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<document id="72FD249CD5B792C4C9058E85DBF14451" ID-CLB-Dataset="31619" ID-DOI="10.11646/zootaxa.4344.1.4" ID-GBIF-Dataset="3c782ba4-ed7e-4064-a613-2c95aae16bbe" ID-ISSN="1175-5326" ID-Zenodo-Dep="1042317" ID-ZooBank="E87548A3-3CDF-43DA-A5F2-5BE87F8F1910" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509955301820" checkinUser="plazi" docAuthor="Tahami, Mohadeseh Sadat, Muilwijk, Jan, Lohaj, Roman &amp; Sadeghi, Saber" docDate="2017" docId="A71687A9E451FFAD84801E0128CFFAA1" docLanguage="en" docName="zootaxa.4344.1.4.pdf" docOrigin="Zootaxa 4344 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laemostenus Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi 2017" docType="key" docVersion="7" lastPageNumber="132" masterDocId="5B2FFFD1E441FFBC841719392D53FFC1" masterDocTitle="Study of Laemostenus species across Zagros and Central zone of Iran, with the description of seven new cavernicolous species and notes on subgenus Iranosphodrus (Coleoptera: Carabidae: Sphodrini)" masterLastPageNumber="136" masterPageNumber="115" pageNumber="131" updateTime="1698480792691" updateUser="ExternalLinkService">
<mods:mods id="3A07A6FA7E159F05A2E1292EEDC15E7D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D99D715FFC5E9AF51068C0AFD12DCED1">Study of Laemostenus species across Zagros and Central zone of Iran, with the description of seven new cavernicolous species and notes on subgenus Iranosphodrus (Coleoptera: Carabidae: Sphodrini)</mods:title>
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<mods:namePart id="E501E29A911ACFEB7E771C0407B55268">Tahami, Mohadeseh Sadat</mods:namePart>
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<mods:namePart id="EDF6C2D1F5BD7527EABCE09C803F2443">Muilwijk, Jan</mods:namePart>
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<mods:namePart id="7D06497408026BDB3786EDAEDEE225A2">Lohaj, Roman</mods:namePart>
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<mods:namePart id="74AC9CD189C3BDAEAAA7F05F25AE8ACC">Sadeghi, Saber</mods:namePart>
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<mods:title id="5DBA63CF492EA60D5DF3171EEDFFEF03">Zootaxa</mods:title>
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<mods:date id="0C75F611E55EDAEEC7857E08B3A6A6C7">2017</mods:date>
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<mods:number id="8CB9BCA549F6086F18C579FEB2CFA565">2017-11-06</mods:number>
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<treatment id="A71687A9E451FFAD84801E0128CFFAA1" ID-DOI="http://doi.org/10.5281/zenodo.6020146" ID-GBIF-Taxon="137239338" ID-Zenodo-Dep="6020146" LSID="urn:lsid:plazi:treatment:A71687A9E451FFAD84801E0128CFFAA1" httpUri="http://treatment.plazi.org/id/A71687A9E451FFAD84801E0128CFFAA1" lastPageId="17" lastPageNumber="132" pageId="16" pageNumber="131">
<subSubSection id="67A56534E451FFAC84801E012C97F8B4" pageId="16" pageNumber="131" type="nomenclature">
<paragraph id="2F0036BFE451FFAC84801E012C97F8B4" blockId="16.[151,1429,1848,1909]" pageId="16" pageNumber="131">
<heading id="744881D3E451FFAC84801E012C97F8B4" bold="true" fontSize="11" level="1" pageId="16" pageNumber="131" reason="1">
<emphasis id="1DCBEAADE451FFAC84801E012C97F8B4" bold="true" pageId="16" pageNumber="131">
Key to the Iranian
<taxonomicName id="E8BF4D3CE451FFAC85951E012F72F893" ID-CoL="8KWKS" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[386,545,1848,1874]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="1DCBEAADE451FFAC85951E012F72F893" bold="true" box="[386,545,1848,1874]" italics="true" pageId="16" pageNumber="131">Laemostenus</emphasis>
</taxonomicName>
species from Zagros and Central zone (Adapted from
<bibRefCitation id="4B2E4B4EE451FFAC80D31E012833F893" author="Casale" box="[1220,1376,1848,1874]" pageId="16" pageNumber="131" refString="Casale, A. (1988) Revisione degli Sphodrina (Coleoptera, Carabidae, Sphodrini). Museo regionale di Scienze naturali, Torino, Monografie 5, 1 - 1024." type="journal article" year="1988">Casale, 1988</bibRefCitation>
and
<bibRefCitation id="4B2E4B4EE451FFAC84801E622CEFF8B4" author="Lohaj" box="[151,444,1883,1909]" pageId="16" pageNumber="131" refString="Lohaj, R. &amp; Mlejnek, R. (2007) Two new species of Laemostenus (Antisphodrus) (Coleoptera: Carabidae) from Turkey and Syria. Acta Societatis Zoologicae Bohemicae, 71, 7 - 14." type="journal article" year="2007">Lohaj &amp; Mlejnek, 2007</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67A56534E451FFAD84801E9828CFFAA1" lastPageId="17" lastPageNumber="132" pageId="16" pageNumber="131" type="key">
<paragraph id="2F0036BFE451FFAC84801E9828CFF812" blockId="16.[151,1436,1953,2033]" pageId="16" pageNumber="131">
<keyStep id="944B2E1AE451FFAC84801E9828CFF812" pageId="16" pageNumber="131">
<keyLead id="944E958AE451FFAC84801E9828CFF812" pageId="16" pageNumber="131">
1 Elytral base strongly concave, humeri strongly pointed forwards. Pronotum cordiform, strongly narrowed to base, which is as wide as half of elytral width.........................................
<taxonomicName id="E8BF4D3CE451FFAC87F11E8628CFF812" ID-CoL="6NTSH" authority="(Chaudoir, 1854)" authorityName="Chaudoir" authorityYear="1854" baseAuthorityName="Chaudoir" baseAuthorityYear="1854" box="[998,1436,1982,2004]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="131" phylum="Arthropoda" rank="species" species="cordicollis" subGenus="Sphodroides">
<emphasis id="1DCBEAADE451FFAC87F11E862EA9F815" box="[998,1018,1983,2004]" italics="true" pageId="16" pageNumber="131">L.</emphasis>
(
<emphasis id="1DCBEAADE451FFAC80101E87292EF815" box="[1031,1149,1982,2004]" italics="true" pageId="16" pageNumber="131">Sphodroides</emphasis>
)
<emphasis id="1DCBEAADE451FFAC809B1E8729A1F815" box="[1164,1266,1982,2004]" italics="true" pageId="16" pageNumber="131">cordicollis</emphasis>
(Chaudoir, 1854)
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="2F0036BFE451FFAC84801EE528C8F830" blockId="16.[151,1436,1953,2033]" box="[151,1435,2012,2033]" pageId="16" pageNumber="131">
<keyLead id="944E958AE451FFAC84801EE528C8F830" box="[151,1435,2012,2033]" pageId="16" pageNumber="131">- Elytral base straight or slightly concave. Pronotum elongate-cordiform, rectangular or subquadrate; pronotal base distinctly</keyLead>
</paragraph>
<paragraph id="2F0036BFE450FFAD84F619AE28CFFF26" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
wider than half of elytral width.......................................................................... 2
<keyStep id="944B2E1AE450FFAD8480198C28CFFF26" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD8480198C28CFFF26" pageId="17" pageNumber="132">
2 Meso- and metatibiae without apical brush of setae; upperside black. Pronotum slightly shining. Male mesotibiae straight, not curved. Claws with denticulation.........................................
<taxonomicName id="E8BF4D3CE450FFAD801E19EB28CFFF26" ID-CoL="6NTQR" authority="Casale, 1988" authorityName="Casale" authorityYear="1988" box="[1033,1436,209,231]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="luristanus" subGenus="Laemostenus">
<emphasis id="1DCBEAADE450FFAD801E19EB284AFF26" box="[1033,1305,209,231]" italics="true" pageId="17" pageNumber="132">L. (Laemostenus) luristanus</emphasis>
<bibRefCitation id="4B2E4B4EE450FFAD813719EB28CFFF26" author="Casale" box="[1312,1436,210,231]" pageId="17" pageNumber="132" refString="Casale, A. (1988) Revisione degli Sphodrina (Coleoptera, Carabidae, Sphodrini). Museo regionale di Scienze naturali, Torino, Monografie 5, 1 - 1024." type="journal article" year="1988">Casale, 1988</bibRefCitation>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="2F0036BFE450FFAD848019D628C8FEC5" blockId="17.[151,1436,151,1376]" box="[151,1435,239,260]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD848019D628C8FEC5" box="[151,1435,239,260]" pageId="17" pageNumber="132">- Not this combination of characters....................................................................... 3</keyLead>
</paragraph>
<keyStep id="944B2E1AE450FFAD8480183428C8FEBB" pageId="17" pageNumber="132">
<paragraph id="2F0036BFE450FFAD8480183428C8FE9D" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD8480183428C8FE9D" pageId="17" pageNumber="132">
3 Head elongated, with only posterior pair of supraorbital setae; pronotum with only antero-lateral pair of setae; elytra more or less parallel; upperside dark brown to black, sometimes with slight blue shine; aedeagus with median lobe short and arcuated (subgenus
<emphasis id="1DCBEAADE450FFAD855F187E2C86FE9C" box="[328,469,327,349]" italics="true" pageId="17" pageNumber="132">Iranosphodrus</emphasis>
). Central zone of
<collectingCountry id="57A8762FE450FFAD8694187E2FFDFE9D" box="[643,686,327,348]" name="Iran" pageId="17" pageNumber="132">Iran</collectingCountry>
............................................................. 4
</keyLead>
</paragraph>
<paragraph id="2F0036BFE450FFAD8480185C28C8FEBB" blockId="17.[151,1436,151,1376]" box="[151,1435,356,378]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD8480185C28C8FEBB" box="[151,1435,356,378]" pageId="17" pageNumber="132">
- Not this combination of characters (subgenus
<taxonomicName id="E8BF4D3CE450FFAD868E185D2E4BFEBB" authorityName="Schaufuss" authorityYear="1865" box="[665,792,356,378]" class="Insecta" family="Carabidae" genus="Antisphodrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="1DCBEAADE450FFAD868E185D2E4BFEBB" box="[665,792,356,378]" italics="true" pageId="17" pageNumber="132">Antisphodrus</emphasis>
</taxonomicName>
)................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="944B2E1AE450FFAD848018BB28C8FE13" pageId="17" pageNumber="132">
<paragraph id="2F0036BFE450FFAD848018BB28C8FE75" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD848018BB28C8FE75" pageId="17" pageNumber="132">
4 Profemori with tooth; antennomere 3 stout, shorter than antennomeres 1+2.
<collectingRegion id="ED7BF85DE450FFAD87AE18BB2955FE56" box="[953,1030,386,407]" country="Iran" name="Kerman" pageId="17" pageNumber="132">Kerman</collectingRegion>
.......................................................................................................
<taxonomicName id="E8BF4D3CE450FFAD8025189928C8FE75" ID-CoL="6NTPH" authority="Lohaj &amp; Casale, 2011" authorityName="Lohaj &amp; Casale" authorityYear="2011" box="[1074,1435,415,437]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="rudichae" subGenus="Iranosphodrus">
<emphasis id="1DCBEAADE450FFAD80251899296CFE74" box="[1074,1087,416,437]" italics="true" pageId="17" pageNumber="132">L</emphasis>
. (
<emphasis id="1DCBEAADE450FFAD804418992908FE74" box="[1107,1115,416,437]" italics="true" pageId="17" pageNumber="132">I</emphasis>
.)
<emphasis id="1DCBEAADE450FFAD807818A62997FE74" box="[1135,1220,415,437]" italics="true" pageId="17" pageNumber="132">rudichae</emphasis>
<bibRefCitation id="4B2E4B4EE450FFAD80DD18A628C8FE75" author="Lohaj" box="[1226,1435,415,436]" pageId="17" pageNumber="132" refString="Lohaj, R. &amp; Casale A. (2011) Laemostenus (Iranosphodrus) rudichae, new subgenus and new species of Sphodrine beetle from Iran (Coleoptera: Carabidae: Sphodrini). Acta Entomologica Slovenica, 19, 43 - 50. http: // www. zobodat. at / pdf / ActaEntSlov _ 19 _ 0043 - 0050. pdf" type="journal article" year="2011">Lohaj &amp; Casale, 2011</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2F0036BFE450FFAD8480188428C8FE13" blockId="17.[151,1436,151,1376]" box="[151,1435,445,466]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD8480188428C8FE13" box="[151,1435,445,466]" pageId="17" pageNumber="132">- Profemori without tooth; antennomere 3 slender, as long as antennomeres 1+ 2. Yazd................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="944B2E1AE450FFAD848018E328CFFD89" pageId="17" pageNumber="132">
<paragraph id="2F0036BFE450FFAD848018E328CFFDCC" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD848018E328CFFDCC" pageId="17" pageNumber="132">
5 Mesosternum without tooth; metepisternum as long as wide (
<figureCitation id="B7842A3AE450FFAD873C18E32E20FE2E" box="[811,883,474,495]" captionStart="FIGURES 22 – 23" captionStartId="13.[151,264,1821,1844]" captionTargetBox="[197,1297,967,1777]" captionTargetId="figure@13.[158,1428,914,1797]" captionTargetPageId="13" captionText="FIGURES 22 – 23. Habitus in dorsal view of Laemostenus yazdensis (holotype) (22), Laemostenus troglophilus (holotype) (23)." httpUri="https://zenodo.org/record/1042331/files/figure.png" pageId="17" pageNumber="132">Fig. 23</figureCitation>
). Pronotum elongate-cordiform................................................................................
<taxonomicName id="E8BF4D3CE450FFAD87B018C1281AFDCD" ID-CoL="8MMJB" authority="Muilwijk, Tahami &amp; Lohaj" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[935,1353,503,525]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="yazdensis" status="sp. nov." subGenus="Iranosphodrus">
<emphasis id="1DCBEAADE450FFAD87B018C12EE7FDCC" box="[935,948,504,525]" italics="true" pageId="17" pageNumber="132">L</emphasis>
. (
<emphasis id="1DCBEAADE450FFAD87DF18C12E83FDCC" box="[968,976,504,525]" italics="true" pageId="17" pageNumber="132">I</emphasis>
.)
<emphasis id="1DCBEAADE450FFAD87F318CE2913FDCC" box="[996,1088,503,525]" italics="true" pageId="17" pageNumber="132">yazdensis</emphasis>
Muilwijk, Tahami &amp; Lohaj
</taxonomicName>
<taxonomicNameLabel id="06F857D6E450FFAD815818CE28CFFDCC" box="[1359,1436,503,525]" pageId="17" pageNumber="132" rank="species">
<emphasis id="1DCBEAADE450FFAD815818CE28CFFDCC" bold="true" box="[1359,1436,503,525]" pageId="17" pageNumber="132">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="2F0036BFE450FFAD84801B2C28CFFD89" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801B2C28CFFD89" pageId="17" pageNumber="132">
- Mesosternum with small tooth; metepisternum longer than wide (
<figureCitation id="B7842A3AE450FFAD87431B2C2EC8FDEB" box="[852,923,533,554]" captionStart="FIGURES 26 – 29" captionStartId="15.[151,264,1898,1921]" captionTargetBox="[190,1423,1111,1837]" captionTargetId="figure@15.[151,1436,1022,1874]" captionTargetPageId="15" captionText="FIGURES 26 – 29. Aedeagus. Iin lateral view, Laemostenus yazdensis (26), troglophilus speleophilus (27). In dorsal view,. Laemostenus yazdensis (28), Laemostenus troglophilus (29)." httpUri="https://zenodo.org/record/1042335/files/figure.png" pageId="17" pageNumber="132">Fig. 28</figureCitation>
), pronotum cordiform..................................................................................
<taxonomicName id="E8BF4D3CE450FFAD879A1B0B281AFD86" ID-CoL="8M7QS" authority="Muilwijk, Tahami &amp; Lohaj" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[909,1353,561,583]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="troglophilus" status="sp. nov." subGenus="Iranosphodrus">
<emphasis id="1DCBEAADE450FFAD879A1B0B2EC9FD86" box="[909,922,562,583]" italics="true" pageId="17" pageNumber="132">L</emphasis>
. (
<emphasis id="1DCBEAADE450FFAD87B81B0B2EE4FD86" box="[943,951,562,583]" italics="true" pageId="17" pageNumber="132">I</emphasis>
.)
<emphasis id="1DCBEAADE450FFAD87DB1B082913FD86" box="[972,1088,561,583]" italics="true" pageId="17" pageNumber="132">troglophilus</emphasis>
Muilwijk, Tahami &amp; Lohaj
</taxonomicName>
<taxonomicNameLabel id="06F857D6E450FFAD81581B0B28CFFD89" box="[1359,1436,562,584]" pageId="17" pageNumber="132" rank="species">
<emphasis id="1DCBEAADE450FFAD81581B0B28CFFD89" bold="true" box="[1359,1436,562,584]" pageId="17" pageNumber="132">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="944B2E1AE450FFAD84801B7628C8FD43" pageId="17" pageNumber="132">
<paragraph id="2F0036BFE450FFAD84801B7628CFFDA5" blockId="17.[151,1436,151,1376]" box="[151,1436,591,613]" pageId="17" pageNumber="132">
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<specimenCount id="39B9FD36E450FFAD84801B762C41FDA5" box="[151,274,591,612]" pageId="17" pageNumber="132" type="male">6 Male</specimenCount>
protarsi not dilated (
<taxonomicName id="E8BF4D3CE450FFAD85CC1B762F7FFDA4" ID-CoL="6NTMF" authorityName="Muilwijk &amp; Malek Hosseini" authorityYear="2016" box="[475,556,591,613]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="1DCBEAADE450FFAD85CC1B762F7FFDA4" box="[475,556,591,613]" italics="true" pageId="17" pageNumber="132">aequalis</emphasis>
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species-group).
<collectingMunicipality id="CF64ACC5E450FFAD86DC1B762E6FFDA5" box="[715,828,591,612]" pageId="17" pageNumber="132">Kohgiluyeh</collectingMunicipality>
and
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and
<collectingRegion id="ED7BF85DE450FFAD80301B762903FDA5" box="[1063,1104,591,612]" country="Iran" name="Fars" pageId="17" pageNumber="132">Fars</collectingRegion>
..........................7
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</materialsCitation>
</paragraph>
<paragraph id="2F0036BFE450FFAD84801B5428C8FD43" blockId="17.[151,1436,151,1376]" box="[151,1435,621,642]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801B5428C8FD43" box="[151,1435,621,642]" pageId="17" pageNumber="132">- Male protarsi dilated. Zagros Mts........................................................................ 8</keyLead>
</paragraph>
</keyStep>
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<paragraph id="2F0036BFE450FFAD84801BB328CFFD7D" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
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7 Both supraorbital setae present. Body more convex, larger,
<quantity id="E8479B5AE450FFAD870D1BB32EF4FD5E" box="[794,935,650,671]" metricMagnitude="-2" metricUnit="m" metricValue="1.465" metricValueMax="1.68" metricValueMin="1.25" pageId="17" pageNumber="132" unit="mm" value="14.65" valueMax="16.8" valueMin="12.5">12.5–16.8 mm</quantity>
. Kohgiluyeh and Boyer–Ahmad............................................................................
<taxonomicName id="E8BF4D3CE450FFAD87A21B9128CFFD7D" ID-CoL="6NTMF" authority="Muilwijk &amp; Malek Hosseini, 2016" authorityName="Muilwijk &amp; Malek Hosseini" authorityYear="2016" box="[949,1436,679,701]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="aequalis" subGenus="Antisphodrus">
<emphasis id="1DCBEAADE450FFAD87A21B912E9AFD7C" box="[949,969,680,701]" italics="true" pageId="17" pageNumber="132">L.</emphasis>
(
<emphasis id="1DCBEAADE450FFAD87C01B9E2EB6FD7C" box="[983,997,679,701]" italics="true" pageId="17" pageNumber="132">A</emphasis>
.)
<emphasis id="1DCBEAADE450FFAD87ED1B9E2918FD7C" box="[1018,1099,679,701]" italics="true" pageId="17" pageNumber="132">aequalis</emphasis>
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</paragraph>
<paragraph id="2F0036BFE450FFAD84801BFC28CFFD39" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801BFC28CFFD39" pageId="17" pageNumber="132">
- Anterior supraorbital setae missing. Body less convex; smaller,
<quantity id="E8479B5AE450FFAD87531BFC2E83FD1B" box="[836,976,709,730]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" metricValueMax="1.15" metricValueMin="1.13" pageId="17" pageNumber="132" unit="mm" value="11.4" valueMax="11.5" valueMin="11.3">11.3–11.5 mm</quantity>
.
<collectingRegion id="ED7BF85DE450FFAD87CE1BFC295BFD1B" box="[985,1032,709,730]" country="Iran" name="Fars" pageId="17" pageNumber="132">Fars</collectingRegion>
..........................................................................................
<taxonomicName id="E8BF4D3CE450FFAD87921BDB281AFD36" ID-CoL="8MG9V" authority="Muilwijk Tahami &amp; Lohaj" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[901,1353,737,759]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="zakariyiensis" status="sp. nov." subGenus="Antisphodrus">
<emphasis id="1DCBEAADE450FFAD87921BDB2EC1FD36" box="[901,914,738,759]" italics="true" pageId="17" pageNumber="132">L</emphasis>
. (
<emphasis id="1DCBEAADE450FFAD87B11BD82EE7FD36" box="[934,948,737,759]" italics="true" pageId="17" pageNumber="132">A</emphasis>
.)
<emphasis id="1DCBEAADE450FFAD87DE1BD82915FD36" box="[969,1094,737,759]" italics="true" pageId="17" pageNumber="132">zakariyiensis</emphasis>
Muilwijk Tahami &amp; Lohaj
</taxonomicName>
<emphasis id="1DCBEAADE450FFAD81581BDB28CFFD39" bold="true" box="[1359,1436,738,760]" pageId="17" pageNumber="132">
<taxonomicNameLabel id="06F857D6E450FFAD81581BDB28CFFD39" box="[1359,1436,738,760]" pageId="17" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="2F0036BFE450FFAD84801BC628C8FCF3" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801BC628C8FCF3" pageId="17" pageNumber="132">
8 Mesosternum with teeth before mesocoxae. Body flattened. Metatibiae with scattered pubescence at ventral sides. Elytral striae superficial, without or with very fine punctation (
<taxonomicName id="E8BF4D3CE450FFAD86E31A252E04FCF3" ID-CoL="6NTNP" baseAuthorityName="Semenov" baseAuthorityYear="1895" box="[756,855,796,818]" class="Insecta" family="Carabidae" genus="Antisphodrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="glasunowi">
<emphasis id="1DCBEAADE450FFAD86E31A252E04FCF3" box="[756,855,796,818]" italics="true" pageId="17" pageNumber="132">glasunowi</emphasis>
</taxonomicName>
species-group (part.). Zagros Mts................... 9
</keyLead>
</paragraph>
<paragraph id="2F0036BFE450FFAD84801A0328C8FCAD" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801A0328C8FCAD" pageId="17" pageNumber="132">- Mesosternum without teeth before mesocoxae. Body more convex; metatibiae with apical brush of setae at ventral side; elytral striae impressed and punctate.......................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="944B2E1AE450FFAD84801A4C28CFFC3E" pageId="17" pageNumber="132">
<paragraph id="2F0036BFE450FFAD84801A4C28CFFC05" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801A4C28CFFC05" pageId="17" pageNumber="132">
9 Eyes larger, slightly reduced, almost as long as temples; pronotum subquadrate, slightly sinuated toward hind angles. Elytral striae without punctation. Body larger,
<quantity id="E8479B5AE450FFAD864F1AAB2F80FC66" box="[600,723,914,935]" metricMagnitude="-2" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="17" pageNumber="132" unit="mm" value="13.75" valueMax="14.5" valueMin="13.0">13–14.5 mm</quantity>
.
<collectingRegion id="ED7BF85DE450FFAD86CA1AAB2E60FC66" box="[733,819,914,935]" country="Iran" name="Lorestan" pageId="17" pageNumber="132">Lorestan</collectingRegion>
, Zardeh–Kuh Kurang Mts........................................................................................................
<taxonomicName id="E8BF4D3CE450FFAD804D1A8928CFFC05" ID-CoL="6NTN8" authority="(Morvan, 1974)" authorityName="Morvan" authorityYear="1974" baseAuthorityName="Morvan" baseAuthorityYear="1974" box="[1114,1436,943,965]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="mirzayani" subGenus="Antisphodrus">
<emphasis id="1DCBEAADE450FFAD804D1A8929ADFC04" box="[1114,1278,943,965]" italics="true" pageId="17" pageNumber="132">L. (A.) mirzayani</emphasis>
(Morvan, 1974)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2F0036BFE450FFAD84801AF428CFFC3E" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801AF428CFFC3E" pageId="17" pageNumber="132">
- Eyes smaller, strongly reduced, as long as about half of the width of temples; pronotum slightly cordiforme. Elytral striae with fine punctation. Body smaller,
<quantity id="E8479B5AE450FFAD861E1AD32F32FC3E" box="[521,609,1002,1023]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="17" pageNumber="132" unit="mm" value="11.5">11.5 mm</quantity>
. Kuhe Dena Mts..............................
<taxonomicName id="E8BF4D3CE450FFAD807D1AD328CFFC3E" ID-CoL="6NTNL" authority="(Dvorak, 1982)" authorityName="Dvorak" authorityYear="1982" baseAuthorityName="Dvorak" baseAuthorityYear="1982" box="[1130,1436,1001,1023]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Antisphodrus">
<emphasis id="1DCBEAADE450FFAD807D1AD32851FC3E" box="[1130,1282,1001,1023]" italics="true" pageId="17" pageNumber="132">L. (A.) iranicus</emphasis>
(
<bibRefCitation id="4B2E4B4EE450FFAD81181AD328C6FC3E" author="Dvorak" box="[1295,1429,1002,1023]" pageId="17" pageNumber="132" refString="Dvorak, M. (1982) Neue Antisphodrus - Arten ans Mittelasien and Iran (Coleoptera, Carabidae, Sphodrini). Acta Entomologica Bohemoslovaca, 79, 274 - 280." type="journal article" year="1982">Dvořák, 1982</bibRefCitation>
)
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="944B2E1AE450FFAD84801D3E28CFFB96" pageId="17" pageNumber="132">
<paragraph id="2F0036BFE450FFAD84801D3E28C8FBFB" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801D3E28C8FBFB" pageId="17" pageNumber="132">
10 Humeri not pronounced. Elytra ellipsoid. Median lobe of aedeagus inflated, without prominent apex (
<taxonomicName id="E8BF4D3CE450FFAD80C01D312815FBDC" ID-CoL="8MGZD" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[1239,1350,1032,1053]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="1DCBEAADE450FFAD80C01D312815FBDC" box="[1239,1350,1032,1053]" italics="true" pageId="17" pageNumber="132">zagrosensis</emphasis>
</taxonomicName>
speciesgroup). Chaharmahal and Bakhtiari and Fars............................................................... 11
</keyLead>
</paragraph>
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<keyLead id="944E958AE450FFAD84801D7B28CFFB96" box="[151,1436,1089,1111]" pageId="17" pageNumber="132">
- Humeri more pronounced. Elytra oblong-elliptical. Median lobe with prominent apex (
<taxonomicName id="E8BF4D3CE450FFAD80351D7829D6FB96" ID-CoL="6NTNP" baseAuthorityName="Semenov" baseAuthorityYear="1895" box="[1058,1157,1089,1111]" class="Insecta" family="Carabidae" genus="Antisphodrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="glasunowi">
<emphasis id="1DCBEAADE450FFAD80351D7829D6FB96" box="[1058,1157,1089,1111]" italics="true" pageId="17" pageNumber="132">glasunowi</emphasis>
</taxonomicName>
species-group (part.).. 13
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<paragraph id="2F0036BFE450FFAD84801D6628CFFB53" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801D6628CFFB53" pageId="17" pageNumber="132">
11 Only posterior supraorbital setae present. Elytra small ellipsoid. Pronotum cordiform;
<collectingRegion id="ED7BF85DE450FFAD800C1D662914FBB5" box="[1051,1095,1119,1140]" country="Iran" name="Fars" pageId="17" pageNumber="132">Fars</collectingRegion>
.......................................................................................
<taxonomicName id="E8BF4D3CE450FFAD87A71D44281AFB53" ID-CoL="8MBHH" authority="Muilwijk, Tahami &amp; Lohaj" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[944,1353,1148,1170]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="farsicus" status="sp. nov." subGenus="Antisphodrus">
<emphasis id="1DCBEAADE450FFAD87A71D442913FB53" box="[944,1088,1148,1170]" italics="true" pageId="17" pageNumber="132">L. (A.) farsicus</emphasis>
Muilwijk, Tahami &amp; Lohaj
</taxonomicName>
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<emphasis id="1DCBEAADE450FFAD81581D4528CFFB53" bold="true" box="[1359,1436,1148,1170]" pageId="17" pageNumber="132">sp. nov.</emphasis>
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</paragraph>
<paragraph id="2F0036BFE450FFAD84801DA328C8FB6E" blockId="17.[151,1436,151,1376]" box="[151,1435,1178,1199]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801DA328C8FB6E" box="[151,1435,1178,1199]" pageId="17" pageNumber="132">- Both supraorbital setae present. Chaharmahal and Bakhtiari...................................................12</keyLead>
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<paragraph id="2F0036BFE450FFAD84801D8E28CFFB2B" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801D8E28CFFB2B" pageId="17" pageNumber="132">
12 Temples prominent, eyes reduced, slightly convex, longer than half of width of temples........................................................................................
<taxonomicName id="E8BF4D3CE450FFAD879A1DEC281AFB2B" ID-CoL="8MGZD" authority="Muilwijk, Tahami &amp; Lohaj" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[909,1353,1236,1258]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="zagrosensis" status="sp. nov." subGenus="Antisphodrus">
<emphasis id="1DCBEAADE450FFAD879A1DEC2913FB2B" box="[909,1088,1236,1258]" italics="true" pageId="17" pageNumber="132">L. (A.) zagrosensis</emphasis>
Muilwijk, Tahami &amp; Lohaj
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<emphasis id="1DCBEAADE450FFAD81581DED28CFFB2B" bold="true" box="[1359,1436,1236,1258]" pageId="17" pageNumber="132">sp. nov.</emphasis>
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<paragraph id="2F0036BFE450FFAD84801DCB28CFFAE4" blockId="17.[151,1436,151,1376]" pageId="17" pageNumber="132">
<keyLead id="944E958AE450FFAD84801DCB28CFFAE4" pageId="17" pageNumber="132">
- Temples not prominent, eyes strongly reduced, flat, shorter than half of width of temples......................................................................................
<taxonomicName id="E8BF4D3CE450FFAD87931C29281AFAE5" ID-CoL="8M72Q" authority="Muilwijk, Tahami &amp; Lohaj" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[900,1353,1295,1317]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="speleophilus" status="sp. nov." subGenus="Antisphodrus">
<emphasis id="1DCBEAADE450FFAD87931C292913FAE4" box="[900,1088,1295,1317]" italics="true" pageId="17" pageNumber="132">L. (A.) speleophilus</emphasis>
Muilwijk, Tahami &amp; Lohaj
</taxonomicName>
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<emphasis id="1DCBEAADE450FFAD81581C3628CFFAE4" bold="true" box="[1359,1436,1295,1317]" pageId="17" pageNumber="132">sp. nov.</emphasis>
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13 Both supraorbital setae present.
<collectingRegion id="ED7BF85DE450FFAD86021C142F3AFA83" box="[533,617,1325,1346]" country="Iran" name="Lorestan" pageId="17" pageNumber="132">Lorestan</collectingRegion>
...............................
<taxonomicName id="E8BF4D3CE450FFAD87E31C1428CFFA83" ID-CoL="6NTLP" authority="Casale &amp; Queinnec, 2001" authorityName="Casale &amp; Queinnec" authorityYear="2001" box="[1012,1436,1324,1346]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="deconincki" subGenus="Antisphodrus">
<emphasis id="1DCBEAADE450FFAD87E31C1429CDFA83" box="[1012,1182,1324,1346]" italics="true" pageId="17" pageNumber="132">L. (A.) deconincki</emphasis>
<bibRefCitation id="4B2E4B4EE450FFAD80B21C1428CFFA83" author="Casale" box="[1189,1436,1325,1346]" pageId="17" pageNumber="132" refString="Casale, A. &amp; Queinnec E. (2001) Un nouveau sphodride cavernicole des hautes montagnes du Zagros, Iran (Coleoptera, Carabidae, Sphodrina). Revue Francaise d'Entomologie, New Series, 23, 177 - 180." type="journal article" year="2001">Casale &amp; Quéinnec, 2001</bibRefCitation>
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</paragraph>
<paragraph id="2F0036BFE450FFAD84801C7328CFFAA1" blockId="17.[151,1436,151,1376]" box="[151,1436,1353,1376]" pageId="17" pageNumber="132">
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<keyLead id="944E958AE450FFAD84801C7328CFFAA1" box="[151,1436,1353,1376]" pageId="17" pageNumber="132">
- Only posterior supraorbital setae present.
<collectingRegion id="ED7BF85DE450FFAD86661C732FC9FA9E" box="[625,666,1354,1375]" country="Iran" name="Fars" pageId="17" pageNumber="132">Fars</collectingRegion>
...................
<taxonomicName id="E8BF4D3CE450FFAD87861C73281AFA9E" ID-CoL="8MLSX" authority="Muilwijk, Tahami &amp; Lohaj" authorityName="Tahami &amp; Muilwijk &amp; Lohaj &amp; Sadeghi" authorityYear="2017" box="[913,1353,1353,1375]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="shirazensis" status="sp. nov." subGenus="Antisphodrus">
<emphasis id="1DCBEAADE450FFAD87861C732913FA9E" box="[913,1088,1353,1375]" italics="true" pageId="17" pageNumber="132">L. (A.) shirazensis</emphasis>
Muilwijk, Tahami &amp; Lohaj
</taxonomicName>
<taxonomicNameLabel id="06F857D6E450FFAD81581C7328CFFAA1" box="[1359,1436,1354,1376]" pageId="17" pageNumber="132" rank="species">
<emphasis id="1DCBEAADE450FFAD81581C7328CFFAA1" bold="true" box="[1359,1436,1354,1376]" pageId="17" pageNumber="132">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</materialsCitation>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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