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<mods:title id="DFD1642C668E8BFB8F887D31035AAF63">Glyptothorax botius (Hamilton, 1822), a valid species of catfish (Teleostei: Sisoridae) from northeast India, with notes on the identity of G. telchitta (Hamilton, 1822).</mods:title>
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<mods:namePart id="825275CEAD74E5EEF54BC6AA6CA226B9">Heok Hee Ng</mods:namePart>
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<treatment id="08DFF8CCA8B80DCBCA5AC9C2F9B001EC" ID-DOI="http://doi.org/10.5281/zenodo.6265328" ID-GBIF-Taxon="100124112" ID-Zenodo-Dep="6265328" LSID="urn:lsid:plazi:treatment:08DFF8CCA8B80DCBCA5AC9C2F9B001EC" httpUri="http://treatment.plazi.org/id/08DFF8CCA8B80DCBCA5AC9C2F9B001EC" lastPageNumber="16" pageNumber="8">
<subSubSection id="D9DBB97259820A9252D82838AB58CAFA" type="nomenclature">
<paragraph id="A283E5AC34D12968B0FBF14E860FC958" pageNumber="8">
<taxonomicName id="18703410077446B3B4CB7C954EC2F80B" ID-CoL="3GJFD" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="8" phylum="Chordata" rank="species" species="telchitta">Glyptothorax telchitta (Hamilton, 1822)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D69FA25B218BFC242C81561B1721DD3A" type="description">
<paragraph id="0617D017208422E5EC673F27D8F3C181" pageNumber="8">(Figs. 5-7)</paragraph>
</subSubSection>
<subSubSection id="AB4895AD66147466CFF813D9DAF4B679" type="reference_group">
<paragraph id="5132B8D185363ACC8D5096AC483B7D57" pageNumber="8">
<taxonomicName id="0B51DF5A2E2E420973D362DDA31A9EB6" LSID-ZBK="urn:lsid:zoobank.org:act:F2EC45B7-1BB6-44B8-8961-D9A43F991DDE" family="Pimelodidae" genus="Pimelodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pimelodus telchitta Hamilton 1822:185, 378" order="Siluriformes" pageNumber="8" phylum="Chordata" rank="species" species="telchitta">Pimelodus telchitta Hamilton, 1822</taxonomicName>
: 185
<materialsCitation id="28757939DD1DB1B5EE54D14AD0EDBD6C" ID-GBIF-Occurrence="923889511" country="India">(type locality: Jungipur, Nathpur)</materialsCitation>
.
</paragraph>
<paragraph id="074A217A62937A996F1E86B8FC8C4483" pageNumber="8">
<taxonomicName id="275FDC5DFA5BFEDBE74086C3191EA70D" family="Sisoridae" genus="Glyptosternum" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="8" phylum="Chordata" rank="species" species="botia">Glyptosternum botia</taxonomicName>
(non Hamilton) Day, 1877: 497, Pl. CXIII Figs. 4 &amp; 4a.
</paragraph>
<paragraph id="5EA2A627C2E66C16572ECAD6EC661E39" pageNumber="8">
<taxonomicName id="C7F4845BC116C01EB280B60D13F427D9" family="Sisoridae" genus="Glyptosternum" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="8" phylum="Chordata" rank="species" species="botium">Glyptosternum botium</taxonomicName>
(non Hamilton) Day, 1889: 198.
</paragraph>
<paragraph id="B0C4C5A35F652CA0F373EAA83ACFF42B" pageNumber="8">
<taxonomicName id="108B1FFF9100910C6A2E7AC78E6F4975" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="8" phylum="Chordata" rank="species" species="botia">Glyptothorax botia</taxonomicName>
(non Hamilton) Hora, 1923: 27.
</paragraph>
<paragraph id="261AF45332E88C2DA64A070DAE42CFCC" pageNumber="8">
<taxonomicName id="33A0D8613415565A139C0C28BEAE15FC" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="8" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
sp. Hora, 1923: 26, Pl. IV Fig. 2.
</paragraph>
<paragraph id="927AC81B4D9CD550A34AE03E889FE092" pageNumber="8">
<taxonomicName id="97C89F6653DC9AB61A1C2137CE90940F" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="8" phylum="Chordata" rank="species" species="telchitta">Glyptothorax telchitta</taxonomicName>
Hora &amp; Menon, 1949: 57, Pl. II Figs. 1-3 (in part); Menon, 1954: 42 (in part); Misra, 1976: 281, Pl. XII Figs. 2-4; Jayaram &amp; Singh, 1977: 267; Jayaram, 1979: 47.
</paragraph>
</subSubSection>
<subSubSection id="02BEBD3969AFBE36DD1894F44021CB38" type="nomenclature">
<paragraph id="EDC2F1229E9AC44C5559BC67B5296A94" pageNumber="10">
Material examined.
<materialsCitation id="659E5C8E7854C6536AD5D9AD475C03FC" ID-GBIF-Occurrence="923889498" country="India" stateProvince="West Bengal">
<collectionCode id="8AF5180C7B3EC98ECCA44974D3CAB555" collectionName="USA, Michigan, Ann Arbor, University of Michigan, Museum of Zoology">UMMZ</collectionCode>
244946,
<typeStatus id="EF9D8A7411DD43DC803F7A5C72B4C930">neotype</typeStatus>
(herein designated), 69.2 mm SL
</materialsCitation>
;
<materialsCitation id="D21DEB4A7040C77BE419C5BB2F19E146" ID-GBIF-Occurrence="923889496" country="India" stateProvince="West Bengal">
<collectionCode id="3ED19319B500422A74594328BD003CDC" collectionName="USA, Michigan, Ann Arbor, University of Michigan, Museum of Zoology">UMMZ</collectionCode>
244944 (13), 36.5-63.8 mm SL
</materialsCitation>
;
<materialsCitation id="067DA859C09D09C31F22D76197AE0588" ID-GBIF-Occurrence="923889499" country="India" stateProvince="West Bengal">
<collectionCode id="B4760E96E2C28B6068F209F54E398AEE" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
49130 (2), 35.4-44.3 mm SL;
<collectingCountry id="B580B0DA9E6B4600B22DDAB65F82368E">India</collectingCountry>
:
<collectingRegion id="2046806505EFF9B02EC70FE746904089">West Bengal</collectingRegion>
, Hooghly River at Kalna,
<geoCoordinate id="D075C0996FF334396114B63598536E76" direction="north" orientation="latitude" value="23.225">23°13'30.0&quot;N</geoCoordinate>
<geoCoordinate id="C58379388637A44F28BFCB7253CA6533" direction="east" orientation="longitude" value="88.3775">88°22'39.0&quot;E</geoCoordinate>
</materialsCitation>
.
<materialsCitation id="F61027DFEF56DC4CB6DF4D11C58856BD" ID-GBIF-Occurrence="923889512" country="India" stateProvince="West Bengal">
<collectionCode id="08F509C514556D63F832236D9FA6150C" collectionName="USA, Michigan, Ann Arbor, University of Michigan, Museum of Zoology">UMMZ</collectionCode>
244951 (2), 58.7-79.0 mm SL;
<collectingCountry id="A4EC96C5C32F84634683A2CB2711D2F2">India</collectingCountry>
:
<collectingRegion id="47EA069EAF2D70F772279B2697401BFE">West Bengal</collectingRegion>
, Hooghly River at Ranaghat
</materialsCitation>
.
<materialsCitation id="D70DF9881035C06A74AB2FC11D4BCC65" ID-GBIF-Occurrence="923889501" country="India" stateProvince="West Bengal">
<collectionCode id="583D9CFEE1F0D4F05D9AEF730B6E11EE" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
49128 (7), 43.5-56.5 mm SL;
<collectingCountry id="8664E5D783DC22784A74E6C325E2BEDD">India</collectingCountry>
:
<collectingRegion id="3154B89CBB3E79FA7ABBA4C22A2877EE">West Bengal</collectingRegion>
, Simurali
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7B4D142FC10802AA44379A8A994FC200" type="diagnosis">
<paragraph id="B789EE3A6D833349A0A57040B12D8D79" pageNumber="10">
Diagnosis.
<taxonomicName id="67B9489DA3F62719D6DD6E1BA1ECE5F5" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="10" phylum="Chordata" rank="species" species="botius">Glyptothorax botius</taxonomicName>
and
<taxonomicName id="C0644F466BEEC95A834C9CFCE46B7375" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="10" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
can be distinguished from congeners in northeast India in having a combination of large, prominent tubercles on the head and body, a thoracic adhesive apparatus without a median depression, and a very slender body and caudal peduncle (sometimes described as “spindle shaped”).
<taxonomicName id="3B44A12C50E3E49201C17EA80CE9712C" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="10" phylum="Chordata" rank="species" species="botius">Glyptothorax botius</taxonomicName>
differs from
<taxonomicName id="05FAB133BF855A2F69E5272C903654CE" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="10" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
in having a more triangular snout when viewed laterally (Fig. 2), the absence (vs. presence) of dark saddles on the body, a thoracic adhesive apparatus with narrower folds of skin (Fig. 3), a shorter adipose-fin base (9.5-11.5% SL vs. 12.0-16.4) and a deeper caudal peduncle (4.7-5.9% SL vs. 3.1-4.2).
</paragraph>
</subSubSection>
<subSubSection id="C27E58688D07BDC7D1EFD0386F5C2A07" type="description">
<paragraph id="9264BAA6317884FEF0A6275572DEF314" pageNumber="10">Description. Morphometric data in Table 2. Head depressed, body moderately compressed. Dorsal profile rising evenly from tip of snout to origin of dorsal fin, then sloping gently ventrally from there to end of caudal peduncle. Ventral profile flat to anal-fin base, then sloping gently dorsally from there to end of caudal peduncle. Caudal peduncle long and moderately slender. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Skin prominently tuberculate, with ovoid tubercles particularly visible on sides of body. Lateral line complete and midlateral, delineated by series of closely spaced, conical tubercles. Vertebrae 13+21 (1), 14+20 (2), 15+19 (1), 13+22 (1), 14+21 (13), 14+22 (6) or 15+20 (1)</paragraph>
<paragraph id="A5CFDEA57E163D12779CD7FCDE6B9F79" pageNumber="10">Head depressed and narrow, with triangular snout margin when viewed laterally (Fig. 2b). Snout prominent. Anterior and posterior nares large and separated only by base of nasal barbel. Gill openings broad, extending from immediately ventral to posttemporal to isthmus. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eye ovoid, horizontal axis longest; located entirely in dorsal half of head. Orbit with free margin.</paragraph>
<paragraph id="C73E9A5DD8AE7705BEB6129AC61F69E8" pageNumber="10">Barbels in four pairs. Maxillary barbel long and slender, extending almost to base of pectoral spine. Nasal barbel slender, extending to one-third of distance between its base and anterior orbital margin. Inner mandibular-barbel origin close to midline, extending to midway between its base and that of pectoral spine. Outer mandibular barbel originates posterolateral of inner mandibular barbel, extending to two-thirds of distance between its base and that of pectoral spine.</paragraph>
<paragraph id="9D0BB4E244F85B7599F4116F6E43AE42" pageNumber="10">Mouth inferior, premaxillary tooth band almost wholly exposed when mouth is closed. Oral teeth small and villiform, in irregular rows on all tooth-bearing surfaces. Premaxillary teeth in single broad semilunate band. Dentary teeth in two narrow crescentic bands separated at midline.</paragraph>
<paragraph id="99BB5E90C4FA170B727CDC585E9388BF" lastPageNumber="12" pageNumber="11">Dorsal fin located above anterior two fifths of body, with I,5,i (21) or I,6 (4) rays; posterior fin margin straight or slightly concave; spine moderately long, slender and gently curved, without serrations on anterior or posterior edges. Adipose fin with anterior margin slightly concave and posterior margin angular. Caudal fin strongly forked, with i,7,7,i (1), i,7,8,i (17) or i,8,7,i (7) principal rays, and subequal upper and lower lobes; lower lobe slightly longer and broader than upper lobe. Procurrent rays symmetrical and extending only slightly anterior to fin base. Anal-fin base slightly anterior to adipose-fin origin. Anal fin with straight anterior margin and slightly concave posterior margin; with iv,8 (1), iv,9 (11), iv,9,i (3), iv,10 (9) or iv,11,i (1) rays. Pelvic-fin origin immediately posterior to vertical through posterior end of dorsal-fin base. Pelvic fin with slightly convex margin and i,5 (25) rays; tip of adpressed fin not reaching anal-fin origin. Pectoral fin with I,7,i (5), I,8 (6) or I,8,i (14) rays; posterior fin margin slightly concave; anterior spine margin smooth, posterior margin with 8-10 serrations. Thoracic adhesive apparatus present, located on belly and consisting of narrow longitudinal pleats of skin in elliptical field. Adhesive apparatus without median depression and extending from just behind gill opening to immediately posterior to base of last pectoral-fin ray.</paragraph>
<paragraph id="D40660AAD974909CFBEECC781207CF39" pageNumber="12">Coloration. In 70% ethanol: Dorsal and lateral surfaces of head, and dorsal surface of body brown to brownish gray, fading to a lighter color ventrally. A paler but distinct stripe running laterally along dorsal second quarter of body (immediately above lateral line). Body with small dark brown spots randomly and sparsely scattered throughout body in some individuals. Ventral surface of head and belly yellowish. Dorsal-fin base brown to brownish gray; dorsal-fin rays with evenly-distributed brown chromatophores along middle third of fin rays, imparting appearance of irregular band across fin; all other parts of dorsal fin hyaline. Pectoral, pelvic and anal fins hyaline. Pectoral spine with slightly darker coloration on dorsal surface throughout most of its length. Adipose fin brown to brownish gray, with hyaline distal margin. Caudal fin hyaline, with dark brown chromatophores on median fin rays of each caudal lobe, imparting appearance of a broad dark brown band along each lobe. Brown bands coalescent with dark brown spot at base of caudal peduncle. Other parts of caudal fin hyaline. Barbels brown to brownish gray dorsally, yellowish ventrally.</paragraph>
</subSubSection>
<subSubSection id="3FB2A5552B4819E11EF590C96C2235A0" type="distribution">
<paragraph id="0023F8F8A6EA0BDFF220B48D5CC4AE88" pageNumber="12">Distribution. Known from the Ganges River drainage. Only material from the Hooghly River was examined for this study (Fig. 4).</paragraph>
</subSubSection>
<subSubSection id="CDB8260292D47FE8DA02E8A75B7A58E7" type="discussion">
<paragraph id="A6EACC1369FE9647298D90D4A3628BEF" pageNumber="12">Discussion</paragraph>
<paragraph id="462B938B8358FB8EDBBD55CE587CD20B" lastPageNumber="13" pageNumber="12">
Hamilton (1822) described
<taxonomicName id="15939E78CF4C9D893F7BBD48FF96E407" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="species" species="botius">Glyptothorax botius</taxonomicName>
from “Pargong, fluviis Kamrupa borealis” (=Patgaon in Assam) and
<taxonomicName id="994657EC3441C0D3AECB08DDF1ADF559" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
from Jungipur (in West Bengal) and Nathpur (in Uttar Pradesh). Subsequent workers (e.g. Day, 1877) had considered the two species distinct, until Hora &amp; Menon (1949) synonymized them (retaining
<taxonomicName id="89A202B6531EE811AA40DCD6B3D6E287" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
as the valid name). Since then, this convention has been followed and
<taxonomicName id="1635A881BCDE3DCCF1AB6FDA5CAD12BD" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
is generally acknowledged as the name for the slender-bodied
<taxonomicName id="0A6F4A53E21971FB7319448270471D7F" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
species from northeast India (e.g. Talwar &amp; Jhingran, 1991);
<taxonomicName id="CD24737145973C21290CADD1DBA68581" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
is considered distinct enough that it is never confused with any other Indian
<taxonomicName id="73C08FADBAEFF31D40FB9F8AF8B319A0" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
. During recent ichthyological surveys of northeast India, I encountered two clearly distinct species of slender-bodied
<taxonomicName id="6254A60BBA19EB94231B8B78C3BCCD0B" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
that were found syntopically in the Ganges River drainage in West Bengal. One species had a deeper caudal peduncle, shorter adipose-fin base, a more triangular snout, and more uniform coloration without any distinct saddles; this was referable to
<taxonomicName id="315558DB11168E0434634C87AB77177C" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
as currently understood. The second species had a more slender caudal peduncle, longer adipose-fin base, a more rounded snout, and a series of dark saddles on the dorsal surfaces of the body. The identity of this species was not immediately obvious. It then became necessary to reinvestigate the validity of
<taxonomicName id="D22B4C287C8C557D6BD960D1A4852EA7" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
(as the only nominal species considered a synonym of
<taxonomicName id="2B7E5393F07427F71427F38B697BA9C6" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="12" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
).
</paragraph>
<paragraph id="A647088F99C2B952526A77CD10736044" pageNumber="13">
The original descriptions of
<taxonomicName id="64A9AA06E216E9F94B4ED54B5F1C9498" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="13" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
and
<taxonomicName id="1B8D6657FE6E5226FF6867EA864DBC36" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="13" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
are not very informative. A comparison of the descriptions reveals that no significant difference could be found in morphology and coloration; this led Hora &amp; Menon (1949) to consider the two species synonyms. Comparison with the figure prepared for the original description of
<taxonomicName id="C7FB70C7D1DF6ACE37DA533B2E2575B1" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="13" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
(published as Pl. 21 Fig. 3 in Hora, 1929; Fig. 6) indicates that
<taxonomicName id="B94283472D50052C1C35EAD9FB9F7DF8" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="13" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
is the species with a deeper caudal peduncle, shorter adipose-fin base, a more triangular snout, and more uniform coloration without any distinct saddles. Day (1877) considered
<taxonomicName id="A8DAADABB53A01F74AA130B0F8BDB5BB" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="13" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
to be distinct from
<taxonomicName id="D9149339F66A79358B4DAA1F882887F2" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="13" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
, and illustrated both species. His illustration of
<taxonomicName id="738377A972CDC3B9781B0D06CD5C428A" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="13" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
(Pl. CXIII Figs. 4 &amp; 4a; Fig 7) matches that of
<taxonomicName id="A033BC7DAF34173AB19985B6B2533235" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="13" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
, and his illustration of
<taxonomicName id="0E9CAFC28B2BA61E7A3596C0F3B9925E" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="13" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
(Pl. CXVI Figs. 2 &amp; 2a; Fig 8), matches that of the species with a more slender caudal peduncle, longer adipose-fin base, a more rounded snout, and a series of dark saddles on the dorsal surfaces of the body, even showing the broader folds of the skin on the thoracic adhesive apparatus.
</paragraph>
<paragraph id="E960CD396BC973D7D0BED3AFDCA2A241" lastPageNumber="15" pageNumber="14">
Therefore, there is some confusion regarding the identity of
<taxonomicName id="6DC6D2DD02874EF56322F7A69AC07027" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="14" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
. No illustration was prepared for the original description (Hora, 1929), nor is any mention made of the distinctive color pattern (the presence of dark saddles) in the original description (the color pattern is merely described as “...lurid brown, with a little yellowness intermixed...”). In fact, the description is sufficiently ambiguous that it could refer to any of the
<taxonomicName id="A00049CEB91DA1CFF27318D754252BC6" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="14" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
species occurring in northeast India, although the statement that it is “...long in proportion to its breadth...” (this statement was also used in the description of
<taxonomicName id="D52E64A90B630AD60130F9EE678210C5" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="14" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
but not in
<taxonomicName id="0C2D5437582FED8FE4D0BA547FEB3A65" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="14" phylum="Chordata" rank="species" species="cavia">G. cavia</taxonomicName>
, the only other
<taxonomicName id="4E207386F2436D259396665B6525F14C" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="14" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
described by Hamilton) suggests that it is similar in overall morphology to
<taxonomicName id="8FC8EC6B7A3781B0C7A2284BB1714475" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="14" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
. This confusion is heightened by the misidentification of
<taxonomicName id="CC97B55ECA64190F504A1B5BD5DCFE44" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="14" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
as
<taxonomicName id="548C09C1FDDD7477AABA0181161846C0" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="14" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
and vice versa by Day (1877), as explained above. The limited evidence available suggests that
<taxonomicName id="85D2BD8C5B6433933AD05D3164D383EC" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="14" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
is the oldest (and only) available name for the
<taxonomicName id="6B200D542D67BE5ABD43AFFB4995494A" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="14" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
with a more slender caudal peduncle, longer adipose-fin base, a more rounded snout, and a series of dark saddles on the dorsal surfaces of the body, and the name is herein defined as such.
</paragraph>
<paragraph id="F7E4CBBFCFFEFE7DCC0CDF0F3923677D" pageNumber="15">
Given the confusion surrounding the identity of
<taxonomicName id="DB280D5D7DF0997F8E11FEA309636D11" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
and the fact that type material does not exist for any of the Hamilton species as he did not retain any specimens used for his descriptions (Gudger, 1924; Hora, 1929), it is necessary to designate
<typeStatus id="9A4D192857F80C4BD1306BA4F1AF8645">neotypes</typeStatus>
for
<taxonomicName id="2247B3D9A5B71C6EEB16053E69C62969" LSID-ZBK="urn:lsid:zoobank.org:act:39B07C09-5C32-47B8-A3A0-2C8124D11CC1" family="Pimelodidae" genus="Pimelodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pimelodus botius Hamilton 1822:192, 378" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="botius">Pimelodus botius</taxonomicName>
and
<taxonomicName id="2458799DCF92CA40DCF427E8893502CD" LSID-ZBK="urn:lsid:zoobank.org:act:F2EC45B7-1BB6-44B8-8961-D9A43F991DDE" family="Pimelodidae" genus="Pimelodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pimelodus telchitta Hamilton 1822:185, 378" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="telchitta">P. telchitta</taxonomicName>
, and in accordance with the conditions stipulated in Article 75 of the International Code of Zoological Nomenclature,
<collectionCode id="3E24BA82FD470D7D532F062CD0E55670" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection">ZRC</collectionCode>
50223, a specimen 74.6 mm SL collected from the Hooghly River at Kalna, West Bengal, India by H. H. Ng &amp; D. C. J. Yeo on
<date id="BE158808CF5E21B3772BFACA13A1ED63" day="24" month="01" value="1999-01-24" year="1999">24 January 1999</date>
is hereby designated the
<typeStatus id="D83200D706EEFC8D5356158AB801DC7A">neotype</typeStatus>
of
<taxonomicName id="6F6C5ED15D8FFDDAB70CE03D528530A0" LSID-ZBK="urn:lsid:zoobank.org:act:39B07C09-5C32-47B8-A3A0-2C8124D11CC1" family="Pimelodidae" genus="Pimelodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pimelodus botius Hamilton 1822:192, 378" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="botius">Pimelodus botius Hamilton, 1822</taxonomicName>
, and
<collectionCode id="A07B7E71AB88DC06FD6981C2C8997D86" collectionName="USA, Michigan, Ann Arbor, University of Michigan, Museum of Zoology">UMMZ</collectionCode>
244946, a specimen 69.2 mm SL collected from the Hooghly River at Kalna, West Bengal, India by H. H. Ng et al. on
<date id="6B96BFB6F5ECC7E1AC8CEC83F71B7FD4" day="20" month="04" value="2004-04-20" year="2004">20 April 2004</date>
is hereby designated the
<typeStatus id="1C3051141B35BE946E6B68D1584F0F3C">neotype</typeStatus>
of
<taxonomicName id="B08498BDAEA150AE7D0BDD62E8692AB6" LSID-ZBK="urn:lsid:zoobank.org:act:F2EC45B7-1BB6-44B8-8961-D9A43F991DDE" family="Pimelodidae" genus="Pimelodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pimelodus telchitta Hamilton 1822:185, 378" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="telchitta">Pimelodus telchitta Hamilton, 1822</taxonomicName>
. It is possible that
<taxonomicName id="CF5164976DB7F70CA1E260E349EEDED1" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
sensu Hamilton may not be conspecific with the definition of the species used here, but the only alternative would be to make
<taxonomicName id="B6B65EAD15D7030DC814C87148787D42" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
and
<taxonomicName id="830F3EF7F43F7F275C3545833844EDAB" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
objective synonyms by designating a common
<typeStatus id="C3FBAB0181EE38BC80670E9F627B1E3A">neotype</typeStatus>
and describing the species defined as
<taxonomicName id="0F3B1C0A3188916022BF03E1046B3CEA" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
here as new. However, the naming of another
<taxonomicName id="196D9576AACB1829121D400B8F719521" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
species does nothing to resolve the confused taxonomy of the Indian
<taxonomicName id="78BA867DA3D271E012010E8C894B90EB" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
; it is felt that stability in nomenclature can be best served by the course of action here.
</paragraph>
<paragraph id="1C7322399B435FEC78FFD52923F7FAAB" lastPageNumber="16" pageNumber="15">
As a result of the confused taxonomy of the Indian
<taxonomicName id="CE419A1EC9D436886165B6FB4F914908" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
, the exact number of valid
<taxonomicName id="EFFDAFE51586CA292E5F746A66E76986" LSID-ZBK="urn:lsid:zoobank.org:act:C4304186-4FD4-4672-BDF1-AC51D2C01419" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax Blyth 1860:154" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
species occurring in the Ganges River drainage cannot be ascertained. Besides
<taxonomicName id="AE5BD3D781DBCB8FDA5C800A759DAA4A" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
and
<taxonomicName id="6149B080661B8377E7B2496080CBD3FA" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
, 12 other nominal species have been described from the Ganges River drainage:
<taxonomicName id="132044C4F2951EEF0C92FE887B23C4C8" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="cavia">G. cavia (Hamilton, 1822)</taxonomicName>
,
<taxonomicName id="678DED56CDE6FD52F33145C0CD9E1D9A" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="pectinopterus">G. pectinopterus (MClelland, 1842)</taxonomicName>
,
<taxonomicName id="5B48FE97D346B0E6A741AFD89DA2D25F" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="gracilis">
G. gracilis (
<normalizedToken id="9587CFD83AD6A31A89A9E48639BCAE83" originalValue="Günther">Guenther</normalizedToken>
, 1864)
</taxonomicName>
,
<taxonomicName id="443C58FAB4F58D75E37FC99855D11E80" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="conirostre">G. conirostre (Steindachner, 1867)</taxonomicName>
,
<taxonomicName id="BC11C37E41983815714D2616FFD7CFA4" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="stolickae">G. stolickae (Steindachner, 1867)</taxonomicName>
,
<taxonomicName id="552A8FEECCA42EE77F92823C7614F6E8" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="modestum">G. modestum (Day, 1872)</taxonomicName>
,
<taxonomicName id="3DAE30ADD84C5E71718C456047E14FFB" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="lineatum">G. lineatum (Day, 1877)</taxonomicName>
,
<taxonomicName id="3B123B31BE7BC94FCB4352B769ED2566" LSID-ZBK="urn:lsid:zoobank.org:act:AD401153-2B78-47C8-AED3-80E04A92B3E5" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax brevipinnis Hora 1923:16" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="brevipinnis">G. brevipinnis Hora, 1923</taxonomicName>
,
<taxonomicName id="932490955234C74717DDA6002708F2DB" LSID-ZBK="urn:lsid:zoobank.org:act:0A7C1D18-18BA-46B2-A0AE-1E9C4518B4D2" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax brevipinnis alaknandi Tilak 1969:42" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="alaknandi">G. alaknandi Tilak, 1969</taxonomicName>
,
<taxonomicName id="95E01E9CEA017F0B452EB116166F72A0" LSID-ZBK="urn:lsid:zoobank.org:act:8354A501-8FAB-4F5A-8A61-8C00E42C2EFD" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax garhwali Tilak 1969:37" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="garhwali">G. garhwali Tilak, 1969</taxonomicName>
,
<taxonomicName id="81071A121C648B9B4D967B6080A02729" LSID-ZBK="urn:lsid:zoobank.org:act:A25FDBA9-C689-4ADD-AA52-25E311E1E7F6" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax dakpathari Tilak &amp; Husain 1976:229" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="dakpathari">G. dakpathari Tilak &amp; Husain, 1976</taxonomicName>
, and
<taxonomicName id="B677D38BB749FE494916C429DBAEB1CD" LSID-ZBK="urn:lsid:zoobank.org:act:958309ED-A0E7-4D6A-8DFD-0813602920C3" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax indicus Talwar 1991:654" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="indicus">G. indicus Talwar in Talwar &amp; Jhingran, 1991</taxonomicName>
. I tentatively consider only seven of these to be valid:
<taxonomicName id="7E4C8793BB7C2E623EBAA8DB481B87A3" LSID-ZBK="urn:lsid:zoobank.org:act:AD401153-2B78-47C8-AED3-80E04A92B3E5" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax brevipinnis Hora 1923:16" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="brevipinnis">G. brevipinnis</taxonomicName>
(with
<taxonomicName id="CDC4CB36652707824F3590203734D581" LSID-ZBK="urn:lsid:zoobank.org:act:0A7C1D18-18BA-46B2-A0AE-1E9C4518B4D2" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax brevipinnis alaknandi Tilak 1969:42" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="alaknandi">G. alaknandi</taxonomicName>
as a junior synonym),
<taxonomicName id="CBE075F78A51C78011D05AE0A3F6A420" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="cavia">G. cavia</taxonomicName>
(with
<taxonomicName id="0CD52A1DD85F5D053C901C2F87708D66" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="lineatum">G. lineatum</taxonomicName>
as a junior synonym),
<taxonomicName id="BD35DC73B9A2D7C9F0AFD91171AF93D1" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="conirostre">G. conirostre</taxonomicName>
,
<taxonomicName id="35E6FC62EBDB9B714447112F7029ED05" LSID-ZBK="urn:lsid:zoobank.org:act:8354A501-8FAB-4F5A-8A61-8C00E42C2EFD" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax garhwali Tilak 1969:37" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="garhwali">G. garhwali</taxonomicName>
,
<taxonomicName id="B1D51D7110CF460BE2541167AAB3D083" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="gracilis">G. gracilis</taxonomicName>
(with
<taxonomicName id="1E73432EC1D97B3BD9E951425FF19B81" LSID-ZBK="urn:lsid:zoobank.org:act:A25FDBA9-C689-4ADD-AA52-25E311E1E7F6" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax dakpathari Tilak &amp; Husain 1976:229" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="dakpathari">G. dakpathari</taxonomicName>
,
<taxonomicName id="3FF43074EF820504F5B55C532A55DAE4" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="modestum">G. modestum</taxonomicName>
and
<taxonomicName id="A24E5B16961FFB3234C50872B6970C61" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="stolickae">G. stolickae</taxonomicName>
as junior synonyms),
<taxonomicName id="100464F7106AE26F3D8EF9E254BC27C2" LSID-ZBK="urn:lsid:zoobank.org:act:958309ED-A0E7-4D6A-8DFD-0813602920C3" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax indicus Talwar 1991:654" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="indicus">G. indicus</taxonomicName>
, and
<taxonomicName id="87AD3B78912DF3FBE43AB6560C97414C" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="pectinopterus">G. pectinopterus</taxonomicName>
. Besides the characters mentioned in the diagnosis, both
<taxonomicName id="E9453E558CE5FF001D38C7DE1ED5D64D" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
and
<taxonomicName id="D43F0EDF1C709498F50FB0B31B8C94CC" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
can be distinguished from
<taxonomicName id="6F58E8A2A1B96F676678B0EDF749AF5B" LSID-ZBK="urn:lsid:zoobank.org:act:AD401153-2B78-47C8-AED3-80E04A92B3E5" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax brevipinnis Hora 1923:16" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="brevipinnis">G. brevipinnis</taxonomicName>
and
<taxonomicName id="1339A466377047E827BC9F1BC7A70E12" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="pectinopterus">G. pectinopterus</taxonomicName>
in having a longer thoracic adhesive apparatus (reaching immediately posterior to base of last pectoral-fin ray vs. to middle of pectoral-fin base), from
<taxonomicName id="10CFC698B4C08653C85254B9106AEDA9" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="cavia">G. cavia</taxonomicName>
in having a shorter head (22.0-25.7% SL vs. 27.6-29.9), and from
<taxonomicName id="0C703A194E08832BB7C56F94CA1ABDB7" LSID-ZBK="urn:lsid:zoobank.org:act:8354A501-8FAB-4F5A-8A61-8C00E42C2EFD" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax garhwali Tilak 1969:37" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="garhwali">G. garhwali</taxonomicName>
in having a narrower head (14.5-17.1% SL vs. 18.0-19.9). Both
<taxonomicName id="A6C2986E1B9F47720EDD690B8B284F1B" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
and
<taxonomicName id="6C6420560F93FD7F340844377C90A37E" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
further differ from both
<taxonomicName id="455FBBC223748376B40F53F50303EF03" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="conirostre">G. conirostre</taxonomicName>
and
<taxonomicName id="DD55B632C71FD069E80BD9D445989D16" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="gracilis">G. gracilis</taxonomicName>
in having a smooth (vs. serrated) posterior edge of the dorsal spine, and from
<taxonomicName id="BD7C0477D98B1D1541987EEBFAFB695E" LSID-ZBK="urn:lsid:zoobank.org:act:958309ED-A0E7-4D6A-8DFD-0813602920C3" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Glyptothorax indicus Talwar 1991:654" order="Siluriformes" pageNumber="15" phylum="Chordata" rank="species" species="indicus">G. indicus</taxonomicName>
in having a shorter head (22.0-25.7% SL vs. 27.3-29.1) and the thoracic adhesive apparatus extending (vs. not extending) to the gular region.
</paragraph>
<paragraph id="11543ED8B4A101505656E78F2620DDDA" pageNumber="16">
<taxonomicName id="4F2AAE9A150FF70B03AAE2EDAB83C718" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="botius">Glyptothorax botius</taxonomicName>
and
<taxonomicName id="3D570FECE725F40643C0F78AE7E06F97" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
occur syntopically in the Ganges River drainage. Both species were collected from the bottom of a large, fairly swift river (Hooghly River) with a mixed sand/mud bottom, along with the many other sisorids (
<taxonomicName id="0D48E93E41B90B1D8CD54DA5B176B34D" family="Sisoridae" genus="Bagarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="yarrelli">Bagarius yarrelli</taxonomicName>
,
<taxonomicName id="854898EF18034643D3C238806CCE3C4B" family="Sisoridae" genus="Gagata" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="cenia">Gagata cenia</taxonomicName>
,
<taxonomicName id="72107A676B892A198ACFDF881E03798D" LSID-ZBK="urn:lsid:zoobank.org:act:C237C044-129B-45D0-9009-9F1542FEA4A1" family="Sisoridae" genus="Gagata" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Gagata sexualis Tilak 1970:207" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="sexualis">G. sexualis</taxonomicName>
,
<taxonomicName id="D918C39B070B942D35C7C8699F9120E8" LSID-ZBK="urn:lsid:zoobank.org:act:E4EAE479-7B8D-428E-9AAE-DEDC64579A30" family="Sisoridae" genus="Nangra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Nangra assamensis Sen &amp; Biswas 1994:441" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="assamensis">Nangra assamensis</taxonomicName>
,
<taxonomicName id="A4AFFBE50B71E4DC6C4F42FC67A44D1E" LSID-ZBK="urn:lsid:zoobank.org:act:F3B32855-E7E7-4FA0-A5D8-3DCC93C966FC" family="Sisoridae" genus="Nangra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pimelodus nangra Hamilton 1822:193, 378" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="nangra">N. nangra</taxonomicName>
,
<taxonomicName id="C2D42D4ED7BEA5D66384F82084F28917" family="Sisoridae" genus="Gogangra" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="viridescens">Gogangra viridescens</taxonomicName>
and
<taxonomicName id="EFD53137150A1E4E57D91C76BC0F593B" LSID-ZBK="urn:lsid:zoobank.org:act:AEDF2A42-D8B1-42F6-8545-3C6315C8D747" family="Sisoridae" genus="Sisor" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Sisor rabdophorus Hamilton 1822:208, 379" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="rabdophorus">Sisor rabdophorus</taxonomicName>
).
</paragraph>
<paragraph id="A5B3B964EF7C85C0704207FEEE4E13F7" pageNumber="16">
The differences in biometrics between
<taxonomicName id="E945F670C547C66AF393339D33241A59" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="botius">G. botius</taxonomicName>
and
<taxonomicName id="EE29FC9DD4B4005DE936DD824D09C1E1" family="Sisoridae" genus="Glyptothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="16" phylum="Chordata" rank="species" species="telchitta">G. telchitta</taxonomicName>
are not solely due to ontogeny. The biplots of caudal peduncle depth (Fig. 9) and length of the adipose-fin base (Fig. 10) against SL show that the regression lines are significantly different (ANCOVA; P&lt;0.00000005).
</paragraph>
</subSubSection>
</treatment>
</document>