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<mods:url id="79667EC313B5494FF4F5074C97F96A79">http://dx.doi.org/10.3897/phytokeys.240.101716</mods:url>
</mods:location>
<mods:classification id="E33E60185632D29ADFBC778F84D41129">journal article</mods:classification>
<mods:identifier id="FC20617DC75636580CAE2F479D7C8811" type="DOI">http://dx.doi.org/10.3897/phytokeys.240.101716</mods:identifier>
<mods:identifier id="F9F0796C41EA6DB6DEF37F3978ED1066" type="Pensoft-Pub">1314-2003-240-1</mods:identifier>
<mods:identifier id="5BE538A681E87C05919129E52BDB15EE" type="Pensoft-UUID">B699D9DE2B435B1093DE3C38C703D430</mods:identifier>
</mods:mods>
<treatment id="7CD668CF757BE5698446212A3621B20F" LSID="urn:lsid:plazi:treatment:7CD668CF757BE5698446212A3621B20F" httpUri="http://treatment.plazi.org/id/7CD668CF757BE5698446212A3621B20F" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="46C61571139101C733685EE5C68ECED3" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="2FB966216511E252302AD3D3D2DD5A00" pageId="0" pageNumber="1">
Tribe 
<taxonomicName id="C9EDE66139B98092686DD20D39DD19AB" authority="Rchb., Fl. Germ. Excurs. 2 (2): 544. 1832." authorityName="Rchb., Fl. Germ. Excurs. 2 (2): 544. 1832." family="Fabaceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Ceratonieae">Ceratonieae Rchb., Fl. Germ. Excurs. 2(2): 544. 1832.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="0567FE1EBA41E337B2DC5E1EE5FBD19D" pageId="0" pageNumber="1" type="description">
<paragraph id="F9F0592DAE3E879526831D204D67B15C" pageId="0" pageNumber="1">
<figureCitation id="FB84EA659D923E204F6B97E4FE277B39" captionStart="Figure 6" captionStartId="F13" captionText="Figure 6. Generic relationships in tribe Ceratonieae. Left part of figure shows complete genus-level Caesalpinioideae phylogeny with the Ceratonieae indicated with a red rectangle. Branch lengths are expressed in coalescent units and terminal branches were assigned an arbitrary uniform length for visual clarity. Support for relationships is based on fractions of supporting and conflicting gene trees: pie charts show gene tree support and conflict per node (blue representing supporting gene trees, green gene trees supporting the most common alternative topology, red gene trees supporting further alternative topologies, grey gene trees uninformative for this node), and numbers above pie charts are Internode Certainty All support values [both calculated with PhyParts (Smith et al. 2015)]. If present, red numbers below pie charts are non-significant (i. e.&gt; 0.05) outcomes of ASTRAL's polytomy test (Sayyari and Mirarab 2018), which tests for each node whether the polytomy null model can be rejected. Monophyletic genera are represented by single branches; see Suppl. material 2 for a phylogeny with all accessions. See Suppl. material 3 for gene tree support across the phylogeny. The phylogeny is a pruned version of the backbone phylogeny of Ringelberg et al. (2023), where full details of the data and phylogenomic analysis methods are presented." figureDoi="10.3897/phytokeys.240.101716.figure6" httpUri="https://binary.pensoft.net/fig/1014679" pageId="0" pageNumber="1">Figs 6</figureCitation>
<figureCitation id="3D16D26E0AB06218EF6AB9B5FDE50B89" captionStart="Figure 7" captionStartId="F14" captionText="Figure 7. Flower, fruit, and vegetative characters of tribe Ceratonieae A Arcoa gonavensis Urb., foliage and fruits, Dominican Republic B Tetrapterocarpon geayi Humbert, fruits and part of bipinnate leaf, Madagascar (Du Puy M 410) C-E Acrocarpus fraxinifolius Wight &amp; Arn., India, cultivated tree C inflorescence D dehisced fruits and new flush foliage E part of a bipinnate leaf F-J Ceratonia siliqua L. F tree, Jordan, semi-desert near Petra G fruits, Israel, Mt. Scopus Botanical Gardens, Jerusalem H pistillate flowers, Crete I staminate flowers, Crete J staminate inflorescences, Israel, Jerusalem Botanical Gardens. Photo credits A F Jimenez R. B D Du Puy C-E https: // efloraofindia. com / 2011 / 02 / 01 / acrocarpus-fraxinifolius / E D Valke F, G, J O Fragman-Sapir H, I G Lewis." figureDoi="10.3897/phytokeys.240.101716.figure7" httpUri="https://binary.pensoft.net/fig/1014680" pageId="0" pageNumber="1">, 7</figureCitation>
<figureCitation id="92E2350E68BEAADF82E4C26F98F0880A" captionStart="Figure 8" captionStartId="F15" captionText="Figure 8. Distribution of Arcoa based on quality-controlled digitised herbarium records. See Suppl. material 1 for the source of occurrence data." figureDoi="10.3897/phytokeys.240.101716.figure8" httpUri="https://binary.pensoft.net/fig/1014681" pageId="0" pageNumber="1">, 8</figureCitation>
<figureCitation id="39A6E129AD63426F5C14BEC4ADD19DA9" captionStart="Figure 9" captionStartId="F16" captionText="Figure 9. Distribution of Tetrapterocarpon based on quality-controlled digitised herbarium records. See Suppl. material 1 for the source of occurrence data." figureDoi="10.3897/phytokeys.240.101716.figure9" httpUri="https://binary.pensoft.net/fig/1014682" pageId="0" pageNumber="1">, 9</figureCitation>
<figureCitation id="42CA1F7A23CE47EB68EF01555F7B001F" captionStart="Figure 10" captionStartId="F17" captionText="Figure 10. Distribution of Acrocarpus based on quality-controlled digitised herbarium records. See Suppl. material 1 for the source of occurrence data." figureDoi="10.3897/phytokeys.240.101716.figure10" httpUri="https://binary.pensoft.net/fig/1014683" pageId="0" pageNumber="1">, 10</figureCitation>
<figureCitation id="BC8DFBDA5EB3B3E3E9550C7C6B5C1363" captionStart="Figure 11" captionStartId="F18" captionText="Figure 11. Distribution of Ceratonia based on quality-controlled digitised herbarium records. Note that many distribution points observed here are from cultivated Ceratonia specimens. The true native range of the genus Ceratonia is likely to be more restricted than depicted on the map. See Suppl. material 1 for the source of occurrence data." figureDoi="10.3897/phytokeys.240.101716.figure11" httpUri="https://binary.pensoft.net/fig/1014684" pageId="0" pageNumber="1">, 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="11C52BDDFA866F9AAA9A67AD3ABDBE18" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="2CA02AF582615935609218CB6F7D8BF9" pageId="0" pageNumber="1">
<taxonomicName id="2B1C4678DF547B19EB467FE5D5945798" family="Fabaceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="subTribe" subTribe="Ceratoniinae">Ceratoniinae</taxonomicName>
H.S. Irwin &amp; Barneby in R.M. Polhill &amp; P.H. Raven, Adv. Legume System. 1: 98. 1981.
</paragraph>
</subSubSection>
<subSubSection id="E6F64FFB04701065F34D06C27072A74A" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="E7E7C0DF807F528B713DE7D25C682A5E" pageId="0" pageNumber="1">
<typeStatus id="760556BB39D7B09FBBBF394ADEFF39E0">Type</typeStatus>
.
</paragraph>
<paragraph id="546634C12A1C21C607D99D7A476B1CA9" pageId="0" pageNumber="1">
<taxonomicName id="AA7591F02EEB07D424EB83FEC789CA8A" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="36F1F4A554CFEAFE97AC5A4528B985E2" italics="true" pageId="0" pageNumber="1">Ceratonia</emphasis>
</taxonomicName>
L.
</paragraph>
</subSubSection>
<subSubSection id="451729848F295EFB0E9DAC5C10806D68" pageId="0" pageNumber="1" type="included genera">
<paragraph id="DE4BCCEA670D7B54461829D64A971424" pageId="0" pageNumber="1">Included genera</paragraph>
<paragraph id="131BA63AC04D91836E92B8517FAEE27A" pageId="0" pageNumber="1">
<emphasis id="B79EEC11FF59E608A632AC54AE4339A2" bold="true" pageId="0" pageNumber="1">(4).</emphasis>
<taxonomicName id="FA2BDB6F7B9DA67A4E3F5BF2C3EE80BC" class="Magnoliopsida" family="Fabaceae" genus="Acrocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acrocarpus" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="9A0B705E95853721729DB1F54173C888" italics="true" pageId="0" pageNumber="1">Acrocarpus</emphasis>
</taxonomicName>
Wight ex Arn. (1 species), 
<taxonomicName id="7378FCFB903CB972DB80DA97359D1BF1" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="642FD3D206F580DDF4A3B091A9B5320A" italics="true" pageId="0" pageNumber="1">Arcoa</emphasis>
</taxonomicName>
Urb. (1), 
<taxonomicName id="69A128610FE76B0435253B567B1F273B" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="BCF77F957C95A5D9B99FF5BDC46FB2F5" italics="true" pageId="0" pageNumber="1">Ceratonia</emphasis>
</taxonomicName>
L. (2), 
<taxonomicName id="E8D426CA8C06225C318BFE39284D2EFE" class="Magnoliopsida" family="Fabaceae" genus="Tetrapterocarpon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tetrapterocarpon" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="F84F9C765FF95D1C9B9E688444816FE3" italics="true" pageId="0" pageNumber="1">Tetrapterocarpon</emphasis>
</taxonomicName>
Humbert (2).
</paragraph>
</subSubSection>
<subSubSection id="FDC17349ADC90F4F25EC7E471F3B3DA7" pageId="0" pageNumber="1" type="description">
<paragraph id="0DD694E11C24A45519220F7D9EFC4808" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="FCFB9FE21F690BB9A59FE816BAF127D4" pageId="0" pageNumber="1">
Unarmed (rarely the stipules of juvenile leaves spinescent) shrubs or trees. 
<emphasis id="07B9D6820468A0997533D5D97E114B6E" bold="true" pageId="0" pageNumber="1">Stipules</emphasis>
spinescent (
<taxonomicName id="8FB6A81E96D30E39338E53081E1FDC54" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="A9F7A35408837A6FB1BBA445297044E1" italics="true" pageId="0" pageNumber="1">Arcoa</emphasis>
</taxonomicName>
), minute, caducous or lacking. 
<emphasis id="74499D23DCCAD0C3E9A90B659498E4C8" bold="true" pageId="0" pageNumber="1">Leaves</emphasis>
pinnate or bipinnate with a terminal pinna. 
<emphasis id="562CAC168A56F8C6150FE9B69E902A63" bold="true" pageId="0" pageNumber="1">Inflorescences</emphasis>
erect or pendent racemes or panicles, sometimes clustered on short shoots or ramiflorous. 
<emphasis id="402351E158F923108FEED790DEE0CF60" bold="true" pageId="0" pageNumber="1">Flowers</emphasis>
unisexual or bisexual, a short hypanthium usually present, and sometimes a prominent central pulviniform disk; sepals valvate to slightly imbricate; petals absent or 4-5 (6) per flower; androecium haplo- to diplostemonous, or occasionally stamens more than 2 
<normalizedToken id="1BEF657D86C76ADFA159BD9F62461EA7" originalValue="×">x</normalizedToken>
petal number. 
<emphasis id="3BA3FB3FD403366C813F7B1B5123F822" bold="true" pageId="0" pageNumber="1">Fruits</emphasis>
dehiscent or indehiscent, linear to linear-oblong with an adaxial narrow wing, or oblong-ellipsoid, or laterally compressed and 4-winged (the wings in two unequal pairs), 1-several-seeded. 
<emphasis id="29E2EE78829E8D1D339B4F1770C81541" bold="true" pageId="0" pageNumber="1">Seeds</emphasis>
compressed, usually separated by areas of pulpy mesocarp, pleurogram lacking.
</paragraph>
</subSubSection>
<subSubSection id="6C03F30A60F6C05BCCC1C0C5B29D959F" pageId="0" pageNumber="1" type="distribution">
<paragraph id="A03B7F28F47147F0184D7ACB9BDE965F" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="062162DD92256DA7B060310D079B19A3" pageId="0" pageNumber="1">
Highly disjunct in Hispaniola (
<taxonomicName id="66F09883CEF7BFB00486B6105EEFF9A6" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="779843085C1998789E4BFFB8AC96D516" italics="true" pageId="0" pageNumber="1">Arcoa</emphasis>
</taxonomicName>
), Madagascar (
<taxonomicName id="7B8B3BDBAB10B4644D08CCBE66CA64B8" class="Magnoliopsida" family="Fabaceae" genus="Tetrapterocarpon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tetrapterocarpon" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="4EB6AA5689FFFC6DC9EA557819E3D526" italics="true" pageId="0" pageNumber="1">Tetrapterocarpon</emphasis>
</taxonomicName>
), tropical (South-)East Asia (
<taxonomicName id="327B4B06F593F223166B11A327059FEB" class="Magnoliopsida" family="Fabaceae" genus="Acrocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acrocarpus" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="E07A7EFC03AD6D6E03D11BA68B7F1FE5" italics="true" pageId="0" pageNumber="1">Acrocarpus</emphasis>
</taxonomicName>
), and north-eastern Africa, the Mediterranean, Oman, Yemen and Somalia (
<taxonomicName id="4DF5C1E90DC1568FB20C202BA45F1A2F" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="1F7F9F6A9A8D3DC9A59D6B9F090F04DC" italics="true" pageId="0" pageNumber="1">Ceratonia</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="80D5DA14804C8DFC86C9C73368BA251B" pageId="0" pageNumber="1" type="clade-based definition">
<paragraph id="B566047D0962ED0D6279BF2B2AA59366" pageId="0" pageNumber="1">Clade-based definition.</paragraph>
<paragraph id="7316EDE1BD383D291105DE3328AA5695" pageId="0" pageNumber="1">
The most inclusive crown clade containing 
<taxonomicName id="B19EED6ABDE73433BE37AD773E4765A3" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa gonavensis" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="gonavensis">
<emphasis id="C855A838FBD368556082FA9E562CCB69" italics="true" pageId="0" pageNumber="1">Arcoa gonavensis</emphasis>
</taxonomicName>
Urb. and 
<taxonomicName id="9BD8F0A18E233B71A81649D14F91272F" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia siliqua" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="siliqua">
<emphasis id="DB33968F49D079CD6943B314BAC00B03" italics="true" pageId="0" pageNumber="1">Ceratonia siliqua</emphasis>
</taxonomicName>
L., but not 
<taxonomicName id="3DD0FFEE71DC48DF0E3E1DABF0BEEE00" class="Magnoliopsida" family="Fabaceae" genus="Umtiza" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Umtiza listeriana" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="listeriana">
<emphasis id="02A9FF0D201F5D139C66E817B8828AC7" italics="true" pageId="0" pageNumber="1">Umtiza listeriana</emphasis>
</taxonomicName>
Sim, 
<taxonomicName id="51DCAF4FD0ED7E8BD34E732ADFE23B7A" class="Magnoliopsida" family="Caesalpiniaceae" genus="Dimorphandra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dimorphandra conjugata" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="conjugata">
<emphasis id="A9E4D1F601EC6E6A5FC364933E920DC1" italics="true" pageId="0" pageNumber="1">Dimorphandra conjugata</emphasis>
</taxonomicName>
(Splitg.) Sandwith or 
<taxonomicName id="05FFF9ECF2E67E9E3AF3A0AACF585273" class="Magnoliopsida" family="Leguminosae" genus="Mimosa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Mimosa sensitiva" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sensitiva">
<emphasis id="081E078A37709A480AF7ADF4A1CA5EC7" italics="true" pageId="0" pageNumber="1">Mimosa sensitiva</emphasis>
</taxonomicName>
L. (Fig. 
<figureCitation id="F20D0625D673954E25706A6972DAD581" captionStart="Figure 6" captionStartId="F13" captionText="Figure 6. Generic relationships in tribe Ceratonieae. Left part of figure shows complete genus-level Caesalpinioideae phylogeny with the Ceratonieae indicated with a red rectangle. Branch lengths are expressed in coalescent units and terminal branches were assigned an arbitrary uniform length for visual clarity. Support for relationships is based on fractions of supporting and conflicting gene trees: pie charts show gene tree support and conflict per node (blue representing supporting gene trees, green gene trees supporting the most common alternative topology, red gene trees supporting further alternative topologies, grey gene trees uninformative for this node), and numbers above pie charts are Internode Certainty All support values [both calculated with PhyParts (Smith et al. 2015)]. If present, red numbers below pie charts are non-significant (i. e.&gt; 0.05) outcomes of ASTRAL's polytomy test (Sayyari and Mirarab 2018), which tests for each node whether the polytomy null model can be rejected. Monophyletic genera are represented by single branches; see Suppl. material 2 for a phylogeny with all accessions. See Suppl. material 3 for gene tree support across the phylogeny. The phylogeny is a pruned version of the backbone phylogeny of Ringelberg et al. (2023), where full details of the data and phylogenomic analysis methods are presented." figureDoi="10.3897/phytokeys.240.101716.figure6" httpUri="https://binary.pensoft.net/fig/1014679" pageId="0" pageNumber="1">6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D09C8A7EDCC7EF8D1F373545251B8C84" pageId="0" pageNumber="1" type="notes">
<paragraph id="A63B547C92FBDE22903FEC429B812FB8" pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph id="05D03A4F1655773AA0B7BB0B22F30AF4" pageId="0" pageNumber="1">
The tribal name 
<taxonomicName id="A847C1618FBBF53371751029695BCC85" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Ceratonieae">Ceratonieae</taxonomicName>
was first published by 
<bibRefCitation id="E19ED370E92DAB05F12082BF5B930639" author="Reichenbach, HGL" journalOrPublisher="Flora Germanica Excursoria" pageId="0" pageNumber="1" pagination=": 544" refId="B697" refString="Reichenbach, HGL, 1832. Tribe Caesalpinieae Rchb., Tribe Ceratonieae Rchb. Flora Germanica Excursoria 2 (2): 544" title="Tribe Caesalpinieae Rchb., Tribe Ceratonieae Rchb." volume="2" year="1832">Reichenbach (1832)</bibRefCitation>
to accommodate the genus 
<taxonomicName id="22AF1239C191AABB44315DADE3EE8862" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="B41BA956FF4BBADBEA35E15BD296A652" italics="true" pageId="0" pageNumber="1">Ceratonia</emphasis>
</taxonomicName>
. 
<bibRefCitation id="7F1D5523FE448A33D875EB620DF11A35" author="Irwin, HS" editor="Polhill, RM" journalOrPublisher="Royal Botanic Gardens, Kew, United Kingdom" pageId="0" pageNumber="1" pagination="97 - 106" refId="B406" refString="Irwin, HS, Barneby, RC, 1981. Tribe Cassieae Bronn (1822). In: Polhill, RM, Raven, PH, Eds., Advances in Legume Systematics, Part 1. Royal Botanic Gardens, Kew, United Kingdom: 97 - 106" title="Tribe Cassieae Bronn (1822)." volumeTitle="Advances in Legume Systematics, Part 1." year="1981">Irwin and Barneby (1981)</bibRefCitation>
placed 
<taxonomicName id="58E0C82B4F325CA7AABE1AF97D580FB5" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="142B02BAD268DD55F8F6D2C18ADE1232" italics="true" pageId="0" pageNumber="1">Ceratonia</emphasis>
</taxonomicName>
in its own new subtribe 
<taxonomicName id="BF65E7902D5670D5BC42F53F40E99AA6" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="subTribe" subTribe="Ceratoniinae">Ceratoniinae</taxonomicName>
H.S. Irwin &amp; Barneby of tribe 
<taxonomicName id="0794F359BD4F23A6DC2DA3DE3695743B" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Cassieae">Cassieae</taxonomicName>
, and they would have accorded the genus tribal rank, i.e., a reinstatement of 
<normalizedToken id="096FF066F0E99957FDA70BDB04318C60" originalValue="Reichenbachʼs">Reichenbachʼs</normalizedToken>
(1832) taxon, &quot;if not dissuaded by others&quot; (
<bibRefCitation id="C1D99BC8C30BF05E3A67DBB1FDF09890" author="Irwin, HS" editor="Polhill, RM" journalOrPublisher="Royal Botanic Gardens, Kew, United Kingdom" pageId="0" pageNumber="1" pagination="97 - 106" refId="B406" refString="Irwin, HS, Barneby, RC, 1981. Tribe Cassieae Bronn (1822). In: Polhill, RM, Raven, PH, Eds., Advances in Legume Systematics, Part 1. Royal Botanic Gardens, Kew, United Kingdom: 97 - 106" title="Tribe Cassieae Bronn (1822)." volumeTitle="Advances in Legume Systematics, Part 1." year="1981">Irwin and Barneby 1981</bibRefCitation>
: 98). 
<bibRefCitation id="5DF96BA2117ADA3E4BBEEF96E50C42FD" DOI="https://doi.org/10.1086/376881" author="Herendeen, PS" journalOrPublisher="Royal Botanic Gardens, Kew, United Kingdom" pageId="0" pageNumber="1" refId="B357" refString="Herendeen, PS, Lewis, GP, Bruneau, A, 2003b. Floral morphology in caesalpinioid legumes: testing the monophyly of the &quot;Umtiza clade&quot;. International Journal of Plant Sciences 164(Supplement): S393-S407. https://doi.org/10.1086/376881" title="Floral morphology in caesalpinioid legumes: testing the monophyly of the &quot; Umtiza clade &quot;. International Journal of Plant Sciences 164 (Supplement): S 393 - S 407." url="https://doi.org/10.1086/376881" year="2003 b">Herendeen et al. (2003b)</bibRefCitation>
, based on molecular and morphological data, grouped seven disparate caesalpinioid genera in their &quot; 
<taxonomicName id="7312E7AC1385274B7890BB36B5F90CE3" class="Magnoliopsida" family="Fabaceae" genus="Umtiza" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Umtiza" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Umtiza</taxonomicName>
clade&quot;, which included two major subclades: the 
<taxonomicName id="B57C95963194F3FB2E2C6E25033DE991" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Arcoa</taxonomicName>
- 
<taxonomicName id="A69378F602D49BF91507272D1B4620DD" class="Magnoliopsida" family="Fabaceae" genus="Tetrapterocarpon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tetrapterocarpon" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Tetrapterocarpon</taxonomicName>
- 
<taxonomicName id="C6DB941AC9A11F0552CA40D5D264BB73" class="Magnoliopsida" family="Fabaceae" genus="Acrocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acrocarpus" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Acrocarpus</taxonomicName>
- 
<taxonomicName id="B0B0F013D120A1DB94C7967FE8C783B2" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Ceratonia</taxonomicName>
clade and the 
<taxonomicName id="4AF6737578644C74F2F5668E13EE58BE" class="Magnoliopsida" family="Fabaceae" genus="Gymnocladus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Gymnocladus" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Gymnocladus</taxonomicName>
- 
<taxonomicName id="5CCB390C89A484D1CF254EB9816BA250" class="Magnoliopsida" family="Fabaceae" genus="Umtiza" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Umtiza" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Umtiza</taxonomicName>
- 
<taxonomicName id="A72A27620A82AF4D25663372CFF59E4C" class="Magnoliopsida" family="Leguminosae" genus="Gleditsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Gleditsia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Gleditsia</taxonomicName>
clade. More recent studies do not support the monophyly of these two clades together (
<bibRefCitation id="E979CA38F26EDD6702A12380BBDC811F" DOI="https://doi.org/10.1016/j.ympev.2012.05.035" author="Manzanilla, V" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="1" pagination="149 - 162" refId="B526" refString="Manzanilla, V, Bruneau, A, 2012. Phylogeny reconstruction in the Caesalpinieae grade (Leguminosae) based on duplicated copies of the sucrose synthase gene and plastid markers. Molecular Phylogenetics and Evolution 65 (1): 149 - 162, DOI: https://doi.org/10.1016/j.ympev.2012.05.035" title="Phylogeny reconstruction in the Caesalpinieae grade (Leguminosae) based on duplicated copies of the sucrose synthase gene and plastid markers." url="https://doi.org/10.1016/j.ympev.2012.05.035" volume="65" year="2012">Manzanilla and Bruneau 2012</bibRefCitation>
; 
<bibRefCitation id="E9DCCC703836A89CD423F0106A41E16F" DOI="https://doi.org/10.12705/622.8" author="LPWG, [Legume Phylogeny Working Group]" journalOrPublisher="Taxon" pageId="0" pageNumber="1" pagination="217 - 248" refId="B504" refString="LPWG, [Legume Phylogeny Working Group], 2013. Legume phylogeny and classification in the 21st century: progress, prospects and lessons for other species-rich clades. Taxon 62: 217 - 248, DOI: https://doi.org/10.12705/622.8" title="Legume phylogeny and classification in the 21 st century: progress, prospects and lessons for other species-rich clades." url="https://doi.org/10.12705/622.8" volume="62" year="2013">LPWG 2013</bibRefCitation>
, 
<bibRefCitation id="761CAEDC7413141C166488B6FBBDA1D9" DOI="https://doi.org/10.12705/661.3" author="LPWG, [Legume Phylogeny Working Group]" journalOrPublisher="Taxon" pageId="0" pageNumber="1" pagination="44 - 77" refId="B505" refString="LPWG, [Legume Phylogeny Working Group], 2017. A new subfamily classification of the Leguminosae based on a taxonomically comprehensive phylogeny. Taxon 66: 44 - 77, DOI: https://doi.org/10.12705/661.3" title="A new subfamily classification of the Leguminosae based on a taxonomically comprehensive phylogeny." url="https://doi.org/10.12705/661.3" volume="66" year="2017">2017</bibRefCitation>
; 
<bibRefCitation id="CDE7199A3AB03482D413B9596170FAB0" DOI="https://doi.org/10.3897/phytokeys.205.85866" author="Ringelberg, JJ" editor="Hughes, CE" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="3 - 58" refId="B712" refString="Ringelberg, JJ, Koenen, EJM, Iganci, JR, Queiroz, LP, Murphy, DJ, Gaudeul, M, Bruneau, A, Luckow, M, Lewis, GP, Hughes, CE, 2022. Phylogenomic analysis of 997 nuclear genes reveals the need for extensive generic re-delimitation in Caesalpinioideae (Leguminosae). In: Hughes, CE, Queiroz, LP, Lewis, GP, Eds., Advances in Legume Systematics 14. Classification of Caesalpinioideae Part 1: New generic delimitations. PhytoKeys 205: 3 - 58, DOI: https://doi.org/10.3897/phytokeys.205.85866" title="Phylogenomic analysis of 997 nuclear genes reveals the need for extensive generic re-delimitation in Caesalpinioideae (Leguminosae)." url="https://doi.org/10.3897/phytokeys.205.85866" volume="205" volumeTitle="Advances in Legume Systematics 14. Classification of Caesalpinioideae Part 1: New generic delimitations." year="2022">Ringelberg et al. 2022</bibRefCitation>
) and they are here recognised under the two reinstated tribes 
<taxonomicName id="3BEBC460C6DE12C194EC36CF0A9D2B13" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Ceratonieae">Ceratonieae</taxonomicName>
and 
<taxonomicName id="69A580C09D98C35FDE50296ADC4C0995" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Gleditsieae">Gleditsieae</taxonomicName>
, respectively. Tribe 
<taxonomicName id="1058CBDFB717BD939FF70237B92111BE" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Ceratonieae">Ceratonieae</taxonomicName>
is resolved as sister to the rest of 
<taxonomicName id="69B7F17135F984F9652E9836BE0994B7" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="subFamily" subFamily="Caesalpinioideae">Caesalpinioideae</taxonomicName>
(Fig. 
<figureCitation id="A7691AC4AC816C0FCBFD2928BDE8DE60" captionStart="Figure 6" captionStartId="F13" captionText="Figure 6. Generic relationships in tribe Ceratonieae. Left part of figure shows complete genus-level Caesalpinioideae phylogeny with the Ceratonieae indicated with a red rectangle. Branch lengths are expressed in coalescent units and terminal branches were assigned an arbitrary uniform length for visual clarity. Support for relationships is based on fractions of supporting and conflicting gene trees: pie charts show gene tree support and conflict per node (blue representing supporting gene trees, green gene trees supporting the most common alternative topology, red gene trees supporting further alternative topologies, grey gene trees uninformative for this node), and numbers above pie charts are Internode Certainty All support values [both calculated with PhyParts (Smith et al. 2015)]. If present, red numbers below pie charts are non-significant (i. e.&gt; 0.05) outcomes of ASTRAL's polytomy test (Sayyari and Mirarab 2018), which tests for each node whether the polytomy null model can be rejected. Monophyletic genera are represented by single branches; see Suppl. material 2 for a phylogeny with all accessions. See Suppl. material 3 for gene tree support across the phylogeny. The phylogeny is a pruned version of the backbone phylogeny of Ringelberg et al. (2023), where full details of the data and phylogenomic analysis methods are presented." figureDoi="10.3897/phytokeys.240.101716.figure6" httpUri="https://binary.pensoft.net/fig/1014679" pageId="0" pageNumber="1">6</figureCitation>
) and comprises six species in four genera (two of the genera are monospecific).
</paragraph>
<caption id="704B1173097B44A7D32EFE1F8151550E" doi="10.3897/phytokeys.240.101716.figure6" httpUri="https://binary.pensoft.net/fig/1014679" pageId="0" pageNumber="1" start="Figure 6" startId="F13">
<paragraph id="D587084CDE8B20A3364F0EC44D185EF8" pageId="0" pageNumber="1">
<emphasis id="55C349D7DEB26F4EECD093AE9963C508" bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
Generic relationships in tribe 
<taxonomicName id="70A24BF5D33F46D8F686942DA19433FD" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Ceratonieae">Ceratonieae</taxonomicName>
. Left part of figure shows complete genus-level 
<taxonomicName id="A24985165DBDD7710E379A467C161994" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="subFamily" subFamily="Caesalpinioideae">Caesalpinioideae</taxonomicName>
phylogeny with the 
<taxonomicName id="31CA5F52E8A816F5267B5530FB08E268" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Ceratonieae">Ceratonieae</taxonomicName>
indicated with a red rectangle. Branch lengths are expressed in coalescent units and terminal branches were assigned an arbitrary uniform length for visual clarity. Support for relationships is based on fractions of supporting and conflicting gene trees: pie charts show gene tree support and conflict per node (blue representing supporting gene trees, green gene trees supporting the most common alternative topology, red gene trees supporting further alternative topologies, grey gene trees uninformative for this node), and numbers above pie charts are Internode Certainty All support values [both calculated with PhyParts (
<bibRefCitation id="2572D512D2AF6218C4723AAD30718A00" DOI="https://doi.org/10.1186/s12862-015-0423-0" author="Smith, SA" journalOrPublisher="Taxon" pageId="0" pageNumber="1" refId="B798" refString="Smith, SA, Moore, MJ, Brown, JW, Yang, Y, 2015. Analysis of phylogenomic datasets reveals conflict, concordance, and gene duplications with examples from animals and plants. BMC Evolutionary Biology 15: 150. https://doi.org/10.1186/s12862-015-0423-0" title="Analysis of phylogenomic datasets reveals conflict, concordance, and gene duplications with examples from animals and plants. BMC Evolutionary Biology 15: 150." url="https://doi.org/10.1186/s12862-015-0423-0" year="2015">Smith et al. 2015</bibRefCitation>
)]. If present, red numbers below pie charts are non-significant (i.e.&gt; 0.05) outcomes of 
<normalizedToken id="90FCC6C8399BAECB1615553622FAF221" originalValue="ASTRAL’s">ASTRAL's</normalizedToken>
polytomy test (
<bibRefCitation id="BD2B02236035BB8E9DD9BDFECF8647E6" DOI="https://doi.org/10.3390/genes9030132" author="Sayyari, E" journalOrPublisher="Omagh, Tyrone, Ireland" pageId="0" pageNumber="1" refId="B746" refString="Sayyari, E, Mirarab, S, 2018. Testing for polytomies in phylogenetic species trees using quartet frequencies. Genes 9(3): e132. https://doi.org/10.3390/genes9030132" title="Testing for polytomies in phylogenetic species trees using quartet frequencies. Genes 9 (3): e 132." url="https://doi.org/10.3390/genes9030132" year="2018">Sayyari and Mirarab 2018</bibRefCitation>
), which tests for each node whether the polytomy null model can be rejected. Monophyletic genera are represented by single branches; see Suppl. material 2 for a phylogeny with all accessions. See Suppl. material 3 for gene tree support across the phylogeny. The phylogeny is a pruned version of the backbone phylogeny of 
<bibRefCitation id="2B5776D24EC72CC14D28D4BB00810A5F" DOI="https://doi.org/10.1126/sciadv.ade4954" author="Ringelberg, JJ" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" refId="B713" refString="Ringelberg, JJ, Koenen, EJM, Sauter, B, Aebli, A, Rando, JG, Iganci, JR, Queiroz, LP, Murphy, DJ, Gaudeul, M Bruneau A, Luckow, M, Lewis, GP, Miller, JT, Simon, MF, Jordao, LSB, Morales, M, Bailey, CD, Nageswara-Rao, M, Nicholls, JA, Loiseau, O, Pennington, RT, Dexter, KG, Zimmermann, NE, Hughes, CE, 2023. Precipitation is the main axis of tropical plant phylogenetic turnover across space and time. Science Advances 9: eade4954. https://doi.org/10.1126/sciadv.ade4954" title="Precipitation is the main axis of tropical plant phylogenetic turnover across space and time. Science Advances 9: eade 4954." url="https://doi.org/10.1126/sciadv.ade4954" year="2023">Ringelberg et al. (2023)</bibRefCitation>
, where full details of the data and phylogenomic analysis methods are presented.
</paragraph>
</caption>
<paragraph id="EDC08FA2369DEB1107A1C0A841F0D5B7" pageId="0" pageNumber="1">
Only two characters have been found to be shared by members of tribe 
<taxonomicName id="334EB8284891E08F83D33DFA58467356" family="Leguminosae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Ceratonieae">Ceratonieae</taxonomicName>
: bipinnate leaves with a single terminal pinna (Fig. 
<figureCitation id="B9DACCAD64CDA7E25448C83CADADB294" captionStart="Figure 7" captionStartId="F14" captionText="Figure 7. Flower, fruit, and vegetative characters of tribe Ceratonieae A Arcoa gonavensis Urb., foliage and fruits, Dominican Republic B Tetrapterocarpon geayi Humbert, fruits and part of bipinnate leaf, Madagascar (Du Puy M 410) C-E Acrocarpus fraxinifolius Wight &amp; Arn., India, cultivated tree C inflorescence D dehisced fruits and new flush foliage E part of a bipinnate leaf F-J Ceratonia siliqua L. F tree, Jordan, semi-desert near Petra G fruits, Israel, Mt. Scopus Botanical Gardens, Jerusalem H pistillate flowers, Crete I staminate flowers, Crete J staminate inflorescences, Israel, Jerusalem Botanical Gardens. Photo credits A F Jimenez R. B D Du Puy C-E https: // efloraofindia. com / 2011 / 02 / 01 / acrocarpus-fraxinifolius / E D Valke F, G, J O Fragman-Sapir H, I G Lewis." figureDoi="10.3897/phytokeys.240.101716.figure7" httpUri="https://binary.pensoft.net/fig/1014680" pageId="0" pageNumber="1">7E</figureCitation>
; although 
<taxonomicName id="E90FD5CC4AE64E685EA19E34C5D7C57D" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="D400BDE801889E3AEF45918BBC5EC916" italics="true" pageId="0" pageNumber="1">Ceratonia</emphasis>
</taxonomicName>
leaves are usually singly pinnate) and three of the four genera (all except 
<taxonomicName id="3C245389E791AF87484ADF399B181E44" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="F4646277A490A031F9147147FEC84DC4" italics="true" pageId="0" pageNumber="1">Arcoa</emphasis>
</taxonomicName>
, which is sister to the three other genera) have a large deletion in the plastid 
<emphasis id="FF57AEB9C731D4FDF15D8838996ABDBA" italics="true" pageId="0" pageNumber="1">trnL</emphasis>
intron (
<bibRefCitation id="C7F4E9460504AFB9F445A5D3C0B5B1C5" author="Bruneau, A" journalOrPublisher="Systematic Botany" pageId="0" pageNumber="1" pagination="487 - 514" refId="B130" refString="Bruneau, A, Forest, F, Herendeen, PS, Klitgaard, BB, Lewis, GP, 2001. Phylogenetic relationships in the Caesalpinioideae (Leguminosae) as inferred from chloroplast trn L intron sequences. Systematic Botany 26: 487 - 514" title="Phylogenetic relationships in the Caesalpinioideae (Leguminosae) as inferred from chloroplast trn L intron sequences." volume="26" year="2001">Bruneau et al. 2001</bibRefCitation>
). Three of the genera (
<taxonomicName id="3DEFF0AFA03F3E50FB8EE60D4BB2DA9D" class="Magnoliopsida" family="Fabaceae" genus="Acrocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acrocarpus" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="6483809D9EDE5427FB8104D03A97FA6A" italics="true" pageId="0" pageNumber="1">Acrocarpus</emphasis>
</taxonomicName>
is the exception) are dioecious (although occasional bisexual flowers occur). 
<taxonomicName id="2271921005D0534A6296D1CC34A08C88" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="0FE062D121B55D9DEF2521857618FF9F" italics="true" pageId="0" pageNumber="1">Ceratonia</emphasis>
</taxonomicName>
and 
<taxonomicName id="2B3DFB95A740EA11C5929FE9D170FC96" class="Magnoliopsida" family="Fabaceae" genus="Acrocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acrocarpus" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="E6D8AF11A9D183774922A0626CF645D8" italics="true" pageId="0" pageNumber="1">Acrocarpus</emphasis>
</taxonomicName>
are sister genera, both have simple inflorescences, a haplostemonous androecium, glabrous stamen filaments, alternate first seedling leaves, septate wood fibres, and a chromosome base number of 
<emphasis id="36FBD9DF2C87A6CFA39AD613FD6F34C4" italics="true" pageId="0" pageNumber="1">x</emphasis>
= 12. They differ in unisexual (
<taxonomicName id="8F4C9DF584D0FB4C95113FBA44325C3F" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="4689B8BC0BB9FA6804AF84CCB32BAEDB" italics="true" pageId="0" pageNumber="1">Ceratonia</emphasis>
</taxonomicName>
) versus bisexual (
<taxonomicName id="395050F830B0AD5993AE1FB8A78338F4" class="Magnoliopsida" family="Fabaceae" genus="Acrocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acrocarpus" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="EAF699ED3AF6BF2B95244B18AECADA09" italics="true" pageId="0" pageNumber="1">Acrocarpus</emphasis>
</taxonomicName>
) flowers, absence of petals (
<taxonomicName id="247A60DD0476BB674EF680DC8AFFF9DA" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="DC4735458B4EC8DD9A805B5EF4C3CB49" italics="true" pageId="0" pageNumber="1">Ceratonia</emphasis>
</taxonomicName>
) versus petals present (
<taxonomicName id="D152A851CB00C5E7E8C94732933ADB99" class="Magnoliopsida" family="Fabaceae" genus="Acrocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acrocarpus" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="CED7E065DD38779ED2B0A0F6DE280D0B" italics="true" pageId="0" pageNumber="1">Acrocarpus</emphasis>
</taxonomicName>
), and fruit type: indehiscent (
<taxonomicName id="FF6454EE06688FA981590B08D0F93593" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="CF88101D91C90F4309668E2A56A747E0" italics="true" pageId="0" pageNumber="1">Ceratonia</emphasis>
</taxonomicName>
) versus dehiscent (
<taxonomicName id="EFB048F16D9B53D7D7293D1377BE1A6B" class="Magnoliopsida" family="Fabaceae" genus="Acrocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acrocarpus" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="814AF4C4C6F52C6066E3D0009C136C92" italics="true" pageId="0" pageNumber="1">Acrocarpus</emphasis>
</taxonomicName>
). Three of the four genera are confirmed as non-nodulating and the status of 
<taxonomicName id="AF90D666A0CB1AF63BDD1B74C592B85F" class="Magnoliopsida" family="Fabaceae" genus="Tetrapterocarpon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tetrapterocarpon" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="E47D2C91D97ACD7F853B283B9555A70A" italics="true" pageId="0" pageNumber="1">Tetrapterocarpon</emphasis>
</taxonomicName>
is not known (
<bibRefCitation id="77C57F5ED9CD67DCC68E626043DEE1CD" author="Sprent, JI" journalOrPublisher="Bulletin of Miscellaneous Information, Royal Botanic Gardens, Kew," pageId="0" pageNumber="1" refId="B826" refString="Sprent, JI, 2001. Nodulation in Legumes. Royal Botanic Gardens, Kew, United Kingdom, i-xii, 1-146." title="Nodulation in Legumes. Royal Botanic Gardens, Kew, United Kingdom, i-xii, 1 - 146." year="2001">Sprent 2001</bibRefCitation>
; 
<bibRefCitation id="5F728019EBCBCD15F7E1F13EA4CA33F2" DOI="https://doi.org/10.1111/nph.18321" author="Faria, SM" journalOrPublisher="New Phytologist" pageId="0" pageNumber="1" pagination="2365 - 2377" refId="B268" refString="Faria, SM, Ringelberg, JJ, Gross, E, Koenen, EJM, Cardoso, D, Ametsitsi, GKD, Akomatey, J, Maluk, M, Tak, N, Gehlot, HS, Wright, KM, Teaumroong, N, Songwattana, P, Lima, HC, Prin, Y, Zartman, CE, Sprent, JI, Ardley, J, Hughes, CE, James, EK, 2022. The innovation of the symbiosome has enhanced the evolutionary stability of nitrogen fixation in legumes. New Phytologist 235: 2365 - 2377, DOI: https://doi.org/10.1111/nph.18321" title="The innovation of the symbiosome has enhanced the evolutionary stability of nitrogen fixation in legumes." url="https://doi.org/10.1111/nph.18321" volume="235" year="2022">Faria et al. 2022</bibRefCitation>
).
</paragraph>
<caption id="1518C98286769B54EDE86EECCBF6420C" doi="10.3897/phytokeys.240.101716.figure7" httpUri="https://binary.pensoft.net/fig/1014680" pageId="0" pageNumber="1" start="Figure 7" startId="F14">
<paragraph id="20276886AC57250486F14C53935E8EA6" pageId="0" pageNumber="1">
<emphasis id="23D9A6BC3A7D2386667DB850720B1FB4" bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
Flower, fruit, and vegetative characters of tribe 
<taxonomicName id="AF3D7D00F2CC125F0EE39A30885EA1F8" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Ceratonieae">Ceratonieae</taxonomicName>
<emphasis id="A28A5AF1E8E8D8D73AE46B9CE9C2742A" bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName id="E5D6C8DA9CC3BF67C58A38D5D0743433" class="Magnoliopsida" family="Leguminosae" genus="Arcoa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arcoa gonavensis" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="gonavensis">
<emphasis id="4ED80ECF9AA768F8150D5355E325BA3F" italics="true" pageId="0" pageNumber="1">Arcoa gonavensis</emphasis>
</taxonomicName>
Urb., foliage and fruits, Dominican Republic 
<emphasis id="4E43F9F84AFBD87B323798D9F7F23A17" bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName id="7FEADB8B625618C0716E54FEFD0D1160" class="Magnoliopsida" family="Fabaceae" genus="Tetrapterocarpon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tetrapterocarpon geayi" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="geayi">
<emphasis id="3BCFA6DE1278A72EEDDB9165FD46D1C6" italics="true" pageId="0" pageNumber="1">Tetrapterocarpon geayi</emphasis>
</taxonomicName>
Humbert, fruits and part of bipinnate leaf, Madagascar (
<emphasis id="CB042E512AF06FD9ABE31110F439D936" italics="true" pageId="0" pageNumber="1">Du Puy M410</emphasis>
) 
<emphasis id="F5BF8836DBB6771190B97D8BB9E93078" bold="true" pageId="0" pageNumber="1">C-E</emphasis>
<taxonomicName id="1D325BA8E6855354633781BC482C3E30" class="Magnoliopsida" family="Fabaceae" genus="Acrocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acrocarpus fraxinifolius" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="fraxinifolius">
<emphasis id="E14D5598ECBEDC035C9C4669AA807114" italics="true" pageId="0" pageNumber="1">Acrocarpus fraxinifolius</emphasis>
</taxonomicName>
Wight &amp; Arn., India, cultivated tree 
<emphasis id="B061B0C29DBA4934EE1629AE733BFF98" bold="true" pageId="0" pageNumber="1">C</emphasis>
inflorescence 
<emphasis id="EED46FF3EA3281926215B97235E612CF" bold="true" pageId="0" pageNumber="1">D</emphasis>
dehisced fruits and new flush foliage 
<emphasis id="8DEC700380C1B0737228C55C031ADF84" bold="true" pageId="0" pageNumber="1">E</emphasis>
part of a bipinnate leaf 
<emphasis id="A07933909EBEFE65F85FEA91B2D4A313" bold="true" pageId="0" pageNumber="1">F-J</emphasis>
<taxonomicName id="78EE081F84E6B0942EB119D42D21A89E" class="Magnoliopsida" family="Leguminosae" genus="Ceratonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratonia siliqua" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="siliqua">
<emphasis id="61155CA1F49E546A1966E02ECC1B0D35" italics="true" pageId="0" pageNumber="1">Ceratonia siliqua</emphasis>
</taxonomicName>
L. 
<emphasis id="A96702CC7ACB2D9101464A2814F81AD6" bold="true" pageId="0" pageNumber="1">F</emphasis>
tree, Jordan, semi-desert near Petra 
<emphasis id="0799CF5CC04415A6EF55D25A830178BC" bold="true" pageId="0" pageNumber="1">G</emphasis>
fruits, Israel, Mt. Scopus Botanical Gardens, Jerusalem 
<emphasis id="6F32D246FBAE03F8413978DA86309BEA" bold="true" pageId="0" pageNumber="1">H</emphasis>
pistillate flowers, Crete 
<emphasis id="8FBD061E553E25BBA67C45856BCCAF47" bold="true" pageId="0" pageNumber="1">I</emphasis>
staminate flowers, Crete 
<emphasis id="177CCAFA4A8195FF822483815AD0E215" bold="true" pageId="0" pageNumber="1">J</emphasis>
staminate inflorescences, Israel, Jerusalem Botanical Gardens. Photo credits 
<emphasis id="C3E61B9D6A240A51F379D446BC0BD098" bold="true" pageId="0" pageNumber="1">A</emphasis>
F Jimenez R. 
<emphasis id="BD9CAFCF884FA38D9BAF6942E297B6EC" bold="true" pageId="0" pageNumber="1">B</emphasis>
D Du Puy 
<emphasis id="AC7F0C4EA168DEF76DBEC8998BFCBB67" bold="true" pageId="0" pageNumber="1">C-E</emphasis>
https://efloraofindia.com/2011/02/01/acrocarpus-fraxinifolius/ 
<emphasis id="1BC46AE9DE1F382374EBCFBCA09B3610" bold="true" pageId="0" pageNumber="1">E</emphasis>
D Valke 
<emphasis id="194757B7D111E84EC5CB7CE256403AFF" bold="true" pageId="0" pageNumber="1">F, G, J</emphasis>
O Fragman-Sapir 
<emphasis id="766CDA8513715428BCDF66A377EFD5B1" bold="true" pageId="0" pageNumber="1">H, I</emphasis>
G Lewis.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>