treatments-xml/data/A7/66/C1/A766C1831E285503AD45DF7712F3DC64.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.981.53583" ID-GBIF-Dataset="8e14a5b5-aa20-4050-b2b4-f393764fc6c7" ID-PMC="PMC7644702" ID-Pensoft-Pub="1313-2970-981-1" ID-Pensoft-UUID="7C44C797C4125A71BAE032A55E6FA5DC" ID-PubMed="33199958" ID-ZooBank="5194AAC86B8A473F8A41470A60182A0B" ModsDocID="1313-2970-981-1" checkinTime="1604011718171" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Sajan, Sheikh, Tripathy, Basudev, Meng, Kaibaryer, Asami, Takahiro &amp; Ablett, Jonathan D." docDate="2020" docId="A766C1831E285503AD45DF7712F3DC64" docLanguage="en" docName="ZooKeys 981: 1-220" docOrigin="ZooKeys 981" docSource="http://dx.doi.org/10.3897/zookeys.981.53583" docTitle="Dicharax birugosus" docType="treatment" docVersion="6" id="7C44C797C4125A71BAE032A55E6FA5DC" lastPageNumber="1" masterDocId="7C44C797C4125A71BAE032A55E6FA5DC" masterDocTitle="Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue" masterLastPageNumber="220" masterPageNumber="1" pageNumber="1" updateTime="1668169683351" updateUser="ExternalLinkService">
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<mods:title>Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
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<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sajan, Sheikh</mods:namePart>
<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India &amp; Wildlife Institute of India, Chandrabani, Dehradun 248 002, Uttarakhand, India</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tripathy, Basudev</mods:namePart>
<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Meng, Kaibaryer</mods:namePart>
<mods:affiliation>National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Asami, Takahiro</mods:namePart>
<mods:affiliation>Department of Biology, Shinshu University, Matsumoto 390 - 8621, Japan</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5706-0272</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ablett, Jonathan D.</mods:namePart>
<mods:affiliation>Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, United Kingdom</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>981</mods:number>
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<mods:end>220</mods:end>
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<treatment ID-GBIF-Taxon="168715026" LSID="urn:lsid:plazi:treatment:A766C1831E285503AD45DF7712F3DC64" httpUri="http://treatment.plazi.org/id/A766C1831E285503AD45DF7712F3DC64" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="A766C183-1E28-5503-AD45-DF7712F3DC64" authority="(Godwin-Austen, 1893)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicharax birugosus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="birugosus">Dicharax (?) birugosus (Godwin-Austen, 1893)</taxonomicName>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shells of Dicharax (?) birugosus (Godwin-Austen, 1893) A Alycaeus canaliculus Godwin-Austen, 1914, (NHMUK 1903.7.1.2764) B A. birugosus Godwin-Austen, 1893, syntype (NHMUK 1903.7.1.2628). Photographs: Kevin Webb (NHM)." figureDoi="10.3897/zookeys.981.53583.figure19" httpUri="https://binary.pensoft.net/fig/468664" pageId="0" pageNumber="1">Fig. 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus bi-rugosus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bi-rugosus">Alycaeus bi-rugosus</taxonomicName>
Godwin-Austen, 1893: 593.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus bi-rugosus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bi-rugosus">Alycaeus bi-rugosus</taxonomicName>
- Godwin-Austen 1897: 387, pl. 63, figs 5, 5a.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Godwin-Austen" authorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Dicharax) birugosus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="birugosus" subGenus="Dicharax">Alycaeus (Dicharax) birugosus</taxonomicName>
- 
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/1000211" refId="B97" refString="Kobelt, W, 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlaender und Sohn, Berlin, https://biodiversitylibrary.org/page/1000211" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://biodiversitylibrary.org/page/1000211" year="1902">Kobelt 1902</bibRefCitation>
: 366.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Godwin-Austen" authorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus birugosus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="birugosus">Alycaeus birugosus</taxonomicName>
- Godwin-Austen 1914: 370; Godwin-Austen 1914: 387.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus canaliculus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="canaliculus">Alycaeus canaliculus</taxonomicName>
Godwin-Austen, 1914: 371, pl. 154, fig. 11. syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Godwin-Austen" authorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus birugosus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="birugosus">Alycaeus birugosus</taxonomicName>
var. - Godwin-Austen 1914: 370, pl. 154, figs 7, 7a.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus birugosus subsp. var. var. minor" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="birugosus" subSpecies="var." variety="minor">Alycaeus birugosus var. minor</taxonomicName>
Godwin-Austen, 1914: 370, pl. 155, figs 9, 9a.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) canaliculus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="canaliculus" subGenus="Chamalycaeus">Alycaeus (Chamalycaeus) canaliculus</taxonomicName>
- 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12890" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B74" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.12890" title="The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae)." url="https://doi.org/10.5962/bhl.title.12890" year="1921">Gude 1921</bibRefCitation>
: 225.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Godwin-Austen" authorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Dicharax) birugosus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="birugosus" subGenus="Dicharax">Alycaeus (Dicharax) birugosus</taxonomicName>
- 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12890" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B74" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.12890" title="The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae)." url="https://doi.org/10.5962/bhl.title.12890" year="1921">Gude 1921</bibRefCitation>
: 239-240.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Dicharax) birugosus subsp. var. var. minor" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="birugosus" subGenus="Dicharax" subSpecies="var." variety="minor">Alycaeus (Dicharax) birugosus var. minor</taxonomicName>
- 
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12890" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B74" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.12890" title="The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae)." url="https://doi.org/10.5962/bhl.title.12890" year="1921">Gude 1921</bibRefCitation>
: 240.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus (Chamalycaeus) canaliculus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="canaliculus" subGenus="Chamalycaeus">Chamalycaeus (Chamalycaeus) canaliculus</taxonomicName>
- Ramakrishna et al. 2010: 53.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus (Dicharax) birugosus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="birugosus" subGenus="Dicharax">Chamalycaeus (Dicharax) birugosus</taxonomicName>
- Ramakrishna et al. 2010: 57.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;Khasi Hills and Manipur&quot; (
<taxonomicName lsidName="A. birugosus" pageId="0" pageNumber="1" rank="species" species="birugosus">
<emphasis italics="true" pageId="0" pageNumber="1">A. birugosus</emphasis>
</taxonomicName>
); &quot;Garo Hills&quot; (
<taxonomicName lsidName="A. birugosus subsp. var. var. minor" pageId="0" pageNumber="1" rank="variety" species="birugosus" subSpecies="var." variety="minor">A. birugosus var. minor</taxonomicName>
); &quot;Teria Ghat, foot of the Khasi Hills&quot; (
<taxonomicName lsidName="A. canaliculus" pageId="0" pageNumber="1" rank="species" species="canaliculus">
<emphasis italics="true" pageId="0" pageNumber="1">A. canaliculus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
Khasi Hills, leg. Godwin-Austen, NHMUK 1903.7.1.2628 (2 syntypes of 
<taxonomicName lsidName="A. birugosus" pageId="0" pageNumber="1" rank="species" species="birugosus">
<emphasis italics="true" pageId="0" pageNumber="1">A. birugosus</emphasis>
</taxonomicName>
, Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shells of Dicharax (?) birugosus (Godwin-Austen, 1893) A Alycaeus canaliculus Godwin-Austen, 1914, (NHMUK 1903.7.1.2764) B A. birugosus Godwin-Austen, 1893, syntype (NHMUK 1903.7.1.2628). Photographs: Kevin Webb (NHM)." figureDoi="10.3897/zookeys.981.53583.figure19" httpUri="https://binary.pensoft.net/fig/468664" pageId="0" pageNumber="1">19B</figureCitation>
); Jawai, Jiantia Hills, 282a, leg. Godwin-Austen, NHMUK 1903.7.1.2571 (7 shells, labelled as &quot; 
<emphasis italics="true" pageId="0" pageNumber="1">birugosus</emphasis>
var.&quot;); Garo Hills, NHMUK 1903.7.1.2755 (1 syntype of 
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus birugosus subsp. var. var. minor" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="birugosus" subSpecies="var." variety="minor">Alycaeus birugosus var. minor</taxonomicName>
, labelled as 
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="duorugosus subsp. var. var. minor" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="duorugosus" subSpecies="var." variety="minor">duorugosus var. minor</taxonomicName>
); Teria Ghat, Khasi, leg. Godwin-Austen, NHMUK 1903.7.1.2764 (1 syntype of 
<taxonomicName lsidName="A. canaliculus" pageId="0" pageNumber="1" rank="species" species="canaliculus">
<emphasis italics="true" pageId="0" pageNumber="1">A. canaliculus</emphasis>
</taxonomicName>
, Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Shells of Dicharax (?) birugosus (Godwin-Austen, 1893) A Alycaeus canaliculus Godwin-Austen, 1914, (NHMUK 1903.7.1.2764) B A. birugosus Godwin-Austen, 1893, syntype (NHMUK 1903.7.1.2628). Photographs: Kevin Webb (NHM)." figureDoi="10.3897/zookeys.981.53583.figure19" httpUri="https://binary.pensoft.net/fig/468664" pageId="0" pageNumber="1">19A</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.981.53583.figure19" httpUri="https://binary.pensoft.net/fig/468664" pageId="0" pageNumber="1" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 19.</emphasis>
Shells of 
<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1893" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicharax birugosus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="birugosus">Dicharax (?) birugosus</taxonomicName>
(Godwin-Austen, 1893) 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus canaliculus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="canaliculus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus canaliculus</emphasis>
</taxonomicName>
Godwin-Austen, 1914, (NHMUK 1903.7.1.2764) 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName lsidName="A. birugosus" pageId="0" pageNumber="1" rank="species" species="birugosus">
<emphasis italics="true" pageId="0" pageNumber="1">A. birugosus</emphasis>
</taxonomicName>
Godwin-Austen, 1893, syntype (NHMUK 1903.7.1.2628). Photographs: Kevin Webb (NHM).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Godwin-Austen" authorityYear="1893" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus birugosus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="birugosus">
<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus birugosus</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="A. canaliculus" pageId="0" pageNumber="1" rank="species" species="canaliculus">
<emphasis italics="true" pageId="0" pageNumber="1">A. canaliculus</emphasis>
</taxonomicName>
are practically identical and both of them inhabit the Khasi Hills. Thus, the latter is moved to the synonymy of the former.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Protoconch low, rather glossy, without spiral lines; R1 without spiral lines; R2 short, with blunt, straight ribs. Specimens labelled as &quot; 
<emphasis italics="true" pageId="0" pageNumber="1">birugosus</emphasis>
var.&quot; are smooth on R1, whereas typical shells are more strongly sculptured.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Comments relating to &quot;var. 
<emphasis italics="true" pageId="0" pageNumber="1">minor</emphasis>
&quot;: protoconch low, rather glossy, without spiral lines; R1 glossy, without spiral lines; R2 short, with regular, blunt, not bent ribs.
</paragraph>
</subSubSection>
</treatment>
</document>