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<document id="1C96747413715CBCE2DDC03C63D5A9AF" ID-DOI="10.11606/1807-0205/2020.60.57" ID-ISSN="1807-0205" ID-Zenodo-Dep="4637241" ID-ZooBank="25B25A8B-CED0-4284-A799-5383998BE481" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616691998284" checkinUser="carolina" docAuthor="Simone, Luiz Ricardo L., Belz, Carlos Eduardo &amp; Gernet, Marcos de Vasconcellos" docDate="2020" docId="3B3A87CCFA02880917C45F63FD36FC07" docLanguage="en" docName="PapAvZoo.60.e20206057.pdf" docOrigin="Papéis Avulsos de Zoologia 60" docSource="http://dx.doi.org/10.11606/1807-0205/2020.60.57" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Drymaeus currais Simone &amp; Belz &amp; Gernet 2020, new species" docType="treatment" docUuid="D17AD2E5-D03C-447A-95E0-DFAB83F34F12" docUuidSource="ZooBank" docVersion="3" lastPageNumber="10" masterDocId="C703FFB4FA03880014675B79FFA2FFD4" masterDocTitle="A new species of Drymaeus endemic from Currais Archipelago, Paraná, Brazil (Pulmonata, Bulimulidae)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="2" updateTime="1698893482505" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
<mods:mods id="F50CFCE567A64D7A661D5024C592FF58" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D9D2D4BFD5AF00C7A3D1630F9603FB17">A new species of Drymaeus endemic from Currais Archipelago, Paraná, Brazil (Pulmonata, Bulimulidae)</mods:title>
</mods:titleInfo>
<mods:name id="EEBD9DFD15D0EDF8B8D5560DB7184B90" type="personal">
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<mods:namePart id="20CE221EF4A9A399DF56BA19FEE691B6">Simone, Luiz Ricardo L.</mods:namePart>
</mods:name>
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<mods:role id="DCBCCE35A7C538D0B2C5B6A03610854E">
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<mods:namePart id="C665A5C54437BFE3C1497348E290AEFF">Belz, Carlos Eduardo</mods:namePart>
</mods:name>
<mods:name id="C325C25212F980C741717B9C0EFF86AC" type="personal">
<mods:role id="4E1958A8B787EB5414516139B2265FDD">
<mods:roleTerm id="3943065D6E848D50FEEDE84D23218511">Author</mods:roleTerm>
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<mods:namePart id="DA497CCA13430DEF340BAF34EF12ACB5">Gernet, Marcos de Vasconcellos</mods:namePart>
</mods:name>
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<mods:title id="C4257AD5EFBB74CCBD75EF67E2D8CC2C">Papéis Avulsos de Zoologia</mods:title>
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<mods:part id="4A26F598CBE5F54BA6818DA2C1381F13">
<mods:date id="B4A5A692A7AC4E47B6C4465422B5C4E6">2020</mods:date>
<mods:detail id="7984D9E5D2F5F28AC032CDE9083E82FA" type="pubDate">
<mods:number id="F865A028B170851E7647AA872B244C3C">2020-11-16</mods:number>
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<mods:detail id="0559CDF24E9DF6F9EE59D9B8DA4898AC" type="volume">
<mods:number id="A11E704C9DA500F69C433F69309078CA">60</mods:number>
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<mods:url id="AAD93A9B1B76DCBB7CF5C45C6F6BF782">http://dx.doi.org/10.11606/1807-0205/2020.60.57</mods:url>
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<mods:classification id="4945A3A1A94B4C9AEF123D6A726087BA">journal article</mods:classification>
<mods:identifier id="60A2E0439E24F26DC6995D2F3A1CC61F" type="DOI">10.11606/1807-0205/2020.60.57</mods:identifier>
<mods:identifier id="8D9EA734BE4E3D9B92D7B3E3EF5D84FB" type="ISSN">1807-0205</mods:identifier>
<mods:identifier id="5C1D833E13DC67BD1400C6EF1754C140" type="Zenodo-Dep">4637241</mods:identifier>
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<treatment id="3B3A87CCFA02880917C45F63FD36FC07" LSID="urn:lsid:zoobank.org:act:D17AD2E5-D03C-447A-95E0-DFAB83F34F12" httpUri="http://treatment.plazi.org/id/3B3A87CCFA02880917C45F63FD36FC07" lastPageId="9" lastPageNumber="10" pageId="1" pageNumber="2">
<subSubSection id="FB896551FA02880117C45F63FAA3FBE7" box="[931,1281,1050,1075]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="B32C36DAFA02880117C45F63FAA3FBE7" blockId="1.[871,1340,1050,1171]" box="[931,1281,1050,1075]" pageId="1" pageNumber="2">
<heading id="E86481B6FA02880117C45F63FAA3FBE7" box="[931,1281,1050,1075]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
<emphasis id="81E7EAC8FA02880117C45F63FAA3FBE7" bold="true" box="[931,1281,1050,1075]" pageId="1" pageNumber="2">
<taxonomicName id="74934D59FA02880117C45F63FBCFFBE7" ID-CoL="7R5GS" authority="Simone &amp; Belz &amp; Gernet, 2020" authorityName="Simone &amp; Belz &amp; Gernet" authorityYear="2020" box="[931,1133,1050,1075]" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="2" phylum="Mollusca" rank="species" species="currais" status="sp. nov.">
<emphasis id="81E7EAC8FA02880117C45F63FBCFFBE7" bold="true" box="[931,1133,1050,1075]" italics="true" pageId="1" pageNumber="2">Drymaeus currais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD457B3FA02880110155F63FAA3FBE7" box="[1138,1281,1050,1075]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FB896551FA02880110765F43FB37FAC7" pageId="1" pageNumber="2" type="description">
<paragraph id="B32C36DAFA02880110765F43FA9EFB47" blockId="1.[871,1340,1050,1171]" pageId="1" pageNumber="2">
<emphasis id="81E7EAC8FA02880110765F43FA9EFB47" bold="true" pageId="1" pageNumber="2">
(
<figureCitation id="2BA82A5FFA028801107F5F43FB28FB87" box="[1048,1162,1082,1107]" captionStart-0="Figure 1" captionStart-1="Figures2-8" captionStart-2="Figures 9-13" captionStart-3="Figures 14-22" captionStart-4="Figures 23-26" captionStart-5="Figures 27-31" captionStart-6="Figures 32-36" captionStartId-0="2.[172,231,688,713]" captionStartId-1="3.[92,160,2074,2099]" captionStartId-2="4.[170,238,2010,2035]" captionStartId-3="5.[94,162,2010,2035]" captionStartId-4="6.[170,238,2010,2035]" captionStartId-5="7.[94,162,2010,2035]" captionStartId-6="8.[170,238,2042,2067]" captionTargetBox-0="[171,794,205,658]" captionTargetBox-1="[94,1417,570,2044]" captionTargetBox-2="[170,1493,1054,1980]" captionTargetBox-3="[123,1389,204,1980]" captionTargetBox-4="[174,1489,662,1974]" captionTargetBox-5="[98,1414,568,1976]" captionTargetBox-6="[174,1493,657,2008]" captionTargetId-0="figure-813@2.[170,794,204,658]" captionTargetId-1="figure-178@3.[94,1417,570,2044]" captionTargetId-2="figure-469@4.[170,1493,1054,1980]" captionTargetId-3="figure-1@5.[123,1389,204,1980]" captionTargetId-4="figure-244@6.[170,1493,657,1980]" captionTargetId-5="figure-185@7.[94,1417,563,1980]" captionTargetId-6="figure-217@8.[170,1493,651,2012]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionText-0="Figure 1.Map of Paraná coastal area,showing the Currais archipelago location and its profile in highlight. Occurrence of specimens in the larger island (eastone).A living crawling specimen also shown (shell length ~20 mm)." captionText-1="Figures2-8. Drymaeuscurrais sp.nov.shell oftypes;(2-4)HolotypeMZSP150548(L29.6),frontal,rightand dorsalview.(5-8)ParatypesMZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm)." captionText-2="Figures 9-13. Drymaeus currais sp. nov. shell and anatomical details.(9) Protoconch of paratype MZSP 135683#3 (young specimen of 14.2 mm), arrow showing transition with teleoconch.Scale =1 mm.(10) Anterior region of mouth,ventral view,integument removed.Scale = 0.5 mm.(11) Duct of bursa sectioned longitudinally,showing4 spermatophoresinside.Scale =1 mm.(12)Isolatedspermatophore.Scale =1 mm.(13)Reno-pericardialarea,ventral view,ventral wallofkidney opened along its left edge and deflected right (upwards) to show inner tissue,ventral pericardial wall deflected left (downwards).Scale = 1 mm." captionText-3="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." captionText-4="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." captionText-5="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." captionText-6="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4637243" figureDoi-1="http://doi.org/10.5281/zenodo.4637245" figureDoi-2="http://doi.org/10.5281/zenodo.4637247" figureDoi-3="http://doi.org/10.5281/zenodo.4637251" figureDoi-4="http://doi.org/10.5281/zenodo.4637253" figureDoi-5="http://doi.org/10.5281/zenodo.4637255" figureDoi-6="http://doi.org/10.5281/zenodo.4637257" httpUri-0="https://zenodo.org/record/4637243/files/figure.png" httpUri-1="https://zenodo.org/record/4637245/files/figure.png" httpUri-2="https://zenodo.org/record/4637247/files/figure.png" httpUri-3="https://zenodo.org/record/4637251/files/figure.png" httpUri-4="https://zenodo.org/record/4637253/files/figure.png" httpUri-5="https://zenodo.org/record/4637255/files/figure.png" httpUri-6="https://zenodo.org/record/4637257/files/figure.png" pageId="1" pageNumber="2">Figs. 1‑36</figureCitation>
)
<uri id="C7023AD8FA02880110665F23FA9EFB47" pageId="1" pageNumber="2">
urn:lsid:zoobank. org:act:
<uuid id="C7350C0FFA02880117CC5F03FA9EFB47" box="[939,1340,1146,1171]" pageId="1" pageNumber="2">D17AD2E5-D03C-447A-95E0-DFAB83F34F12</uuid>
</uri>
</emphasis>
</paragraph>
<paragraph id="B32C36DAFA028801177D5FC3FB37FAC7" blockId="1.[794,1419,1210,1299]" pageId="1" pageNumber="2">
<emphasis id="81E7EAC8FA028801177D5FC3FCC4FB07" bold="true" box="[794,870,1210,1235]" pageId="1" pageNumber="2">Types:</emphasis>
<materialsCitation id="03FB3C87FA02880117155FC3FAA5FB07" box="[882,1287,1210,1235]" collectionCode="MZSP" pageId="1" pageNumber="2" specimenCode="MZSP 150548" specimenCount="1" typeStatus="holotype">
<typeStatus id="6C288878FA02880117155FC3FC7EFB07" box="[882,988,1210,1235]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
:
<specimenCode id="E3359EA1FA028801178C5FC3FB2BFB07" box="[1003,1161,1210,1235]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 150548</specimenCode>
(
<figureCitation id="2BA82A5FFA02880110FC5FC3FAA2FB07" box="[1179,1280,1210,1235]" captionStart="Figures2-8" captionStartId="3.[92,160,2074,2099]" captionTargetBox="[94,1417,570,2044]" captionTargetId="figure-178@3.[94,1417,570,2044]" captionTargetPageId="3" captionText="Figures2-8. Drymaeuscurrais sp.nov.shell oftypes;(2-4)HolotypeMZSP150548(L29.6),frontal,rightand dorsalview.(5-8)ParatypesMZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm)." figureDoi="http://doi.org/10.5281/zenodo.4637245" httpUri="https://zenodo.org/record/4637245/files/figure.png" pageId="1" pageNumber="2">Figs. 2-4</figureCitation>
)
</materialsCitation>
.
<typeStatus id="6C288878FA02880111705FC3FA24FB07" box="[1303,1414,1210,1235]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
: MZSP 135683 (19 dissected specimens), LEBIO 577,
<specimenCount id="A595FD53FA028801177D5F83FC04FAC7" box="[794,934,1274,1299]" pageId="1" pageNumber="2" type="generic">5 specimens</specimenCount>
, all from type locality.
</paragraph>
</subSubSection>
<subSubSection id="FB896551FA028801177D5E43FB70F987" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="B32C36DAFA028801177D5E43FA88FA47" blockId="1.[794,1419,1338,1427]" pageId="1" pageNumber="2">
<materialsCitation id="03FB3C87FA028801177D5E43FBD7FA87" box="[794,1141,1338,1363]" country="Brazil" location="Parana" pageId="1" pageNumber="2" specimenCount="1" stateProvince="Parana" typeStatus="holotype">
<emphasis id="81E7EAC8FA028801177D5E43FC62FA87" bold="true" box="[794,960,1338,1363]" pageId="1" pageNumber="2">
<typeStatus id="6C288878FA028801177D5E43FCF1FA87" box="[794,851,1338,1363]" pageId="1" pageNumber="2">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="CB84764AFA02880117A85E43FBAFFA87" box="[975,1037,1338,1363]" name="Brazil" pageId="1" pageNumber="2">Brazil</collectingCountry>
.
<collectingRegion id="7157F838FA02880110785E43FBD7FA87" box="[1055,1141,1338,1363]" country="Brazil" name="Parana" pageId="1" pageNumber="2">
<emphasis id="81E7EAC8FA02880110785E43FBD7FA87" bold="true" box="[1055,1141,1338,1363]" pageId="1" pageNumber="2">Paraná</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="81E7EAC8FA02880110125E43FBDBFA87" bold="true" box="[1141,1145,1338,1363]" pageId="1" pageNumber="2">;</emphasis>
<materialsCitation id="03FB3C87FA02880110EF5E43FA84FA47" collectorName="Belz &amp; Gernet &amp; Colley" country="Brazil" county="Pontal do Parana" latitude="-25.735922" location="Guapira Island" longLatPrecision="1" longitude="-48.36537" municipality="Currais Archipelago" pageId="1" pageNumber="2" specimenCount="1" stateProvince="Parana" typeStatus="holotype">
off
<collectingCounty id="5A4D4E56FA02880110D15E43FA24FA87" box="[1206,1414,1338,1363]" pageId="1" pageNumber="2">Pontal do Paraná</collectingCounty>
,
<collectingMunicipality id="5348ACA0FA028801177D5E23FBA4FAA7" box="[794,1030,1370,1395]" pageId="1" pageNumber="2">Currais Archipelago</collectingMunicipality>
,
<location id="B64C6001FA02880110785E23FB73FAA7" LSID="urn:lsid:plazi:treatment:3B3A87CCFA02880917C45F63FD36FC07:B64C6001FA02880110785E23FB73FAA7" box="[1055,1233,1370,1395]" country="Brazil" county="Pontal do Parana" latitude="-25.735922" longLatPrecision="1" longitude="-48.36537" municipality="Currais Archipelago" name="Guapira Island" pageId="1" pageNumber="2" stateProvince="Parana">Guapirá Island</location>
,
<geoCoordinate id="D6A7501DFA028801108C5E23FA25FAA7" box="[1259,1415,1369,1395]" degrees="25" direction="south" minutes="44" orientation="latitude" pageId="1" pageNumber="2" precision="1" seconds="09.32" value="-25.735922">25°44′09.32″S</geoCoordinate>
,
<geoCoordinate id="D6A7501DFA028801177D5E03FC1EFA47" box="[794,956,1401,1427]" degrees="48" direction="west" minutes="21" orientation="longitude" pageId="1" pageNumber="2" precision="1" seconds="55.34" value="-48.36537">48°21′55.34″W</geoCoordinate>
(
<collectorName id="1E66530CFA02880117AF5E03FC58FA47" box="[968,1018,1402,1427]" pageId="1" pageNumber="2">Belz</collectorName>
,
<collectorName id="1E66530CFA02880110655E03FBEDFA47" box="[1026,1103,1402,1427]" pageId="1" pageNumber="2">Gernet</collectorName>
&amp;
<collectorName id="1E66530CFA028801100D5E03FB0CFA47" box="[1130,1198,1402,1427]" pageId="1" pageNumber="2">Colley</collectorName>
col., 2015)
</materialsCitation>
.
</paragraph>
<paragraph id="B32C36DAFA028801177D5EC3FB70F987" blockId="1.[793,1419,1466,1619]" pageId="1" pageNumber="2">
<emphasis id="81E7EAC8FA028801177D5EC3FC3AFA07" bold="true" box="[794,920,1466,1491]" pageId="1" pageNumber="2">Additional</emphasis>
<materialsCitation id="03FB3C87FA02880117FB5EC3FA24FA27" collectingDate="1988-12-01" collectionCode="MZSP" pageId="1" pageNumber="2" specimenCode="MZSP 29634, 2" specimenCount="2">
<emphasis id="81E7EAC8FA02880117FB5EC3FCC3FA27" bold="true" pageId="1" pageNumber="2">
non‑type material examined from
<typeStatus id="6C288878FA02880111485EC3FAC7FA07" box="[1327,1381,1466,1491]" pageId="1" pageNumber="2">type</typeStatus>
locality:
</emphasis>
<specimenCode id="E3359EA1FA028801170F5EA3FC57FA27" box="[872,1013,1498,1523]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 29634</specimenCode>
,
<specimenCode id="E3359EA1FA02880117995EA3FBAEFA27" box="[1022,1036,1498,1523]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">2</specimenCode>
dissected specimens (
<date id="C72D101AFA02880111625EA3FA22FA27" box="[1285,1408,1498,1523]" pageId="1" pageNumber="2" value="1988-12-01">
<collectingDate id="D769E9F2FA02880111625EA3FA22FA27" box="[1285,1408,1498,1523]" pageId="1" pageNumber="2" value="1988-12-01">01.xii.1988</collectingDate>
</date>
)
</materialsCitation>
;
<materialsCitation id="03FB3C87FA028801177C5E83FB6CF987" collectingDate="2015-03-26" collectionCode="MZSP" pageId="1" pageNumber="2" specimenCode="MZSP 135684, MZSP 135685, MZSP 136031, MZSP 136032, MZSP 143544" specimenCount="26">
<specimenCode id="E3359EA1FA028801177C5E83FC16F9C7" box="[795,948,1530,1555]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 135684</specimenCode>
,
<specimenCount id="A595FD53FA02880117DC5E83FBE5F9C7" box="[955,1095,1530,1555]" pageId="1" pageNumber="2" type="generic">3 specimens</specimenCount>
,
<specimenCode id="E3359EA1FA02880110295E83FB45F9C7" box="[1102,1255,1530,1555]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 135685</specimenCode>
,
<specimenCount id="A595FD53FA02880110885E83FA2AF9C7" box="[1263,1416,1530,1555]" pageId="1" pageNumber="2" type="generic">13 specimens</specimenCount>
,
<specimenCode id="E3359EA1FA028801177E5D63FC16F9E7" box="[793,948,1562,1587]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 136031</specimenCode>
,
<specimenCount id="A595FD53FA02880117DA5D63FBE9F9E7" box="[957,1099,1562,1587]" pageId="1" pageNumber="2" type="generic">2 specimens</specimenCount>
,
<specimenCode id="E3359EA1FA02880110335D63FB4DF9E7" box="[1108,1263,1562,1587]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 136032</specimenCode>
,
<specimenCount id="A595FD53FA028801109E5D63FA24F9E7" box="[1273,1414,1562,1587]" pageId="1" pageNumber="2" type="generic">5 specimens</specimenCount>
,
<specimenCode id="E3359EA1FA028801177C5D43FC17F987" box="[795,949,1594,1619]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="2" type="Museum">MZSP 143544</specimenCode>
,
<specimenCount id="A595FD53FA02880117DA5D43FBE5F987" box="[957,1095,1594,1619]" pageId="1" pageNumber="2" type="generic">3 specimens</specimenCount>
(
<date id="C72D101AFA02880110345D43FB6AF987" box="[1107,1224,1594,1619]" pageId="1" pageNumber="2" value="2015-03-26">
<collectingDate id="D769E9F2FA02880110345D43FB6AF987" box="[1107,1224,1594,1619]" pageId="1" pageNumber="2" value="2015-03-26">26.iii.2015</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FB896551FA028801177D5D03FC05F907" pageId="1" pageNumber="2" type="etymology">
<paragraph id="B32C36DAFA028801177D5D03FC05F907" blockId="1.[794,1417,1658,1747]" pageId="1" pageNumber="2">
<emphasis id="81E7EAC8FA028801177D5D03FC03F947" bold="true" box="[794,929,1658,1683]" pageId="1" pageNumber="2">Etymology:</emphasis>
The epithet is in apposition, as a reference to the location of occurrence, derived from Currais Archipelago.
</paragraph>
</subSubSection>
<subSubSection id="FB896551FA028802177D5D83FD67FC27" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="B32C36DAFA028802177D5D83FD67FC27" blockId="1.[793,1420,1786,2131]" lastBlockId="2.[169,795,826,1011]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<emphasis id="81E7EAC8FA028801177D5D83FC34F8C7" bold="true" box="[794,918,1786,1811]" pageId="1" pageNumber="2">Diagnosis:</emphasis>
shell conical, aperture ample, weakly deflect- ed. Color white-cream, mostly with strong axial, dark brown spots; peri-umbilical area lacking spots; some rare specimens uniformly yellowish beige. Secondary ureter closed only in posterior 2/3, furrow-like in third preceding pneumostome. Pulmonary vases particularly dense, mainly on third preceding pneumostome. Anus external to pneumostome, opened directly outside. Aperture of salivary glands lateral in middle level of buccal cavity. Intrinsic pair of odontophore muscles m7 originating in cartilages close to m6; presence of odontophore pairs m1a and m3. Sigmoid intestinal loop relatively short. Penis slender and long, lacking clear inner chambers. Inner spermoduct of spermoviduct protected by tall fold. Seminal receptacle small, with additional elongated diverticulum called genital appendix. Albumen gland duct single, elongated. Pleural ganglia inconspicuous.
</paragraph>
</subSubSection>
<caption id="E7EC6652FA01880214CB59C9FCA4FCDD" ID-DOI="http://doi.org/10.5281/zenodo.4637243" ID-Zenodo-Dep="4637243" httpUri="https://zenodo.org/record/4637243/files/figure.png" pageId="2" pageNumber="3" startId="2.[172,231,688,713]" targetBox="[171,794,205,658]" targetPageId="2">
<paragraph id="B32C36DAFA01880214CB59C9FCA4FCDD" blockId="2.[170,794,688,777]" pageId="2" pageNumber="3">Figure 1.Map of Paraná coastal area,showing the Currais archipelago location and its profile in highlight. Occurrence of specimens in the larger island (east one).A living crawling specimen also shown (shell length ~20 mm).</paragraph>
</caption>
<subSubSection id="FB896551FA01880914CD5F63FDFCFDC7" lastPageId="9" lastPageNumber="10" pageId="2" pageNumber="3" type="description">
<paragraph id="B32C36DAFA01880214CD5F63FE96FBE7" blockId="2.[170,308,1050,1075]" box="[170,308,1050,1075]" pageId="2" pageNumber="3">
<emphasis id="81E7EAC8FA01880214CD5F63FE96FBE7" bold="true" box="[170,308,1050,1075]" pageId="2" pageNumber="3">Description</emphasis>
</paragraph>
<paragraph id="B32C36DAFA01880214CD5F23FDA0F907" blockId="2.[169,795,1114,1747]" pageId="2" pageNumber="3">
<emphasis id="81E7EAC8FA01880214CD5F23FF4EFBA7" bold="true" box="[170,236,1114,1139]" pageId="2" pageNumber="3">Shell:</emphasis>
(
<figureCitation id="2BA82A5FFA018802149F5F23FEF4FBA7" box="[248,342,1114,1139]" captionStart-0="Figure 1" captionStart-1="Figures2-8" captionStart-2="Figures 9-13" captionStartId-0="2.[172,231,688,713]" captionStartId-1="3.[92,160,2074,2099]" captionStartId-2="4.[170,238,2010,2035]" captionTargetBox-0="[171,794,205,658]" captionTargetBox-1="[94,1417,570,2044]" captionTargetBox-2="[170,1493,1054,1980]" captionTargetId-0="figure-813@2.[170,794,204,658]" captionTargetId-1="figure-178@3.[94,1417,570,2044]" captionTargetId-2="figure-469@4.[170,1493,1054,1980]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figure 1.Map of Paraná coastal area,showing the Currais archipelago location and its profile in highlight. Occurrence of specimens in the larger island (eastone).A living crawling specimen also shown (shell length ~20 mm)." captionText-1="Figures2-8. Drymaeuscurrais sp.nov.shell oftypes;(2-4)HolotypeMZSP150548(L29.6),frontal,rightand dorsalview.(5-8)ParatypesMZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm)." captionText-2="Figures 9-13. Drymaeus currais sp. nov. shell and anatomical details.(9) Protoconch of paratype MZSP 135683#3 (young specimen of 14.2 mm), arrow showing transition with teleoconch.Scale =1 mm.(10) Anterior region of mouth,ventral view,integument removed.Scale = 0.5 mm.(11) Duct of bursa sectioned longitudinally,showing4 spermatophoresinside.Scale =1 mm.(12)Isolatedspermatophore.Scale =1 mm.(13)Reno-pericardialarea,ventral view,ventral wallofkidney opened along its left edge and deflected right (upwards) to show inner tissue,ventral pericardial wall deflected left (downwards).Scale = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4637243" figureDoi-1="http://doi.org/10.5281/zenodo.4637245" figureDoi-2="http://doi.org/10.5281/zenodo.4637247" httpUri-0="https://zenodo.org/record/4637243/files/figure.png" httpUri-1="https://zenodo.org/record/4637245/files/figure.png" httpUri-2="https://zenodo.org/record/4637247/files/figure.png" pageId="2" pageNumber="3">Figs. 1-9</figureCitation>
) adult shell around
<quantity id="746B9B3FFA01880216535F23FD27FBA7" box="[564,645,1114,1139]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="3" unit="mm" value="35.0">35 mm</quantity>
, conical-oval; apex bluntly acuminated; greatest width on last whorl; width ~1/2 shell length. Basal color white to pale-cream; pattern of dark-brown spots arranged in irregular axial bands, slightly more concentrated in middle region of whorls; peri-umbilical area flanked by area lacking spots (
<figureCitation id="2BA82A5FFA01880214D65E63FEB3FAE7" box="[177,273,1306,1331]" captionStart="Figures2-8" captionStartId="3.[92,160,2074,2099]" captionTargetBox="[94,1417,570,2044]" captionTargetId="figure-178@3.[94,1417,570,2044]" captionTargetPageId="3" captionText="Figures2-8. Drymaeuscurrais sp.nov.shell oftypes;(2-4)HolotypeMZSP150548(L29.6),frontal,rightand dorsalview.(5-8)ParatypesMZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm)." figureDoi="http://doi.org/10.5281/zenodo.4637245" httpUri="https://zenodo.org/record/4637245/files/figure.png" pageId="2" pageNumber="3">Figs. 2-6</figureCitation>
); dark spots absent in ~10% of specimens, being purely yellowish-cream (
<figureCitation id="2BA82A5FFA01880215805E43FDEAFA87" box="[487,584,1338,1363]" captionStart="Figures2-8" captionStartId="3.[92,160,2074,2099]" captionTargetBox="[94,1417,570,2044]" captionTargetId="figure-178@3.[94,1417,570,2044]" captionTargetPageId="3" captionText="Figures2-8. Drymaeuscurrais sp.nov.shell oftypes;(2-4)HolotypeMZSP150548(L29.6),frontal,rightand dorsalview.(5-8)ParatypesMZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm)." figureDoi="http://doi.org/10.5281/zenodo.4637245" httpUri="https://zenodo.org/record/4637245/files/figure.png" pageId="2" pageNumber="3">Figs. 7-8</figureCitation>
). Spire angle ~55°. Protoconch of 2 whorls, ~6% of total shell length, uniformly sculptured by delicate reticulate pattern (
<figureCitation id="2BA82A5FFA01880216A85E03FCADFA47" box="[719,783,1402,1427]" captionStart="Figures 9-13" captionStartId="4.[170,238,2010,2035]" captionTargetBox="[170,1493,1054,1980]" captionTargetId="figure-469@4.[170,1493,1054,1980]" captionTargetPageId="4" captionText="Figures 9-13. Drymaeus currais sp. nov. shell and anatomical details.(9) Protoconch of paratype MZSP 135683#3 (young specimen of 14.2 mm), arrow showing transition with teleoconch.Scale =1 mm.(10) Anterior region of mouth,ventral view,integument removed.Scale = 0.5 mm.(11) Duct of bursa sectioned longitudinally,showing4 spermatophoresinside.Scale =1 mm.(12)Isolatedspermatophore.Scale =1 mm.(13)Reno-pericardialarea,ventral view,ventral wallofkidney opened along its left edge and deflected right (upwards) to show inner tissue,ventral pericardial wall deflected left (downwards).Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4637247" httpUri="https://zenodo.org/record/4637247/files/figure.png" pageId="2" pageNumber="3">Fig. 9</figureCitation>
), with ~25 spiral lines in last whorl; transition to teleoconch clear, slightly prosocline (
<figureCitation id="2BA82A5FFA01880216625EC3FDE0FA07" box="[517,578,1466,1491]" captionStart="Figures 9-13" captionStartId="4.[170,238,2010,2035]" captionTargetBox="[170,1493,1054,1980]" captionTargetId="figure-469@4.[170,1493,1054,1980]" captionTargetPageId="4" captionText="Figures 9-13. Drymaeus currais sp. nov. shell and anatomical details.(9) Protoconch of paratype MZSP 135683#3 (young specimen of 14.2 mm), arrow showing transition with teleoconch.Scale =1 mm.(10) Anterior region of mouth,ventral view,integument removed.Scale = 0.5 mm.(11) Duct of bursa sectioned longitudinally,showing4 spermatophoresinside.Scale =1 mm.(12)Isolatedspermatophore.Scale =1 mm.(13)Reno-pericardialarea,ventral view,ventral wallofkidney opened along its left edge and deflected right (upwards) to show inner tissue,ventral pericardial wall deflected left (downwards).Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4637247" httpUri="https://zenodo.org/record/4637247/files/figure.png" pageId="2" pageNumber="3">Fig. 9</figureCitation>
: arrow).Teleoconch smooth, except for growth lines, of ~4 whorls. Whorls profile slightly convex. Suture well-marked, slightly oblique (diagonal) to columellar axis. Aperture prosocline (~20° from longitudinal general axis) (
<figureCitation id="2BA82A5FFA01880216F45D43FD71F987" box="[659,723,1594,1619]" captionStart="Figures2-8" captionStartId="3.[92,160,2074,2099]" captionTargetBox="[94,1417,570,2044]" captionTargetId="figure-178@3.[94,1417,570,2044]" captionTargetPageId="3" captionText="Figures2-8. Drymaeuscurrais sp.nov.shell oftypes;(2-4)HolotypeMZSP150548(L29.6),frontal,rightand dorsalview.(5-8)ParatypesMZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm)." figureDoi="http://doi.org/10.5281/zenodo.4637245" httpUri="https://zenodo.org/record/4637245/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
), oval; ~50% of shell length, ~60% of shell width. Peristome slightly reflected, especially on columellar region, partially covering umbilicus (
<figureCitation id="2BA82A5FFA01880215AD5DE3FDE8F967" box="[458,586,1690,1715]" captionStart="Figures2-8" captionStartId="3.[92,160,2074,2099]" captionTargetBox="[94,1417,570,2044]" captionTargetId="figure-178@3.[94,1417,570,2044]" captionTargetPageId="3" captionText="Figures2-8. Drymaeuscurrais sp.nov.shell oftypes;(2-4)HolotypeMZSP150548(L29.6),frontal,rightand dorsalview.(5-8)ParatypesMZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm)." figureDoi="http://doi.org/10.5281/zenodo.4637245" httpUri="https://zenodo.org/record/4637245/files/figure.png" pageId="2" pageNumber="3">Figs. 2, 5, 7</figureCitation>
). Body whorl ~1/2 shell length. Umbilicus narrow.
</paragraph>
<paragraph id="B32C36DAFA01880214CD5D83FD02F8E7" blockId="2.[170,795,1786,1843]" pageId="2" pageNumber="3">
<emphasis id="81E7EAC8FA01880214CD5D83FE8BF8C7" bold="true" box="[170,297,1786,1811]" pageId="2" pageNumber="3">Head‑foot:</emphasis>
of usual shape. Color uniformly clear. Columellar muscle thick, 1.8 whorls in length.
</paragraph>
<paragraph id="B32C36DAFA01880214CD5C23FBBFFC67" blockId="2.[170,795,1882,2131]" lastBlockId="2.[869,1496,218,947]" pageId="2" pageNumber="3">
<emphasis id="81E7EAC8FA01880214CD5C23FEFDF8A7" bold="true" box="[170,351,1882,1907]" pageId="2" pageNumber="3">Mantle organs:</emphasis>
(
<figureCitation id="2BA82A5FFA018802150A5C23FE4BF8A7" box="[365,489,1882,1907]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Figs. 23-26</figureCitation>
) mantle border thick, lacking pigments. Pneumostome (pn) protected by ventral, right simple flap (mf), with ~1/5 of aperture length. Dorsal fold about double of ventral flap, ~1.5-times longer. Pneumostome (pn) ~1/9 of aperture length, bearing air entrance, urinary escape (
<figureCitation id="2BA82A5FFA01880215955C83FDE2F7C7" box="[498,576,2042,2067]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Fig. 24</figureCitation>
: ua) in left-anterior side (
<figureCitation id="2BA82A5FFA01880214825363FE90F7E7" box="[229,306,2074,2099]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Fig. 23</figureCitation>
: pn), and anus in right-posterior side, separated by transverse fold (
<figureCitation id="2BA82A5FFA01880215A55343FDADF787" box="[450,527,2106,2131]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Fig. 23</figureCitation>
: an). Lung of 1.5 whorls in length, ~tice longer than wide. Pulmonary vessels conspicuous all along right side (
<figureCitation id="2BA82A5FFA01880210845B83FA91FEC7" box="[1251,1331,250,275]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Fig. 24</figureCitation>
), mostly bearing transverse, rather perpendicular vessels, clustered, sometimes bifurcating or anastomosing; in left side of pulmonary vein (cv), visible vessels only in anterior third, occupying right ~half of this area, constituted by transverse vessels of similar fashion as right side, but inserting in longitudinal vessel (
<figureCitation id="2BA82A5FFA01880210035AC3FB10FE07" box="[1124,1202,442,467]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Fig. 24</figureCitation>
: vp), in such anterior end converge with anterior end of pulmonary vessel as part of collar vessel (
<figureCitation id="2BA82A5FFA01880210795A83FBCEFDC7" box="[1054,1132,506,531]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Fig. 26</figureCitation>
: co). Remaining regions of lung almost smooth, with imbricated vessels of difficult visualization. Pulmonary vein (cv) running longitudinally between middle and right thirds of pallial cavity roof, somewhat equidistant from rectum all along its length. Reno-pericardial area triangular, located posteriorly at middle level of posterior end, occupying ~25% of cavity length and ~70% of its width (details below). Rectum (rt) and ureter (ur) narrow, running along right edge. Urinary aperture very elongated (
<figureCitation id="2BA82A5FFA01880210F65863FB4DFCE7" box="[1169,1263,794,819]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Figs. 24</figureCitation>
,
<figureCitation id="2BA82A5FFA01880210995863FABEFCE7" box="[1278,1308,794,819]" captionStart="Figures 9-13" captionStartId="4.[170,238,2010,2035]" captionTargetBox="[170,1493,1054,1980]" captionTargetId="figure-469@4.[170,1493,1054,1980]" captionTargetPageId="4" captionText="Figures 9-13. Drymaeus currais sp. nov. shell and anatomical details.(9) Protoconch of paratype MZSP 135683#3 (young specimen of 14.2 mm), arrow showing transition with teleoconch.Scale =1 mm.(10) Anterior region of mouth,ventral view,integument removed.Scale = 0.5 mm.(11) Duct of bursa sectioned longitudinally,showing4 spermatophoresinside.Scale =1 mm.(12)Isolatedspermatophore.Scale =1 mm.(13)Reno-pericardialarea,ventral view,ventral wallofkidney opened along its left edge and deflected right (upwards) to show inner tissue,ventral pericardial wall deflected left (downwards).Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4637247" httpUri="https://zenodo.org/record/4637247/files/figure.png" pageId="2" pageNumber="3">13</figureCitation>
: ua), occupying ~1/3 of ureter length; its posterior end slightly round- ed, its anterior end just left from anus, T-shaped, being its left branch longer and anteriorly protected by special fold (
<figureCitation id="2BA82A5FFA01880217FA58E3FC4AFC67" box="[925,1000,922,947]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Fig. 26</figureCitation>
: pn).
</paragraph>
<paragraph id="B32C36DAFA018802170258A3FACDFA87" blockId="2.[869,1495,986,1363]" pageId="2" pageNumber="3">
<emphasis id="81E7EAC8FA018802170258A3FBAFFC27" bold="true" box="[869,1037,986,1011]" pageId="2" pageNumber="3">Visceral mass:</emphasis>
(
<figureCitation id="2BA82A5FFA018802107B58A3FBCBFC27" box="[1052,1129,986,1011]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Fig. 25</figureCitation>
) ~3.5 whorls in length. Both digestive gland lobes pale greenish beige in color. Anterior lobe (da) flattened,occupying~1/4 of visceral volume,located just posteriorly to pallial cavity, continuous to kidney. Posterior lobe (dg) with 2.5 spiral whorls, with ~45% of visceral volume. Stomach with ~1/8 of visceral volume, located between both digestive gland lobes, about one whorl posterior to pallial cavity (st). Digestive tubes (described below) surrounding anterior lobe of digestive gland. Gonad clearly multi-lobed, greenish-cream color, encased between posterior lobe of digestive gland and columella, occupying ~1/10 of visceral volume.
</paragraph>
<paragraph id="B32C36DAFA01880217025E03FBAFF867" blockId="2.[868,1495,1402,1971]" pageId="2" pageNumber="3">
<emphasis id="81E7EAC8FA01880217025E03FAAEFA47" bold="true" box="[869,1292,1402,1427]" pageId="2" pageNumber="3">Circulatory and excretory systems:</emphasis>
(
<figureCitation id="2BA82A5FFA018802117A5E03FADAFA47" box="[1309,1400,1402,1427]" captionStart="Figures 9-13" captionStartId="4.[170,238,2010,2035]" captionTargetBox="[170,1493,1054,1980]" captionTargetId="figure-469@4.[170,1493,1054,1980]" captionTargetPageId="4" captionText="Figures 9-13. Drymaeus currais sp. nov. shell and anatomical details.(9) Protoconch of paratype MZSP 135683#3 (young specimen of 14.2 mm), arrow showing transition with teleoconch.Scale =1 mm.(10) Anterior region of mouth,ventral view,integument removed.Scale = 0.5 mm.(11) Duct of bursa sectioned longitudinally,showing4 spermatophoresinside.Scale =1 mm.(12)Isolatedspermatophore.Scale =1 mm.(13)Reno-pericardialarea,ventral view,ventral wallofkidney opened along its left edge and deflected right (upwards) to show inner tissue,ventral pericardial wall deflected left (downwards).Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4637247" httpUri="https://zenodo.org/record/4637247/files/figure.png" pageId="2" pageNumber="3">Figs. 13</figureCitation>
,
<figureCitation id="2BA82A5FFA01880211E15E03FA6DFA47" box="[1414,1487,1402,1427]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">24, 25</figureCitation>
) Pericardium (pc) ~3-times as long as wide, located longitudinally between middle and left thirds of posterior end of pallial roof; occupying ~5% of lung area. Auricle (au) located anteriorly, as continuation from pulmonary vein (cv), slightly larger than ventricle (ve). Kidney (ki) simple, mostly solid, dorso-ventrally flattened; size reported above; somewhat triangular, width ~2/3 of length; inner lobe constituted by longitudinal,tall folds converging anteriorly to middle axis of structure (
<figureCitation id="2BA82A5FFA01880210885DE3FA9EF967" box="[1263,1340,1690,1715]" captionStart="Figures 9-13" captionStartId="4.[170,238,2010,2035]" captionTargetBox="[170,1493,1054,1980]" captionTargetId="figure-469@4.[170,1493,1054,1980]" captionTargetPageId="4" captionText="Figures 9-13. Drymaeus currais sp. nov. shell and anatomical details.(9) Protoconch of paratype MZSP 135683#3 (young specimen of 14.2 mm), arrow showing transition with teleoconch.Scale =1 mm.(10) Anterior region of mouth,ventral view,integument removed.Scale = 0.5 mm.(11) Duct of bursa sectioned longitudinally,showing4 spermatophoresinside.Scale =1 mm.(12)Isolatedspermatophore.Scale =1 mm.(13)Reno-pericardialarea,ventral view,ventral wallofkidney opened along its left edge and deflected right (upwards) to show inner tissue,ventral pericardial wall deflected left (downwards).Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4637247" httpUri="https://zenodo.org/record/4637247/files/figure.png" pageId="2" pageNumber="3">Fig. 13</figureCitation>
). Nephropore small, longitudinal slit in anterior-left corner,turned right (
<figureCitation id="2BA82A5FFA018802170D5DA3FC45F927" box="[874,999,1754,1779]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Figs. 24, 25</figureCitation>
: ne). Primary and secondary ureter complete and closed (tubular); primary ureter (up) lying on right edge of kidney towards posterior and right, after forming strong curve, running afterwards anteriorly, as secondary ureter (ur) along entire left edge of rectum, except for longitudinal urinary aperture of ~1/3 of its length (ua) (details above).
</paragraph>
<paragraph id="B32C36DAFA01880317025CA3FB3EFE67" blockId="2.[869,1494,2010,2131]" lastBlockId="3.[794,1419,218,435]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="81E7EAC8FA01880217025CA3FB9FF827" bold="true" box="[869,1085,2010,2035]" pageId="2" pageNumber="3">Digestive system:</emphasis>
(
<figureCitation id="2BA82A5FFA01880210375CA3FB0CF827" box="[1104,1198,2010,2035]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="2" pageNumber="3">Figs. 25</figureCitation>
,
<figureCitation id="2BA82A5FFA01880210D95CA3FAABF827" box="[1214,1289,2010,2035]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="2" pageNumber="3">27, 28</figureCitation>
) Arrangement of foregut essentially same as that described for
<taxonomicName id="74934D59FA018802110F5C83FC7DF7E7" authorityName="Simone &amp; Amaral" authorityYear="2018" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="castilhensis">
<emphasis id="81E7EAC8FA018802110F5C83FC7DF7E7" italics="true" pageId="2" pageNumber="3">Drymaeus castilhensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="D7024B2BFA018802178D5363FAF6F7E7" author="Simone, L. R. L. &amp; Amaral, V. S." box="[1002,1364,2074,2099]" pageId="2" pageNumber="3" pagination="167 - 187" refId="ref7069" refString="Simone, L. R. L. &amp; Amaral, V. S. 2018. Insular life: new endemic species from Sao Paulo oceanic islands, Brazil (Pulmonata, Bulimulidae), as example of endemicity. Journal of Conchology, 43 (2): 167 - 187." type="journal article" year="2018">Simone &amp; Amaral, 2018: 171-172</bibRefCitation>
, figs. 41-44) except for:
<emphasis id="81E7EAC8FA01880217825343FBB4F787" bold="true" box="[997,1046,2106,2131]" pageId="2" pageNumber="3">m1v</emphasis>
, ventral pair of jugal muscle inserted in middle region of ventral haemocoel surface just posteri- or to buccal mass, running anteriorly, insertion in mouth close to median line;
<emphasis id="81E7EAC8FA00880315215A63FEDBFEE7" bold="true" box="[326,377,282,307]" pageId="3" pageNumber="4">m1a</emphasis>
, small pair of lateral jugal muscles, originated in lateral surface of haemocoel, running short distance, inserting in latero-posterior region of buccal mass;
<emphasis id="81E7EAC8FA00880314895A03FEB6FE47" bold="true" box="[238,276,378,403]" pageId="3" pageNumber="4">m2</emphasis>
, pair of retractor muscles of buccal mass slightly narrower and 1.5-times longer;
<emphasis id="81E7EAC8FA008803166C5AE3FD93FE67" bold="true" box="[523,561,410,435]" pageId="3" pageNumber="4">m3</emphasis>
, pair of superficial muscles running transversally along postero-lateral surface of buccal mass, from dorso-lateral region to region close to radular nucleus at m2 insertion;
<emphasis id="81E7EAC8FA00880316045A83FD34FDC7" bold="true" box="[611,662,506,531]" pageId="3" pageNumber="4">m10</emphasis>
, pair of ventral protractor muscles of buccal mass slightly thinner and narrower; circular muscles of oral tube (mc) not so developed. Radular sac short, not extending beyond odontophore. Jaw plate (
<figureCitation id="2BA82A5FFA00880310555A43FBDCFE87" box="[1074,1150,314,339]" captionStart="Figures 9-13" captionStartId="4.[170,238,2010,2035]" captionTargetBox="[170,1493,1054,1980]" captionTargetId="figure-469@4.[170,1493,1054,1980]" captionTargetPageId="4" captionText="Figures 9-13. Drymaeus currais sp. nov. shell and anatomical details.(9) Protoconch of paratype MZSP 135683#3 (young specimen of 14.2 mm), arrow showing transition with teleoconch.Scale =1 mm.(10) Anterior region of mouth,ventral view,integument removed.Scale = 0.5 mm.(11) Duct of bursa sectioned longitudinally,showing4 spermatophoresinside.Scale =1 mm.(12)Isolatedspermatophore.Scale =1 mm.(13)Reno-pericardialarea,ventral view,ventral wallofkidney opened along its left edge and deflected right (upwards) to show inner tissue,ventral pericardial wall deflected left (downwards).Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4637247" httpUri="https://zenodo.org/record/4637247/files/figure.png" pageId="3" pageNumber="4">Fig. 10</figureCitation>
: jw) with medial slightly constriction, ~12 pairs of transverse folds medially narrow, gradually becoming wider towards lateral. Radular nucleus inlaid inside odontophore.
</paragraph>
<paragraph id="B32C36DAFA008804177D5AA3FC61FEE7" blockId="3.[794,1418,474,531]" lastBlockId="4.[868,1494,218,1011]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="81E7EAC8FA008803177D5AA3FCCFFE27" bold="true" box="[794,877,474,499]" pageId="3" pageNumber="4">Radula</emphasis>
(
<figureCitation id="2BA82A5FFA00880317115AA3FC4FFE27" box="[886,1005,474,499]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="3" pageNumber="4">Figs.14-22</figureCitation>
) as long as odontophore;with rachidi- an teeth,and~55 pairs of lateral teeth;no clear distinction between lateral and marginal teeth (
<figureCitation id="2BA82A5FFA07880416265BA3FCB2FF27" box="[577,784,218,243]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="4" pageNumber="5">Figs. 18, 19, 21, 22</figureCitation>
); all teeth with relative long and flattened base, locat- ed closely from neighboring rows; central set of cusps rather small, located in anterior end of base.
<emphasis id="81E7EAC8FA07880416C15A43FCBEFE87" bold="true" box="[678,796,314,339]" pageId="4" pageNumber="5">Rachidian</emphasis>
tooth (
<figureCitation id="2BA82A5FFA07880414905A23FE07FEA7" box="[247,421,346,371]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="4" pageNumber="5">Figs. 15, 16, 20</figureCitation>
: arrow) small, relatively reduced, ~half of neighboring teeth, ~1/70 of radular width; base ~3-times longer than wide, flattened, barely rectangular; central cusp with ~1/2 of base’s size, tip blunt; pair of basal cusps with ~1/5 of central cusp’s size (
<figureCitation id="2BA82A5FFA07880416A65AA3FCADFE27" box="[705,783,474,499]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="4" pageNumber="5">Fig. 16</figureCitation>
).
<emphasis id="81E7EAC8FA07880414CD5A83FEE2FDC7" bold="true" box="[170,320,506,531]" pageId="4" pageNumber="5">Lateral teeth</emphasis>
similar to rachidian,except in being ~twice larger, asymmetrical, arched towards lateral region, cutting edge ~1.5-times larger than that of rachidian; lateral teeth gradually weakly decreasing towards lateral; set of cusps with ~1/2 of length of base; central cusp bluntly pointed; basal cusps strongly asymmetrical, outer basal cusp almost as large as central cusp in more central teeth, almost originating from base (
<figureCitation id="2BA82A5FFA078804159F59A3FDE6FD27" box="[504,580,730,755]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="4" pageNumber="5">Fig. 16</figureCitation>
), gradually becoming smaller and sometimes separated from central cusp (
<figureCitation id="2BA82A5FFA07880414D55863FF5CFCE7" box="[178,254,794,819]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="4" pageNumber="5">Fig. 15</figureCitation>
); inner cusp ~3-4-times smaller than lateral cusp.
<emphasis id="81E7EAC8FA07880414CD5843FEFDFC87" bold="true" box="[170,351,826,851]" pageId="4" pageNumber="5">Marginal teeth</emphasis>
starting with no clear boundary with lateral teeth; shaped similarly to lateral teeth, except for being weakly smaller and with set of cusps broader and slightly less pointed (
<figureCitation id="2BA82A5FFA07880415F458E3FDFAFC67" box="[403,600,922,947]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="4" pageNumber="5">Figs. 17-19, 21-22</figureCitation>
); inner basal cusp gradually becoming as large as outer basal cusp and bifid, dividing in two cusps (
<figureCitation id="2BA82A5FFA07880415AB58A3FDD1FC27" box="[460,627,986,1011]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="4" pageNumber="5">Figs. 17, 21, 22</figureCitation>
); teeth becom- ing very narrow in margins (
<figureCitation id="2BA82A5FFA07880410C85BA3FB5DFF27" box="[1199,1279,218,243]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="4" pageNumber="5">Fig. 19</figureCitation>
). Each radular row slightly arched disposed from both sides from rachidian (
<figureCitation id="2BA82A5FFA078804170A5A63FC1AFEE7" box="[877,952,282,307]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="4" pageNumber="5">Fig. 14</figureCitation>
).
</paragraph>
<caption id="E7EC6652FA008803143B5363FEEEF787" ID-DOI="http://doi.org/10.5281/zenodo.4637245" ID-Zenodo-Dep="4637245" httpUri="https://zenodo.org/record/4637245/files/figure.png" pageId="3" pageNumber="4" startId="3.[92,160,2074,2099]" targetBox="[94,1417,570,2044]" targetPageId="3">
<paragraph id="B32C36DAFA008803143B5363FEEEF787" blockId="3.[92,1418,2074,2131]" pageId="3" pageNumber="4">
Figures2-8.
<taxonomicName id="74934D59FA00880314AA5363FEF7F7E7" authority="Simone &amp; Belz &amp; Gernet, 2020" authorityName="Simone &amp; Belz &amp; Gernet" authorityYear="2020" box="[205,341,2074,2099]" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="currais" status="sp. nov.">
<emphasis id="81E7EAC8FA00880314AA5363FEF7F7E7" bold="true" box="[205,341,2074,2099]" italics="true" pageId="3" pageNumber="4">Drymaeus currais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD457B3FA008803153E5363FE34F7E7" box="[345,406,2074,2099]" pageId="3" pageNumber="4" rank="species">sp.nov.</taxonomicNameLabel>
shell of types;(2-4)Holotype MZSP150548(L29.6),frontal,right and dorsal view.(5-8)Paratypes MZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm).
</paragraph>
</caption>
<paragraph id="B32C36DAFA07880417EC5A43FBCEFD47" blockId="4.[868,1494,218,1011]" pageId="4" pageNumber="5">
Salivary glands covering esophagus in its region preceding its anterior 1/6 (
<figureCitation id="2BA82A5FFA07880410015A23FB10FEA7" box="[1126,1202,346,371]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="4" pageNumber="5">Fig. 25</figureCitation>
: sg), forming two elongat- ed, white, thin masses. Each salivary duct differentiable in anterior side of glands, relatively broad, with ~1/10 esophageal width (
<figureCitation id="2BA82A5FFA078804105F5AC3FB2DFE07" box="[1080,1167,442,467]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="4" pageNumber="5">Figs. 25</figureCitation>
,
<figureCitation id="2BA82A5FFA07880410FF5AC3FB7FFE07" box="[1176,1245,442,467]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="4" pageNumber="5">27, 28</figureCitation>
: sd). Salivary duct running in both sides of esophageal origin (
<figureCitation id="2BA82A5FFA07880411415AA3FA07FE27" box="[1318,1445,474,499]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="4" pageNumber="5">Figs. 27, 28</figureCitation>
: sd), penetrating buccal mass wall in region close to buccal ganglia (
<figureCitation id="2BA82A5FFA07880417A15963FBB1FDE7" box="[966,1043,538,563]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="4" pageNumber="5">Fig. 27</figureCitation>
: sd), running immersed in buccal dorsal wall along ~1/4 its length (
<figureCitation id="2BA82A5FFA07880410CF5943FB5BFD87" box="[1192,1273,570,595]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="4" pageNumber="5">Fig. 28</figureCitation>
: sd). Salivary ducts opening wide, in middle level of dorsal folds, on their middle side (
<figureCitation id="2BA82A5FFA07880417955903FB9CFD47" box="[1010,1086,634,659]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="4" pageNumber="5">Fig. 28</figureCitation>
: sa).
</paragraph>
<paragraph id="B32C36DAFA07880617EC59E3FD4FFE67" blockId="4.[868,1494,218,1011]" lastBlockId="6.[169,794,218,435]" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
Esophagus 1 whorl long, with thin, flaccid walls lacking clear subdivisions (
<figureCitation id="2BA82A5FFA078804100F59C3FB14FD07" box="[1128,1206,698,723]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="4" pageNumber="5">Fig. 25</figureCitation>
: es). Stomach (
<figureCitation id="2BA82A5FFA078804110759C3FA0DFD07" box="[1376,1455,698,723]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="4" pageNumber="5">Fig. 25</figureCitation>
: st) relatively narrow, curved, not bulged; position and size described above (visceral mass);gastric walls thin,flaccid; inner surface smooth. Esophageal insertion on posterior side, intestinal origin on anterior side, both close to columella. Duct to anterior lobe of digestive gland located at some distance from stomach, highly branched, covering adjacent intestinal origin and loop (
<figureCitation id="2BA82A5FFA078804117F58E3FAC1FC67" box="[1304,1379,922,947]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="4" pageNumber="5">Fig. 25</figureCitation>
: dd-right). Duct to posterior lobe of digestive gland located short distance from esophageal insertion (
<figureCitation id="2BA82A5FFA078804117E58A3FACEFC27" box="[1305,1388,986,1011]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="4" pageNumber="5">Fig. 25</figureCitation>
: dd-left). Intestine ~half of width of esophageal insertion, all along its length,including narrow sigmoid loop in anterior lobe of digestive gland (
<figureCitation id="2BA82A5FFA058806151B5A63FE65FEE7" box="[380,455,282,307]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="6" pageNumber="7">Fig. 25</figureCitation>
: in). Rectum and anus position described above (pallial cavity) (
<figureCitation id="2BA82A5FFA05880616695A43FDF8FE87" box="[526,602,314,339]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="6" pageNumber="7">Fig. 24</figureCitation>
: rt, an). Anus sessile, as slit in right end of mantle edge directly turned outside, but still inside pneumostome (
<figureCitation id="2BA82A5FFA05880616315A03FD00FE47" box="[598,674,378,403]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="6" pageNumber="7">Fig. 23</figureCitation>
); inner surface with 8-10 longitudinal, simple folds (
<figureCitation id="2BA82A5FFA05880616175AE3FD19FE67" box="[624,699,410,435]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="6" pageNumber="7">Fig. 26</figureCitation>
: an).
</paragraph>
<caption id="E7EC6652FA07880414CD5CA3FAE6F787" ID-DOI="http://doi.org/10.5281/zenodo.4637247" ID-Zenodo-Dep="4637247" httpUri="https://zenodo.org/record/4637247/files/figure.png" pageId="4" pageNumber="5" startId="4.[170,238,2010,2035]" targetBox="[170,1493,1054,1980]" targetPageId="4">
<paragraph id="B32C36DAFA07880414CD5CA3FAE6F787" blockId="4.[169,1493,2010,2131]" pageId="4" pageNumber="5">
Figures 9-13.
<taxonomicName id="74934D59FA07880415405CA3FE10F827" authority="Simone &amp; Belz &amp; Gernet, 2020" authorityName="Simone &amp; Belz &amp; Gernet" authorityYear="2020" box="[295,434,2010,2035]" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="currais" status="sp. nov.">
<emphasis id="81E7EAC8FA07880415405CA3FE10F827" bold="true" box="[295,434,2010,2035]" italics="true" pageId="4" pageNumber="5">Drymaeus currais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD457B3FA07880415DF5CA3FE55F827" box="[440,503,2010,2035]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
shell and anatomical details.(9) Protoconch of paratype MZSP 135683#3 (young specimen of 14.2 mm), arrow showing transition with teleoconch.Scale =1 mm.(10) Anterior region of mouth,ventral view,integument removed.Scale = 0.5 mm.(11) Duct of bursa sectioned longitudinally,showing4 spermatophores inside.Scale =1 mm.(12)Isolated spermatophore.Scale =1 mm.(13)Reno-pericardial area,ventral view,ventral wall of kidney opened along its left edge and deflected right (upwards) to show inner tissue,ventral pericardial wall deflected left (downwards).Scale = 1 mm.
</paragraph>
</caption>
<caption id="E7EC6652FA06880514395CA3FCAFF787" ID-DOI="http://doi.org/10.5281/zenodo.4637251" ID-Zenodo-Dep="4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="5" pageNumber="6" startId="5.[94,162,2010,2035]" targetBox="[123,1389,204,1980]" targetPageId="5">
<paragraph id="B32C36DAFA06880514395CA3FCAFF787" blockId="5.[94,1419,2010,2131]" pageId="5" pageNumber="6">
Figures 14-22.
<taxonomicName id="74934D59FA06880514815CA3FED2F827" authority="Simone &amp; Belz &amp; Gernet, 2020" authorityName="Simone &amp; Belz &amp; Gernet" authorityYear="2020" box="[230,368,2010,2035]" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="currais" status="sp. nov.">
<emphasis id="81E7EAC8FA06880514815CA3FED2F827" bold="true" box="[230,368,2010,2035]" italics="true" pageId="5" pageNumber="6">Drymaeus currais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD457B3FA06880515125CA3FE11F827" box="[373,435,2010,2035]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column.
</paragraph>
</caption>
<paragraph id="B32C36DAFA05880814CD5AA3FA74FE27" blockId="6.[170,795,474,627]" lastBlockId="8.[868,1495,218,499]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<emphasis id="81E7EAC8FA05880614CD5AA3FEC0FE27" bold="true" box="[170,354,474,499]" pageId="6" pageNumber="7">Genital system:</emphasis>
(
<figureCitation id="2BA82A5FFA05880615175AA3FE49FE27" box="[368,491,474,499]" captionStart-0="Figures 27-31" captionStart-1="Figures 32-36" captionStartId-0="7.[94,162,2010,2035]" captionStartId-1="8.[170,238,2042,2067]" captionTargetBox-0="[98,1414,568,1976]" captionTargetBox-1="[174,1493,657,2008]" captionTargetId-0="figure-185@7.[94,1417,563,1980]" captionTargetId-1="figure-217@8.[170,1493,651,2012]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." captionText-1="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4637255" figureDoi-1="http://doi.org/10.5281/zenodo.4637257" httpUri-0="https://zenodo.org/record/4637255/files/figure.png" httpUri-1="https://zenodo.org/record/4637257/files/figure.png" pageId="6" pageNumber="7">Figs. 29-34</figureCitation>
) gonad position described above (visceral mass), composed of 4-5 lobes with minute digitiform acini (
<figureCitation id="2BA82A5FFA05880615FF5963FE4BFDE7" box="[408,489,538,563]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="6" pageNumber="7">Fig. 29</figureCitation>
: go). Hermaphroditic duct (
<figureCitation id="2BA82A5FFA05880614D75943FEA5FD87" box="[176,263,570,595]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="6" pageNumber="7">Figs. 29</figureCitation>
,
<figureCitation id="2BA82A5FFA05880615765943FE8DFD87" box="[273,303,570,595]" captionStart="Figures 14-22" captionStartId="5.[94,162,2010,2035]" captionTargetBox="[123,1389,204,1980]" captionTargetId="figure-1@5.[123,1389,204,1980]" captionTargetPageId="5" captionText="Figures 14-22. Drymaeus currais sp. nov.radulae in SEM; (14) Holotype,wide view.Scale = 200 µm. (15) Same,higher magnification.Scale = 30 µm. (16) Same, higher magnification. Scale = 20 µm. (17) Detail of lateral region.Scale = 20 µm. (18) Same, lower magnification. Scale = 50 µm. (19) Detail of marginal region. Scale = 50 µm. (20) Paratype MZSP 135683, detail of central region. Scale = 20 µm. (21) Detail of lateral-marginal region, middle region folded.Scale = 50 µm. (22) Same,higher magnification.Scale = 20 µm.Arrows showing rachidian column." figureDoi="http://doi.org/10.5281/zenodo.4637251" httpUri="https://zenodo.org/record/4637251/files/figure.png" pageId="6" pageNumber="7">19</figureCitation>
: hd) narrow and weakly coiled in both ends, gradually becoming very wider (up to 5-times wider) and more intensely coiled in middle third (
<figureCitation id="2BA82A5FFA058806116F5BA3FAF6FF27" box="[1288,1364,218,243]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="6" pageNumber="7">Fig. 29</figureCitation>
: hd); inserting in left side of receptacle’s base (
<figureCitation id="2BA82A5FFA05880610945B83FAE2FEC7" box="[1267,1344,250,275]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="6" pageNumber="7">Fig. 32</figureCitation>
: hd). Seminal receptacle (
<figureCitation id="2BA82A5FFA05880617885A63FBF2FEE7" box="[1007,1104,282,307]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="6" pageNumber="7">Figs. 29</figureCitation>
,
<figureCitation id="2BA82A5FFA05880610055A63FB22FEE7" box="[1122,1152,282,307]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="6" pageNumber="7">32</figureCitation>
: sr) relatively small, sac-like, ~twice longer than wide, with ~twice hermaphroditic duct width. Fertilization complex simple, located at narrow and elongated base of seminal receptacle (
<figureCitation id="2BA82A5FFA058806110E5A03FA11FE47" box="[1385,1459,378,403]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="6" pageNumber="7">Fig. 32</figureCitation>
) as duct of seminal receptacle; ~as long as length of receptacle. Presence of genital appendix (
<figureCitation id="2BA82A5FFA05880610985AC3FAFAFE07" box="[1279,1368,442,467]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="6" pageNumber="7">Figs. 29</figureCitation>
,
<figureCitation id="2BA82A5FFA05880611045AC3FA23FE07" box="[1379,1409,442,467]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="6" pageNumber="7">32</figureCitation>
: ga), as wide as terminal width of hermaphrodite duct, inserted just posterior to it in seminal receptacle’s base; weakly coiled, ~3-times longer than receptacle. Fertilization complex totally immersed in albumen gland (
<figureCitation id="2BA82A5FFA058806111D5943FA69FD87" box="[1402,1483,570,595]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="6" pageNumber="7">Fig. 32</figureCitation>
), inserting in posterior end of spermoviduct, in base of albumen gland duct (
<figureCitation id="2BA82A5FFA04880715375BA3FE3FFF27" box="[336,413,218,243]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="7" pageNumber="8">Fig. 32</figureCitation>
: ad) relatively wide, simple, ~3-times wider than receptacle’s duct. Albumen gland (
<figureCitation id="2BA82A5FFA04880714035A63FF1DFEE7" box="[100,191,282,307]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="7" pageNumber="8">Figs. 29</figureCitation>
,
<figureCitation id="2BA82A5FFA04880714AC5A63FF4BFEE7" box="[203,233,282,307]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="7" pageNumber="8">32</figureCitation>
: ag) solid, white, elliptical, 1.5-times larger than gonad (~1/3 whorl). Albumen gland duct subterminal, connected to distal end of spermoviduct (
<figureCitation id="2BA82A5FFA04880716345A23FD3CFEA7" box="[595,670,346,371]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="7" pageNumber="8">Fig. 32</figureCitation>
: ad), in lateral, large albumen chamber (
<figureCitation id="2BA82A5FFA04880715915A03FDE4FE47" box="[502,582,378,403]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="7" pageNumber="8">Fig. 32</figureCitation>
: ac); widely connected to distal end of spermoviduct. Spermoviduct (eo) of ~1.5 whorl in length, slightly narrower than albumen gland,
<emphasis id="81E7EAC8FA04880715615AA3FE87FE27" box="[262,293,474,499]" italics="true" pageId="7" pageNumber="8">ca.</emphasis>
20-times longer than wide; intensely coiled and difficulty to rectify. Prostate gland occupy- ing ~1/4 of spermoviduct surface and ~1/5 its volume (
<figureCitation id="2BA82A5FFA04880717475B83FCD0FEC7" box="[800,882,250,275]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="7" pageNumber="8">Fig. 33</figureCitation>
: pt). Uterus occupying ~3/4 of spermoviduct space, external walls thick-glandular (
<figureCitation id="2BA82A5FFA04880710D05A63FA96FEE7" box="[1207,1332,282,307]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="7" pageNumber="8">Figs. 29, 31</figureCitation>
,
<figureCitation id="2BA82A5FFA04880711595A63FAFEFEE7" box="[1342,1372,282,307]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="7" pageNumber="8">32</figureCitation>
: ut), inner surface completely covered by ample transverse folds. Sperm grove simple in posterior 2/3 of spermoviduct (
<figureCitation id="2BA82A5FFA048807173F5A03FC01FE47" box="[856,931,378,403]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="7" pageNumber="8">Fig. 33</figureCitation>
: sp), protected ventrally by tall fold; abruptly becoming tubular becoming, as vas deferens in anteri- or end of spermoviduct (
<figureCitation id="2BA82A5FFA04880710565AC3FBDFFE07" box="[1073,1149,442,467]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="7" pageNumber="8">Fig. 31</figureCitation>
: vd). Free oviduct short, ~1/20 of spermoviduct length, Vagina ~1/10 spermoviduct length (
<figureCitation id="2BA82A5FFA04880717CC5A83FB89FDC7" box="[939,1067,506,531]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="7" pageNumber="8">Figs. 29, 31</figureCitation>
: vg); inner surface simple, with 4-5 longitudinal, low, wide folds (
<figureCitation id="2BA82A5FFA0B8808167A5BA3FDCBFF27" box="[541,617,218,243]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">Fig. 31</figureCitation>
: vg). Bursa copulatrix ~2/3 of spermoviduct length; bursa duct as wide as adjacent spermoviduct in its origin (
<figureCitation id="2BA82A5FFA0B880816305A63FD00FEE7" box="[599,674,282,307]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">Fig. 31</figureCitation>
), gradually narrowing towards posterior end (
<figureCitation id="2BA82A5FFA0B880816465A43FDCEFE87" box="[545,620,314,339]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">Fig. 29</figureCitation>
: bd); bursa oval, ~1/5 of albumen gland size (
<figureCitation id="2BA82A5FFA0B880815835A23FD8DFEA7" box="[484,559,346,371]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">Fig. 29</figureCitation>
: bc), located encased between pericardium and adjacent intestinal loop. Some specimens with 4-5 elongated, simple spermatophores in bursa duct’s base (
<figureCitation id="2BA82A5FFA0B880815FE5AC3FDB9FE07" box="[409,539,442,467]" captionStart="Figures 9-13" captionStartId="4.[170,238,2010,2035]" captionTargetBox="[170,1493,1054,1980]" captionTargetId="figure-469@4.[170,1493,1054,1980]" captionTargetPageId="4" captionText="Figures 9-13. Drymaeus currais sp. nov. shell and anatomical details.(9) Protoconch of paratype MZSP 135683#3 (young specimen of 14.2 mm), arrow showing transition with teleoconch.Scale =1 mm.(10) Anterior region of mouth,ventral view,integument removed.Scale = 0.5 mm.(11) Duct of bursa sectioned longitudinally,showing4 spermatophoresinside.Scale =1 mm.(12)Isolatedspermatophore.Scale =1 mm.(13)Reno-pericardialarea,ventral view,ventral wallofkidney opened along its left edge and deflected right (upwards) to show inner tissue,ventral pericardial wall deflected left (downwards).Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4637247" httpUri="https://zenodo.org/record/4637247/files/figure.png" pageId="8" pageNumber="9">Figs. 11, 12</figureCitation>
,
<figureCitation id="2BA82A5FFA0B880816415AC3FDE6FE07" box="[550,580,442,467]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">29</figureCitation>
: sm). Penis ~1/2 of spermoviduct length, ~2/3 its anterior width (
<figureCitation id="2BA82A5FFA0B880816F95AA3FD4AFE27" box="[670,744,474,499]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">Fig. 29</figureCitation>
: pe); penis muscle inserting terminally,very short (
<figureCitation id="2BA82A5FFA0B880816F15A83FD42FDC7" box="[662,736,506,531]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">Fig. 29</figureCitation>
: pm). Epiphallus ~1/8 penis’ length, located as short terminal continuation of penis (
<figureCitation id="2BA82A5FFA0B880815C65943FE55FD87" box="[417,503,570,595]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">Figs. 29</figureCitation>
,
<figureCitation id="2BA82A5FFA0B880815985943FDBFFD87" box="[511,541,570,595]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="8" pageNumber="9">34</figureCitation>
: eh), inner surface with narrow, irregular longitudinal folds, located close from each other, uniformly sized (
<figureCitation id="2BA82A5FFA0B880810C75B83FB4EFEC7" box="[1184,1260,250,275]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="8" pageNumber="9">Fig. 34</figureCitation>
: eh). Vas deferens inserted subterminally in penis tip (
<figureCitation id="2BA82A5FFA0B880810B75A63FA87FEE7" box="[1232,1317,282,307]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">Figs.29</figureCitation>
,
<figureCitation id="2BA82A5FFA0B8808114B5A63FAE8FEE7" box="[1324,1354,282,307]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="8" pageNumber="9">34</figureCitation>
: vd). Internal penial surface lacking clear sub-chambers (
<figureCitation id="2BA82A5FFA0B880811375A43FA3DFE87" box="[1360,1439,314,339]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="8" pageNumber="9">Fig. 34</figureCitation>
: pe); short smooth middle region posterior region with 5-6 longitudinal, narrow, low folds; anterior region with pair of larger folds (
<figureCitation id="2BA82A5FFA0B8808106E5AE3FBF7FE67" box="[1033,1109,410,435]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="8" pageNumber="9">Fig. 34</figureCitation>
). Penis shield occupying basal ~1/4 of penis length (
<figureCitation id="2BA82A5FFA0B880810705AC3FB37FE07" box="[1047,1173,442,467]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">Figs. 29, 30</figureCitation>
: ps); vas deferens piercing its walls close to its base (
<figureCitation id="2BA82A5FFA0B880810315AA3FB02FE27" box="[1110,1184,474,499]" captionStart="Figures 27-31" captionStartId="7.[94,162,2010,2035]" captionTargetBox="[98,1414,568,1976]" captionTargetId="figure-185@7.[94,1417,563,1980]" captionTargetPageId="7" captionText="Figures 27-31. Drymaeus currais sp.nov.anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="8" pageNumber="9">Fig.30</figureCitation>
).Genital pore round,simple.
</paragraph>
<caption id="E7EC6652FA05880614CD5CA3FDE0F787" ID-DOI="http://doi.org/10.5281/zenodo.4637253" ID-Zenodo-Dep="4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="6" pageNumber="7" startId="6.[170,238,2010,2035]" targetBox="[174,1489,662,1974]" targetPageId="6">
<paragraph id="B32C36DAFA05880614CD5CA3FDE0F787" blockId="6.[170,1494,2010,2131]" pageId="6" pageNumber="7">
Figures 23-26.
<taxonomicName id="74934D59FA05880615565CA3FE19F827" authority="Simone &amp; Belz &amp; Gernet, 2020" authorityName="Simone &amp; Belz &amp; Gernet" authorityYear="2020" box="[305,443,2010,2035]" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="currais" status="sp. nov.">
<emphasis id="81E7EAC8FA05880615565CA3FE19F827" bold="true" box="[305,443,2010,2035]" italics="true" pageId="6" pageNumber="7">Drymaeus currais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD457B3FA05880615D85CA3FE5FF827" box="[447,509,2010,2035]" pageId="6" pageNumber="7" rank="species">sp.nov.</taxonomicNameLabel>
anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflected upwards.Scales = 2 mm.
</paragraph>
</caption>
<caption id="E7EC6652FA04880714395CA3FD90F787" ID-DOI="http://doi.org/10.5281/zenodo.4637255" ID-Zenodo-Dep="4637255" httpUri="https://zenodo.org/record/4637255/files/figure.png" pageId="7" pageNumber="8" startId="7.[94,162,2010,2035]" targetBox="[98,1414,568,1976]" targetPageId="7">
<paragraph id="B32C36DAFA04880714395CA3FD90F787" blockId="7.[94,1418,2010,2131]" pageId="7" pageNumber="8">
Figures 27-31.
<taxonomicName id="74934D59FA04880714805CA3FED3F827" authority="Simone &amp; Belz &amp; Gernet, 2020" authorityName="Simone &amp; Belz &amp; Gernet" authorityYear="2020" box="[231,369,2010,2035]" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="currais" status="sp. nov.">
<emphasis id="81E7EAC8FA04880714805CA3FED3F827" bold="true" box="[231,369,2010,2035]" italics="true" pageId="7" pageNumber="8">Drymaeus currais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD457B3FA04880715105CA3FE14F827" box="[375,438,2010,2035]" pageId="7" pageNumber="8" rank="species">sp.nov.</taxonomicNameLabel>
anatomy; (27) Foregut, left view. (28) Buccal mass, ventral view, odontophore (od) partially sectioned and deflected to left, right buccal ganglion (bg) and salivary duct (sd) (right in Fig.) insertion seen by translucency.(29) Genital system, dorsal view,partially uncoiled, some spermatophores (sm) seen by translucency.(30) Penis,detail of its proximal region,penis sheath (ps) partially sectioned.(31) Genital ducts,detail of its anterior region, some structures sectioned longitudinally.Scales = 2 mm.
</paragraph>
</caption>
<paragraph id="B32C36DAFA0B880817025963FA75FD87" blockId="8.[869,1495,538,595]" pageId="8" pageNumber="9">
<emphasis id="81E7EAC8FA0B880817025963FB27FDE7" bold="true" box="[869,1157,538,563]" pageId="8" pageNumber="9">Central nervous system:</emphasis>
(
<figureCitation id="2BA82A5FFA0B880810F45963FAB1FDE7" box="[1171,1299,538,563]" captionStart="Figures 32-36" captionStartId="8.[170,238,2042,2067]" captionTargetBox="[174,1493,657,2008]" captionTargetId="figure-217@8.[170,1493,651,2012]" captionTargetPageId="8" captionText="Figures 32-36.Drymaeuscurrais sp.nov.anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="8" pageNumber="9">Figs. 35, 36</figureCitation>
) cerebral ganglia located dorsally in middle level of buccal mass, pedal
</paragraph>
<caption id="E7EC6652FA0B880814CD5C83FE8EF787" ID-DOI="http://doi.org/10.5281/zenodo.4637257" ID-Zenodo-Dep="4637257" httpUri="https://zenodo.org/record/4637257/files/figure.png" pageId="8" pageNumber="9" startId="8.[170,238,2042,2067]" targetBox="[174,1493,657,2008]" targetPageId="8">
<paragraph id="B32C36DAFA0B880814CD5C83FE8EF787" blockId="8.[170,1494,2042,2131]" pageId="8" pageNumber="9">
Figures 32-36.
<taxonomicName id="74934D59FA0B880815575C83FE1BF7C7" authority="Simone &amp; Belz &amp; Gernet, 2020" authorityName="Simone &amp; Belz &amp; Gernet" authorityYear="2020" box="[304,441,2042,2067]" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="currais" status="sp. nov.">
<emphasis id="81E7EAC8FA0B880815575C83FE1BF7C7" bold="true" box="[304,441,2042,2067]" italics="true" pageId="8" pageNumber="9">Drymaeus currais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD457B3FA0B880815D95C83FE59F7C7" box="[446,507,2042,2067]" pageId="8" pageNumber="9" rank="species">sp.nov.</taxonomicNameLabel>
anatomy;(32) Middle region of genital system,ventral view,some structures seen if albumen gland(ag)was transparent. (33) Spermoviduct, transverse section in its middle level. (34) Penis, ventral view, opened longitudinally. (35) Nerve ring, dorsal view. (36) Same, ventral view. Scales = 2 mm.
</paragraph>
</caption>
<paragraph id="B32C36DAFA0A880914395BA3FDFCFDC7" blockId="9.[94,718,218,531]" pageId="9" pageNumber="10">
ganglia located more posteriorly. Pair of cerebral ganglia (ce) widely fused with each other; cerebral commissure invisible; each ganglion about as wide as adjacent esophageal section (
<figureCitation id="2BA82A5FFA0A8809149C5A43FEE4FE87" box="[251,326,314,339]" captionStart="Figures 23-26" captionStartId="6.[170,238,2010,2035]" captionTargetBox="[174,1489,662,1974]" captionTargetId="figure-244@6.[170,1493,657,1980]" captionTargetPageId="6" captionText="Figures 23-26. Drymaeus currais sp.nov.anatomy;(23) Mantle border as in situ,frontal view.(24) Pallial(pulmonary) cavity,ventral inner view,inner mantle fold deflected upwards. (25) Digestive tubes and some adjacent structures, mainly from visceral mass and posterior region of pulmonary cavity, buccal mass in dorsal view, remaining ventral view, nephrostome (ne) seen by translucency. (26) Anterior-right region of pulmonary cavity, ventral-inner view, inner mantle fold (mf) sectioned and deflectedupwards.Scales = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4637253" httpUri="https://zenodo.org/record/4637253/files/figure.png" pageId="9" pageNumber="10">Fig. 25</figureCitation>
: nr); several wide nerves originating in cerebral antero-lateral region. No clear cerebral node or gland.Two pairs of parallel connectives between cerebral ganglia and pedal ganglia. Pair of pedal ganglia (pp) forming single mass located opposite to cerebral ganglia, slightly larger sized than cerebral ganglia. No differentiable individual pedal ganglion detectable.
</paragraph>
</subSubSection>
<subSubSection id="FB896551FA0A880914395943FE16FC87" pageId="9" pageNumber="10" type="distribution">
<paragraph id="B32C36DAFA0A880914395943FE98FDA7" blockId="9.[94,718,570,627]" pageId="9" pageNumber="10">
<emphasis id="81E7EAC8FA0A880914395943FF54FD87" bold="true" box="[94,246,570,595]" pageId="9" pageNumber="10">Distribution:</emphasis>
Endemic from Guapirá Island, Currais Archipelago (
<figureCitation id="2BA82A5FFA0A880914965923FE89FDA7" box="[241,299,602,627]" captionStart="Figure 1" captionStartId="2.[172,231,688,713]" captionTargetBox="[171,794,205,658]" captionTargetId="figure-813@2.[170,794,204,658]" captionTargetPageId="2" captionText="Figure 1.Map of Paraná coastal area,showing the Currais archipelago location and its profile in highlight. Occurrence of specimens in the larger island (eastone).A living crawling specimen also shown (shell length ~20 mm)." figureDoi="http://doi.org/10.5281/zenodo.4637243" httpUri="https://zenodo.org/record/4637243/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="B32C36DAFA0A8809143959E3FE16FC87" blockId="9.[94,719,666,851]" pageId="9" pageNumber="10">
<emphasis id="81E7EAC8FA0A8809143959E3FF1CFD67" bold="true" box="[94,190,666,691]" pageId="9" pageNumber="10">Habitat:</emphasis>
The animals were found in a shaded area, inhabiting both the trunks and the abaxial part of the leaves of the tree vegetation of the Ombrophiles Dense Forest that covers the island. They were also located in the bromeliads of the rocky shores that are part of the pioneer rupicolous vegetation.
</paragraph>
</subSubSection>
<subSubSection id="FB896551FA0A880914395803FD36FC07" pageId="9" pageNumber="10" type="description">
<paragraph id="B32C36DAFA0A880914395803FD36FC07" blockId="9.[93,718,890,979]" pageId="9" pageNumber="10">
<emphasis id="81E7EAC8FA0A880914395803FEB0FC47" bold="true" box="[94,274,890,915]" pageId="9" pageNumber="10">Measurements:</emphasis>
(length and width in mm)
<typeStatus id="6C288878FA0A880916405803FD2EFC47" box="[551,652,890,915]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
MZSP 150548 (
<figureCitation id="2BA82A5FFA0A880914DC58E3FEBBFC67" box="[187,281,922,947]" captionStart="Figures2-8" captionStartId="3.[92,160,2074,2099]" captionTargetBox="[94,1417,570,2044]" captionTargetId="figure-178@3.[94,1417,570,2044]" captionTargetPageId="3" captionText="Figures2-8. Drymaeuscurrais sp.nov.shell oftypes;(2-4)HolotypeMZSP150548(L29.6),frontal,rightand dorsalview.(5-8)ParatypesMZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm)." figureDoi="http://doi.org/10.5281/zenodo.4637245" httpUri="https://zenodo.org/record/4637245/files/figure.png" pageId="9" pageNumber="10">Figs. 2-4</figureCitation>
): 29.6 by 10.6;
<typeStatus id="6C288878FA0A880915DA58E3FD8BFC67" box="[445,553,922,947]" pageId="9" pageNumber="10" type="paratype">paratypes</typeStatus>
MZSP 135683: #1 (
<figureCitation id="2BA82A5FFA0A880914E158C3FF43FC07" box="[134,225,954,979]" captionStart="Figures2-8" captionStartId="3.[92,160,2074,2099]" captionTargetBox="[94,1417,570,2044]" captionTargetId="figure-178@3.[94,1417,570,2044]" captionTargetPageId="3" captionText="Figures2-8. Drymaeuscurrais sp.nov.shell oftypes;(2-4)HolotypeMZSP150548(L29.6),frontal,rightand dorsalview.(5-8)ParatypesMZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm)." figureDoi="http://doi.org/10.5281/zenodo.4637245" httpUri="https://zenodo.org/record/4637245/files/figure.png" pageId="9" pageNumber="10">Figs. 5-6</figureCitation>
), 30.7 by 14.5; #2 (
<figureCitation id="2BA82A5FFA0A880915C558C3FE5CFC07" box="[418,510,954,979]" captionStart="Figures2-8" captionStartId="3.[92,160,2074,2099]" captionTargetBox="[94,1417,570,2044]" captionTargetId="figure-178@3.[94,1417,570,2044]" captionTargetPageId="3" captionText="Figures2-8. Drymaeuscurrais sp.nov.shell oftypes;(2-4)HolotypeMZSP150548(L29.6),frontal,rightand dorsalview.(5-8)ParatypesMZSP135683.(5-6)Frontal and dorsal view (L 30.7 mm)." figureDoi="http://doi.org/10.5281/zenodo.4637245" httpUri="https://zenodo.org/record/4637245/files/figure.png" pageId="9" pageNumber="10">Figs.7-8</figureCitation>
), 25.2 by 12.7.
</paragraph>
</subSubSection>
</treatment>
</document>
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