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<document id="3867586B99629809716271E8B13F475E" ID-CLB-Dataset="50813" ID-DOI="10.5281/zenodo.171551" ID-GBIF-Dataset="ba40087f-fcf9-4de2-bb74-ffbf81a27f32" ID-ISSN="1175­5326" ID-Zenodo-Dep="171551" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459754367453" checkinUser="plazi" docAuthor="El-Hawagry, Magdi S. &amp; Greathead, David J." docDate="2006" docId="7C49EB33FFFFC2130371FB1CFB9F0E6E" docLanguage="en" docName="zt01113p032.pdf" docOrigin="Zootaxa 1113" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Villa efflatouni El-Hawagry &amp; Greathead, 2006, sp. nov." docType="treatment" docVersion="10" lastPageNumber="29" masterDocId="8070934BFFF8C21B0279FF90FFC10B70" masterDocTitle="Review of the genus Villa Lioy (Bombyliidae, Diptera) from Egypt, with descriptions of two new species" masterLastPageNumber="32" masterPageNumber="21" pageNumber="28" updateTime="1698217012612" updateUser="ExternalLinkService">
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<mods:title id="B8E79C599068C42B96A8E888CC08A276">Review of the genus Villa Lioy (Bombyliidae, Diptera) from Egypt, with descriptions of two new species</mods:title>
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<mods:namePart id="DD072D56061D25CC32C9C93FB40C4B19">El-Hawagry, Magdi S.</mods:namePart>
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<mods:namePart id="4C36A4EB9D888A59D75DA8742833A297">Greathead, David J.</mods:namePart>
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<mods:date id="927F72DE70C33740C716055CA9C33133">2006</mods:date>
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<subSubSection id="BCFA09AEFFFFC21C0371FB1CFDE50FD6" box="[264,548,1164,1190]" pageId="7" pageNumber="28" type="nomenclature">
<paragraph id="F45F5A25FFFFC21C0371FB1CFDE50FD6" blockId="7.[264,548,1164,1190]" box="[264,548,1164,1190]" pageId="7" pageNumber="28">
<heading id="AF17ED49FFFFC21C0371FB1CFDE50FD6" bold="true" box="[264,548,1164,1190]" fontSize="11" level="1" pageId="7" pageNumber="28" reason="1">
<emphasis id="C6948637FFFFC21C0371FB1CFDE50FD6" bold="true" box="[264,548,1164,1190]" pageId="7" pageNumber="28">
<taxonomicName id="33E021A6FFFFC21C0371FB1CFE7F0FD6" ID-CoL="7FRWG" box="[264,446,1164,1190]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="efflatouni" status="sp. nov.">
<emphasis id="C6948637FFFFC21C0371FB1CFE7F0FD6" bold="true" box="[264,446,1164,1190]" italics="true" pageId="7" pageNumber="28">Villa efflatouni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDA73B4CFFFFC21C03BFFB1CFDE50FD6" box="[454,548,1164,1190]" pageId="7" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="F45F5A25FFFFC21C0371FB47FE180E60" blockId="7.[264,1323,1238,1296]" pageId="7" pageNumber="28">
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<taxonomicName id="33E021A6FFFFC21C0371FB47FD5B0F9D" ID-CoL="7G3MM" authority="Loew, 1869: 188" authorityName="Loew" authorityPageNumber="188" authorityYear="1869" box="[264,666,1238,1262]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="senecio">
<emphasis id="C6948637FFFFC21C0371FB47FE5E0F9E" box="[264,415,1239,1262]" italics="true" pageId="7" pageNumber="28">Villa senecio</emphasis>
(
<treatmentCitation id="75417C34FFFFC21C03C5FB46FD530F9D" author="Loew" box="[444,658,1238,1261]" page="188" pageId="7" pageNumber="28" year="1869">
<bibRefCitation id="907127D4FFFFC21C03C5FB46FD530F9D" author="Loew" box="[444,658,1238,1261]" pageId="7" pageNumber="32" refString="Loew, H (1869) Beschreibung europaischer Dipteren. Von Johann Wilhelm Meigen. Erster Band. Systematischer Beschreibung der bekannten europaischer zweiflugeligen Insekten. Achter oder zweiter Supplementband. H. W. Schmidt, Halle, xvi + 310 + [1] pp." type="journal article" year="1869">Loew, 1869: 188</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
:
<treatmentCitation id="75417C34FFFFC21C00CDFB46FC4B0F9D" author="Bezzi" box="[692,906,1238,1261]" page="226" pageId="7" pageNumber="28" year="1925">
<bibRefCitation id="907127D4FFFFC21C00CDFB46FC870F9D" author="Bezzi" box="[692,838,1238,1261]" pageId="7" pageNumber="32" refString="Bezzi, M. (1925) Quelques notes sur les bombyliides (Dipt.) d'Egypte, avec description d'especes nouvelles. Bulletin de la Societe Royale Entomologique d'Egypte, 8, 159 - 242." type="journal article" year="1925">Bezzi, 1925</bibRefCitation>
: 226
</treatmentCitation>
; Steyskal &amp; El Bialy, 1967: 18 (misidentifications)
</treatmentCitationGroup>
</paragraph>
<paragraph id="F45F5A25FFFFC2130371FAD4FBBC0AA6" blockId="7.[264,1324,1348,1894]" lastBlockId="8.[264,1325,284,1310]" lastPageId="8" lastPageNumber="29" pageId="7" pageNumber="28">
Specimens of this species were misidentified as
<taxonomicName id="33E021A6FFFFC21C0130FAD4FBFE0E2E" ID-CoL="7G3MM" authority="Loew" authorityName="Loew" box="[841,1087,1348,1374]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="senecio">
<emphasis id="C6948637FFFFC21C0130FAD4FC200E2D" box="[841,993,1348,1373]" italics="true" pageId="7" pageNumber="28">Villa senecio</emphasis>
(Loew)
</taxonomicName>
by
<bibRefCitation id="907127D4FFFFC21C0616FAD4FACD0E2E" author="Bezzi" box="[1135,1292,1348,1374]" pageId="7" pageNumber="32" refString="Bezzi, M. (1925) Quelques notes sur les bombyliides (Dipt.) d'Egypte, avec description d'especes nouvelles. Bulletin de la Societe Royale Entomologique d'Egypte, 8, 159 - 242." type="journal article" year="1925">Bezzi (1925)</bibRefCitation>
, E ngel (1937) and Steyskal &amp; El Bialy (1967). Bowden labelled specimens of this species as
<taxonomicName id="33E021A6FFFFC21C0371FA04FDC30EDE" ID-CoL="7G3MM" authority="Loew" authorityName="Loew" box="[264,514,1428,1454]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="senecio">
<emphasis id="C6948637FFFFC21C0371FA04FE600EDD" box="[264,417,1428,1453]" italics="true" pageId="7" pageNumber="28">Villa senecio</emphasis>
(Loew)
</taxonomicName>
as well, but they can not be
<taxonomicName id="33E021A6FFFFC21C011EFA04FC1E0EDD" ID-CoL="7G3MM" box="[871,991,1428,1453]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="senecio">
<emphasis id="C6948637FFFFC21C011EFA04FC1E0EDD" box="[871,991,1428,1453]" italics="true" pageId="7" pageNumber="28">V. senecio</emphasis>
</taxonomicName>
. The two
<typeStatus id="2B5BE487FFFFC21C0627FA04FB780EDE" box="[1118,1209,1428,1454]" pageId="7" pageNumber="28" type="syntype">syntype</typeStatus>
males of
<taxonomicName id="33E021A6FFFFC21C0371FA2CFD840EA6" ID-CoL="7G3MM" authority="Loew, 1869" authorityName="Loew" authorityYear="1869" box="[264,581,1468,1494]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="senecio">
<emphasis id="C6948637FFFFC21C0371FA2CFE5E0EA5" box="[264,415,1468,1493]" italics="true" pageId="7" pageNumber="28">Villa senecio</emphasis>
(
<bibRefCitation id="907127D4FFFFC21C03D6FA2CFDFF0EA6" author="Loew" box="[431,574,1468,1494]" pageId="7" pageNumber="32" refString="Loew, H (1869) Beschreibung europaischer Dipteren. Von Johann Wilhelm Meigen. Erster Band. Systematischer Beschreibung der bekannten europaischer zweiflugeligen Insekten. Achter oder zweiter Supplementband. H. W. Schmidt, Halle, xvi + 310 + [1] pp." type="journal article" year="1869">Loew, 1869</bibRefCitation>
)
</taxonomicName>
in ZMHB were examined and
<taxonomicName id="33E021A6FFFFC21C01C0FA2CFBEE0EA5" ID-CoL="7G3MM" box="[953,1071,1468,1493]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="senecio">
<emphasis id="C6948637FFFFC21C01C0FA2CFBEE0EA5" box="[953,1071,1468,1493]" italics="true" pageId="7" pageNumber="28">V. senecio</emphasis>
</taxonomicName>
is considered to be a synonym of
<taxonomicName id="33E021A6FFFFC21C03D9FA74FD470E8E" ID-CoL="7FRZK" authority="Fabricius" authorityName="Fabricius" box="[416,646,1508,1534]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="ixion">
<emphasis id="C6948637FFFFC21C03D9FA74FE3B0E8D" box="[416,506,1508,1533]" italics="true" pageId="7" pageNumber="28">V. ixion</emphasis>
(Fabricius)
</taxonomicName>
<emphasis id="C6948637FFFFC21C00F6FA74FCE10E8E" bold="true" box="[655,800,1508,1534]" pageId="7" pageNumber="28">
<taxonomicNameLabel id="DDA73B4CFFFFC21C00F6FA74FCE10E8E" box="[655,800,1508,1534]" pageId="7" pageNumber="28">SYN. NOV.</taxonomicNameLabel>
</emphasis>
It differs from it, as does
<taxonomicName id="33E021A6FFFFC21C061DFA74FE8D0D56" authority="Loew, 1869" authorityName="Loew" authorityYear="1869" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="albula">
<emphasis id="C6948637FFFFC21C061DFA74FB0E0E8D" box="[1124,1231,1508,1533]" italics="true" pageId="7" pageNumber="28">V. albula</emphasis>
(
<bibRefCitation id="907127D4FFFFC21C0698FA74FE850D56" author="Loew" pageId="7" pageNumber="32" refString="Loew, H (1869) Beschreibung europaischer Dipteren. Von Johann Wilhelm Meigen. Erster Band. Systematischer Beschreibung der bekannten europaischer zweiflugeligen Insekten. Achter oder zweiter Supplementband. H. W. Schmidt, Halle, xvi + 310 + [1] pp." type="journal article" year="1869">Loew, 1869</bibRefCitation>
)
</taxonomicName>
previously synonymised with
<taxonomicName id="33E021A6FFFFC21C00CFF99CFB430D56" ID-CoL="7FRZK" authority="Evenhuis &amp; Greathead, 2003" authorityName="Evenhuis &amp; Greathead" authorityYear="2003" box="[694,1154,1548,1574]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="ixion">
<emphasis id="C6948637FFFFC21C00CFF99CFCCF0D55" box="[694,782,1548,1573]" italics="true" pageId="7" pageNumber="28">V. ixion</emphasis>
(
<bibRefCitation id="907127D4FFFFC21C0167F99CFBBB0D56" author="Evenhuis" box="[798,1146,1548,1574]" pageId="7" pageNumber="32" refString="Evenhuis, N. L. &amp; Greathead, D. J. (2003) World Catalog of Bee flies (Diptera- Bombyliidae): Corrigenda and Addenda. Zootaxa, 300, 1 - 64." type="journal article" year="2003">Evenhuis &amp; Greathead, 2003</bibRefCitation>
)
</taxonomicName>
, in not having silver scales on the apical abdominal tergites. It is smaller,
<quantity id="3318F7C0FFFFC21C01BCF9A4FBCC0D3E" box="[965,1037,1588,1614]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="28" unit="mm" value="8.0">8 mm</quantity>
, has only one pale band of whitish scales at the base of tergum 4 and dense long whitish hair. However, the E gyptian specimens have sparse hair, pale scale bands on all abdominal tergites and black scales at the sides of tergites 56 and all thoracic and abdominal bristles and hairs pale, except for rows of black hairs on the hind margins of the terminal abdominal tergites. These specimens do not run to any species in Engel's (1937) key, whether or not the black scales on tergites 56 are considered to constitute tufts. They are very similar in appearance to small specimens of
<taxonomicName id="33E021A6FFFFC21C00D0F8DCFCC30C15" ID-CoL="7FS8L" box="[681,770,1868,1893]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="cana">
<emphasis id="C6948637FFFFC21C00D0F8DCFCC30C15" box="[681,770,1868,1893]" italics="true" pageId="7" pageNumber="28">V. cana</emphasis>
</taxonomicName>
but differ most notably from it in having the alula brownish with a dark thick margin; in having the wing base, costal, and subcostal cells brownish; and in having a few black hairs at sides of 3rd tergite. The male genitalia correspond with the figure for '
<emphasis id="C6948637FFF0C2130008FEFCFD200AF5" box="[625,737,364,389]" italics="true" pageId="8" pageNumber="29">V. s e n e c i o</emphasis>
' in
<bibRefCitation id="907127D4FFF0C2130177FEFCFBA90AF6" author="Theodor" box="[782,1128,364,390]" pageId="8" pageNumber="32" refString="Theodor, O. (1983) The genitalia of Bombyliidae (Diptera). Jerusalem, Israel Academy of Science and Humanities. 275 pp." type="book" year="1983">Theodor (1983: 253 Fig. 719)</bibRefCitation>
in having a long aedeagus and no spine­like outgrowths at the apex of the epiphallus. We consider that Theodor misidentified and dissected a specimen of the present new species.
</paragraph>
</subSubSection>
<subSubSection id="BCFA09AEFFF0C2130341FE74FB9F0E6E" pageId="8" pageNumber="29" type="materials_examined">
<paragraph id="F45F5A25FFF0C2130341FE74FC450F26" blockId="8.[264,1325,284,1310]" pageId="8" pageNumber="29">
<emphasis id="C6948637FFF0C2130341FE74FE620A8D" box="[312,419,484,509]" italics="true" pageId="8" pageNumber="29">
<typeStatus id="2B5BE487FFF0C2130341FE74FE620A8D" box="[312,419,484,509]" pageId="8" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
ɗ. Medium to large sized black species. Body length:
<quantity id="3318F7C0FFF0C2130628FE74FB170A8E" box="[1105,1238,484,510]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="8" pageNumber="29" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">1213 mm</quantity>
. Head. Frons covered with black hairs mixed with yellowish scaly hairs. Upper frons narrow, slightly wider than ocellar tubercle. Face entirely covered with long whitish scaly hairs become more yellowish and larger at middle. Antennae black, with long black hairs on scape mixed with some long white ones at inner side. Thorax. All hairs and scaly hairs on scutum yellowish white becoming brownish at anterior margin. Short scaly hairs at posterior half of scutum form a triangular area the same size of scutellum with its base lying along the line of scutellar suture. Pleural vestiture yellowish white. Wings (
<figureCitation id="6CDB46A0FFF0C21306A0FD6CFAE10866" box="[1241,1312,764,790]" captionStart="FIGURES 1, 2" captionStartId="2.[264,382,1800,1824]" captionTargetBox="[374,1088,1034,1799]" captionTargetId="figure@2.[336,1288,1022,1828]" captionTargetPageId="2" captionText="FIGURES 1, 2. Wings: 1. V. efflatouni sp. nov.; 2. V. stenozoides sp. nov." httpUri="https://zenodo.org/record/171552/files/figure.png" pageId="8" pageNumber="29">Fig. 1</figureCitation>
) hyaline; alula brownish with dark thick margin; wing base, costal, and subcostal cells tinged brownish; veins yellowish to brown. Patagium dark brown, covered with whitish scales; costal comb black; halter yellowish brown to orange with a paler tip. Abdomen. All hairs of abdomen whitish except rows of long black bristly hairs on hind margins of 3 anal tergites, and few black hairs at sides of tergites 34; tergites 27 banded with pale scaly hairs becoming darker at 3 anal ones; tergites with black scales reaching sides; sides of tergites 56 with distinct black scale tufts. Hypopygium (
<figureCitation id="6CDB46A0FFF0C21301D3FB84FC310F5E" box="[938,1008,1044,1070]" captionStart="FIGURES 7, 8" captionStartId="10.[264,382,1390,1414]" captionTargetBox="[300,1281,859,1389]" captionTargetId="figure@10.[279,1308,817,1397]" captionTargetPageId="10" captionText="FIGURES 7, 8. Male genitalia: 7. V. efflatouni sp. nov.: a. lateral view, b. tip of epiphallus; 8. V. stenozoides sp. nov.: a. lateral view, b. tip of epiphallus." httpUri="https://zenodo.org/record/171554/files/figure.png" pageId="8" pageNumber="29">Fig. 7</figureCitation>
) with a long aedeagus and no spine­like outgrowths at the apex of the epiphallus.
</paragraph>
<paragraph id="F45F5A25FFF0C2130341FBF4FE040FD6" blockId="8.[264,1325,284,1310]" pageId="8" pageNumber="29">
<emphasis id="C6948637FFF0C2130341FBF4FE620F0D" box="[312,419,1124,1149]" italics="true" pageId="8" pageNumber="29">
<typeStatus id="2B5BE487FFF0C2130341FBF4FE620F0D" box="[312,419,1124,1149]" pageId="8" pageNumber="29" type="paratype">Paratype</typeStatus>
</emphasis>
Ψ. Like male but upper frons wider, about two times or more as wide as ocellar tubercle.
</paragraph>
<paragraph id="F45F5A25FFF0C2130341FB24FB9F0E6E" blockId="8.[264,1325,284,1310]" pageId="8" pageNumber="29">
<emphasis id="C6948637FFF0C2130341FB24FDD90FBD" box="[312,536,1204,1229]" italics="true" pageId="8" pageNumber="29">Material examined</emphasis>
.
<typeStatus id="2B5BE487FFF0C2130053FB24FD560FBE" box="[554,663,1204,1230]" pageId="8" pageNumber="29" type="holotype">Holotype</typeStatus>
ɗ.
<emphasis id="C6948637FFF0C21300BFFB24FCE90FBE" bold="true" box="[710,808,1204,1230]" pageId="8" pageNumber="29">
<collectingCountry id="8CF71AB5FFF0C21300BFFB24FCE90FBE" box="[710,808,1204,1230]" name="Egypt" pageId="8" pageNumber="29">EGYPT</collectingCountry>
</emphasis>
: Mariout,
<date id="805E7CE5FFF0C21301D3FB24FBE70FBE" box="[938,1062,1204,1230]" pageId="8" pageNumber="29" value="1925-04-18">18.iv.1925</date>
(Efflatoun) [genitalia dissected].
<typeStatus id="2B5BE487FFF0C21303F7FB4CFDC20F86" box="[398,515,1244,1270]" pageId="8" pageNumber="29" type="paratype">Paratypes</typeStatus>
, 1ɗ, same data as
<typeStatus id="2B5BE487FFF0C2130093FB4CFC900F86" box="[746,849,1244,1270]" pageId="8" pageNumber="29" type="holotype">holotype</typeStatus>
[head broken];
<emphasis id="C6948637FFF0C2130665FB4CFBBE0F86" bold="true" box="[1052,1151,1244,1270]" pageId="8" pageNumber="29">
<collectingCountry id="8CF71AB5FFF0C2130665FB4CFBBE0F86" box="[1052,1151,1244,1270]" name="Egypt" pageId="8" pageNumber="29">EGYPT</collectingCountry>
</emphasis>
: 1Ψ, Ismailia,
<date id="805E7CE5FFF0C2130371FA94FE440E6E" box="[264,389,1284,1310]" pageId="8" pageNumber="29" value="1925-06-14">14.vi.1925</date>
(Efflatoun); 1Ψ, same except
<date id="805E7CE5FFF0C213009FFA94FCA60E6E" box="[742,871,1284,1310]" pageId="8" pageNumber="29" value="1925-09-14">14.ix.1925</date>
. [deposited in
<collectionCode id="92F1C2E0FFF0C213066BFA94FB920E6E" box="[1042,1107,1284,1310]" httpUri="http://grbio.org/cool/e8gh-2tfg" name="Cairo University" pageId="8" pageNumber="29">CUE</collectionCode>
].
</paragraph>
</subSubSection>
</treatment>
</document>