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<document id="D6C9F55EFDF3FEAF2ED1823E71BE4E68" ID-CLB-Dataset="24707" ID-DOI="10.11646/zootaxa.4706.1.1" ID-GBIF-Dataset="ec3e13ba-b3ea-4f5f-b9bc-9816967a8561" ID-ISSN="1175-5326" ID-Zenodo-Dep="3564999" ID-ZooBank="68EE7CB3-4DCE-4502-9895-C6C624E51A11" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575618127180" checkinUser="plazi" docAuthor="Naruse, Tohru" docDate="2019" docId="7206B76E3B35C513308DFF69E632F870" docLanguage="en" docName="zootaxa..4706.1.1.pdf" docOrigin="Zootaxa 4706 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Conchoecetes pembawa Naruse 2019, n. sp." docType="treatment" docVersion="8" lastPageNumber="38" masterDocId="8E3FCF163B10C536301AFFF0E300FFAF" masterDocTitle="Revision of the shell-carrying crab genus Conchoecetes Stimpson, 1858 (Crustacea: Brachyura: Dromiidae)" masterLastPageNumber="47" masterPageNumber="1" pageNumber="38" updateTime="1698773417815" updateUser="ExternalLinkService">
<mods:mods id="44FA6EBE8B1866D330DBC865D774B6C2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="FD49B491CB0A866CE60AFFCF79CF33CE">Revision of the shell-carrying crab genus Conchoecetes Stimpson, 1858 (Crustacea: Brachyura: Dromiidae)</mods:title>
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<mods:namePart id="43C36787E0FD46316C7F5D4A564A7F81">Naruse, Tohru</mods:namePart>
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<mods:title id="7E8B3E576AA97AF1B547053ED600C73B">Zootaxa</mods:title>
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<mods:part id="A7BA7BF0D611F234FE2C38F0870A15A0">
<mods:date id="6B1E4F1AC8854B5140DED501EEA588AD">2019</mods:date>
<mods:detail id="F1C7E8B334D52DF835233676E4E630F4" type="pubDate">
<mods:number id="ED7982AB804FA8E214CC76849709D9F2">2019-12-06</mods:number>
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<mods:number id="A8DFE02D0C93D0298CF6D7C72CF11E67">4706</mods:number>
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<mods:number id="8E74F35B3FF5A5D8575A5C6ECA350102">1</mods:number>
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<mods:classification id="4C4A933B88CBA16BC39B38B6FB1B286D">journal article</mods:classification>
<mods:identifier id="2DBCF542494D16135D50902F02B2871A" type="CLB-Dataset">24707</mods:identifier>
<mods:identifier id="484CFC1310DB5E4556A6BBAFA74C4C8E" type="DOI">10.11646/zootaxa.4706.1.1</mods:identifier>
<mods:identifier id="433891AF04A28C470C80271D066DC8EF" type="GBIF-Dataset">ec3e13ba-b3ea-4f5f-b9bc-9816967a8561</mods:identifier>
<mods:identifier id="A28E48F3B399E613E211EE78A51B501D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DB216783346CD4968A56A020A2BEAA7F" type="Zenodo-Dep">3564999</mods:identifier>
<mods:identifier id="CD531E6AC600F9010E624C00E8A45161" type="ZooBank">68EE7CB3-4DCE-4502-9895-C6C624E51A11</mods:identifier>
</mods:mods>
<treatment id="7206B76E3B35C513308DFF69E632F870" ID-DOI="http://doi.org/10.5281/zenodo.5668703" ID-GBIF-Taxon="160671573" ID-Zenodo-Dep="5668703" LSID="urn:lsid:plazi:treatment:7206B76E3B35C513308DFF69E632F870" httpUri="http://treatment.plazi.org/id/7206B76E3B35C513308DFF69E632F870" lastPageNumber="38" pageId="37" pageNumber="38">
<subSubSection id="B2B555F33B35C513308DFF69E101FF1B" box="[151,513,153,180]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="FA1006783B35C513308DFF69E101FF1B" blockId="37.[151,513,153,215]" box="[151,513,153,180]" pageId="37" pageNumber="38">
<heading id="A158B1143B35C513308DFF69E101FF1B" bold="true" box="[151,513,153,180]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
<emphasis id="C8DBDA6A3B35C513308DFF69E101FF1B" bold="true" box="[151,513,153,180]" pageId="37" pageNumber="38">
<taxonomicName id="3DAF7DFB3B35C513308DFF69E2B7FF1B" ID-CoL="84GS4" authority="Naruse, 2019" authorityName="Naruse" authorityYear="2019" box="[151,439,153,180]" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pembawa" status="sp. nov.">
<emphasis id="C8DBDA6A3B35C513308DFF69E2B7FF1B" bold="true" box="[151,439,153,180]" italics="true" pageId="37" pageNumber="38">Conchoecetes pembawa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3E867113B35C51331A4FF6AE101FF1B" box="[446,513,154,180]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B2B555F33B35C513308DFF4DE22EFF78" box="[151,302,189,215]" pageId="37" pageNumber="38" type="description">
<paragraph id="FA1006783B35C513308DFF4DE22EFF78" blockId="37.[151,513,153,215]" box="[151,302,189,215]" pageId="37" pageNumber="38">
(
<figureCitation id="62941AFD3B35C5133085FF4DE226FF78" box="[159,294,189,215]" captionStart-0="FIGURE 22" captionStart-1="FIGURE 23" captionStart-2="FIGURE 24" captionStartId-0="38.[151,250,1960,1985]" captionStartId-1="39.[151,250,1934,1959]" captionStartId-2="40.[151,250,1864,1889]" captionTargetBox-0="[301,1286,166,1940]" captionTargetBox-1="[291,1296,166,1915]" captionTargetBox-2="[217,1371,169,1844]" captionTargetId-0="figure@38.[301,1286,166,1940]" captionTargetId-1="figure@39.[291,1296,166,1915]" captionTargetId-2="figure@40.[215,1372,166,1844]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionTargetPageId-2="40" captionText-0="FIGURE 22. Conchoecetes pembawa n. sp. ZRC 2019.1632, holotype, male, 34.4 × 34.0 mm, Tokong, Penang Is, Malaysia. A, dorsal view; B, frontal view; C, ventral view." captionText-1="FIGURE 23. Conchoecetes pembawa n. sp., (A, B) ZRC 2019.1632, holotype, male, 34.4 × 34.0 mm, Tokong, Penang Is, Malaysia; (C) WAM 85-83, male, 25.5 × 25.8 mm, Point Hedland, Australia. A, frontal view of right chela; B, dorsal view of right chela; C, dorsal view." captionText-2="FIGURE 24. Conchoecetes pembawa n. sp., Tokong, Penang I., Malaysia: (A–C) ZRC 2019. 1665, paratype, ovig. female, 27.0 x 25.8 mm; (D, E) female, 24.6 × 23.2 mm; and (F) female, 25.4 × 24.0 mm. A, dorsal view; B, frontal view of chela and anterolateral margin; C, dorsal view of carapace and chela; D, ventral view of thoracic sternum, with sternal sutures and gonopores; E, front profile view of unblocked sutures; F, ventral view of blocked sternal sutures and pleon and telson. (Labels: Gp—female gonopore; Pq—plaque; Sg—sternal groove; Sp—spermathecal aperture)." figureDoi-0="http://doi.org/10.5281/zenodo.3565043" figureDoi-1="http://doi.org/10.5281/zenodo.3565045" figureDoi-2="http://doi.org/10.5281/zenodo.3565047" httpUri-0="https://zenodo.org/record/3565043/files/figure.png" httpUri-1="https://zenodo.org/record/3565045/files/figure.png" httpUri-2="https://zenodo.org/record/3565047/files/figure.png" pageId="37" pageNumber="38">Figs. 22–24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B2B555F33B35C513308DFEF6E255FE58" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="FA1006783B35C513308DFEF6E2E0FEEC" blockId="37.[151,1437,261,2016]" pageId="37" pageNumber="38">
<emphasis id="C8DBDA6A3B35C513308DFEF6E3DBFE8F" bold="true" box="[151,219,262,288]" pageId="37" pageNumber="38">Type:</emphasis>
<materialsCitation id="4AC70C253B35C51330F8FEF5E00DFE8F" ID-GBIF-Occurrence="2464720923" box="[226,781,261,288]" collectionCode="ZRC" pageId="37" pageNumber="38" specimenCode="ZRC 2019.1632" specimenCount="1" typeStatus="holotype">
<specimenCode id="AA09AE033B35C51330F8FEF5E299FEB0" box="[226,409,261,287]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="37" pageNumber="38">ZRC 2019.1632</specimenCode>
,
<typeStatus id="2514B8DA3B35C51331BEFEF5E109FEB0" box="[420,521,261,287]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
, male, 34.4 × 34.0 mm
</materialsCitation>
,
<materialsCitation id="4AC70C253B35C5133303FEF5E2DCFEEC" ID-GBIF-Occurrence="2464719917" collectionCode="ZRC" country="Malaysia" location="Tokong" pageId="37" pageNumber="38" specimenCode="ZRC 2019.1665" specimenCount="1" stateProvince="Pulau Pinang" typeStatus="paratype">
<specimenCode id="AA09AE033B35C5133303FEF5E0D0FE8F" box="[793,976,261,288]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="37" pageNumber="38">ZRC 2019.1665</specimenCode>
,
<typeStatus id="2514B8DA3B35C51333C1FEF6E73EFE8F" box="[987,1086,262,288]" pageId="37" pageNumber="38" type="paratype">paratype</typeStatus>
, ovig. female, 27.0 ×
<quantity id="3D57AB9D3B35C5133528FEF5E698FE8F" box="[1330,1432,261,288]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="37" pageNumber="38" unit="mm" value="25.8">25.8 mm</quantity>
,
<location id="FF7050A33B35C513308DFED9E3EFFEEC" LSID="urn:lsid:plazi:treatment:7206B76E3B35C513308DFF69E632F870:FF7050A33B35C513308DFED9E3EFFEEC" box="[151,239,297,323]" country="Malaysia" name="Tokong" pageId="37" pageNumber="38" stateProvince="Pulau Pinang">Tokong</location>
,
<collectingRegion id="386BC89A3B35C51330E0FEDAE24CFEEB" box="[250,332,298,324]" country="Malaysia" name="Pulau Pinang" pageId="37" pageNumber="38">Penang</collectingRegion>
I.,
<collectingCountry id="82B846E83B35C5133168FED9E2DCFEEC" box="[370,476,297,323]" name="Malaysia" pageId="37" pageNumber="38">Malaysia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="FA1006783B35C51330DDFEBDE2ADFE00" blockId="37.[151,1437,261,2016]" pageId="37" pageNumber="38">
<emphasis id="C8DBDA6A3B35C51330DDFEBDE2B6FEC7" bold="true" box="[199,438,333,360]" pageId="37" pageNumber="38">Material examined.</emphasis>
<materialsCitation id="4AC70C253B35C51331D8FEBDE2A9FE00" ID-GBIF-Occurrence="2464721914" country="Malaysia" latitude="5.4" location="Tokong" longLatPrecision="1305" longitude="100.23333" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Pulau Pinang">
<emphasis id="C8DBDA6A3B35C51331D8FEBDE138FEC7" bold="true" box="[450,568,333,360]" pageId="37" pageNumber="38">
<collectingCountry id="82B846E83B35C51331D8FEBDE133FEC7" box="[450,563,333,360]" name="Malaysia" pageId="37" pageNumber="38">Malaysia</collectingCountry>
:
</emphasis>
WAM-64-83, ovig. female 24.8 ×
<quantity id="3D57AB9D3B35C51333C7FEBDE748FEC7" box="[989,1096,333,360]" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="37" pageNumber="38" unit="mm" value="24.3">24.3 mm</quantity>
,
<location id="FF7050A33B35C5133442FEBDE7B0FEC8" LSID="urn:lsid:plazi:treatment:7206B76E3B35C513308DFF69E632F870:FF7050A33B35C5133442FEBDE7B0FEC8" box="[1112,1200,333,359]" country="Malaysia" latitude="5.4" longLatPrecision="1305" longitude="100.23333" name="Tokong" pageId="37" pageNumber="38" stateProvince="Pulau Pinang">Tokong</location>
,
<collectingRegion id="386BC89A3B35C51334DAFEBEE612FEC7" box="[1216,1298,334,360]" country="Malaysia" name="Pulau Pinang" pageId="37" pageNumber="38">Penang</collectingRegion>
I.,
<geoCoordinate id="9F9B60BF3B35C5133558FEBDE697FEC7" box="[1346,1431,333,360]" degrees="5" direction="north" minutes="24" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="5.4">5°24’N</geoCoordinate>
,
<geoCoordinate id="9F9B60BF3B35C513308DFE81E206FE24" box="[151,262,369,395]" degrees="100" direction="east" minutes="14" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="100.23333">100°14’E</geoCoordinate>
; NHMB-87551 (was ZMUC-CRU-10106), mature female 32.6 ×
<quantity id="3D57AB9D3B35C51333E0FE81E766FE23" box="[1018,1126,369,396]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="37" pageNumber="38" unit="mm" value="31.8">31.8 mm</quantity>
., exact origin unknown but probably from Malaysia
</materialsCitation>
.
</paragraph>
<paragraph id="FA1006783B35C51330DDFE4AE255FE58" blockId="37.[151,1437,261,2016]" pageId="37" pageNumber="38">
<materialsCitation id="4AC70C253B35C51330DDFE4AE251FE58" ID-GBIF-Occurrence="2464718919" collectingDate="1982-02-28" collectionCode="CSIRO" elevation="150" latitude="-18.683332" location="Western Australia" longLatPrecision="1274" longitude="117.833336" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Western Australia">
<emphasis id="C8DBDA6A3B35C51330DDFE4AE29FFE7B" bold="true" box="[199,415,441,468]" pageId="37" pageNumber="38">
<collectingRegion id="386BC89A3B35C51330DDFE4AE29FFE7B" box="[199,415,441,468]" country="Australia" name="Western Australia" pageId="37" pageNumber="38">Western Australia</collectingRegion>
</emphasis>
: WAM-85-83, male 25.5 ×
<quantity id="3D57AB9D3B35C51332CBFE49E036FE7B" box="[721,822,441,468]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="37" pageNumber="38" unit="mm" value="25.8">25.8 mm</quantity>
,
<collectionCode id="9CBE9EBD3B35C513335AFE49E092FE7C" box="[832,914,441,467]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="37" pageNumber="38" type="Museum">CSIRO</collectionCode>
“Soela”,
<geoCoordinate id="9F9B60BF3B35C51333E6FE49E75EFE7C" box="[1020,1118,441,467]" degrees="18" direction="south" minutes="41" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-18.683332">18º 41’S</geoCoordinate>
,
<geoCoordinate id="9F9B60BF3B35C5133473FE49E7D0FE7B" box="[1129,1232,441,468]" degrees="117" direction="east" minutes="50" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="117.833336">117º50’E</geoCoordinate>
(Pt Hedland),
<quantity id="3D57AB9D3B35C5133568FE49E3AFFE57" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="38" unit="m" value="150.0">
<elevation id="7182E14B3B35C5133568FE49E3AFFE57" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="38" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="8E1120B83B35C51330A1FE2DE251FE58" box="[187,337,477,503]" pageId="37" pageNumber="38" value="1982-02-28">
<collectingDate id="9E55D9503B35C51330A1FE2DE251FE58" box="[187,337,477,503]" pageId="37" pageNumber="38" value="1982-02-28">28 Feb. 1982</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B555F33B35C51330DDFDF1E3E0FA0F" pageId="37" pageNumber="38" type="description">
<paragraph id="FA1006783B35C51330DDFDF1E659FC2C" blockId="37.[151,1437,261,2016]" pageId="37" pageNumber="38">
<emphasis id="C8DBDA6A3B35C51330DDFDF1E259FDB3" bold="true" box="[199,345,513,540]" pageId="37" pageNumber="38">Description.</emphasis>
Carapace about as wide as long; surface smooth covered with sparse, short, soft tomentum; regions well defined by prominent, cervical, and branchial grooves; rostrum tridentate, lateral teeth longer than median tooth, projecting well forward; median tooth on lower level, deflexed; teeth edges finely granulate; supraorbital tooth well developed, obscuring similar suborbital tooth beneath; no post-orbital tooth. Anterolateral margin complex, sculptured; behind orbit carapace surface concave sloping ventrally without forming an edge; further back margin is edged armed with several small tubercles and 2 larger ones sloping down below; edge continues posteriorly without tubercles followed by concave section preceding anterolateral tooth behind cervical groove; prominent blunt tooth is curved upwards; posteriorly margin is convergent to branchial groove preceding posterolateral tooth which is prominent directed upwards; posterior lateral margin may have several small tubercles. Sub-hepatic area has prominent blunt shelf at lower level and anterior to anterolateral tooth; shelf variable in size bearing several unequal granules at about same level. All structures on anterolateral margin and hepatic area visible dorsally.
</paragraph>
<paragraph id="FA1006783B35C51330DDFC7DE7EEFC08" blockId="37.[151,1437,261,2016]" box="[199,1262,909,936]" pageId="37" pageNumber="38">Sternal sutures 7/8 end apart on scroll-like structures, tube-like in profile, between bases of P2.</paragraph>
<paragraph id="FA1006783B35C51330DDFC41E7F9FC64" blockId="37.[151,1437,261,2016]" box="[199,1273,945,972]" pageId="37" pageNumber="38">Third maxillipeds operculiform, crista dentate has 7 to 10 stout acute teeth, directed proximally.</paragraph>
<paragraph id="FA1006783B35C51330DDFC25E124FA9C" blockId="37.[151,1437,261,2016]" pageId="37" pageNumber="38">Cheliped carpus mostly smooth except for granules along inner dorsal margin and 4 or 5 scattered granules on superior face and 2 large distal tubercles; propodus mostly smooth except for granules along dorsal margins and several prominent granules arranged in row-like on outer propodal face, several unequal, large distal tubercles at articulation with dactyl; dorsal surface of dactylus covered in small closely spaced granules; margins of fingers setose, armed with 9 or 10 blunt interlocking white teeth. P2 and P3 meri have few granules along distal border, otherwise unarmed; dactyli fringed with two rows of dorsal setae and row of 20 to 30 tiny adpressed spines on inner margin. Anterior margin of P4 carpus has 8 or 9 small, irregularly distributed, blunt granules; propodus has stout quadrangular proximal projection that engages with strong curved dactylus which folds back to make shell grasping mechanism; talon-like dactylus over-reaches quadrangular process. P5 shortest pereopod, sub-dorsal, flattened, dactylus short and curved upwards.</paragraph>
<paragraph id="FA1006783B35C51330DDFACDE3E0FA0F" blockId="37.[151,1437,261,2016]" pageId="37" pageNumber="38">Female telson triangular, posterior margin convex; uropods large and visible externally. Male telson sub-triangular, wider than long, posterior margin convex; uropods visible externally, locking in front of tubercles on P2 coxae.</paragraph>
</subSubSection>
<subSubSection id="B2B555F33B35C51330DDFA5AE7D2F9FB" pageId="37" pageNumber="38" type="etymology">
<paragraph id="FA1006783B35C51330DDFA5AE74BFA48" blockId="37.[151,1437,261,2016]" pageId="37" pageNumber="38">
<emphasis id="C8DBDA6A3B35C51330DDFA5AE248FA6B" bold="true" box="[199,328,1450,1476]" pageId="37" pageNumber="38">Etymology</emphasis>
. The specific name “pembawa” is the Malay word for “carrier”, alluding to the habit of this crab carrying a bivalve shell as a mobile shelter and the
<typeStatus id="2514B8DA3B35C51332CFFA3EE005FA47" box="[725,773,1486,1512]" pageId="37" pageNumber="38">type</typeStatus>
locality,
<collectingRegion id="386BC89A3B35C5133376FA3EE0BEFA47" box="[876,958,1486,1512]" country="Malaysia" name="Pulau Pinang" pageId="37" pageNumber="38">Penang</collectingRegion>
Is., Malaya.
</paragraph>
<paragraph id="FA1006783B35C51330DDFA01E7D2F9FB" blockId="37.[151,1437,261,2016]" pageId="37" pageNumber="38">
<emphasis id="C8DBDA6A3B35C51330DDFA01E22AF9A3" bold="true" box="[199,298,1521,1548]" pageId="37" pageNumber="38">Habitat.</emphasis>
The only verified depth is
<quantity id="3D57AB9D3B35C513327DFA01E1B0F9A3" box="[615,688,1521,1548]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="38" unit="m" value="150.0">150 m</quantity>
from Point Hedland, Western Australia, otherwise the specimens from Peninsula
<collectingCountry id="82B846E83B35C5133153F9E5E2B0F980" box="[329,432,1557,1583]" name="Malaysia" pageId="37" pageNumber="38">Malaysia</collectingCountry>
came from coastal waters of uncertain depth. They were recovered from refuse delivered to the fish port by shrimp fishermen operating in depths less than
<quantity id="3D57AB9D3B35C513336DF9C9E0BEF9FB" box="[887,958,1593,1620]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="38" unit="m" value="100.0">100 m</quantity>
(Peter Ng, pers. comm.)
</paragraph>
</subSubSection>
<subSubSection id="B2B555F33B35C51330DDF9ADE3D8F934" pageId="37" pageNumber="38" type="distribution">
<paragraph id="FA1006783B35C51330DDF9ADE3D8F934" blockId="37.[151,1437,261,2016]" pageId="37" pageNumber="38">
<emphasis id="C8DBDA6A3B35C51330DDF9ADE261F9D7" bold="true" box="[199,353,1629,1656]" pageId="37" pageNumber="38">Distribution.</emphasis>
Known only from Malaysian. Peninsula in vicinity of Penang and Point Hedland,
<collectingRegion id="386BC89A3B35C513351FF9AEE3D4F934" country="Australia" name="Western Australia" pageId="37" pageNumber="38">Western Australia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B555F33B35C51330DDF956E632F870" pageId="37" pageNumber="38" type="discussion">
<paragraph id="FA1006783B35C51330DDF956E632F870" blockId="37.[151,1437,261,2016]" pageId="37" pageNumber="38">
<emphasis id="C8DBDA6A3B35C51330DDF956E23BF96F" bold="true" box="[199,315,1702,1728]" pageId="37" pageNumber="38">Remarks.</emphasis>
<taxonomicName id="3DAF7DFB3B35C5133158F955E14FF910" authority="Naruse, 2019" authorityName="Naruse" authorityYear="2019" box="[322,591,1701,1727]" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pembawa" status="sp. nov.">
<emphasis id="C8DBDA6A3B35C5133158F955E14FF910" box="[322,591,1701,1727]" italics="true" pageId="37" pageNumber="38">Conchoecetes pembawa</emphasis>
</taxonomicName>
<emphasis id="C8DBDA6A3B35C513324FF956E194F96F" bold="true" box="[597,660,1702,1728]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D3E867113B35C513324FF956E194F96F" box="[597,660,1702,1728]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="62941AFD3B35C51332B9F955E000F910" box="[675,768,1701,1727]" captionStart="FIGURE 22" captionStartId="38.[151,250,1960,1985]" captionTargetBox="[301,1286,166,1940]" captionTargetId="figure@38.[301,1286,166,1940]" captionTargetPageId="38" captionText="FIGURE 22. Conchoecetes pembawa n. sp. ZRC 2019.1632, holotype, male, 34.4 × 34.0 mm, Tokong, Penang Is, Malaysia. A, dorsal view; B, frontal view; C, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3565043" httpUri="https://zenodo.org/record/3565043/files/figure.png" pageId="37" pageNumber="38">Figs. 22</figureCitation>
,
<figureCitation id="62941AFD3B35C5133311F955E028F910" box="[779,808,1701,1727]" captionStart="FIGURE 24" captionStartId="40.[151,250,1864,1889]" captionTargetBox="[217,1371,169,1844]" captionTargetId="figure@40.[215,1372,166,1844]" captionTargetPageId="40" captionText="FIGURE 24. Conchoecetes pembawa n. sp., Tokong, Penang I., Malaysia: (A–C) ZRC 2019. 1665, paratype, ovig. female, 27.0 x 25.8 mm; (D, E) female, 24.6 × 23.2 mm; and (F) female, 25.4 × 24.0 mm. A, dorsal view; B, frontal view of chela and anterolateral margin; C, dorsal view of carapace and chela; D, ventral view of thoracic sternum, with sternal sutures and gonopores; E, front profile view of unblocked sutures; F, ventral view of blocked sternal sutures and pleon and telson. (Labels: Gp—female gonopore; Pq—plaque; Sg—sternal groove; Sp—spermathecal aperture)." figureDoi="http://doi.org/10.5281/zenodo.3565047" httpUri="https://zenodo.org/record/3565047/files/figure.png" pageId="37" pageNumber="38">24</figureCitation>
) is sympatric with two other species of
<taxonomicName id="3DAF7DFB3B35C51334F9F955E67EF910" authorityName="Stimpson" authorityYear="1858" box="[1251,1406,1701,1727]" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="C8DBDA6A3B35C51334F9F955E67EF910" box="[1251,1406,1701,1727]" italics="true" pageId="37" pageNumber="38">Conchoecetes</emphasis>
</taxonomicName>
: it can be distinguished from
<taxonomicName id="3DAF7DFB3B35C51331A6F93AE111F94C" authority="Naruse, 2019" authorityName="Naruse" authorityYear="2019" box="[444,529,1737,1764]" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="atlas" status="sp. nov.">
<emphasis id="C8DBDA6A3B35C51331A6F93AE111F94C" box="[444,529,1737,1764]" italics="true" pageId="37" pageNumber="38">C. atlas</emphasis>
</taxonomicName>
<emphasis id="C8DBDA6A3B35C513320DF93AE155F94B" bold="true" box="[535,597,1738,1764]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D3E867113B35C513320DF93AE155F94B" box="[535,597,1738,1764]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="62941AFD3B35C5133279F939E1B1F94C" box="[611,689,1737,1763]" captionStart="FIGURE 4" captionStartId="11.[151,250,1966,1991]" captionTargetBox="[292,1295,166,1946]" captionTargetId="figure@11.[292,1295,166,1946]" captionTargetPageId="11" captionText="FIGURE 4. Conchoecetes atlas n. sp., ZRC 2019.1631, holotype, male, 32.5 × 31.1 mm, Camiguin Island, Bohol Sea, Philippines. A, dorsal view; B, frontal view; C, ventral view of anterior region." figureDoi="http://doi.org/10.5281/zenodo.3565007" httpUri="https://zenodo.org/record/3565007/files/figure.png" pageId="37" pageNumber="38">Figs. 4</figureCitation>
,
<figureCitation id="62941AFD3B35C51332A6F939E1CAF94B" box="[700,714,1737,1764]" captionStart="FIGURE 5" captionStartId="12.[151,250,1924,1949]" captionTargetBox="[393,1192,167,1905]" captionTargetId="figure@12.[393,1193,166,1905]" captionTargetPageId="12" captionText="FIGURE 5. Conchoecetes atlas n. sp., dorsal views. A, SUF Cat. No. 9161, male, 29.7 × 29.2 mm, Ohshima, Japan; B, ZRC1999.0463, male, 30.0 × 29.8 mm, Guangdong, Nanao Island, China; C, NHMB-87555 (was ZMUC-CRU-10110), male, 28 × 27.5 mm, Taiwan." figureDoi="http://doi.org/10.5281/zenodo.3565009" httpUri="https://zenodo.org/record/3565009/files/figure.png" pageId="37" pageNumber="38">5</figureCitation>
) by having carapace regions clearly defined by deep grooves (regions not defined by deep grooves), posterolateral carapace margins sub-parallel (margins convergent), anterolateral tooth well defined, strong, blunt directed anterolaterally (tooth short, shoulder—like, wedge-shaped), posterolateral tooth well developed, similar to anterolateral tooth (no posterolateral tooth). Using the same characters
<taxonomicName id="3DAF7DFB3B35C513350BF8C6E69CF8E0" authorityName="Naruse" authorityYear="2019" box="[1297,1436,1845,1872]" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pembawa">
<emphasis id="C8DBDA6A3B35C513350BF8C6E69CF8E0" box="[1297,1436,1845,1872]" italics="true" pageId="37" pageNumber="38">C. pembawa</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="3DAF7DFB3B35C5133181F8A9E1C1F8DC" baseAuthorityName="Adams, in Belcher" baseAuthorityYear="1848" box="[411,705,1881,1907]" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pectenicola">
<emphasis id="C8DBDA6A3B35C5133181F8A9E1C1F8DC" box="[411,705,1881,1907]" italics="true" pageId="37" pageNumber="38">Conchoecetes pectenicola</emphasis>
</taxonomicName>
(
<figureCitation id="62941AFD3B35C51332C8F8A9E031F8DC" box="[722,817,1881,1907]" captionStart="FIGURE 19" captionStartId="34.[151,250,1744,1769]" captionTargetBox="[152,1436,445,1724]" captionTargetId="figure@34.[151,1436,445,1724]" captionTargetPageId="34" captionText="FIGURE 19. Conchoecetes pectenicola (Adams, in Belcher, 1848), (A, B) BMNH 44.106, holotype, dried ovigerous female, 6.2 × 6.2 mm, Sunda Strait, Java; (C–E) ZRC 1969.10.4.3, ovig. female, 12.8 × 12.9 mm, Gulf of Thailand. A, dorsal view (scale in mm only applies to A, photograph by Harry Taylor); B, drawing of holotype from Wood (1867: 225); C, dorsal view; D, frontal view; E, right P4 carpus, propodus and dactylus." figureDoi="http://doi.org/10.5281/zenodo.3565037" httpUri="https://zenodo.org/record/3565037/files/figure.png" pageId="37" pageNumber="38">Figs. 19</figureCitation>
,
<figureCitation id="62941AFD3B35C5133324F8A9E05BF8DC" box="[830,859,1881,1907]" captionStart="FIGURE 20" captionStartId="35.[151,250,1932,1957]" captionTargetBox="[391,1196,166,1912]" captionTargetId="figure@35.[391,1196,166,1912]" captionTargetPageId="35" captionText="FIGURE 20. Conchoecetes pectenicola (Adams, in Belcher, 1848), NHMB-87556 (was ZMUC-CR-10111), ovig. female, 19.8 × 19.4 mm, exact origin uncertain, forwarded to Copenhagen Museum by Swedish Consul in Singapore, 4 Jun. 1903. A, dorsal view; B frontal view; C, right chela and anterolateral margin." figureDoi="http://doi.org/10.5281/zenodo.3565039" httpUri="https://zenodo.org/record/3565039/files/figure.png" pageId="37" pageNumber="38">20</figureCitation>
) which has carapace regions only weakly defined, posterolateral margins convergent, anterolateral corner without a tooth, only a blunt lobe, and no posterolateral tooth.
<taxonomicName id="3DAF7DFB3B35C51330FAF852E26CF814" authority="Naruse, 2019" authorityName="Naruse" authorityYear="2019" box="[224,364,1953,1980]" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pembawa" status="sp. nov.">
<emphasis id="C8DBDA6A3B35C51330FAF852E26CF814" box="[224,364,1953,1980]" italics="true" pageId="37" pageNumber="38">C. pembawa</emphasis>
</taxonomicName>
<emphasis id="C8DBDA6A3B35C513316EF852E2B4F813" bold="true" box="[372,436,1954,1980]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D3E867113B35C513316EF852E2B4F813" box="[372,436,1954,1980]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has well-developed anterolateral as well as posterolateral teeth, a feature shared with
<taxonomicName id="3DAF7DFB3B35C5133599F852E3E6F870" authority="Naruse, 2019" authorityName="Naruse" authorityYear="2019" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="avikele" status="sp. nov.">
<emphasis id="C8DBDA6A3B35C5133599F852E3E6F870" italics="true" pageId="37" pageNumber="38">C. avikele</emphasis>
</taxonomicName>
<emphasis id="C8DBDA6A3B35C51330F7F836E22CF84F" bold="true" box="[237,300,1990,2016]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="D3E867113B35C51330F7F836E22CF84F" box="[237,300,1990,2016]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from
<collectingCountry id="82B846E83B35C513316BF835E102F870" box="[369,514,1989,2015]" name="South Africa" pageId="37" pageNumber="38">South Africa</collectingCountry>
, but this species is densely covered with a shaggy tomentum (
<figureCitation id="62941AFD3B35C51334A3F835E608F84F" box="[1209,1288,1989,2016]" captionStart="FIGURE 7" captionStartId="15.[151,250,1936,1961]" captionTargetBox="[168,1419,166,1914]" captionTargetId="figure@15.[168,1419,166,1916]" captionTargetPageId="15" captionText="FIGURE 7. Conchoecetes avikele n. sp., ZRC2009.0838, holotype, male, 29.2 × 29.3 mm, Tugela Reef, Kwa Zulu Coast, South Africa. A, dorsal view; B, ventral view of right anterolateral area; C, frontal view of right anterolateral area; D, outer face of right chela; E, dorsal face of right chela." figureDoi="http://doi.org/10.5281/zenodo.3565013" httpUri="https://zenodo.org/record/3565013/files/figure.png" pageId="37" pageNumber="38">Figs. 7</figureCitation>
,
<figureCitation id="62941AFD3B35C513350EF835E624F870" box="[1300,1316,1989,2015]" captionStart="FIGURE 8" captionStartId="16.[151,250,1951,1976]" captionTargetBox="[189,1350,215,1808]" captionTargetId="figure@16.[151,1436,166,1913]" captionTargetPageId="16" captionText="FIGURE 8. Conchoecetes avikele n. sp., fresh male specimen which has some stalked barnacles attached on right-hand side, dimensions not available, collected off Durban Coast, South Africa. A, dorsal view; B, frontal view (after Emmerson, 2016)." figureDoi="http://doi.org/10.5281/zenodo.3565015" httpUri="https://zenodo.org/record/3565015/files/figure.png" pageId="37" pageNumber="38">8</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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