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<document id="1D66B95A507DA4DA2476245C7DB4EC0C" ID-CLB-Dataset="107006" ID-DOI="10.11646/zootaxa.5169.1.7" ID-GBIF-Dataset="b1752d3a-a019-4519-a993-08a3f3611f3c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6911323" ID-ZooBank="0B2D6D6D-DB7D-42A4-9D53-238C0D512EA9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658901473937" checkinUser="plazi" docAuthor="Yin, Zi-Wei, Kurbatov, Sergey A. &amp; Ку, A." docDate="2022" docId="78046D27677DB13E488DFCEF29FB2CA8" docLanguage="en" docName="zootaxa.5169.1.7.pdf" docOrigin="Zootaxa 5169 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Batrisini Reitter 1882" docType="treatment" docVersion="5" lastPageNumber="84" masterDocId="843D155F6778B138481AFFAD29372C51" masterDocTitle="Schmidtinus annapurnensis, a new genus and species of Batrisitae from Nepal (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="86" masterPageNumber="79" pageNumber="84" updateTime="1699362183017" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="366F65C54A8704DB3263A1301135F205">Schmidtinus annapurnensis, a new genus and species of Batrisitae from Nepal (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="864282D287588409252D9402839D6E44">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="0A4B4A02DF0D1B2FEF0CFD29E9B8C826">College of Life Sciences, Shanghai Normal University, Shanghai 200234, China</mods:affiliation>
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<mods:namePart id="9EBCF956A2C046890F50C2E4596DC03E">Kurbatov, Sergey A.</mods:namePart>
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<mods:namePart id="40673749E3712CFAF98E3B5C7841A8E4">Ку, A.</mods:namePart>
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<mods:date id="8EE51F660B75B457A2C02B7759F5B9A3">2022</mods:date>
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<treatment id="78046D27677DB13E488DFCEF29FB2CA8" ID-DOI="http://doi.org/10.5281/zenodo.6915071" ID-GBIF-Taxon="197861495" ID-Zenodo-Dep="6915071" LSID="urn:lsid:plazi:treatment:78046D27677DB13E488DFCEF29FB2CA8" httpUri="http://treatment.plazi.org/id/78046D27677DB13E488DFCEF29FB2CA8" lastPageId="6" lastPageNumber="84" pageId="5" pageNumber="84">
<subSubSection id="B8B78FBA677DB13D488DFCEF282C2F0D" box="[151,283,834,860]" pageId="5" pageNumber="84" type="nomenclature">
<paragraph id="F012DC31677DB13D488DFCEF282C2F0D" blockId="5.[151,283,834,860]" box="[151,283,834,860]" pageId="5" pageNumber="84">
<heading id="AB5A6B5D677DB13D488DFCEF282C2F0D" bold="true" box="[151,283,834,860]" fontSize="11" level="1" pageId="5" pageNumber="84" reason="1">
<taxonomicName id="37ADA7B2677DB13D488DFCEF282C2F0D" ID-CoL="8VR76" authorityName="Reitter" authorityYear="1882" box="[151,283,834,860]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="tribe" tribe="Batrisini">
<emphasis id="C2D90023677DB13D488DFCEF282C2F0D" bold="true" box="[151,283,834,860]" pageId="5" pageNumber="84">Discussion</emphasis>
</taxonomicName>
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<subSubSection id="B8B78FBA677DB13D488DFC26280B2840" pageId="5" pageNumber="84" type="description">
<paragraph id="F012DC31677DB13D488DFC26280B2840" blockId="5.[151,1436,907,1041]" pageId="5" pageNumber="84">
<taxonomicName id="37ADA7B2677DB13D488DFC2628172FF4" authority="Yin &amp; Kurbatov &amp; Ку, 2022" authorityName="Yin &amp; Kurbatov &amp; Ку" authorityYear="2022" box="[151,288,907,933]" class="Insecta" family="Staphylinidae" genus="Schmidtinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C2D90023677DB13D488DFC2628172FF4" box="[151,288,907,933]" italics="true" pageId="5" pageNumber="84">Schmidtinus</emphasis>
</taxonomicName>
<emphasis id="C2D90023677DB13D4932FC2628A52FF4" bold="true" box="[296,402,907,933]" pageId="5" pageNumber="84">
<taxonomicNameLabel id="D9EABD58677DB13D4932FC2628A52FF4" box="[296,402,907,933]" pageId="5" pageNumber="84" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is morphologically similar to a number of East Asian genera that are known or putative inquilines of
<taxonomicName id="37ADA7B2677DB13D4909FC02286B2F99" authorityName="Fabricius" authorityYear="1804" box="[275,348,943,968]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4909FC02286B2F99" box="[275,348,943,968]" italics="true" pageId="5" pageNumber="84">Lasius</emphasis>
</taxonomicName>
ants,
<emphasis id="C2D90023677DB13D4987FC0228882F99" box="[413,447,943,968]" italics="true" pageId="5" pageNumber="84">i.e.</emphasis>
,
<taxonomicName id="37ADA7B2677DB13D49D6FC022B122F98" box="[460,549,943,969]" class="Insecta" family="Staphylinidae" genus="Songius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D49D6FC022B122F98" box="[460,549,943,969]" italics="true" pageId="5" pageNumber="84">Songius</emphasis>
</taxonomicName>
,
<taxonomicName id="37ADA7B2677DB13D4A28FC022BBD2F99" authorityName="Yin &amp; Li" authorityYear="2012" box="[562,650,943,968]" class="Insecta" family="Staphylinidae" genus="Tangius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4A28FC022BBD2F99" box="[562,650,943,968]" italics="true" pageId="5" pageNumber="84">Tangius</emphasis>
</taxonomicName>
, and
<taxonomicName id="37ADA7B2677DB13D4ADFFC022AA32F98" box="[709,916,943,969]" class="Insecta" family="Staphylinidae" genus="Dendrolasiophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4ADFFC022AA32F98" box="[709,916,943,969]" italics="true" pageId="5" pageNumber="84">Dendrolasiophilus</emphasis>
</taxonomicName>
. These genera share a similar general structure of the aedeagus, and may form a monophyletic group within Asian
<taxonomicName id="37ADA7B2677DB13D4B80FC7E2AC92FBC" authorityName="Reitter" authorityYear="1882" box="[922,1022,979,1005]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
. They can be separated by using the following key:
</paragraph>
</subSubSection>
<subSubSection id="B8B78FBA677DB13D488DFB912CAB2908" pageId="5" pageNumber="84" type="key">
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<paragraph id="F012DC31677DB13D488DFB912CAB2800" blockId="5.[151,1437,1083,1370]" box="[151,1436,1083,1106]" pageId="5" pageNumber="84">
<keyLead id="4B5C7F04677DB13D488DFB912CAB2800" box="[151,1436,1083,1106]" pageId="5" pageNumber="84">
1 Head strongly transverse; each elytron with one basal fovea........................
<taxonomicName id="37ADA7B2677DB13D4C4FFB962CAB2800" ID-CoL="8XKV3" authority="Nomura, 2010" authorityName="Nomura" authorityYear="2010" box="[1109,1436,1083,1105]" class="Insecta" family="Staphylinidae" genus="Dendrolasiophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4C4FFB962C3B2800" bold="true" box="[1109,1292,1083,1105]" italics="true" pageId="5" pageNumber="84">Dendrolasiophilus</emphasis>
<bibRefCitation id="943CA1C0677DB13D4D08FB912CAB2800" author="Nomura, S." box="[1298,1436,1084,1105]" pageId="5" pageNumber="84" pagination="53 - 60" refId="ref4244" refString="Nomura, S. (2010) A new genus Maajappia and its new species of the subtribe Batrisina, tribe Batrisini (Coleoptera, Staphylinidae, Pselaphinae) from Japan, with a note on the genus Dendrolasiophilus. Elytra, 38, 53 - 60." type="journal article" year="2010">Nomura, 2010</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F012DC31677DB13D488DFBF42CAB283F" blockId="5.[151,1437,1083,1370]" box="[151,1436,1113,1135]" pageId="5" pageNumber="84">
<keyLead id="4B5C7F04677DB13D488DFBF42CAB283F" box="[151,1436,1113,1135]" pageId="5" pageNumber="84">- Head almost as long as to slightly longer than wide; each elytron with three basal foveae............................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="4B59C494677DB13D488DFBDB2CAB28B2" pageId="5" pageNumber="84">
<paragraph id="F012DC31677DB13D488DFBDB2CAB28F8" blockId="5.[151,1437,1083,1370]" pageId="5" pageNumber="84">
<keyLead id="4B5C7F04677DB13D488DFBDB2CAB28F8" pageId="5" pageNumber="84">
2 Sides of tergite 1 (IV) clearly convergent from middle toward base, so base of tergite distinctly..narrower than elytra in their widest part; tergite
<quantity id="375571D4677DB13D4981FB3928F728FB" box="[411,448,1172,1194]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="84" unit="in" value="1.0">1 in</quantity>
dorsal view markedly longer than 2 (V) and 3 (VI) combined............
<taxonomicName id="37ADA7B2677DB13D4CAFFB3E2CAB28F8" ID-CoL="8XQ5X" authority="Yin &amp; Li, 2012" authorityName="Yin &amp; Li" authorityYear="2012" box="[1205,1436,1171,1194]" class="Insecta" family="Staphylinidae" genus="Tangius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4CAFFB3E2C3328F9" bold="true" box="[1205,1284,1171,1192]" italics="true" pageId="5" pageNumber="84">Tangius</emphasis>
<bibRefCitation id="943CA1C0677DB13D4D10FB392CAB28F8" author="Yin, Z. - W. &amp; Li, L. - Z. &amp; Zhao, M. - J." box="[1290,1436,1172,1194]" pageId="5" pageNumber="84" pagination="55 - 61" refId="ref4995" refString="Yin, Z. - W., Li, L. - Z. &amp; Zhao, M. - J. (2012) Contributions to the knowledge of the myrmecophilous pselaphines (Coleoptera, Staphylinidae, Pselaphinae) from China. I. Tangius glabellus gen. et sp. nov. from Tibet. Zootaxa, 3397 (1), 55 - 61. https: // doi. org / 10.11646 / zootaxa. 3397.1.4" type="journal article" year="2012">Yin &amp; Li, 2012</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F012DC31677DB13D488DFB1C2CAB28B2" blockId="5.[151,1437,1083,1370]" pageId="5" pageNumber="84">
<keyLead id="4B5C7F04677DB13D488DFB1C2CAB28B2" pageId="5" pageNumber="84">- Sides of tergite 1 (IV) broadest at or near base, base of tergite of equal width with or slightly..... broader than elytra at their widest part; tergite 1 (IV) in dorsal view as long as or shorter than 2 (V) and 3 (VI) combined........................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="4B59C494677DB13D488DFB412CAB2908" pageId="5" pageNumber="84">
<paragraph id="F012DC31677DB13D488DFB412CAB294F" blockId="5.[151,1437,1083,1370]" pageId="5" pageNumber="84">
<keyLead id="4B5C7F04677DB13D488DFB412CAB294F" pageId="5" pageNumber="84">
3 Vertical heights of postocular margins clearly longer than those of eyes; pronotum trapezoidal, broadest at basal 1/3; elytra lacking discal stria; habitus typically stouter.............................................
<taxonomicName id="37ADA7B2677DB13D4CAEFAA42CAB294F" ID-CoL="8X7VN" authority="Yin &amp; Li, 2010" authorityName="Yin &amp; Li" authorityYear="2010" box="[1204,1436,1289,1311]" class="Insecta" family="Staphylinidae" genus="Songius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4CAEFAA42C33294F" bold="true" box="[1204,1284,1289,1310]" italics="true" pageId="5" pageNumber="84">Songius</emphasis>
<bibRefCitation id="943CA1C0677DB13D4D10FAA42CAB294F" author="Yin, Z. - W. &amp; Li, L. - Z. &amp; Zhao, M. - J." box="[1290,1436,1289,1311]" pageId="5" pageNumber="84" pagination="241 - 253" refId="ref4835" refString="Yin, Z. - W., Li, L. - Z. &amp; Zhao, M. - J. (2010) Contributions to the knowledge of the myrmecophilous pselaphines (Coleoptera, Staphylinidae, Pselaphinae) from China. III. Two new genera and two new species of the subtribe Batrisina (Staphylinidae, Pselaphinae, Batrisitae) from a colony of Lasius niger (Hymeptera (sic!), Formicidae, Formicinae) in East China. Sociobiology, 55, 241 - 253." type="journal article" year="2010">Yin &amp; Li, 2010</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F012DC31677DB13D488DFA8A2CAB2908" blockId="5.[151,1437,1083,1370]" pageId="5" pageNumber="84">
<keyLead id="4B5C7F04677DB13D488DFA8A2CAB2908" pageId="5" pageNumber="84">
- Vertical heights of postocular margins shorter than those of eyes; pronotum roundly explanate laterally, broadest near middle; elytra with discal stria; habitus more elongate.............................................
<emphasis id="C2D90023677DB13D4CDAFAEE2CAB2908" bold="true" box="[1216,1436,1347,1369]" pageId="5" pageNumber="84">
<taxonomicName id="37ADA7B2677DB13D4CDAFAEE2C0B2908" ID-CoL="9GDKV" authority="Yin &amp; Kurbatov &amp; Ку, 2022" authorityName="Yin &amp; Kurbatov &amp; Ку" authorityYear="2022" box="[1216,1340,1347,1369]" class="Insecta" family="Staphylinidae" genus="Schmidtinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C2D90023677DB13D4CDAFAEE2C0B2908" bold="true" box="[1216,1340,1347,1369]" italics="true" pageId="5" pageNumber="84">Schmidtinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9EABD58677DB13D4D58FAE92CAB2908" box="[1346,1436,1348,1369]" pageId="5" pageNumber="84" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="B8B78FBA677DB13E48DDFA2429FB2CA8" lastPageId="6" lastPageNumber="85" pageId="5" pageNumber="84" type="discussion">
<paragraph id="F012DC31677DB13D48DDFA242D032B27" blockId="5.[151,1437,1416,2018]" pageId="5" pageNumber="84">
In addition, the new genus differs from
<taxonomicName id="37ADA7B2677DB13D4A9AFA252BEE29F3" box="[640,729,1416,1442]" class="Insecta" family="Staphylinidae" genus="Songius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4A9AFA252BEE29F3" box="[640,729,1416,1442]" italics="true" pageId="5" pageNumber="84">Songius</emphasis>
</taxonomicName>
also by a different pattern of male sexual characters, i.e., modified antennae, protibiae, and posterior margin of the metaventrite, which have never been observed in the latter genus. More specifically, all
<taxonomicName id="37ADA7B2677DB13D49FEFA7D2BA029BB" box="[484,663,1488,1514]" pageId="5" pageNumber="84">
<emphasis id="C2D90023677DB13D49FEFA7D2B0A29BB" box="[484,573,1488,1514]" italics="true" pageId="5" pageNumber="84">Songius</emphasis>
species
</taxonomicName>
have the aedeagus with a more-or-less serrate apical margin of the dorsal lobe, and there is a short apophysis on the lateral margin of the ventral stalk, both of which are lacking in
<taxonomicName id="37ADA7B2677DB13D488DF9B528172A63" authority="Yin &amp; Kurbatov &amp; Ку, 2022" authorityName="Yin &amp; Kurbatov &amp; Ку" authorityYear="2022" box="[151,288,1560,1586]" class="Insecta" family="Staphylinidae" genus="Schmidtinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C2D90023677DB13D488DF9B528172A63" box="[151,288,1560,1586]" italics="true" pageId="5" pageNumber="84">Schmidtinus</emphasis>
</taxonomicName>
<emphasis id="C2D90023677DB13D4933F9B528A42A63" bold="true" box="[297,403,1560,1586]" pageId="5" pageNumber="84">
<taxonomicNameLabel id="D9EABD58677DB13D4933F9B528A42A63" box="[297,403,1560,1586]" pageId="5" pageNumber="84" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="37ADA7B2677DB13D4986F9B428C32A63" authorityName="Yin &amp; Li" authorityYear="2012" box="[412,500,1561,1586]" class="Insecta" family="Staphylinidae" genus="Tangius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4986F9B428C32A63" box="[412,500,1561,1586]" italics="true" pageId="5" pageNumber="84">Tangius</emphasis>
</taxonomicName>
contains two Himalayan species from
<collectingRegion id="326912D3677DB13D4BAAF9B52ADD2A63" box="[944,1002,1560,1586]" country="China" name="Xizang" pageId="5" pageNumber="84">Tibet</collectingRegion>
(
<collectingCountry id="88BA9CA1677DB13D4BE0F9B52D772A63" box="[1018,1088,1560,1586]" name="China" pageId="5" pageNumber="84">China</collectingCountry>
) and northern
<collectingCountry id="88BA9CA1677DB13D4CF3F9B52C142A63" box="[1257,1315,1560,1586]" name="India" pageId="5" pageNumber="84">India</collectingCountry>
(
<bibRefCitation id="943CA1C0677DB13D4D28F9B529D52A07" author="Yin, Z. - W. &amp; Li, L. - Z." pageId="5" pageNumber="84" pagination="23 - 27" refId="ref4656" refString="Yin, Z. - W. &amp; Li, L. - Z. (2013 b) A new species of Tangius from north India (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys, 346, 23 - 27. https: // doi. org / 10.3897 / zookeys. 346.6233" type="journal article" year="2013">Yin &amp; Li 2013b</bibRefCitation>
);
<taxonomicName id="37ADA7B2677DB13D48EFF99128792A07" box="[245,334,1596,1622]" class="Insecta" family="Staphylinidae" genus="Songius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D48EFF99128792A07" box="[245,334,1596,1622]" italics="true" pageId="5" pageNumber="84">Songius</emphasis>
</taxonomicName>
consists of four species from
<collectingRegion id="326912D3677DB13D4A85F9912A312A07" box="[671,774,1596,1622]" country="China" name="Zhejiang" pageId="5" pageNumber="84">Zhejiang</collectingRegion>
,
<collectingRegion id="326912D3677DB13D4B0AF9902A6A2A06" box="[784,861,1597,1623]" country="China" name="Hunan" pageId="5" pageNumber="84">Hunan</collectingRegion>
, Guanxi, and was shown to inhabit a much broader distributional range in
<collectingCountry id="88BA9CA1677DB13D498CF9CD28EE2A2B" box="[406,473,1632,1658]" name="China" pageId="5" pageNumber="84">China</collectingCountry>
(
<bibRefCitation id="943CA1C0677DB13D49F2F9CD2BA32A2A" author="Yin, Z. - W. &amp; Li, L. - Z." box="[488,660,1632,1659]" pageId="5" pageNumber="84" pagination="293 - 300" refId="ref4775" refString="Yin, Z. - W. &amp; Li, L. - Z. (2015) New and little known species of the genus Songius (Coleoptera: Staphylinidae: Pselaphinae) in China. Zootaxa, 3905 (2), 293 - 300. https: // doi. org / 10.11646 / zootaxa. 3905.2.10" type="journal article" year="2015">Yin &amp; Li 2015</bibRefCitation>
); and
<taxonomicName id="37ADA7B2677DB13D4ACDF9CD2A912A2B" box="[727,934,1632,1658]" class="Insecta" family="Staphylinidae" genus="Dendrolasiophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4ACDF9CD2A912A2B" box="[727,934,1632,1658]" italics="true" pageId="5" pageNumber="84">Dendrolasiophilus</emphasis>
</taxonomicName>
is represented by four species from Far East
<collectingCountry id="88BA9CA1677DB13D488DF92929D22ACF" box="[151,229,1668,1694]" name="Russia" pageId="5" pageNumber="84">Russia</collectingCountry>
,
<collectingCountry id="88BA9CA1677DB13D48F5F92828052ACE" box="[239,306,1669,1695]" name="Japan" pageId="5" pageNumber="84">Japan</collectingCountry>
, and
<collectingCountry id="88BA9CA1677DB13D4973F929289B2ACF" box="[361,428,1668,1694]" name="China" pageId="5" pageNumber="84">China</collectingCountry>
(
<bibRefCitation id="943CA1C0677DB13D49A3F9292B6F2ACF" author="Nomura, S." box="[441,600,1668,1694]" pageId="5" pageNumber="84" pagination="133 - 148" refId="ref4204" refString="Nomura, S. (2008) A new genus Dendrolasiophilus and a new synonym in the subtribe Batrisina, tribe Batrisini (Coleoptera, Staphylinidae, Pselaphinae) from Japan. Elytra, 36, 133 - 148." type="journal article" year="2008">Nomura 2008</bibRefCitation>
,
<bibRefCitation id="943CA1C0677DB13D4A7BF9292BAA2ACF" author="Nomura, S." box="[609,669,1668,1694]" pageId="5" pageNumber="84" pagination="53 - 60" refId="ref4244" refString="Nomura, S. (2010) A new genus Maajappia and its new species of the subtribe Batrisina, tribe Batrisini (Coleoptera, Staphylinidae, Pselaphinae) from Japan, with a note on the genus Dendrolasiophilus. Elytra, 38, 53 - 60." type="journal article" year="2010">2010</bibRefCitation>
;
<bibRefCitation id="943CA1C0677DB13D4ABCF9292A612ACF" author="Yin, Z. - W. &amp; Li, L. - Z." box="[678,854,1668,1694]" pageId="5" pageNumber="84" pagination="183 - 187" refId="ref4566" refString="Yin, Z. - W. &amp; Li, L. - Z. (2013 a) Dendrolasiophilus wenhsini (Coleoptera: Staphylinidae: Pselaphinae), a new myrmecophilous batrisine associated with Lasius (Dendrolasius) spathepus from Guangxi, South China. In: Lin, M. - Y. &amp; Chen, C. - C. (Eds.). In memory of Mr. Wenhsin Lin. Formosa Ecological Company, New Taipei City, pp. 183 - 187." type="book chapter" year="2013">Yin &amp; Li 2013a</bibRefCitation>
;
<bibRefCitation id="943CA1C0677DB13D4B7AF9292DBB2ACF" author="Kurbatov, S. A. &amp; Kovalev, A. V." box="[864,1164,1668,1694]" pageId="5" pageNumber="84" pagination="296 - 300" refId="ref4069" refString="Kurbatov, S. A. &amp; Kovalev, A. V. (2022) A new species of the genus Dendrolasiophilus Nomura, 2010 from the south of the Russian Far East (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa, 5100 (2), 296 - 300. https: // doi. org / 10.11646 / zootaxa. 5100.2.9" type="journal article" year="2022">Kurbatov &amp; Kovalev 2022</bibRefCitation>
). All members of the latter three genera variably possess a suite of ‘reductive’ (
<bibRefCitation id="943CA1C0677DB13D4AE5F9052ACA2A93" author="Parker, J. &amp; Owens, B." box="[767,1021,1704,1730]" pageId="5" pageNumber="84" pagination="219 - 229" refId="ref4292" refString="Parker, J. &amp; Owens, B. (2018) Batriscydmaenus Parker and Owens, new genus, and convergent evolution of a &quot; reductive &quot; ecomorph in socially symbiotic Pselaphinae (Coleoptera: Staphylinidae). Coleopterists Bulletin, 72, 219 - 229. https: // doi. org / 10.1649 / 0010 - 065 X- 72.2.219" type="journal article" year="2018">Parker &amp; Owens 2018</bibRefCitation>
;
<bibRefCitation id="943CA1C0677DB13D4C12F9052D462A93" author="Yin, Z. - W." box="[1032,1137,1704,1730]" pageId="5" pageNumber="84" pagination="233 - 240" refId="ref4458" refString="Yin, Z. - W. (2018) Loeblibatrus Yin, a new genus of myrmecophilous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Coleopterists Bulletin, 72, 233 - 240. https: // doi. org / 10.1649 / 0010 - 065 X- 72.2.233" type="journal article" year="2018">Yin 2018</bibRefCitation>
) morphological traits such as a smooth and glabrous appearance, moniliform antennomeres, a reduced number of foveae, carinae and sulcus on the body surface, and have developed gland-associated trichomes at the posterior corners of the head. Some of these characters are found in
<taxonomicName id="37ADA7B2677DB13D49C4F8B92B502B7F" authority="Yin &amp; Kurbatov &amp; Ку, 2022" authorityName="Yin &amp; Kurbatov &amp; Ку" authorityYear="2022" box="[478,615,1812,1838]" class="Insecta" family="Staphylinidae" genus="Schmidtinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C2D90023677DB13D49C4F8B92B502B7F" box="[478,615,1812,1838]" italics="true" pageId="5" pageNumber="84">Schmidtinus</emphasis>
</taxonomicName>
<emphasis id="C2D90023677DB13D4A74F8B92BE02B7F" bold="true" box="[622,727,1812,1838]" pageId="5" pageNumber="84">
<taxonomicNameLabel id="D9EABD58677DB13D4A74F8B92BE02B7F" box="[622,727,1812,1838]" pageId="5" pageNumber="84" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, which suggests that it can be likewise a myrmecophilic genus. Almost each species of this group is represented by such few specimens in collections that continuing field efforts are needed to further explore the biology and evolutionary history of these beetles.
</paragraph>
<paragraph id="F012DC31677DB13E48DDF82D29FB2CA8" blockId="5.[151,1437,1416,2018]" lastBlockId="6.[151,1437,151,249]" lastPageId="6" lastPageNumber="85" pageId="5" pageNumber="84">
There are a few other recently described batrisine genera from
<collectingCountry id="88BA9CA1677DB13D4B9AF82C2AD02BCB" box="[896,999,1920,1947]" name="China" pageId="5" pageNumber="84">East Asia</collectingCountry>
that host different ant genera and show morphological peculiarities related to myrmecophily, e.g.,
<taxonomicName id="37ADA7B2677DB13D4B2BF8092C212BEF" authority="Yin &amp; Li" authorityName="Yin &amp; Li" box="[817,1302,1956,1982]" class="Insecta" family="Staphylinidae" genus="Pengzhongiella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4B2BF8092AE82BEF" box="[817,991,1956,1982]" italics="true" pageId="5" pageNumber="84">Pengzhongiella</emphasis>
Yin &amp; Li (
<emphasis id="C2D90023677DB13D4C78F8092C3B2BEF" box="[1122,1292,1956,1982]" italics="true" pageId="5" pageNumber="84">Odontomachus</emphasis>
)
</taxonomicName>
,
<taxonomicName id="37ADA7B2677DB13D4D3EF80929F52BB3" authority="Yin" authorityName="Yin" class="Insecta" family="Staphylinidae" genus="Lisubatrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4D3EF8092CAB2BEF" box="[1316,1436,1956,1982]" italics="true" pageId="5" pageNumber="84">Lisubatrus</emphasis>
Yin
</taxonomicName>
,
<taxonomicName id="37ADA7B2677DB13D48D4F8652B622BB3" authority="Yin" authorityName="Yin" box="[206,597,1992,2018]" class="Insecta" family="Staphylinidae" genus="Loeblibatrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D48D4F865286A2BB3" box="[206,349,1992,2018]" italics="true" pageId="5" pageNumber="84">Loeblibatrus</emphasis>
Yin (
<emphasis id="C2D90023677DB13D4986F8642B7D2BB3" box="[412,586,1993,2018]" italics="true" pageId="5" pageNumber="84">Ectomomyrmex</emphasis>
)
</taxonomicName>
, and
<taxonomicName id="37ADA7B2677DB13D4A8BF8652D2E2BB3" authority="Yin &amp; Li" authorityName="Yin &amp; Li" box="[657,1049,1992,2018]" class="Insecta" family="Staphylinidae" genus="Myrmicophila" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677DB13D4A8BF8652A072BB3" box="[657,816,1992,2018]" italics="true" pageId="5" pageNumber="84">Myrmicophila</emphasis>
Yin &amp; Li (
<emphasis id="C2D90023677DB13D4BB1F8642D272BB3" box="[939,1040,1993,2018]" italics="true" pageId="5" pageNumber="84">Myrmica</emphasis>
)
</taxonomicName>
(
<bibRefCitation id="943CA1C0677DB13D4C32F8652DEA2BB3" author="Yin, Z. - W. &amp; Li, L. - Z." box="[1064,1245,1992,2018]" pageId="5" pageNumber="84" pagination="17 - 26" refId="ref4711" refString="Yin, Z. - W. &amp; Li, L. - Z. (2013 c) Pengzhongiella daicongchaoi gen. et sp. n., a remarkable myrmecophile (Staphylinidae, Pselaphinae, Batrisitae) from the Gaoligong Mountains. ZooKeys, 326, 17 - 26. https: // doi. org / 10.3897 / zookeys. 326.5933" type="journal article" year="2013">Yin &amp; Li 2013c</bibRefCitation>
;
<bibRefCitation id="943CA1C0677DB13D4CF2F8652C652BB3" author="Yin, Z. - W." box="[1256,1362,1992,2018]" pageId="5" pageNumber="84" pagination="593 - 601" refId="ref4400" refString="Yin, Z. - W. (2017) Lisubatrus dongzhiweii gen. et sp. nov. from the Gaoligong Mountains, China (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae, 57, 593 - 601. https: // doi. org / 10.1515 / aemnp- 2017 - 0091" type="journal article" year="2017">Yin 2017</bibRefCitation>
,
<bibRefCitation id="943CA1C0677DB13D4D47F8652CAF2BB3" author="Yin, Z. - W." box="[1373,1432,1992,2018]" pageId="5" pageNumber="84" pagination="233 - 240" refId="ref4458" refString="Yin, Z. - W. (2018) Loeblibatrus Yin, a new genus of myrmecophilous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Coleopterists Bulletin, 72, 233 - 240. https: // doi. org / 10.1649 / 0010 - 065 X- 72.2.233" type="journal article" year="2018">2018</bibRefCitation>
,
<bibRefCitation id="943CA1C0677EB13E488DFF3A29E52CE0" author="Yin, Z. - W." box="[151,210,151,177]" pageId="6" pageNumber="85" pagination="684 - 688" refId="ref4509" refString="Yin, Z. - W. (2021) A new species of the rare myrmecophilous genus Myrmicophila Yin and Li from Xizang, China (Coleoptera: Staphylinidae: Pselaphinae). Coleopterists Bulletin, 75, 684 - 688. https: // doi. org / 10.1649 / 0010 - 065 X- 75.3.684" type="journal article" year="2021">2021</bibRefCitation>
;
<bibRefCitation id="943CA1C0677EB13E48C6FF3A28B52CE0" author="Yin, Z. - W. &amp; Li, L. - Z. &amp; Zhao, M. - J." box="[220,386,151,177]" pageId="6" pageNumber="85" pagination="175 - 183" refId="ref4928" refString="Yin, Z. - W., Li, L. - Z. &amp; Zhao, M. - J. (2011) Contributions to the knowledge of the myrmecophilous pselaphines (Coleoptera, Staphylinidae, Pselaphinae) from China. VIII. Myrmicophila tangliangi, a new genus and species of Batrisini from Yunnan. Sociobiology, 58, 175 - 183." type="journal article" year="2011">
Yin
<emphasis id="C2D90023677EB13E4911FF3A280C2CE0" box="[267,315,151,177]" italics="true" pageId="6" pageNumber="85">et al</emphasis>
. 2011
</bibRefCitation>
;
<bibRefCitation id="943CA1C0677EB13E4997FF3A2B562CE0" author="Zhang, W. - X. &amp; Yin, Z. - W." box="[397,609,151,177]" pageId="6" pageNumber="85" pagination="129 - 136" refId="ref5076" refString="Zhang, W. - X. &amp; Yin, Z. - W. (2022) Two new species of myrmecophilous Pselaphinae from China (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 129, 129 - 136. https: // doi. org / 10.35929 / RSZ. 0067" type="journal article" year="2022">Zhang &amp; Yin 2022</bibRefCitation>
). They are morphologically distinct from the
<taxonomicName id="37ADA7B2677EB13E4C4DFF3A2C112CE0" box="[1111,1318,151,177]" class="Insecta" family="Staphylinidae" genus="Dendrolasiophilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="C2D90023677EB13E4C4DFF3A2C112CE0" box="[1111,1318,151,177]" italics="true" pageId="6" pageNumber="85">Dendrolasiophilus</emphasis>
</taxonomicName>
group and may have separately evolved in accordance with the post-Cretaceous diversification (Moreau
<emphasis id="C2D90023677EB13E4CBFFF162DE12C84" box="[1189,1238,187,213]" italics="true" pageId="6" pageNumber="85">et al</emphasis>
. 2016) of modern ants.
</paragraph>
</subSubSection>
</treatment>
</document>
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