219 lines
26 KiB
XML
219 lines
26 KiB
XML
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<mods:title id="B2DF7B8A9448338A4BCF4EE0C852A3D0">Study of Laemostenus species across Zagros and Central zone of Iran, with the description of seven new cavernicolous species and notes on subgenus Iranosphodrus (Coleoptera: Carabidae: Sphodrini)</mods:title>
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<mods:namePart id="6DD6BBA4F9B62E3F688A88ABE7F2CC86">Tahami, Mohadeseh Sadat</mods:namePart>
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<mods:namePart id="F0C20C5F2E607A7E2EE875D97AB1673A">Muilwijk, Jan</mods:namePart>
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<mods:namePart id="EC87F6A6D6043DE589F167CF6DB91293">Lohaj, Roman</mods:namePart>
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<mods:namePart id="458E81574F5D76FE7B317510C5421292">Sadeghi, Saber</mods:namePart>
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<mods:title id="DDE3CDC7F014EF4D2502235C65A11939">Zootaxa</mods:title>
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<mods:date id="1F7450ADEF704BE12E425B26BDCE7DFF">2017</mods:date>
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<mods:number id="1859793221B5FA571DC926CACD60AC3B">2017-11-06</mods:number>
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<mods:number id="A38AD8C94C7AAACB94CA1EF397EF2346">4344</mods:number>
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<treatment id="A71687A9E445FFBA84801DD42EE5FE6D" ID-DOI="http://doi.org/10.5281/zenodo.6020098" ID-GBIF-Taxon="137239348" ID-Zenodo-Dep="6020098" LSID="urn:lsid:plazi:treatment:A71687A9E445FFBA84801DD42EE5FE6D" httpUri="http://treatment.plazi.org/id/A71687A9E445FFBA84801DD42EE5FE6D" lastPageId="6" lastPageNumber="121" pageId="4" pageNumber="119">
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<subSubSection id="67A56534E445FFB884801DD42C41FAE9" pageId="4" pageNumber="119" type="nomenclature">
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<paragraph id="2F0036BFE445FFB884801DD4290DFAC6" blockId="4.[151,1118,1261,1320]" box="[151,1118,1261,1287]" pageId="4" pageNumber="119">
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<heading id="744881D3E445FFB884801DD4290DFAC6" bold="true" box="[151,1118,1261,1287]" fontSize="11" level="1" pageId="4" pageNumber="119" reason="1">
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<emphasis id="1DCBEAADE445FFB884801DD4290DFAC6" bold="true" box="[151,1118,1261,1287]" pageId="4" pageNumber="119">
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<taxonomicName id="E8BF4D3CE445FFB884801DD42EA7FAC6" ID-CoL="8MG9V" authority="Muilwijk, Tahami & Lohaj" authorityName="Tahami & Muilwijk & Lohaj & Sadeghi" authorityYear="2017" box="[151,1012,1261,1287]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="zakariyiensis" status="sp. nov." subGenus="Antisphodrus">
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<emphasis id="1DCBEAADE445FFB884801DD42C64FAC6" bold="true" box="[151,311,1261,1287]" italics="true" pageId="4" pageNumber="119">Laemostenus</emphasis>
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(
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<emphasis id="1DCBEAADE445FFB885501DD42CB9FAC6" bold="true" box="[327,490,1261,1287]" italics="true" pageId="4" pageNumber="119">Antisphodrus</emphasis>
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)
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<emphasis id="1DCBEAADE445FFB885EC1DD42FCBFAC6" bold="true" box="[507,664,1261,1287]" italics="true" pageId="4" pageNumber="119">zakariyiensis</emphasis>
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Muilwijk, Tahami & Lohaj
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</taxonomicName>
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,
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<taxonomicNameLabel id="06F857D6E445FFB880171DD4290DFAC6" box="[1024,1118,1261,1287]" pageId="4" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2F0036BFE445FFB884801C362C41FAE9" blockId="4.[151,1118,1261,1320]" box="[151,274,1295,1320]" pageId="4" pageNumber="119">
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(
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<figureCitation id="B7842A3AE445FFB884881C362C5AFAE9" box="[159,265,1295,1320]" captionStart="FIGURES 4 – 7" captionStartId="5.[151,264,1496,1519]" captionTargetBox="[155,1382,167,1416]" captionTargetId="figure@5.[151,1436,167,1472]" captionTargetPageId="5" captionText="FIGURES 4 – 7. Habitus in dorsal view of Laemostenus zakariyiensis (paratype) (4). Aedeagus of Laemostenus zakariyiensis in lateral view (5), dorsal view (6). Ovipositor of Laemostenus zakariyiensis (7)." httpUri="https://zenodo.org/record/1042321/files/figure.png" pageId="4" pageNumber="119">Figs. 4–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="67A56534E445FFB884801C6E2CA2FA3E" pageId="4" pageNumber="119" type="materials_examined">
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<paragraph id="2F0036BFE445FFB884801C6E2CA2FA3E" blockId="4.[151,1437,1367,2005]" pageId="4" pageNumber="119">
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<emphasis id="1DCBEAADE445FFB884801C6E2C15FAB1" bold="true" box="[151,326,1367,1392]" pageId="4" pageNumber="119">Type material.</emphasis>
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Holotype male, labelled: “Iran, Fars, Estahban, Zakariya Cave;
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<date id="5B01107FE445FFB880351C6E29F6FAB1" box="[1058,1189,1367,1392]" pageId="4" pageNumber="119" value="2015-09-10">10.IX.2015</date>
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; leg. M.S. Tahami, S. Sadeghi, Sh. Amrollahi, S. Shahabi”/
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<taxonomicName id="E8BF4D3CE445FFB8865C1C442E2BFA54" authority="Tahami & Muilwijk & Lohaj & Sadeghi, 2017" authorityName="Tahami & Muilwijk & Lohaj & Sadeghi" authorityYear="2017" box="[587,888,1404,1429]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="zakariyiensis" status="sp. nov.">
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<emphasis id="1DCBEAADE445FFB8865C1C442E2BFA54" box="[587,888,1404,1429]" italics="true" pageId="4" pageNumber="119">Laemostenus zakariyiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="06F857D6E445FFB887951C452E8DFA54" box="[898,990,1404,1429]" pageId="4" pageNumber="119" rank="species">
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<emphasis id="1DCBEAADE445FFB887951C452E8DFA54" bold="true" box="[898,990,1404,1429]" pageId="4" pageNumber="119">sp. nov.</emphasis>
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</taxonomicNameLabel>
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Muilwijk, Tahami & Lohaj, 2016 (red label Holotype). Paratypes: one male and one female, the same data as Holotype ”/
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<taxonomicName id="E8BF4D3CE445FFB880501C982820FA79" authority="Tahami & Muilwijk & Lohaj & Sadeghi, 2017" authorityName="Tahami & Muilwijk & Lohaj & Sadeghi" authorityYear="2017" box="[1095,1395,1439,1464]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="zakariyiensis" status="sp. nov.">
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<emphasis id="1DCBEAADE445FFB880501C982820FA79" box="[1095,1395,1439,1464]" italics="true" pageId="4" pageNumber="119">Laemostenus zakariyiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="06F857D6E445FFB8816D1CA62D9BFA1C" pageId="4" pageNumber="119" rank="species">
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<emphasis id="1DCBEAADE445FFB8816D1CA62D9BFA1C" bold="true" pageId="4" pageNumber="119">sp. nov.</emphasis>
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</taxonomicNameLabel>
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Muilwijk, Tahami & Lohaj, 2016 (red label Paratype). Holotype and female paratype are deposited in ZM– CBSU, male paratype in cMU.
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</paragraph>
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</subSubSection>
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<subSubSection id="67A56534E445FFB884D01F352FE0F9AD" pageId="4" pageNumber="119" type="diagnosis">
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<paragraph id="2F0036BFE445FFB884D01F352FE0F9AD" blockId="4.[151,1437,1367,2005]" pageId="4" pageNumber="119">
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<emphasis id="1DCBEAADE445FFB884D01F352C12F9E4" bold="true" box="[199,321,1548,1573]" pageId="4" pageNumber="119">Diagnosis.</emphasis>
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Small, brachypterous
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<taxonomicName id="E8BF4D3CE445FFB886291F342F83F9E5" authorityName="Tahami & Muilwijk & Lohaj & Sadeghi" authorityYear="2017" box="[574,720,1549,1572]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="1DCBEAADE445FFB886291F342F83F9E5" box="[574,720,1549,1572]" italics="true" pageId="4" pageNumber="119">Laemostenus</emphasis>
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</taxonomicName>
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sg.
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<taxonomicName id="E8BF4D3CE445FFB887171F352EC6F9E4" authorityName="Schaufuss" authorityYear="1865" box="[768,917,1548,1573]" class="Insecta" family="Carabidae" genus="Antisphodrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis id="1DCBEAADE445FFB887171F352EC6F9E4" box="[768,917,1548,1573]" italics="true" pageId="4" pageNumber="119">Antisphodrus</emphasis>
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</taxonomicName>
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species of the
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<taxonomicName id="E8BF4D3CE445FFB8802A1F3529CFF9E4" authorityName="Muilwijk & Malek Hosseini" authorityYear="2016" box="[1085,1180,1548,1573]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="aequalis">
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<emphasis id="1DCBEAADE445FFB8802A1F3529CFF9E4" box="[1085,1180,1548,1573]" italics="true" pageId="4" pageNumber="119">aequalis</emphasis>
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</taxonomicName>
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species-group. Colour black or dark-brown, antennae and legs reddish-brown. Head and pronotum shiny, with very fine microsculpture, elytra dull, with distinct microsculpture (
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<figureCitation id="B7842A3AE445FFB886761F6D2FF5F9AD" box="[609,678,1620,1645]" captionStart="FIGURES 4 – 7" captionStartId="5.[151,264,1496,1519]" captionTargetBox="[155,1382,167,1416]" captionTargetId="figure@5.[151,1436,167,1472]" captionTargetPageId="5" captionText="FIGURES 4 – 7. Habitus in dorsal view of Laemostenus zakariyiensis (paratype) (4). Aedeagus of Laemostenus zakariyiensis in lateral view (5), dorsal view (6). Ovipositor of Laemostenus zakariyiensis (7)." httpUri="https://zenodo.org/record/1042321/files/figure.png" pageId="4" pageNumber="119">Fig. 4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="67A56534E445FFBA84D01F4E2C84FF6E" lastPageId="6" lastPageNumber="121" pageId="4" pageNumber="119" type="description">
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<paragraph id="2F0036BFE445FFB884D01F4E299FF93C" blockId="4.[151,1437,1367,2005]" pageId="4" pageNumber="119">
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<emphasis id="1DCBEAADE445FFB884D01F4E2C0AF951" bold="true" box="[199,345,1655,1680]" pageId="4" pageNumber="119">Description.</emphasis>
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BL
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<quantity id="E8479B5AE445FFB8859D1F4E2F7DF94E" box="[394,558,1655,1680]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" metricValueMax="1.15" metricValueMin="1.13" pageId="4" pageNumber="119" unit="mm" value="11.4" valueMax="11.5" valueMin="11.3">11.3–11.5 mm</quantity>
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. Head narrow, almost parallel-sided, as long as wide (HL/HW: 1.0), with only posterior pair of supraorbital setae; frontal furrows shallow, eyes reduced (
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<quantity id="E8479B5AE445FFB887C11FA52962F975" box="[982,1073,1692,1717]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="119" unit="mm" value="0.5">0.5 mm</quantity>
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), shorter than temples (
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<quantity id="E8479B5AE445FFB881221FA528C3F975" box="[1333,1424,1692,1716]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="119" unit="mm" value="0.8">0.8 mm</quantity>
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). Clypeus with two long setae; labrum with six setae. Antennae slender, reaching the anterior 1/3 of elytral length, antennomere 3 without accessory setae except a few apical ones; antennomeres 4–11 pubescent.
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</paragraph>
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<paragraph id="2F0036BFE445FFB884D01E312DB7F814" blockId="4.[151,1437,1367,2005]" pageId="4" pageNumber="119">Pronotum elongate, distinctly sinuate before hind angles (more in male HT, less in female PT), longer than wide, (PL/PW: 1.1), with maximum width in apical third, slightly wider than head. Basal impressions short, sparsely punctuated. Anterior angles obtuse, slightly protruding in HT, more rounded in female PT, posterior angles acute, sharp. Lateral margins with some shallow punctures and with two pairs of setae present in lateral furrow, pair of baso-lateral setae situated near posterior angles, pair of antero-lateral setae situated in anterior third of pronotal length.</paragraph>
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<caption id="7BC06637E444FFB984801CE12EF8F9CC" httpUri="https://zenodo.org/record/1042321/files/figure.png" pageId="5" pageNumber="120" targetBox="[155,1382,167,1416]" targetPageId="5">
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<paragraph id="2F0036BFE444FFB984801CE12EF8F9CC" blockId="5.[151,1436,1496,1550]" pageId="5" pageNumber="120">
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<emphasis id="1DCBEAADE444FFB984801CE12C69FA2F" bold="true" box="[151,314,1496,1519]" pageId="5" pageNumber="120">FIGURES 4–7.</emphasis>
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Habitus in dorsal view of
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<taxonomicName id="E8BF4D3CE444FFB986531CE32E02FA2E" authorityName="Tahami & Muilwijk & Lohaj & Sadeghi" authorityYear="2017" box="[580,849,1497,1519]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="zakariyiensis">
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<emphasis id="1DCBEAADE444FFB986531CE32E02FA2E" box="[580,849,1497,1519]" italics="true" pageId="5" pageNumber="120">Laemostenus zakariyiensis</emphasis>
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</taxonomicName>
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(paratype) (4). Aedeagus of
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<taxonomicName id="E8BF4D3CE444FFB980631CE329ABFA2E" authorityName="Tahami & Muilwijk & Lohaj & Sadeghi" authorityYear="2017" box="[1140,1272,1498,1519]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="120" phylum="Arthropoda" rank="genus">
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<emphasis id="1DCBEAADE444FFB980631CE329ABFA2E" box="[1140,1272,1498,1519]" italics="true" pageId="5" pageNumber="120">Laemostenus</emphasis>
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</taxonomicName>
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<taxonomicName id="E8BF4D3CE444FFB980E91CE028D1FA2E" authorityName="Tahami & Muilwijk & Lohaj & Sadeghi" authorityYear="2017" box="[1278,1410,1497,1519]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="zakariyiensis">zakariyiensis</taxonomicName>
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in lateral view (5), dorsal view (6). Ovipositor of
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<taxonomicName id="E8BF4D3CE444FFB986651CC02ED3F9CC" authorityName="Tahami & Muilwijk & Lohaj & Sadeghi" authorityYear="2017" box="[626,896,1527,1550]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="zakariyiensis">
|
||
<emphasis id="1DCBEAADE444FFB986651CC02ED3F9CC" box="[626,896,1527,1550]" italics="true" pageId="5" pageNumber="120">Laemostenus zakariyiensis</emphasis>
|
||
</taxonomicName>
|
||
(7).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2F0036BFE444FFB984D01F032C0BF923" blockId="5.[151,1437,1594,2015]" pageId="5" pageNumber="120">Elytra elongate, nearly parallel-sided, ratio EL/EW 1.6–1.7, with maximum width in posterior third; disc flattened, with clear depression along suture. Base slightly wider than pronotal base. Humeri moderately rounded; with small tooth. Striae fine, with superficial punctures, intervals flat. Scutellar striae present; scutellar setiferous punctures situated in stria 1. Umbilicate series consists of 18 setiferous punctures, in interrupted in middle; 2 setae at apex of stria 7.</paragraph>
|
||
<paragraph id="2F0036BFE444FFB984D01FD62C14F8EA" blockId="5.[151,1437,1594,2015]" pageId="5" pageNumber="120">Mesosternum unarmed, without teeth before mesocoxae. Abdominal sternites smooth, brownish, with dense microsculpture.</paragraph>
|
||
<paragraph id="2F0036BFE444FFB984D01E0E2824F856" blockId="5.[151,1437,1594,2015]" pageId="5" pageNumber="120">Legs elongated, very slender. Protibiae with sparse, short pubescence at apex of internal anterior sides. Meso- and metatibiae not curved, metatibiae with brush of setae on ventral side of apical third, mesotibiae pubescent on ventral side of apical third. Tarsi dorsally with decumbent pubescence, claws smooth. Male protarsi not dilated.</paragraph>
|
||
<paragraph id="2F0036BFE444FFB984D01E9B28CEF81E" blockId="5.[151,1437,1594,2015]" pageId="5" pageNumber="120">
|
||
Aedeagus (
|
||
<figureCitation id="B7842A3AE444FFB985501E9B2CE4F87A" box="[327,439,1954,1979]" captionStart="FIGURES 4 – 7" captionStartId="5.[151,264,1496,1519]" captionTargetBox="[155,1382,167,1416]" captionTargetId="figure@5.[151,1436,167,1472]" captionTargetPageId="5" captionText="FIGURES 4 – 7. Habitus in dorsal view of Laemostenus zakariyiensis (paratype) (4). Aedeagus of Laemostenus zakariyiensis in lateral view (5), dorsal view (6). Ovipositor of Laemostenus zakariyiensis (7)." httpUri="https://zenodo.org/record/1042321/files/figure.png" pageId="5" pageNumber="120">Figs. 5–6</figureCitation>
|
||
): median lobe short (
|
||
<quantity id="E8479B5AE444FFB986BA1E9A2E4AF87A" box="[685,793,1955,1979]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="5" pageNumber="120" unit="mm" value="2.07">2.07 mm</quantity>
|
||
); basal bulb about one third of aedeagus length, ventral margin almost straight, dorsal margin curved in lateral view, apex elongated in lateral view, rounded in dorsal view.
|
||
</paragraph>
|
||
<paragraph id="2F0036BFE447FFBA84D019AE2C84FF6E" blockId="6.[151,1436,151,429]" box="[199,471,151,176]" pageId="6" pageNumber="121">
|
||
Ovipositor: as on
|
||
<figureCitation id="B7842A3AE447FFBA859819AE2C80FF6E" box="[399,467,151,176]" captionStart="FIGURES 4 – 7" captionStartId="5.[151,264,1496,1519]" captionTargetBox="[155,1382,167,1416]" captionTargetId="figure@5.[151,1436,167,1472]" captionTargetPageId="5" captionText="FIGURES 4 – 7. Habitus in dorsal view of Laemostenus zakariyiensis (paratype) (4). Aedeagus of Laemostenus zakariyiensis in lateral view (5), dorsal view (6). Ovipositor of Laemostenus zakariyiensis (7)." httpUri="https://zenodo.org/record/1042321/files/figure.png" pageId="6" pageNumber="121">Fig. 7</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67A56534E447FFBA84D019852EA9FE81" pageId="6" pageNumber="121" type="diagnosis">
|
||
<paragraph id="2F0036BFE447FFBA84D019852EA9FE81" blockId="6.[151,1436,151,429]" pageId="6" pageNumber="121">
|
||
<emphasis id="1DCBEAADE447FFBA84D019852C9DFF14" bold="true" box="[199,462,188,213]" pageId="6" pageNumber="121">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="E8BF4D3CE447FFBA85C119842FEAFF14" authority="Tahami & Muilwijk & Lohaj & Sadeghi, 2017" authorityName="Tahami & Muilwijk & Lohaj & Sadeghi" authorityYear="2017" box="[470,697,188,213]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="zakariyiensis" status="sp. nov." subGenus="Antisphodrus">
|
||
<emphasis id="1DCBEAADE447FFBA85C119842FEAFF14" box="[470,697,188,213]" italics="true" pageId="6" pageNumber="121">L. (A.) zakariyiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06F857D6E447FFBA86D619852E4FFF14" box="[705,796,188,213]" pageId="6" pageNumber="121" rank="species">
|
||
<emphasis id="1DCBEAADE447FFBA86D619852E4FFF14" bold="true" box="[705,796,188,213]" pageId="6" pageNumber="121">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from closely related
|
||
<taxonomicName id="E8BF4D3CE447FFBA804919842CC3FF39" authority="Muilwijk & Malek Hosseini, 2016" authorityName="Muilwijk & Malek Hosseini" authorityYear="2016" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="aequalis" subGenus="Antisphodrus">
|
||
<emphasis id="1DCBEAADE447FFBA80491984285DFF14" box="[1118,1294,188,213]" italics="true" pageId="6" pageNumber="121">L. (A.) aequalis</emphasis>
|
||
<bibRefCitation id="4B2E4B4EE447FFBA810119852CC3FF39" author="Muilwijk" pageId="6" pageNumber="121" refString="Malek Hosseini, M. J., Muilwijk, J., Sadeghi, S. & Bakhshi, Y. (2016) The Carabid fauna of caves in the southern Zagros Mountains and description of Laemostenus (Antisphodrus) aequalis nov. sp. and Duvalius kileri nov. sp. from Kohgiluyeh and Boyer-Ahmad Province, Iran (Coleoptera: Carabidae). Entomofauna, 37, 185 - 204." type="journal article" year="2016">Muilwijk & Malek Hosseini, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
by its smaller body size, less convex body, more parallel elytra, presence of only posterior pair of supraorbital setae and shape of pronotum and aedeagus. Shape of pronotum in
|
||
<taxonomicName id="E8BF4D3CE447FFBA807F183C29B6FEDC" authorityName="Muilwijk & Malek Hosseini" authorityYear="2016" box="[1128,1253,260,285]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="aequalis">
|
||
<emphasis id="1DCBEAADE447FFBA807F183C29B6FEDC" box="[1128,1253,260,285]" italics="true" pageId="6" pageNumber="121">L. aequalis</emphasis>
|
||
</taxonomicName>
|
||
is very variable and some specimens have a similar pronotum to
|
||
<taxonomicName id="E8BF4D3CE447FFBA86AE18102EC9FE81" authority="Tahami & Muilwijk & Lohaj & Sadeghi, 2017" authorityName="Tahami & Muilwijk & Lohaj & Sadeghi" authorityYear="2017" box="[697,922,295,320]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="zakariyiensis" status="sp. nov." subGenus="Antisphodrus">
|
||
<emphasis id="1DCBEAADE447FFBA86AE18102EC9FE81" box="[697,922,295,320]" italics="true" pageId="6" pageNumber="121">L. (A.) zakariyiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06F857D6E447FFBA87B6181E2EA9FE81" box="[929,1018,295,320]" pageId="6" pageNumber="121" rank="species">
|
||
<emphasis id="1DCBEAADE447FFBA87B6181E2EA9FE81" bold="true" box="[929,1018,295,320]" pageId="6" pageNumber="121">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67A56534E447FFBA84D018752C2AFE49" pageId="6" pageNumber="121" type="distribution">
|
||
<paragraph id="2F0036BFE447FFBA84D018752C2AFE49" blockId="6.[151,1436,151,429]" pageId="6" pageNumber="121">
|
||
<emphasis id="1DCBEAADE447FFBA84D018752CA7FEA4" bold="true" box="[199,500,332,357]" pageId="6" pageNumber="121">Distribution and habitat.</emphasis>
|
||
So far only known from the twilight and dark zones of the
|
||
<typeStatus id="F004881DE447FFBA80881874299CFEA4" box="[1183,1231,333,357]" pageId="6" pageNumber="121">type</typeStatus>
|
||
locality, Zakariya cave,
|
||
<collectingRegion id="ED7BF85DE447FFBA84CF18492C26FE49" box="[216,373,367,392]" country="Iran" name="Fars" pageId="6" pageNumber="121">Fars province</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67A56534E447FFBA84D018AD2EE5FE6D" box="[199,950,404,429]" pageId="6" pageNumber="121" type="etymology">
|
||
<paragraph id="2F0036BFE447FFBA84D018AD2EE5FE6D" blockId="6.[151,1436,151,429]" box="[199,950,404,429]" pageId="6" pageNumber="121">
|
||
<emphasis id="1DCBEAADE447FFBA84D018AD2C1DFE6C" bold="true" box="[199,334,404,429]" pageId="6" pageNumber="121">Etymology.</emphasis>
|
||
Topotypic, referred to the
|
||
<typeStatus id="F004881DE447FFBA866E18AC2FFAFE6C" box="[633,681,405,429]" pageId="6" pageNumber="121">type</typeStatus>
|
||
locality, Zakariya cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |