,960,1446,1473]" pageId="2" pageNumber="63" refId="ref18875" refString="Bates, H. W. (1887) Tribe Lamellicornia. In: Bates, H. W. (1886 - 1889) Biologia Centrali-Americana. Insecta. Coleoptera. Volume II. Part 2. Pectinicornia and Lamellicornia. Taylor and Francis, London, United Kingdom, pp. 25 - 64, pls. 2 - 3. [publication date following Bousquet (2016)]" type="journal volume" year="1887">Bates, 1887</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C339653AFFFFFFAD71F5FA17B8C71980" pageId="2" pageNumber="63" type="key">
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<keyStep id="30D72E14FFFFFFAD71F5FA17B8C61849" pageId="2" pageNumber="63">
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<paragraph id="8B9C36B1FFFFFFAD71F5FA17B8C61856" blockId="2.[151,1437,1516,2007]" box="[151,1436,1516,1538]" pageId="2" pageNumber="63">
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<keyLead id="30D29584FFFFFFAD71F5FA17B8C61856" box="[151,1436,1516,1538]" pageId="2" pageNumber="63">
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||
1. Pronotum centrally with dense, granulate, and well-impressed microsculpture (
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||
<figureCitation id="13182A34FFFFFFAD72BAFA17B9791855" box="[984,1059,1516,1538]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2A–D); 0.2 mm (2E–H)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="2" pageNumber="63">Fig. 2A</figureCitation>
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)............................. 2
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</keyLead>
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</paragraph>
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<paragraph id="8B9C36B1FFFFFFAD71F5F9F2B8C61849" blockId="2.[151,1437,1516,2007]" box="[151,1436,1545,1567]" pageId="2" pageNumber="63">
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<keyLead id="30D29584FFFFFFAD71F5F9F2B8C61849" box="[151,1436,1545,1567]" pageId="2" pageNumber="63">
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<emphasis id="B957EAA3FFFFFFAD71F5F9F2BDC51849" bold="true" box="[151,159,1545,1566]" pageId="2" pageNumber="63">-</emphasis>
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Pronotum centrally with inconspicuous microsculpture (
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<figureCitation id="13182A34FFFFFFAD739CF9F2BE1D1848" box="[766,839,1545,1567]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2A–D); 0.2 mm (2E–H)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="2" pageNumber="63">Fig. 2B</figureCitation>
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)................................................ 4
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30D72E14FFFFFFAD71F5F9DDB8C618E6" pageId="2" pageNumber="63">
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<paragraph id="8B9C36B1FFFFFFAD71F5F9DDB8C71820" blockId="2.[151,1437,1516,2007]" pageId="2" pageNumber="63">
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<keyLead id="30D29584FFFFFFAD71F5F9DDB8C71820" pageId="2" pageNumber="63">
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2. Eighth elytral stria absent (
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||
<figureCitation id="13182A34FFFFFFAD7080F9DDBF71186B" box="[482,555,1574,1596]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2A–D); 0.2 mm (2E–H)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="2" pageNumber="63">Fig. 2C</figureCitation>
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) or reduced to basal one-fourth of elytra. Posterior edge of metafemora regularly curved, not projected towards the abdominal ventrites (
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||
<figureCitation id="13182A34FFFFFFAD73E0F9BFBF94180D" box="[642,718,1604,1626]" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="2" pageNumber="63">Fig. 4C</figureCitation>
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).
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<collectingCountry id="F3347621FFFFFFAD73BCF9BFBE43180D" box="[734,793,1604,1626]" name="Brazil" pageId="2" pageNumber="63">Brazil</collectingCountry>
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(
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<collectingRegion id="49E7F853FFFFFFAD7244F9BFBEFF180D" box="[806,933,1604,1626]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="63">Mato Grosso</collectingRegion>
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) and
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<collectingCountry id="F3347621FFFFFFAD72BBF9BFB963180D" box="[985,1081,1604,1626]" name="Argentina" pageId="2" pageNumber="63">Argentina</collectingCountry>
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...................................................................................
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<emphasis id="B957EAA3FFFFFFAD721AF99BB85D1820" bold="true" box="[888,1287,1632,1655]" pageId="2" pageNumber="63">
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<taxonomicName id="4C234D32FFFFFFAD721AF99BB9D11821" ID-CoL="8NCCK" authority="Nazaré-Silva & Silva, 2021" authorityName="Nazaré-Silva & Silva" authorityYear="2021" box="[888,1163,1632,1654]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="vazdemelloi" status="sp. nov.">
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<emphasis id="B957EAA3FFFFFFAD721AF99BB9D11821" bold="true" box="[888,1163,1632,1654]" italics="true" pageId="2" pageNumber="63">Pseudocanthon vazdemelloi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26457D8FFFFFFAD75F3F99AB85D1820" box="[1169,1287,1633,1655]" pageId="2" pageNumber="63" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="13182A34FFFFFFAD7476F99AB8321821" box="[1300,1384,1633,1655]" captionStart="FIGURE 3" captionStartId="12.[151,250,1927,1951]" captionTargetBox="[292,1300,191,1893]" captionTargetId="figure-30@12.[280,1307,181,1903]" captionTargetPageId="12" captionText="FIGURE 3. South American species of Pseudocanthon Bates, 1887, dorsal view. A, Pseudocanthon perplexus (LeConte, 1847); B–C, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448094" httpUri="https://zenodo.org/record/5448094/files/figure.png" pageId="2" pageNumber="63">Figs. 3C</figureCitation>
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,
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<figureCitation id="13182A34FFFFFFAD7410F99AB8C81820" box="[1394,1426,1633,1655]" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="2" pageNumber="63">4C</figureCitation>
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).
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</keyLead>
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</paragraph>
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<paragraph id="8B9C36B1FFFFFFAD71F5F985B8C618E6" blockId="2.[151,1437,1516,2007]" pageId="2" pageNumber="63">
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<keyLead id="30D29584FFFFFFAD71F5F985B8C618E6" pageId="2" pageNumber="63">
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<emphasis id="B957EAA3FFFFFFAD71F5F985BDC518C4" bold="true" box="[151,159,1662,1683]" pageId="2" pageNumber="63">-</emphasis>
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Eighth elytral stria present and extending across approximately the basal half of elytra (
|
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<figureCitation id="13182A34FFFFFFAD7546F985B92B18C3" box="[1060,1137,1662,1684]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2A–D); 0.2 mm (2E–H)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="2" pageNumber="63">Fig. 2D</figureCitation>
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). Posterior edge of metafemora slightly angulated at basal third, with expansion towards the abdominal ventrites (
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<figureCitation id="13182A34FFFFFFAD7296F967B90018E5" box="[1012,1114,1692,1714]" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="2" pageNumber="63">Fig. 4D–E</figureCitation>
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)......................... 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30D72E14FFFFFFAD71F5F942B8C71913" pageId="2" pageNumber="63">
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<paragraph id="8B9C36B1FFFFFFAD71F5F942BCB4195D" blockId="2.[151,1437,1516,2007]" pageId="2" pageNumber="63">
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<keyLead id="30D29584FFFFFFAD71F5F942BCB4195D" pageId="2" pageNumber="63">
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<materialsCitation id="3B4B3CECFFFFFFAD71F5F942BCB0195D" country="Brazil" location="Mato Grosso" pageId="2" pageNumber="63" specimenCount="1" stateProvince="Mato Grosso">
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3. Pygidium metallic green, with orange band at posterior edge (
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<figureCitation id="13182A34FFFFFFAD7228F942BEC01898" box="[842,922,1721,1743]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2A–D); 0.2 mm (2E–H)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="2" pageNumber="63">Fig. 2G</figureCitation>
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). Apex of parameres pointed in lateral view, curved towards the ventral edge of aedeagus (
|
||
<figureCitation id="13182A34FFFFFFAD7330F92DBFC718BB" box="[594,669,1750,1772]" captionStart="FIGURE 5" captionStartId="16.[151,250,1919,1943]" captionTargetBox="[175,1410,894,1887]" captionTargetId="figure-346@16.[164,1422,885,1896]" captionTargetPageId="16" captionText="FIGURE 5. Aedeagus in South American species of Pseudocanthon Bates, 1887, lateral and ventral views, respectively. A, Pseudocanthon perplexus (LeConte, 1847); B–C, P. xanthurus (Blanchard, 1847) (arrows indicate variation at apex of parameres); D. P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5448098" httpUri="https://zenodo.org/record/5448098/files/figure.png" pageId="2" pageNumber="63">Fig. 5E</figureCitation>
|
||
).
|
||
<collectingCountry id="F3347621FFFFFFAD73CCF92DBFB318BB" box="[686,745,1750,1772]" name="Brazil" pageId="2" pageNumber="63">Brazil</collectingCountry>
|
||
(
|
||
<collectingRegion id="49E7F853FFFFFFAD7397F92DBE2918BC" box="[757,883,1750,1771]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="63">Mato Grosso</collectingRegion>
|
||
and
|
||
<collectingRegion id="49E7F853FFFFFFAD72C0F92CB93918BB" box="[930,1123,1750,1772]" country="Brazil" name="Mato Grosso do Sul" pageId="2" pageNumber="63">Mato Grosso do Sul</collectingRegion>
|
||
)..
|
||
<emphasis id="B957EAA3FFFFFFAD75E4F92DBC02195E" bold="true" pageId="2" pageNumber="63">
|
||
<taxonomicName id="4C234D32FFFFFFAD75E4F92DB8C718BC" ID-CoL="8NCCB" authority="Nazaré-Silva & Silva, 2021" authorityName="Nazaré-Silva & Silva" authorityYear="2021" box="[1158,1437,1750,1772]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="pantanensis" status="sp. nov.">
|
||
<emphasis id="B957EAA3FFFFFFAD75E4F92DB8C718BC" bold="true" box="[1158,1437,1750,1772]" italics="true" pageId="2" pageNumber="63">Pseudocanthon pantanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26457D8FFFFFFAD7180F90FBC02195E" box="[226,344,1779,1801]" pageId="2" pageNumber="63" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13182A34FFFFFFAD7007F90FBCE2195E" box="[357,440,1780,1802]" captionStart="FIGURE 3" captionStartId="12.[151,250,1927,1951]" captionTargetBox="[292,1300,191,1893]" captionTargetId="figure-30@12.[280,1307,181,1903]" captionTargetPageId="12" captionText="FIGURE 3. South American species of Pseudocanthon Bates, 1887, dorsal view. A, Pseudocanthon perplexus (LeConte, 1847); B–C, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448094" httpUri="https://zenodo.org/record/5448094/files/figure.png" pageId="2" pageNumber="63">Figs. 3E</figureCitation>
|
||
,
|
||
<figureCitation id="13182A34FFFFFFAD70A0F90FBCB8195D" box="[450,482,1780,1802]" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="2" pageNumber="63">4D</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9C36B1FFFFFFAD71F5F8EAB8C71913" blockId="2.[151,1437,1516,2007]" pageId="2" pageNumber="63">
|
||
<keyLead id="30D29584FFFFFFAD71F5F8EAB8C71913" pageId="2" pageNumber="63">
|
||
<emphasis id="B957EAA3FFFFFFAD71F5F8EABDC51971" bold="true" box="[151,159,1809,1830]" pageId="2" pageNumber="63">-</emphasis>
|
||
Pygidium completely orange (
|
||
<figureCitation id="13182A34FFFFFFAD7361F8EABF0A1970" box="[515,592,1809,1831]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2A–D); 0.2 mm (2E–H)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="2" pageNumber="63">Fig. 2H</figureCitation>
|
||
). Apex of parameres truncate, with small and obtuse projection at ventral edge (
|
||
<figureCitation id="13182A34FFFFFFAD742BF8EAB8CB1970" box="[1353,1425,1809,1831]" captionStart="FIGURE 5" captionStartId="16.[151,250,1919,1943]" captionTargetBox="[175,1410,894,1887]" captionTargetId="figure-346@16.[164,1422,885,1896]" captionTargetPageId="16" captionText="FIGURE 5. Aedeagus in South American species of Pseudocanthon Bates, 1887, lateral and ventral views, respectively. A, Pseudocanthon perplexus (LeConte, 1847); B–C, P. xanthurus (Blanchard, 1847) (arrows indicate variation at apex of parameres); D. P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5448098" httpUri="https://zenodo.org/record/5448098/files/figure.png" pageId="2" pageNumber="63">Fig. 5F</figureCitation>
|
||
).
|
||
<collectingCountry id="F3347621FFFFFFAD7180F8D5BC411913" box="[226,283,1838,1860]" name="Brazil" pageId="2" pageNumber="63">Brasil</collectingCountry>
|
||
(
|
||
<collectingRegion id="49E7F853FFFFFFAD704AF8D5BCFD1913" box="[296,423,1838,1860]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="63">Mato Grosso</collectingRegion>
|
||
) and
|
||
<collectingCountry id="F3347621FFFFFFAD70B9F8D5BF611913" box="[475,571,1838,1860]" name="Argentina" pageId="2" pageNumber="63">Argentina</collectingCountry>
|
||
............................
|
||
<emphasis id="B957EAA3FFFFFFAD72EBF8D5B8491913" bold="true" box="[905,1299,1838,1860]" pageId="2" pageNumber="63">
|
||
<taxonomicName id="4C234D32FFFFFFAD72EBF8D5B9CD1913" ID-CoL="8NCC9" authority="Nazaré-Silva & Silva, 2021" authorityName="Nazaré-Silva & Silva" authorityYear="2021" box="[905,1175,1838,1860]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="chaquensis" status="sp. nov.">
|
||
<emphasis id="B957EAA3FFFFFFAD72EBF8D5B9CD1913" bold="true" box="[905,1175,1838,1860]" italics="true" pageId="2" pageNumber="63">Pseudocanthon chaquensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26457D8FFFFFFAD75FFF8D5B8491913" box="[1181,1299,1838,1860]" pageId="2" pageNumber="63" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13182A34FFFFFFAD747DF8D5B8301914" box="[1311,1386,1838,1860]" captionStart="FIGURE 3" captionStartId="12.[151,250,1927,1951]" captionTargetBox="[292,1300,191,1893]" captionTargetId="figure-30@12.[280,1307,181,1903]" captionTargetPageId="12" captionText="FIGURE 3. South American species of Pseudocanthon Bates, 1887, dorsal view. A, Pseudocanthon perplexus (LeConte, 1847); B–C, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448094" httpUri="https://zenodo.org/record/5448094/files/figure.png" pageId="2" pageNumber="63">Figs 3F</figureCitation>
|
||
,
|
||
<figureCitation id="13182A34FFFFFFAD7416F8D5B8C81913" box="[1396,1426,1838,1860]" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="2" pageNumber="63">4E</figureCitation>
|
||
).
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D72E14FFFFFFAD71F5F8B7B8C71980" pageId="2" pageNumber="63">
|
||
<paragraph id="8B9C36B1FFFFFFAD71F5F8B7B8C71928" blockId="2.[151,1437,1516,2007]" pageId="2" pageNumber="63">
|
||
<keyLead id="30D29584FFFFFFAD71F5F8B7B8C71928" pageId="2" pageNumber="63">
|
||
4. Dorsal surface of body almost completely black or metallic blue, orange portion restricted to the clypeal surface (
|
||
<figureCitation id="13182A34FFFFFFAD7420F8B7B8C81935" box="[1346,1426,1868,1890]" captionStart="FIGURE 3" captionStartId="12.[151,250,1927,1951]" captionTargetBox="[292,1300,191,1893]" captionTargetId="figure-30@12.[280,1307,181,1903]" captionTargetPageId="12" captionText="FIGURE 3. South American species of Pseudocanthon Bates, 1887, dorsal view. A, Pseudocanthon perplexus (LeConte, 1847); B–C, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448094" httpUri="https://zenodo.org/record/5448094/files/figure.png" pageId="2" pageNumber="63">Fig. 3A</figureCitation>
|
||
). Internal edge of protibia strongly arched (
|
||
<figureCitation id="13182A34FFFFFFAD7310F892BFE71928" box="[626,701,1897,1919]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2A–D); 0.2 mm (2E–H)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="2" pageNumber="63">Fig. 2E</figureCitation>
|
||
).
|
||
<collectingCountry id="F3347621FFFFFFAD73AFF892BE6A1928" box="[717,816,1897,1919]" name="Venezuela" pageId="2" pageNumber="63">Venezuela</collectingCountry>
|
||
...
|
||
<taxonomicName id="4C234D32FFFFFFAD7203F893B8511928" ID-CoL="4NQPK" authority="(LeConte, 1847)" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[865,1291,1896,1919]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="perplexus">
|
||
<emphasis id="B957EAA3FFFFFFAD7203F893B8511928" bold="true" box="[865,1291,1896,1919]" pageId="2" pageNumber="63">
|
||
<emphasis id="B957EAA3FFFFFFAD7203F893B93A1929" bold="true" box="[865,1120,1896,1918]" italics="true" pageId="2" pageNumber="63">Pseudocanthon perplexus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB24B40FFFFFFAD750FF892B8591928" author="LeConte, J." box="[1133,1283,1897,1919]" pageId="2" pageNumber="63" pagination="71 - 93" refId="ref20977" refString="LeConte, J. (1847) Fragmenta entomologica. Journal of the Academy of Natural Sciences of Philadelphia, 2, 71 - 93." type="journal article" year="1847">LeConte, 1847</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13182A34FFFFFFAD7475F892B83D1929" box="[1303,1383,1897,1919]" captionStart="FIGURE 3" captionStartId="12.[151,250,1927,1951]" captionTargetBox="[292,1300,191,1893]" captionTargetId="figure-30@12.[280,1307,181,1903]" captionTargetPageId="12" captionText="FIGURE 3. South American species of Pseudocanthon Bates, 1887, dorsal view. A, Pseudocanthon perplexus (LeConte, 1847); B–C, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448094" httpUri="https://zenodo.org/record/5448094/files/figure.png" pageId="2" pageNumber="63">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="13182A34FFFFFFAD7413F892B8C81928" box="[1393,1426,1897,1919]" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="2" pageNumber="63">4A</figureCitation>
|
||
).
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9C36B1FFFFFFAD71F5F87CB8C71980" blockId="2.[151,1437,1516,2007]" pageId="2" pageNumber="63">
|
||
<keyLead id="30D29584FFFFFFAD71F5F87CB8C71980" pageId="2" pageNumber="63">
|
||
- Surface of clypeus, lateral edges of pronotum, humeral angle, lateral and apex of elytra with orange spots (
|
||
<figureCitation id="13182A34FFFFFFAD75B8F87DB81B19CB" box="[1242,1345,1926,1948]" captionStart="FIGURE 3" captionStartId="12.[151,250,1927,1951]" captionTargetBox="[292,1300,191,1893]" captionTargetId="figure-30@12.[280,1307,181,1903]" captionTargetPageId="12" captionText="FIGURE 3. South American species of Pseudocanthon Bates, 1887, dorsal view. A, Pseudocanthon perplexus (LeConte, 1847); B–C, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448094" httpUri="https://zenodo.org/record/5448094/files/figure.png" pageId="2" pageNumber="63">Fig. 3B–C</figureCitation>
|
||
). Internal edge of protibia substraight (
|
||
<figureCitation id="13182A34FFFFFFAD7094F85FBF1A19ED" box="[502,576,1956,1978]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2A–D); 0.2 mm (2E–H)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="2" pageNumber="63">Fig. 2F</figureCitation>
|
||
). Widespread in South America..............................................................................................
|
||
<taxonomicName id="4C234D32FFFFFFAD7245F83BB9B11980" ID-CoL="4NQPN" authority="(Blanchard, 1847)" baseAuthorityName="Blanchard" baseAuthorityYear="1847" box="[807,1259,1984,2007]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="xanthurus">
|
||
<emphasis id="B957EAA3FFFFFFAD7245F83BB9B11980" bold="true" box="[807,1259,1984,2007]" pageId="2" pageNumber="63">
|
||
<emphasis id="B957EAA3FFFFFFAD7245F83BB9771981" bold="true" box="[807,1069,1984,2006]" italics="true" pageId="2" pageNumber="63">Pseudocanthon xanthurus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB24B40FFFFFFAD755BF83AB9BE1980" author="Blanchard, E." box="[1081,1252,1985,2007]" pageId="2" pageNumber="63" refId="ref19068" refString="Blanchard, E. (1847) livraison 79. In: Blanchard, E. & Brulle, G. A. (1835 - 1847) Voyage dans L'Amerique Meridionale (le Bresil, la Republique orientale de L'Uruguay, la Republique Argentine, la Patagonie, la Republique du Chili, la Republique de Bolivia, la Republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833, par Alcide d'Orbigny. Tome Sixieme. 2. e partie: Insectes. P. Bertrand, Paris, pp. 153 - 168. [publication date following Evenhuis (1997: 573)]" type="journal volume" year="1847">Blanchard, 1847</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13182A34FFFFFFAD759AF83AB8321981" box="[1272,1384,1985,2007]" captionStart="FIGURE 3" captionStartId="12.[151,250,1927,1951]" captionTargetBox="[292,1300,191,1893]" captionTargetId="figure-30@12.[280,1307,181,1903]" captionTargetPageId="12" captionText="FIGURE 3. South American species of Pseudocanthon Bates, 1887, dorsal view. A, Pseudocanthon perplexus (LeConte, 1847); B–C, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448094" httpUri="https://zenodo.org/record/5448094/files/figure.png" pageId="2" pageNumber="63">Figs. 3B–C</figureCitation>
|
||
,
|
||
<figureCitation id="13182A34FFFFFFAD7410F83AB8CB1980" box="[1394,1425,1985,2007]" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3A–E); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="2" pageNumber="63">4B</figureCitation>
|
||
).
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |