269 lines
30 KiB
XML
269 lines
30 KiB
XML
<document id="4F1DF2369382BAA148377198569DA650" ID-DOI="10.11646/zootaxa.3719.1.1" ID-GBIF-Dataset="523becce-8715-4b42-bab0-a66766621afd" ID-ISSN="1175-5326" ID-Zenodo-Dep="246657" ID-ZooBank="1F275DB9-0D3B-4322-9F21-A402F1384D8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460687712109" checkinUser="plazi" docAuthor="Hughes, Lauren E. & Peart, Rachael A." docDate="2013" docId="E574A473FF907E4A7BC48143FB7DFA7C" docLanguage="en" docName="zt03719p102.pdf" docOrigin="Zootaxa 3719 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cymadusa Hughes & Peart, 2013, comb. nov." docType="treatment" docVersion="7" lastPageNumber="5" masterDocId="194DDC0BFF937E4E7B53843AFFD8FF98" masterDocTitle="New species and new records of Ampithoidae (Peracarida: Amphipoda) from Australian Waters" masterLastPageNumber="102" masterPageNumber="1" pageNumber="4" updateTime="1698351413782" updateUser="plazi">
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<mods:mods id="4E2EEC78D645E4BA9A7DB92664DE7E25" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A59419F80F9D88F32DDFC49B8FD2FA99">
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<mods:title id="1754BC1E36D2EA6220AA22E4007E3E91">New species and new records of Ampithoidae (Peracarida: Amphipoda) from Australian Waters</mods:title>
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<mods:name id="69F08DBB47187B37BAB8C13BDDF34769" type="personal">
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<mods:roleTerm id="2EEE887D1BEA830778AADACAC8B21880">Author</mods:roleTerm>
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<mods:namePart id="E19F2124DB4E7FD861DC8F41DE1D3D08">Hughes, Lauren E.</mods:namePart>
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<mods:name id="0C6BEAEEEE4AE291B23E6B735471150F" type="personal">
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<mods:roleTerm id="AC146419DDC35553F446C735CDD56DAD">Author</mods:roleTerm>
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<mods:namePart id="3003CF896F120AB372CB8C80745B3739">Peart, Rachael A.</mods:namePart>
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<mods:typeOfResource id="2BD44FAAB91E211C8C7B3AB9DD68F2D7">text</mods:typeOfResource>
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<mods:title id="FD0CA0B6B90CD8E655E1536B662AAC9D">Zootaxa</mods:title>
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<mods:part id="96BC41246A14566D052D2B8043368E8D">
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<mods:date id="B243BCF49C3322C30A9D71360BA76B67">2013</mods:date>
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<mods:detail id="0C5FB12C4C3DFFE23280CDAA9BF55AF3" type="volume">
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<mods:number id="F47264945852E691B1F1DF094270A0C2">3719</mods:number>
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<mods:number id="7BBE89DD4BE966A5202BEC4B7FE79C88">1</mods:number>
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<mods:classification id="9257CA0826E0785DFA0778D3877ABEEA">journal article</mods:classification>
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<mods:identifier id="C705F0AD3F3C269E258AF1DF69EA64D9" type="DOI">10.11646/zootaxa.3719.1.1</mods:identifier>
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<mods:identifier id="BB16336538D17284971180D4E854B637" type="ISSN">1175-5326</mods:identifier>
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<treatment id="E574A473FF907E4A7BC48143FB7DFA7C" ID-DOI="http://doi.org/10.5281/zenodo.6154802" ID-GBIF-Taxon="119576722" ID-Zenodo-Dep="6154802" LSID="urn:lsid:plazi:treatment:E574A473FF907E4A7BC48143FB7DFA7C" httpUri="http://treatment.plazi.org/id/E574A473FF907E4A7BC48143FB7DFA7C" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<subSubSection id="25C746EEFF907E4A7BC48143FB7DFA7C" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="6D621565FF907E4D7BC48143FE07FA0B" blockId="3.[151,479,1401,1427]" box="[151,479,1401,1427]" pageId="3" pageNumber="4">
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<heading id="362AA209FF907E4D7BC48143FE07FA0B" bold="true" box="[151,479,1401,1427]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<emphasis id="5FA9C977FF907E4D7BC48143FE07FA0B" bold="true" box="[151,479,1401,1427]" pageId="3" pageNumber="4">
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<taxonomicName id="AADD6EE6FF907E4D7BC48143FECEFA0B" ID-CoL="7NV6K" box="[151,278,1401,1427]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D7BC48143FECEFA0B" bold="true" box="[151,278,1401,1427]" italics="true" pageId="3" pageNumber="4">Cymadusa</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="AADD6EE6FF907E4D7A078143FE07FA0B" box="[340,479,1401,1427]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D7A078143FE07FA0B" bold="true" box="[340,479,1401,1427]" italics="true" pageId="3" pageNumber="4">Paragrubia</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6D621565FF907E4D7BC48185FD46F9F0" blockId="3.[151,1436,1471,2001]" pageId="3" pageNumber="4">
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The genus
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<taxonomicName id="AADD6EE6FF907E4D7A4A8185FE44FA40" box="[281,412,1471,1496]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D7A4A8185FE44FA40" box="[281,412,1471,1496]" italics="true" pageId="3" pageNumber="4">Paragrubia</emphasis>
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</taxonomicName>
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was established by monotypy for
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<taxonomicName id="AADD6EE6FF907E4D78628185FB0DFA40" authority="Chevreux, 1901" authorityName="Chevreux" authorityYear="1901" box="[817,1237,1471,1496]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vorax">
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<emphasis id="5FA9C977FF907E4D78628185FC26FA40" box="[817,1022,1471,1496]" italics="true" pageId="3" pageNumber="4">Paragrubia vorax</emphasis>
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(Chevreux, 1901)
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</taxonomicName>
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.
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<taxonomicName id="AADD6EE6FF907E4D7FB08185FABEFA40" box="[1251,1382,1471,1496]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D7FB08185FABEFA40" box="[1251,1382,1471,1496]" italics="true" pageId="3" pageNumber="4">Paragrubia</emphasis>
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</taxonomicName>
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was originally diagnosed by the following features: gnathopod 1 is more robust than gnathopod 2, the antenna 1 accessory flagellum is multi-articulate and the telson is subovate. Three characters separated
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<taxonomicName id="AADD6EE6FF907E4D7FE2823DFAECF9B8" box="[1201,1332,1543,1568]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D7FE2823DFAECF9B8" box="[1201,1332,1543,1568]" italics="true" pageId="3" pageNumber="4">Paragrubia</emphasis>
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</taxonomicName>
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from the closely related genus
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<taxonomicName id="AADD6EE6FF907E4D7ADC8216FDDDF9DD" box="[399,517,1580,1605]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D7ADC8216FDDDF9DD" box="[399,517,1580,1605]" italics="true" pageId="3" pageNumber="4">Cymadusa</emphasis>
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</taxonomicName>
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in which gnathopod 1 is smaller than gnathopod 2, the accessory flagellum has only 1-article and the telson was less rounded.
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</paragraph>
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<paragraph id="6D621565FF907E4D7B94824EFE29F849" blockId="3.[151,1436,1471,2001]" pageId="3" pageNumber="4">
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Conlan (1983) expanded the diagnosis of
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<taxonomicName id="AADD6EE6FF907E4D79EE824EFCEBF915" box="[701,819,1652,1677]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D79EE824EFCEBF915" box="[701,819,1652,1677]" italics="true" pageId="3" pageNumber="4">Cymadusa</emphasis>
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</taxonomicName>
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to include a multi-articulate accessory flagellum to accommodate
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<taxonomicName id="AADD6EE6FF907E4D7A1382A2FDADF928" authority="Stout, 1912" authorityName="Stout" authorityYear="1912" box="[320,629,1688,1712]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="uncinata">
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<emphasis id="5FA9C977FF907E4D7A1382A2FE11F928" box="[320,457,1688,1712]" italics="true" pageId="3" pageNumber="4">C. uncinata</emphasis>
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(Stout, 1912)
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</taxonomicName>
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. However, this was later overlooked in the synonymy and generic diagnoses proposed by Barnard & Karaman (1991) and also Poore & Lowry (1997). In Poore & Lowry (1997), two species with a multi-articulate accessory flagellum on antenna 1,
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<taxonomicName id="AADD6EE6FF907E4D782F82DBFC23F960" box="[892,1019,1761,1784]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="uncinata">
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<emphasis id="5FA9C977FF907E4D782F82DBFC23F960" box="[892,1019,1761,1784]" italics="true" pageId="3" pageNumber="4">P. uncinata</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="AADD6EE6FF907E4D7F6782DBFB79F960" box="[1076,1185,1761,1784]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="variata">
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<emphasis id="5FA9C977FF907E4D7F6782DBFB79F960" box="[1076,1185,1761,1784]" italics="true" pageId="3" pageNumber="4">P. variata</emphasis>
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</taxonomicName>
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, were transferred into
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<taxonomicName id="AADD6EE6FF907E4D7BC4833EFEC2F885" box="[151,282,1796,1821]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D7BC4833EFEC2F885" box="[151,282,1796,1821]" italics="true" pageId="3" pageNumber="4">Paragrubia</emphasis>
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</taxonomicName>
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. Consequently, the diagnosis of
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<taxonomicName id="AADD6EE6FF907E4D79D8833EFCD6F885" box="[651,782,1796,1821]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D79D8833EFCD6F885" box="[651,782,1796,1821]" italics="true" pageId="3" pageNumber="4">Paragrubia</emphasis>
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</taxonomicName>
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was expanded to include gnathopod 1 larger or stouter or subequal in size to gnathopod 2 to accommodate these two species.
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<taxonomicName id="AADD6EE6FF907E4D78EF831DFBEAF8D8" box="[956,1074,1831,1856]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D78EF831DFBEAF8D8" box="[956,1074,1831,1856]" italics="true" pageId="3" pageNumber="4">Cymadusa</emphasis>
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</taxonomicName>
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was also diagnosed to include gnathopod 1 as smaller than or subequal to gnathopod 2, which created overlap between gnathopod characterization of
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<taxonomicName id="AADD6EE6FF907E4D7A298355FE25F810" box="[378,509,1903,1928]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D7A298355FE25F810" box="[378,509,1903,1928]" italics="true" pageId="3" pageNumber="4">Paragrubia</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="AADD6EE6FF907E4D796D8355FC0EF810" authority="Poore & Lowry 1997" authorityName="Poore & Lowry" authorityYear="1997" box="[574,982,1903,1928]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D796D8355FD6CF810" box="[574,692,1903,1928]" italics="true" pageId="3" pageNumber="4">Cymadusa</emphasis>
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(Poore & Lowry 1997)
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</taxonomicName>
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. Currently, the two characters used to separate
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<taxonomicName id="AADD6EE6FF907E4D7BA983AEFEA8F835" box="[250,368,1940,1965]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D7BA983AEFEA8F835" box="[250,368,1940,1965]" italics="true" pageId="3" pageNumber="4">Cymadusa</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="AADD6EE6FF907E4D7AE783AEFDEFF835" box="[436,567,1940,1965]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF907E4D7AE783AEFDEFF835" box="[436,567,1940,1965]" italics="true" pageId="3" pageNumber="4">Paragrubia</emphasis>
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</taxonomicName>
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are the relative sizes of gnathopods 1 and 2, and the number of articles in the antenna 1 accessory flagellum.
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</paragraph>
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<paragraph id="6D621565FF977E4A7B9484A2FCDAFE35" blockId="4.[151,1436,151,1509]" pageId="4" pageNumber="5">
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In an assessment of new and known species that best correspond to the ampithoid genera
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<taxonomicName id="AADD6EE6FF977E4A7FA184ADFAB0FF28" box="[1266,1384,151,176]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF977E4A7FA184ADFAB0FF28" box="[1266,1384,151,176]" italics="true" pageId="4" pageNumber="5">Cymadusa</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="AADD6EE6FF977E4A7BC48486FEC2FF4D" box="[151,282,188,213]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="5FA9C977FF977E4A7BC48486FEC2FF4D" box="[151,282,188,213]" italics="true" pageId="4" pageNumber="5">Paragrubia</emphasis>
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</taxonomicName>
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, the number of articles in the antenna 1 accessory flagellum range from scale-like to 1, 3, 4, 6, 7 and 10, forming an almost continuous series.
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<collectingCountry id="15CA55F5FF977E4A793F84DAFD53FF60" box="[620,651,224,248]" name="American Samoa" pageId="4" pageNumber="5">As</collectingCountry>
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there is no clear break in the range of articles present, delimiting the two genera based on the number of articles would be largely arbitrary. A mutli-articulate accessory flagellum was initially thought to be only associated with gnathopod 1 dominance over gnathopod 2; however, new species as well as previously described species do not conform to this pattern. The opposing character combinations of a uniarticulate flagellum with dominant gnathopod 1 and a multi-articulate flagellum with dominant gnathopod 2 are now recognised among the new species identified here.
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</paragraph>
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<paragraph id="6D621565FF977E4A7B94858DFDC0FD1D" blockId="4.[151,1436,151,1509]" pageId="4" pageNumber="5">
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Presently gnathopod 1 and 2 sizes are assessed in relation to one another, gnathopod 1 is larger than gnathopod 2, gnathopod 1 is smaller than gnathopod 2, gnathopod 1 is subequal to gnathopod 2. This character delineation does not acknowledge the independence in development of these limbs. Within the newly described species, gnathopod 1 is enlarged (
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<taxonomicName id="AADD6EE6FF977E4A7AE2861FFDFAFDA4" box="[433,546,549,572]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="apoorei">
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<emphasis id="5FA9C977FF977E4A7AE2861FFDFAFDA4" box="[433,546,549,572]" italics="true" pageId="4" pageNumber="5">P. apoorei</emphasis>
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</taxonomicName>
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), but also gnathopod 2 is reduced (
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<taxonomicName id="AADD6EE6FF977E4A78F8861FFB86FDA5" box="[939,1118,548,573]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="springthorpei">
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<emphasis id="5FA9C977FF977E4A78F8861FFB86FDA5" box="[939,1118,548,573]" italics="true" pageId="4" pageNumber="5">P. springthorpei</emphasis>
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</taxonomicName>
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). Equally, for species where gnathopod 1 and 2 are of similar size, they may both be enlarged or both reduced (
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<taxonomicName id="AADD6EE6FF977E4A7FDE867DFABFFDF8" box="[1165,1383,583,608]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pilipes">
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<emphasis id="5FA9C977FF977E4A7FDE867DFABFFDF8" box="[1165,1383,583,608]" italics="true" pageId="4" pageNumber="5">Paragrubia pilipes</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName id="AADD6EE6FF977E4A7BC48656FEA9FD1D" box="[151,369,620,645]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bilobata">
|
||
<emphasis id="5FA9C977FF977E4A7BC48656FEA9FD1D" box="[151,369,620,645]" italics="true" pageId="4" pageNumber="5">Cymadusa bilobata</emphasis>
|
||
</taxonomicName>
|
||
, respectively).
|
||
</paragraph>
|
||
<paragraph id="6D621565FF977E4A7B9486AAFCF6FC50" blockId="4.[151,1436,151,1509]" pageId="4" pageNumber="5">
|
||
We propose that the enlargement of gnathopod 1 and 2 be assessed separately.
|
||
<collectingCountry id="15CA55F5FF977E4A7F3586AAFB5DFD30" box="[1126,1157,656,680]" name="American Samoa" pageId="4" pageNumber="5">As</collectingCountry>
|
||
independent characters, gnathopods may be either feeble/diminutive or stout/robust. This new character concept more clearly defines the generic diagnosis of
|
||
<taxonomicName id="AADD6EE6FF977E4A7ACA86EDFDD7FD68" box="[409,527,727,752]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A7ACA86EDFDD7FD68" box="[409,527,727,752]" italics="true" pageId="4" pageNumber="5">Cymadusa</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="AADD6EE6FF977E4A793686EDFD30FD68" box="[613,744,727,752]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A793686EDFD30FD68" box="[613,744,727,752]" italics="true" pageId="4" pageNumber="5">Paragrubia</emphasis>
|
||
</taxonomicName>
|
||
within the
|
||
<taxonomicName id="AADD6EE6FF977E4A78D286EDFBCFFD68" box="[897,1047,727,752]" class="Malacostraca" family="Ampithoidae" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Ampithoidae</taxonomicName>
|
||
. The relative size difference of gnathopod 1, in comparision to gnathopod 2, is now considered to be of secondary importance. This subtle but important shift in character emphasis removes previous taxonomic ambiguity between
|
||
<taxonomicName id="AADD6EE6FF977E4A7F368725FB03FCA0" box="[1125,1243,799,824]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A7F368725FB03FCA0" box="[1125,1243,799,824]" italics="true" pageId="4" pageNumber="5">Cymadusa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AADD6EE6FF977E4A7E418725FA4DFCA0" box="[1298,1429,799,824]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A7E418725FA4DFCA0" box="[1298,1429,799,824]" italics="true" pageId="4" pageNumber="5">Paragrubia</emphasis>
|
||
</taxonomicName>
|
||
. The enlargement or reduction of gnathopods 1 and 2 is recognised as having generic level importance elsewhere in the Corophiidira (
|
||
<taxonomicName id="AADD6EE6FF977E4A7A0D875DFE66FC18" box="[350,446,871,896]" class="Malacostraca" family="Aoridae" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Aoridae</taxonomicName>
|
||
), but limited/poor character description and assessment has hampered the recognition of this generic level character within the
|
||
<taxonomicName id="AADD6EE6FF977E4A791687B6FD03FC3D" box="[581,731,908,933]" class="Malacostraca" family="Ampithoidae" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Ampithoidae</taxonomicName>
|
||
. The enlargement or reduction in the gnathopods is considered here, also, to be of generic importance for the
|
||
<taxonomicName id="AADD6EE6FF977E4A79CF8795FCF6FC50" box="[668,814,943,968]" class="Malacostraca" family="Ampithoidae" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Ampithoidae</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="6D621565FF977E4A7B9487EEFE4CFBE5" blockId="4.[151,1436,151,1509]" pageId="4" pageNumber="5">
|
||
Based on the above character information gained from the new species, the current generic diagnosis suggested by Lowry & Poore (1997) is no longer appropriate.
|
||
<taxonomicName id="AADD6EE6FF977E4A798487CDFC95FB88" box="[727,845,1015,1040]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A798487CDFC95FB88" box="[727,845,1015,1040]" italics="true" pageId="4" pageNumber="5">Cymadusa</emphasis>
|
||
</taxonomicName>
|
||
is rediagnosed to accommodate species with either a scale-like, 1, 2 or multi-articulate accessory flagellum; gnathopod 1 not stout or robust; gnathopod 1 smaller or subequal to 2; gnathopod 1 palm acute; pereopods 5–7 normal to prehensile and uropod 1 with distoventral spine; and telson trapezoidal.
|
||
</paragraph>
|
||
<paragraph id="6D621565FF977E4A7B9480BDFA89FB70" blockId="4.[151,1436,151,1509]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="AADD6EE6FF977E4A7B9480BDFD85FB38" authority="Chevreux (1901)" authorityName="Chevreux" authorityYear="1901" box="[199,605,1159,1184]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A7B9480BDFE92FB38" box="[199,330,1159,1184]" italics="true" pageId="4" pageNumber="5">Paragrubia</emphasis>
|
||
sensu Chevreux (1901)
|
||
</taxonomicName>
|
||
is reinstated here and expanded: gnathopod 1 enlarged to appear stout/ robust; gnathopod 1 is more robust than gnathopod 2. The antenna 1 accessory flagellum is present, and either single or multi-articulate. The diagnosis is expanded to include both subovate and trapezoidal telson shapes.
|
||
</paragraph>
|
||
<paragraph id="6D621565FF977E4A7B9480CEFB7DFA7C" blockId="4.[151,1436,151,1509]" pageId="4" pageNumber="5">
|
||
This new diagnosis for
|
||
<taxonomicName id="AADD6EE6FF977E4A7A8680CEFD80FA95" box="[469,600,1268,1293]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A7A8680CEFD80FA95" box="[469,600,1268,1293]" italics="true" pageId="4" pageNumber="5">Paragrubia</emphasis>
|
||
</taxonomicName>
|
||
affects the placement of three previously described species.
|
||
<emphasis id="5FA9C977FF977E4A7E4380CEFA44FA95" box="[1296,1436,1268,1293]" italics="true" pageId="4" pageNumber="5">Ampithoides</emphasis>
|
||
<taxonomicName id="AADD6EE6FF977E4A7BC4812DFE0BFAA8" authority="Ledoyer, 1967" authorityName="Ledoyer" authorityYear="1967" box="[151,467,1303,1328]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mahafalensis">
|
||
<emphasis id="5FA9C977FF977E4A7BC4812DFEF3FAA8" box="[151,299,1303,1328]" italics="true" pageId="4" pageNumber="5">mahafalensis</emphasis>
|
||
Ledoyer, 1967
|
||
</taxonomicName>
|
||
previously in
|
||
<taxonomicName id="AADD6EE6FF977E4A7921812DFD30FAA8" box="[626,744,1303,1328]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A7921812DFD30FAA8" box="[626,744,1303,1328]" italics="true" pageId="4" pageNumber="5">Cymadusa</emphasis>
|
||
</taxonomicName>
|
||
is transferred to
|
||
<taxonomicName id="AADD6EE6FF977E4A78F7812DFBFFFAA8" box="[932,1063,1303,1328]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A78F7812DFBFFFAA8" box="[932,1063,1303,1328]" italics="true" pageId="4" pageNumber="5">Paragrubia</emphasis>
|
||
</taxonomicName>
|
||
based on the robust gnathopod 1.
|
||
<taxonomicName id="AADD6EE6FF977E4A7BC48106FDE0FACC" authority="Stout, 1912" authorityName="Stout" authorityYear="1912" box="[151,568,1340,1365]" class="Malacostraca" family="Ampithoidae" genus="Acanthogrubia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="uncinata">
|
||
<emphasis id="5FA9C977FF977E4A7BC48106FE72FACC" box="[151,426,1340,1365]" italics="true" pageId="4" pageNumber="5">Acanthogrubia uncinata</emphasis>
|
||
Stout, 1912
|
||
</taxonomicName>
|
||
, placed in
|
||
<taxonomicName id="AADD6EE6FF977E4A79E58106FCE1FACD" box="[694,825,1340,1365]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A79E58106FCE1FACD" box="[694,825,1340,1365]" italics="true" pageId="4" pageNumber="5">Paragrubia</emphasis>
|
||
</taxonomicName>
|
||
by Shoemaker (1941) and Poore & Lowry (1997), is now transferred back to the
|
||
<taxonomicName id="AADD6EE6FF977E4A7A848165FD95FAE0" box="[471,589,1375,1400]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A7A848165FD95FAE0" box="[471,589,1375,1400]" italics="true" pageId="4" pageNumber="5">Cymadusa</emphasis>
|
||
</taxonomicName>
|
||
based on the gnathopod 1 being smaller than gnathopod 2. Lastly,
|
||
<taxonomicName id="AADD6EE6FF977E4A7E188165FE5EFA05" authority="Sheard, 1936" authorityName="Sheard" authorityYear="1936" class="Malacostraca" family="Ampithoidae" genus="Grubia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="variata">
|
||
<emphasis id="5FA9C977FF977E4A7E188165FF30FA04" italics="true" pageId="4" pageNumber="5">Grubia variata</emphasis>
|
||
Sheard, 1936
|
||
</taxonomicName>
|
||
is transferred from
|
||
<taxonomicName id="AADD6EE6FF977E4A793881BEFD36FA05" box="[619,750,1412,1437]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A793881BEFD36FA05" box="[619,750,1412,1437]" italics="true" pageId="4" pageNumber="5">Paragrubia</emphasis>
|
||
</taxonomicName>
|
||
as designated by Poore & Lowry (1997) back to
|
||
<taxonomicName id="AADD6EE6FF977E4A7E7581BEFA44FA05" box="[1318,1436,1412,1437]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5FA9C977FF977E4A7E7581BEFA44FA05" box="[1318,1436,1412,1437]" italics="true" pageId="4" pageNumber="5">Cymadusa</emphasis>
|
||
</taxonomicName>
|
||
based on the feeble/diminutive gnathopod 1, which is also smaller than gnathopod 2. The new combinations for these taxa are
|
||
<taxonomicName id="AADD6EE6FF977E4A7A6581F7FE3EFA7D" box="[310,486,1484,1509]" class="Malacostraca" family="Ampithoidae" genus="Paragrubia" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mahafalensis" status="comb. nov.">
|
||
<emphasis id="5FA9C977FF977E4A7A6581F7FE3EFA7D" box="[310,486,1484,1509]" italics="true" pageId="4" pageNumber="5">P. mahafalensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="449A740CFF977E4A7ABF81F6FDB3FA7C" box="[492,619,1484,1509]" pageId="4" pageNumber="5">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="AADD6EE6FF977E4A792581F7FD20FA7C" box="[630,760,1485,1508]" class="Malacostraca" family="Ampithoidae" genus="Acanthogrubia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="uncinata" status="comb. nov.">
|
||
<emphasis id="5FA9C977FF977E4A792581F7FD20FA7C" box="[630,760,1485,1508]" italics="true" pageId="4" pageNumber="5">C. uncinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="449A740CFF977E4A79AC81F6FCA3FA7C" box="[767,891,1484,1509]" pageId="4" pageNumber="5">comb. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="AADD6EE6FF977E4A78E281F7FBFBFA7C" box="[945,1059,1485,1508]" class="Malacostraca" family="Ampithoidae" genus="Grubia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="variata" status="comb. nov.">
|
||
<emphasis id="5FA9C977FF977E4A78E281F7FBFBFA7C" box="[945,1059,1485,1508]" italics="true" pageId="4" pageNumber="5">C. variata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="449A740CFF977E4A7F7981F6FB7DFA7C" box="[1066,1189,1484,1509]" pageId="4" pageNumber="5">comb. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |