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<document id="D6FBEE909D708BA0DAC1B5382B133D3F" ID-DOI="10.11646/zootaxa.4039.1.1" ID-GBIF-Dataset="4ecc5394-2569-4cbd-8790-a3a4301d304a" ID-ISSN="1175-5326" ID-Zenodo-Dep="289932" ID-ZooBank="CB081EA2-3F93-4FD3-8AC8-B24F772833FC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461319549927" checkinUser="plazi" docAuthor="Plant, Adrian R." docDate="2015" docId="D96487F99918FFA107E94C95AC398117" docLanguage="en" docName="zt04039p056.pdf" docOrigin="Zootaxa 4039 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hemerodromia etalutea Plant, 2015, sp. nov." docType="treatment" docVersion="6" lastPageNumber="27" masterDocId="255DFF819901FFBB077E4E17A842800A" masterDocTitle="Diversity of Hemerodromia Meigen, 1822 (Diptera: Empididae) in Thailand, the tip of a tropical iceberg?" masterLastPageNumber="56" masterPageNumber="1" pageNumber="26" updateTime="1698618296215" updateUser="plazi">
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<mods:title id="0CA23ADD6D3CF60BA64F23D44B47B470">Diversity of Hemerodromia Meigen, 1822 (Diptera: Empididae) in Thailand, the tip of a tropical iceberg?</mods:title>
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<mods:namePart id="8717DAB7F34923F106CB8051B4EEDB82">Plant, Adrian R.</mods:namePart>
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<mods:part id="10FA3E7150D6C276588166DFE86C85CB">
<mods:date id="9991E966F793786B62402B956081576F">2015</mods:date>
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<mods:number id="BA366245DF098D657A91C7F10BE96063">4039</mods:number>
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<mods:identifier id="7D4905CA15C1E982C645891F2B3587B6" type="DOI">10.11646/zootaxa.4039.1.1</mods:identifier>
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<treatment id="D96487F99918FFA107E94C95AC398117" ID-DOI="http://doi.org/10.5281/zenodo.6121222" ID-GBIF-Taxon="127693989" ID-Zenodo-Dep="6121222" LSID="urn:lsid:plazi:treatment:D96487F99918FFA107E94C95AC398117" httpUri="http://treatment.plazi.org/id/D96487F99918FFA107E94C95AC398117" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<subSubSection id="19D765649918FFA207E94C95A91282B4" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="517236EF9918FFA207E94C95AA588296" blockId="25.[151,538,642,702]" box="[151,538,642,668]" pageId="25" pageNumber="26">
<heading id="0A3A81839918FFA207E94C95AA588296" bold="true" box="[151,538,642,668]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="63B9EAFD9918FFA207E94C95AA588296" bold="true" box="[151,538,642,668]" pageId="25" pageNumber="26">
<taxonomicName id="96CD4D6C9918FFA207E94C95A9F48296" ID-CoL="9HTJH" box="[151,438,642,668]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="etalutea" status="sp. nov.">
<emphasis id="63B9EAFD9918FFA207E94C95A9F48296" bold="true" box="[151,438,642,668]" italics="true" pageId="25" pageNumber="26">Hemerodromia etalutea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="788A57869918FFA206C34C95AA588296" box="[445,538,642,668]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="517236EF9918FFA207E94CB2A91282B4" blockId="25.[151,538,642,702]" box="[151,336,677,702]" pageId="25" pageNumber="26">
(
<figureCitation id="C9F62A6A9918FFA207E14CB2A96282B4" box="[159,288,677,702]" captionStart="FIGURES 37 – 39" captionStartId="26.[151,264,1117,1140]" captionTargetBox="[163,1427,353,1093]" captionTargetId="figure@26.[150,1437,339,1096]" captionTargetPageId="26" captionText="FIGURES 37 – 39. Male terminalia of Hemerodromia etalutea sp. nov. 37, lateral view of terminalia; 38, dorsal view of cerci; 39, apex of surstylus. Abbreviations: cer = cerci, epa = epandrium, hyp = hypandrium, sur = surstylus." httpUri="https://zenodo.org/record/289945/files/figure.png" pageId="25" pageNumber="26">Figs 37–39</figureCitation>
,
<figureCitation id="C9F62A6A9918FFA206554CB1A90A82B4" box="[299,328,678,702]" captionStart="FIGURES 82 – 90" captionStartId="48.[151,264,1913,1936]" captionTargetBox="[292,1274,205,1822]" captionTargetId="figure@48.[277,1305,193,1856]" captionTargetPageId="48" captionText="FIGURES 82 – 90. Distribution of Hemerodromia spp. in Thailand at a resolution of 0.25 ° and symbols sized to 0.5 ° of latitude and longitude. 82, H. demissa sp. nov.; 83, H. epsilutea sp. nov.; 84, H. etalutea sp. nov.; 85, H. flaviventris Yang &amp; Yang; 86, H. furcata Grootaert, Yang &amp; Saigusa; 87; H. fusca Yang &amp; Yang; 88, H. gammalutea sp. nov.; 89, H. isochita sp. nov.; 90, H. namtokhinpoon sp. nov." httpUri="https://zenodo.org/record/289960/files/figure.png" pageId="25" pageNumber="26">84</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="19D765649918FFA207E94CF1ADCB8329" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="517236EF9918FFA207E94CF1ADCB8329" blockId="25.[151,1437,742,2002]" pageId="25" pageNumber="26">
<emphasis id="63B9EAFD9918FFA207E94CF1A94882F5" bold="true" box="[151,266,742,767]" pageId="25" pageNumber="26">Diagnosis</emphasis>
. A species with yellow thorax with only sutures about laterotergite blackish. Sct present. Cell bm+dm long. Wing veins entirely yellow. Abdomen with tergite 2 yellow. Epandrium strongly concave on dorsal margin.
</paragraph>
</subSubSection>
<subSubSection id="19D765649918FFA207B94D39AD518714" pageId="25" pageNumber="26" type="description">
<paragraph id="517236EF9918FFA207B94D39AD518714" blockId="25.[151,1437,742,2002]" pageId="25" pageNumber="26">
<emphasis id="63B9EAFD9918FFA207B94D39A913834D" bold="true" box="[199,337,814,839]" pageId="25" pageNumber="26">Description</emphasis>
.
<emphasis id="63B9EAFD9918FFA2061E4D39A9DF834D" bold="true" box="[352,413,814,839]" pageId="25" pageNumber="26">Male</emphasis>
: Body length 2.6–3.0 mm; wing length
<quantity id="96359B0A9918FFA2041B4D39ABFD834C" box="[869,959,814,839]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="25" pageNumber="26" unit="mm" value="2.5">2.5 mm</quantity>
.
<emphasis id="63B9EAFD9918FFA204B44D39AC4B834D" bold="true" box="[970,1033,814,839]" pageId="25" pageNumber="26">Head</emphasis>
. Black, dusted greyish; frons paler, brownish in front of ocelli, becoming whitish near base of antenna; antenna and mouthparts pale whitish yellow, all setae pale; one pair proclinate ocl; postocular setae uniseriate, merging above with ~1–2 pairs small vtl and several fine setulae on vertex; frontal setulae apparently absent. Antenna with postpedicel ~2.5–3.0X long as wide, a few minute outstanding setulae subapically; stylus shorter, ~0.4–0.45X long as postpedicel; scape with distinct fine dorsal seta.
<emphasis id="63B9EAFD9918FFA206604DF5A93583F1" bold="true" box="[286,375,994,1019]" pageId="25" pageNumber="26">Thorax</emphasis>
. Dusted greyish. Ground colour clear yellow including mediotergite and prothoracic ‘collar’; only sutures about laterotergite blackish. All setae yellowish; minute apart from a pair of distinct sct and one npl.
<emphasis id="63B9EAFD9918FFA207E94A3DA88D8449" bold="true" box="[151,207,1066,1091]" pageId="25" pageNumber="26">Legs</emphasis>
. Pale yellow. C1 ~1.2X longer than distance between C1 and C2, a few distinct setulae dorsally and apically. F1 ~ 1.2X long as C1, ~
<date id="2573102F9918FFA206D64A59A988846D" box="[424,458,1102,1127]" pageId="25" pageNumber="26">6X</date>
long as wide, evenly inflated, hardly constricted on proximal 0.3; femoral formula ~ 7–
<date id="2573102F9918FFA207E94A65A8E68481" box="[151,164,1138,1163]" pageId="25" pageNumber="26" value="1918-08-18" valueMax="1918-08-19" valueMin="1918-08-18">8</date>
/18–19/18–19+1–2/6–8, denticles black, rows converging apically; spines yellow, pv and av rows rather short, 0.3–0.5X long as limb is deep, becoming somewhat longer proximally with basal spine of pv row much stronger. T1 ~ 0.6–0.7X long as F1, evenly curved, ventral face shallowly concave; with one row of ~ 15 sharply pointed spinose setae ventrally; ventroapical spur extremely small with strong erect black spinose seta arising from near base; a ciliation of short setulae dorsally on distal 0.3. Mid and hind legs slender with only small pale setulae.
<emphasis id="63B9EAFD9918FFA207E94B31A89A8535" bold="true" box="[151,216,1318,1343]" pageId="25" pageNumber="26">Wing</emphasis>
. Membrane with yellow microtrichia but hyaline basally. Veins yellow, paler near base of wing. Marginal setulae pale. R2+3 almost linear, joining C ~ 0.6X distance between end of R1 and R4; length of C between ends of R2+3 and R4 ~1.8–2.0X long as R4; R4 slightly S-shaped, angle with R5 at extreme base ~ 85°, becoming ~70–80° thereafter, joining C at ~ 85°. R5 ~ 2.0–2.5X long as R4, almost linear; R4+5 fork distal to M1+2 fork by ~
<date id="2573102F9918FFA202714B8FAD7385BA" box="[1295,1329,1432,1456]" pageId="25" pageNumber="26">1X</date>
length of R4. M1 slightly curved, convergent distally with R5; cell bm+dm long, ending beyond tip of R1. Halter yellowish white. Squama with margin yellowish.
<emphasis id="63B9EAFD9918FFA205254BF2AA9285F4" bold="true" box="[603,720,1509,1534]" pageId="25" pageNumber="26">Abdomen</emphasis>
. Yellow with tergites 3–5 and 8 blackish; tergite 2 sometimes faintly darker medially. Tergite 8 with dark bristly setae on posterior margin; tergite 7 with a few smaller yellow setae on disc.
<emphasis id="63B9EAFD9918FFA20647483AA9FE864C" bold="true" box="[313,444,1581,1606]" pageId="25" pageNumber="26">Terminalia</emphasis>
. Cercus of complex structure; slightly shorter than epandrium; narrow distally with tip upturned in lateral view (
<figureCitation id="C9F62A6A9918FFA206C64846AA4F8660" box="[440,525,1617,1642]" captionStart="FIGURES 37 – 39" captionStartId="26.[151,264,1117,1140]" captionTargetBox="[163,1427,353,1093]" captionTargetId="figure@26.[150,1437,339,1096]" captionTargetPageId="26" captionText="FIGURES 37 – 39. Male terminalia of Hemerodromia etalutea sp. nov. 37, lateral view of terminalia; 38, dorsal view of cerci; 39, apex of surstylus. Abbreviations: cer = cerci, epa = epandrium, hyp = hypandrium, sur = surstylus." httpUri="https://zenodo.org/record/289945/files/figure.png" pageId="25" pageNumber="26">Fig. 37</figureCitation>
); 2 small subtriangular dorsal process on proximal 0.5 with a short row of very fine setulae adjacent on outer face (
<figureCitation id="C9F62A6A9918FFA205434862AA8A8684" box="[573,712,1653,1678]" captionStart="FIGURES 37 – 39" captionStartId="26.[151,264,1117,1140]" captionTargetBox="[163,1427,353,1093]" captionTargetId="figure@26.[150,1437,339,1096]" captionTargetPageId="26" captionText="FIGURES 37 – 39. Male terminalia of Hemerodromia etalutea sp. nov. 37, lateral view of terminalia; 38, dorsal view of cerci; 39, apex of surstylus. Abbreviations: cer = cerci, epa = epandrium, hyp = hypandrium, sur = surstylus." httpUri="https://zenodo.org/record/289945/files/figure.png" pageId="25" pageNumber="26">Figs 37, 38</figureCitation>
). Epandrium with upper margin distinctly and rather abruptly curved beyond middle; a cluster of 3–4 strong setae apically. Surstylus present, not reaching beyond tip of epandrium; apically inflated with several strong black peg-like modified setae around tip (
<figureCitation id="C9F62A6A9918FFA2030548AAAC8F86DC" box="[1147,1229,1725,1750]" captionStart="FIGURES 37 – 39" captionStartId="26.[151,264,1117,1140]" captionTargetBox="[163,1427,353,1093]" captionTargetId="figure@26.[150,1437,339,1096]" captionTargetPageId="26" captionText="FIGURES 37 – 39. Male terminalia of Hemerodromia etalutea sp. nov. 37, lateral view of terminalia; 38, dorsal view of cerci; 39, apex of surstylus. Abbreviations: cer = cerci, epa = epandrium, hyp = hypandrium, sur = surstylus." httpUri="https://zenodo.org/record/289945/files/figure.png" pageId="25" pageNumber="26">Fig. 39</figureCitation>
). A pale yellowish (almost transparent) dorsoapically pointed process emerging slightly beyond tip of epandrium. Hypandrium small, angular, subquadrate with a few distinct setae proximally.
<emphasis id="63B9EAFD9918FFA204644912AB328714" bold="true" box="[794,880,1797,1822]" pageId="25" pageNumber="26">Female</emphasis>
: Similar to male; terminalia blackish.
</paragraph>
</subSubSection>
<subSubSection id="19D765649918FFA207B9493EAA438780" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="517236EF9918FFA207B9493EAA438780" blockId="25.[151,1437,742,2002]" pageId="25" pageNumber="26">
<emphasis id="63B9EAFD9918FFA207B9493EA92F8748" bold="true" box="[199,365,1833,1858]" pageId="25" pageNumber="26">
<typeStatus id="8E76884D9918FFA207B9493EA9408748" box="[199,258,1833,1858]" pageId="25" pageNumber="26">Type</typeStatus>
material
</emphasis>
.
<emphasis id="63B9EAFD9918FFA20605493EAA588748" bold="true" box="[379,538,1833,1858]" pageId="25" pageNumber="26">
<typeStatus id="8E76884D9918FFA20605493EAA588748" box="[379,538,1833,1858]" pageId="25" pageNumber="26" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
♂,
<collectingCountry id="29DA767F9918FFA2053D493DAA9A8748" box="[579,728,1834,1858]" name="Thailand" pageId="25" pageNumber="26">THAILAND</collectingCountry>
: Nakhon Si Thammarat Province, Namtok Yong NP, TV aerial,
<quantity id="96359B0A9918FFA207E9495AA8A2876C" box="[151,224,1869,1894]" metricMagnitude="2" metricUnit="m" metricValue="9.66" pageId="25" pageNumber="26" unit="m" value="966.0">966 m</quantity>
,
<geoCoordinate id="34F950289918FFA20792495AA92F876C" box="[236,365,1869,1894]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="8.2377">8°14.262'N</geoCoordinate>
,
<geoCoordinate id="34F950289918FFA206074959AA44876C" box="[377,518,1870,1894]" direction="east" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="99.80482">99°48.289'E</geoCoordinate>
,
<date id="2573102F9918FFA2056F495AAAEF876C" box="[529,685,1869,1894]" pageId="25" pageNumber="26" value="2008-10-15" valueMax="2008-10-22" valueMin="2008-10-15">15–22.x.2008</date>
, MT, Paiboon [T4226](
<collectionCode id="37DCAE2A9918FFA204C8495AAC44876C" box="[950,1030,1869,1894]" pageId="25" pageNumber="26">QSBG</collectionCode>
).
<emphasis id="63B9EAFD9918FFA20366495AACF8876C" bold="true" box="[1048,1210,1869,1894]" pageId="25" pageNumber="26">
<typeStatus id="8E76884D9918FFA20366495AACF8876C" box="[1048,1210,1869,1894]" pageId="25" pageNumber="26" type="paratype">PARATYPES</typeStatus>
</emphasis>
:
<specimenCount id="47CBFD669918FFA203B54959ACB2876C" box="[1227,1264,1870,1894]" pageId="25" pageNumber="26" type="generic">3♂</specimenCount>
, 2♀, same data as
<typeStatus id="8E76884D9918FFA207CB4966A9558780" box="[181,279,1905,1930]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
(
<collectionCode id="37DCAE2A9918FFA206584965A9308780" box="[294,370,1906,1930]" pageId="25" pageNumber="26">QSBG</collectionCode>
&amp;
<collectionCode id="37DCAE2A9918FFA206E84965A9B48780" box="[406,502,1906,1930]" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="25" pageNumber="26">NMWC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="19D765649918FFA207B94982AAF787D8" pageId="25" pageNumber="26" type="etymology">
<paragraph id="517236EF9918FFA207B94982AAF787D8" blockId="25.[151,1437,742,2002]" pageId="25" pageNumber="26">
<emphasis id="63B9EAFD9918FFA207B94982A90487A4" bold="true" box="[199,326,1941,1966]" pageId="25" pageNumber="26">Etymology</emphasis>
. The specific epithet is a concatenation of the Latin
<emphasis id="63B9EAFD9918FFA204C84982ABBB87A4" box="[950,1017,1941,1966]" italics="true" pageId="25" pageNumber="26">luteus</emphasis>
and the seventh letter of the Greek alphabet (
<emphasis id="63B9EAFD9918FFA2067849ACA96A87D8" box="[262,296,1979,2002]" italics="true" pageId="25" pageNumber="26">eta</emphasis>
). It is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="19D76564991BFFA107B94E80AC398117" pageId="26" pageNumber="27" type="discussion">
<paragraph id="517236EF991BFFA107B94E80AC398117" blockId="26.[151,1436,151,285]" pageId="26" pageNumber="27">
<emphasis id="63B9EAFD991BFFA107B94E80A97680BA" bold="true" box="[199,308,151,176]" pageId="26" pageNumber="27">Remarks</emphasis>
.
<taxonomicName id="96CD4D6C991BFFA1063C4E80AA0C80BA" box="[322,590,151,176]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="etalutea" status="sp. nov.">
<emphasis id="63B9EAFD991BFFA1063C4E80AA0C80BA" box="[322,590,151,176]" italics="true" pageId="26" pageNumber="27">Hemerodromia etalutea</emphasis>
</taxonomicName>
<emphasis id="63B9EAFD991BFFA1052B4E80AAED80BA" bold="true" box="[597,687,151,176]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="788A5786991BFFA1052B4E80AAED80BA" box="[597,687,151,176]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from other species with an unmarked yellow scutum by having the mediotergite clear yellow and the wing with both veins and membrane yellow but confirmation of identification requires examination of male terminalia. The species is only known from Nakhon Si Thammarat Province in southern
<collectingCountry id="29DA767F991BFFA105704F13AA308117" box="[526,626,260,285]" name="Thailand" pageId="26" pageNumber="27">Thailand</collectingCountry>
(
<figureCitation id="C9F62A6A991BFFA105FE4F13AA908116" box="[640,722,260,285]" captionStart="FIGURES 82 – 90" captionStartId="48.[151,264,1913,1936]" captionTargetBox="[292,1274,205,1822]" captionTargetId="figure@48.[277,1305,193,1856]" captionTargetPageId="48" captionText="FIGURES 82 – 90. Distribution of Hemerodromia spp. in Thailand at a resolution of 0.25 ° and symbols sized to 0.5 ° of latitude and longitude. 82, H. demissa sp. nov.; 83, H. epsilutea sp. nov.; 84, H. etalutea sp. nov.; 85, H. flaviventris Yang &amp; Yang; 86, H. furcata Grootaert, Yang &amp; Saigusa; 87; H. fusca Yang &amp; Yang; 88, H. gammalutea sp. nov.; 89, H. isochita sp. nov.; 90, H. namtokhinpoon sp. nov." httpUri="https://zenodo.org/record/289960/files/figure.png" pageId="26" pageNumber="27">Fig. 84</figureCitation>
) at mid elevation (
<quantity id="96359B0A991BFFA104DA4F13ABAC8116" box="[932,1006,260,285]" metricMagnitude="2" metricUnit="m" metricValue="9.66" pageId="26" pageNumber="27" unit="m" value="966.0">966 m</quantity>
) in October.
</paragraph>
</subSubSection>
</treatment>
</document>
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