170 lines
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170 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.4.60525" ID-GBIF-Dataset="217bea89-5bd1-4ae7-bfce-fb945a933fc4" ID-GBIF-Taxon="176088089" ID-Pensoft-Pub="2535-0730-2-119" ID-Pensoft-UUID="6301B37BBB7D51A185FB132ED24E4E81" ID-ZooBank="8BD62DBF438C46508C4037C837FD573F" LSID="urn:lsid:zoobank.org:act:4D3EE6C7-D220-4324-8279-944C54906E4E" ModsDocID="2535-0730-2-119" checkinTime="1608157809646" checkinUser="pensoft" docAuthor="Ingrisch, Sigfrid" docDate="2020" docId="7CE2240B1EE454DD9311EEF65B34E309" docLanguage="en" docName="EvolutSyst 4(2): 119-132" docOrigin="Evolutionary Systematics 4 (2)" docPubDate="2020-12-15" docSource="http://dx.doi.org/10.3897/evolsyst.4.60525" docTitle="Liara (Indoliara) dividata Ingrisch 2020, sp. nov." docType="treatment" docUuid="4D3EE6C7-D220-4324-8279-944C54906E4E" docUuidSource="ZooBank" docVersion="3" id="6301B37BBB7D51A185FB132ED24E4E81" lastPageNumber="119" masterDocId="6301B37BBB7D51A185FB132ED24E4E81" masterDocTitle="New subgenera and species of Agraeciini (Orthoptera, Tettigoniidae, Conocephalinae) from South Asia found in historical insect collections" masterLastPageNumber="132" masterPageNumber="119" pageId="0" pageNumber="119" updateTime="1643451831480" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New subgenera and species of Agraeciini (Orthoptera, Tettigoniidae, Conocephalinae) from South Asia found in historical insect collections</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ingrisch, Sigfrid</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8624-0472</mods:nameIdentifier>
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<mods:affiliation>Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier type="email">s.ingrisch@leibniz-zfmk.de</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-12-15</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>119</mods:start>
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<mods:end>132</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/evolsyst.4.60525</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.4.60525</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2535-0730-2-119</mods:identifier>
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<mods:identifier type="ZooBank">8BD62DBF438C46508C4037C837FD573F</mods:identifier>
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<mods:identifier type="Pensoft-UUID">6301B37BBB7D51A185FB132ED24E4E81</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="176088089" LSID="urn:lsid:zoobank.org:act:4D3EE6C7-D220-4324-8279-944C54906E4E" httpUri="http://treatment.plazi.org/id/7CE2240B1EE454DD9311EEF65B34E309" lastPageNumber="119" pageId="0" pageNumber="119">
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<subSubSection pageId="0" pageNumber="119" type="nomenclature">
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<paragraph pageId="0" pageNumber="119">
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<taxonomicName LSID="http://zoobank.org/4D3EE6C7-D220-4324-8279-944C54906E4E" authority="Ingrisch, 2020" authorityName="Ingrisch" authorityYear="2020" class="Insecta" family="Tettigoniidae" genus="Liara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liara (Indoliara) dividata" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="dividata" status="sp. nov." subGenus="Indoliara">Liara (Indoliara) dividata</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="119">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Liara (Indoliara) dividata sp. nov. A-G male holotype: A habitus dorsal view; B face; C stridulatory file; D abdominal apex with cerci in ventro-apical view; E titillators of male phallus; F tip of titillators; G subgenital plate. - H-K female: H habitus lateral view; I head in dorsal view; J thoracic sternites; K subgenital plate." httpUri="https://binary.pensoft.net/fig/487062" pageId="0" pageNumber="119">Fig. 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="holotype">
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<paragraph pageId="0" pageNumber="119">Holotype</paragraph>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">(male).</emphasis>
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India: South India, Thekkadi, Periyar Dam (
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<geoCoordinate degrees="9" direction="north" minutes="35" orientation="latitude" precision="925" value="9.583333">9°35'N</geoCoordinate>
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,
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<geoCoordinate degrees="77" direction="east" minutes="9" orientation="longitude" precision="925" value="77.15">77°9'E</geoCoordinate>
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), 6.-10.v.1937, coll. B.M.-C.M. Expedition to South India - depository: BMNH (Lo032S001; B.M.1939-205).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="paratype">
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<paragraph pageId="0" pageNumber="119">Paratype</paragraph>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">(1 female).</emphasis>
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India: South India, Travancore, Pirmed (
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<geoCoordinate degrees="9" direction="north" minutes="40" orientation="latitude" precision="925" value="9.666667">9°40'N</geoCoordinate>
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,
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<geoCoordinate degrees="76" direction="east" minutes="59" orientation="longitude" precision="925" value="76.98333">76°59'E</geoCoordinate>
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), elev. 1036 m, 4.-6.v.1933, coll. B.M.-C.M. Expedition to South India - BMNH (Lo032S002; B.M.1939-205).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="etymology">
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<paragraph pageId="0" pageNumber="119">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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The name of the new species refers to the shape of the fastigium verticis that is divided into two short apical cones; from Latin
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<emphasis italics="true" pageId="0" pageNumber="119">dividere</emphasis>
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to divide.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="diagnosis">
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<paragraph pageId="0" pageNumber="119">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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This is the so far only species in the subgenus. Its differences to the species of the other subgenera of
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<taxonomicName authorityName="Redtenbacher" authorityYear="1891" class="Insecta" family="Tettigoniidae" genus="Liara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liara" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="119">Liara</emphasis>
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</taxonomicName>
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are discussed above under
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<taxonomicName authority="Ingrisch, 2020" authorityName="Ingrisch" authorityYear="2020" class="Insecta" family="Tettigoniidae" genus="Indoliara" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Indoliara" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus">Indoliara</taxonomicName>
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subgen. nov.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="description">
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<paragraph pageId="0" pageNumber="119">Description.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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Fastigium verticis longer than scapus, with a minute tubercle at top; apex bifurcate and slightly S-shaped, separated from fastigium frontis by a shallow furrow, but connected in middle by a lamella (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Liara (Indoliara) dividata sp. nov. A-G male holotype: A habitus dorsal view; B face; C stridulatory file; D abdominal apex with cerci in ventro-apical view; E titillators of male phallus; F tip of titillators; G subgenital plate. - H-K female: H habitus lateral view; I head in dorsal view; J thoracic sternites; K subgenital plate." httpUri="https://binary.pensoft.net/fig/487062" pageId="0" pageNumber="119">5I</figureCitation>
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). Frons rugose (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Liara (Indoliara) dividata sp. nov. A-G male holotype: A habitus dorsal view; B face; C stridulatory file; D abdominal apex with cerci in ventro-apical view; E titillators of male phallus; F tip of titillators; G subgenital plate. - H-K female: H habitus lateral view; I head in dorsal view; J thoracic sternites; K subgenital plate." httpUri="https://binary.pensoft.net/fig/487062" pageId="0" pageNumber="119">5B</figureCitation>
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). Pronotum rugose, funnel-shaped; anterior margin broadly rounded but slightly concave in middle; posterior margin truncate or nearly so; first and second transverse sulci interrupted on disc; shoulders little marked; paranota with anterior and posterior angles rounded; ventral margin very little sinuate, almost straight; tympanal swelling weak but well delimitated; humeral sinus distinct. Prosternum mute; meso- and metasternal lobes conical, short; intermedial plate with small spines at posterior angles (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Liara (Indoliara) dividata sp. nov. A-G male holotype: A habitus dorsal view; B face; C stridulatory file; D abdominal apex with cerci in ventro-apical view; E titillators of male phallus; F tip of titillators; G subgenital plate. - H-K female: H habitus lateral view; I head in dorsal view; J thoracic sternites; K subgenital plate." httpUri="https://binary.pensoft.net/fig/487062" pageId="0" pageNumber="119">5J</figureCitation>
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). Tegmen covering abdomen, regularly narrowing from basal widening to rounded tip (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Liara (Indoliara) dividata sp. nov. A-G male holotype: A habitus dorsal view; B face; C stridulatory file; D abdominal apex with cerci in ventro-apical view; E titillators of male phallus; F tip of titillators; G subgenital plate. - H-K female: H habitus lateral view; I head in dorsal view; J thoracic sternites; K subgenital plate." httpUri="https://binary.pensoft.net/fig/487062" pageId="0" pageNumber="119">5A, H</figureCitation>
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). Femora with the following number of spines on ventral margins: profemur no external, 5-8 internal; mesofemur 6-9 external, no internal; postfemur 10-11 external, no internal; all with irregularly alternating large and minute spines. Knee lobes of profemur obtuse on external, triangular on internal side; of mesofemur obtuse or triangular on external, spinose on internal side; of postfemur spinose on both sides.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/487062" pageId="0" pageNumber="119" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Figure 5.</emphasis>
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<taxonomicName authorityName="Ingrisch" authorityYear="2020" class="Insecta" family="Tettigoniidae" genus="Liara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liara (Indoliara) dividata" order="Orthoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="dividata" subGenus="Indoliara">Liara (Indoliara) dividata</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="119">A-G</emphasis>
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male holotype:
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<emphasis bold="true" pageId="0" pageNumber="119">A</emphasis>
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habitus dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="119">B</emphasis>
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face;
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<emphasis bold="true" pageId="0" pageNumber="119">C</emphasis>
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stridulatory file;
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<emphasis bold="true" pageId="0" pageNumber="119">D</emphasis>
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abdominal apex with cerci in ventro-apical view;
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<emphasis bold="true" pageId="0" pageNumber="119">E</emphasis>
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titillators of male phallus;
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<emphasis bold="true" pageId="0" pageNumber="119">F</emphasis>
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tip of titillators;
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<emphasis bold="true" pageId="0" pageNumber="119">G</emphasis>
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subgenital plate. -
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<emphasis bold="true" pageId="0" pageNumber="119">H-K</emphasis>
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female:
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<emphasis bold="true" pageId="0" pageNumber="119">H</emphasis>
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habitus lateral view;
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<emphasis bold="true" pageId="0" pageNumber="119">I</emphasis>
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head in dorsal view;
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<emphasis bold="true" pageId="0" pageNumber="119">J</emphasis>
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thoracic sternites;
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<emphasis bold="true" pageId="0" pageNumber="119">K</emphasis>
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subgenital plate.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Male.</emphasis>
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Tenth abdominal tergite with short triangular projections above bases of styli; apical margin broadly concave in between. Epiproct wide with rounded lateral lobes [apex hidden]. Paraprocts with compressed, conical projections. Cerci conical, compressed, apex rounded; dorso-external surface concave; ventro-internal surface convex in apical half, with a broad, oblique, transverse furrow in basal half; with an acute spinule behind furrow and a thin process at very base; process conical at base, rectangularly bent dorsad in about middle, strongly compressed thereafter, apex with acute tooth on proximal side (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Liara (Indoliara) dividata sp. nov. A-G male holotype: A habitus dorsal view; B face; C stridulatory file; D abdominal apex with cerci in ventro-apical view; E titillators of male phallus; F tip of titillators; G subgenital plate. - H-K female: H habitus lateral view; I head in dorsal view; J thoracic sternites; K subgenital plate." httpUri="https://binary.pensoft.net/fig/487062" pageId="0" pageNumber="119">5D</figureCitation>
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). Subgenital plate with lateral areas strongly curved dorsad; lateral margins almost straight, approaching each other posteriorly; apex broadly truncate with strong and dense hairs, hairs also on dorsal (= internal) surface in middle of apical area. Styli long, longer than half the length of the subgenital plate, unsymmetrical (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Liara (Indoliara) dividata sp. nov. A-G male holotype: A habitus dorsal view; B face; C stridulatory file; D abdominal apex with cerci in ventro-apical view; E titillators of male phallus; F tip of titillators; G subgenital plate. - H-K female: H habitus lateral view; I head in dorsal view; J thoracic sternites; K subgenital plate." httpUri="https://binary.pensoft.net/fig/487062" pageId="0" pageNumber="119">5G</figureCitation>
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). Titillators separate; rather thin; bases widened and flat, gradually narrowing and approaching to just before apical area; apical area stretched, ovoid, curved laterad, with granular surface (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Liara (Indoliara) dividata sp. nov. A-G male holotype: A habitus dorsal view; B face; C stridulatory file; D abdominal apex with cerci in ventro-apical view; E titillators of male phallus; F tip of titillators; G subgenital plate. - H-K female: H habitus lateral view; I head in dorsal view; J thoracic sternites; K subgenital plate." httpUri="https://binary.pensoft.net/fig/487062" pageId="0" pageNumber="119">5E, F</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Female.</emphasis>
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Tenth abdominal tergite with apical margin concave in middle. Epiproct triangular, with shallow dorsal groove. Cerci conical, apex slightly curved mediad. Subgenital plate wider than long, roughly triangular with apex roundly excised. Ovipositor compressed blade-shaped, highest in middle.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="coloration">
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<paragraph pageId="0" pageNumber="119">Coloration.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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Uniformly light brown (ochre). Frons reddish brown in male; labrum yellow, mandibles black. Tegmen speckled with small brown spots (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Liara (Indoliara) dividata sp. nov. A-G male holotype: A habitus dorsal view; B face; C stridulatory file; D abdominal apex with cerci in ventro-apical view; E titillators of male phallus; F tip of titillators; G subgenital plate. - H-K female: H habitus lateral view; I head in dorsal view; J thoracic sternites; K subgenital plate." httpUri="https://binary.pensoft.net/fig/487062" pageId="0" pageNumber="119">5A, H</figureCitation>
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). Variation: Frons concolorous. Hind area of vertex and anterior area of pronotum with faint brown medial band split by narrow light line. Spines on hind femur and tibia contrasting black.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="description">
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<paragraph pageId="0" pageNumber="119">Measurements.</paragraph>
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<paragraph pageId="0" pageNumber="119">Body w/wings: male 36, female 42; body w/o wings: male 36, female 44; pronotum: male 7.5, female 9.5; tegmen: male 25, female 26.5; hind femur: male 16, female 19.5; ovipositor: female 28 mm long, greatest height 3.2 mm.</paragraph>
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</subSubSection>
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</treatment>
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</document> |