138 lines
15 KiB
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138 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="8831BC71A3445ADD8A6E49F4F37F8A27" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupinella sonlaensis Jirapatrasilp & Sutcharit & Panha 2022, comb. nov." docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1119</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>115</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
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<treatment ID-GBIF-Taxon="199839068" LSID="urn:lsid:plazi:treatment:8831BC71A3445ADD8A6E49F4F37F8A27" httpUri="http://treatment.plazi.org/id/8831BC71A3445ADD8A6E49F4F37F8A27" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="8831BC71-A344-5ADD-8A6E-49F4F37F8A27" authority="(Do, 2017)" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Do" baseAuthorityYear="2017" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella sonlaensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sonlaensis" status="comb. nov.">Pupinella sonlaensis (Do, 2017)</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">Figs 11H</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F)." figureDoi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1">, 12F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina sonlaensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sonlaensis" status="comb. nov.">Pupina sonlaensis</taxonomicName>
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Do, 2017: 300, 302, figs 2a, 3a. Type locality: limestone karst in Muong Bu Commune, Muong La District, Son La Province, Vietnam.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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HNUE-OC 00108 figured in
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<bibRefCitation author="Do, DS" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="299 - 303" refId="B33" refString="Do, DS, 2017. Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam. Raffles Bulletin of Zoology 65: 299 - 303" title="Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam." volume="65" year="2017">Do (2017</bibRefCitation>
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: figs 2a, 3a).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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ZRC.MOL.9377 (3 shells; Figs
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">11H</figureCitation>
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,
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F)." figureDoi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1">12F</figureCitation>
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) from the type locality.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell ovate-fusiform; last whorl ca. 60% of shell height. Apertural lip slightly expanded and reflected, thickened cord-like peristome absent; apertural lip when observed from lateral view almost straight. Parietal callus somewhat distinct and cord-like. Parietal tooth sharp with wide base, thickened and covering posterior canal. Anterior canal funnel-like, appearing as a slit on the inside, widened on outer margin, bordered by a thickened columellar margin. Umbilicus closed.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Do" baseAuthorityYear="2017" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella sonlaensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sonlaensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupinella sonlaensis</emphasis>
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</taxonomicName>
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is most similar to
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="mansuyi">
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<emphasis italics="true" pageId="0" pageNumber="1">P. mansuyi</emphasis>
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</taxonomicName>
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in shell size, but differs in having an ovate-fusiform shell shape with a less thickened parietal tooth, as well as a less thickened, expanded, and reflected apertural lip without a thickened cord-like inner peristome.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Muong La District, Thuan Chau District, and Van Ho District, Son La Province, Vietnam (
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<bibRefCitation author="Do, DS" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="299 - 303" refId="B33" refString="Do, DS, 2017. Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam. Raffles Bulletin of Zoology 65: 299 - 303" title="Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam." volume="65" year="2017">Do 2017</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This taxon is allocated to the genus
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<taxonomicName authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupinella</emphasis>
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</taxonomicName>
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due to the presence of a funnel-like anterior canal, which is the diagnostic character of this genus. The paratype figured in this study is similar to
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. mansuyi" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="mansuyi">
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<emphasis italics="true" pageId="0" pageNumber="1">P. mansuyi</emphasis>
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</taxonomicName>
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in having a triangular parietal tooth covering the posterior canal and an expanded and reflected apertural lip with somewhat cord-like inner peristome, although with less thickening, and the shell has a less elongate shape. However, the holotype of
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. sonlaensis" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="sonlaensis">
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<emphasis italics="true" pageId="0" pageNumber="1">P. sonlaensis</emphasis>
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</taxonomicName>
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figured in
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<bibRefCitation author="Do, DS" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="299 - 303" refId="B33" refString="Do, DS, 2017. Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam. Raffles Bulletin of Zoology 65: 299 - 303" title="Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam." volume="65" year="2017">Do (2017</bibRefCitation>
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: figs 2a, 3a) has an ovate-fusiform shell with a thickened, wide-based parietal tooth not covering the posterior canal, and a slightly expanded and reflected apertural lip without a thickened cord-like inner peristome. A thorough examination of the specimens would clarify whether the type series contain more than one taxon or whether the validity of this taxon should be reassessed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |