242 lines
36 KiB
XML
242 lines
36 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.901.39432" ID-GBIF-Dataset="654566e8-947b-4dfe-8fbc-baea3d254fa8" ID-PMC="PMC6965333" ID-Pensoft-Pub="1313-2970-901-1" ID-Pensoft-UUID="D8C38C1C3844549F8BEC7C1D884CDF8E" ID-PubMed="31975782" ModsDocID="1313-2970-901-1" checkinTime="1578615027095" checkinUser="pensoft" docAuthor="Balkenohl, Michael" docDate="2020" docId="2539ACAEEF2E5F4DA7DF755F030D1F55" docLanguage="en" docName="ZooKeys 901: 1-81" docOrigin="ZooKeys 901" docSource="http://dx.doi.org/10.3897/zookeys.901.39432" docTitle="Salcedia perrieri Fairmaire 1899" docType="treatment" docVersion="5" id="D8C38C1C3844549F8BEC7C1D884CDF8E" lastPageNumber="1" masterDocId="D8C38C1C3844549F8BEC7C1D884CDF8E" masterDocTitle="A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini)" masterLastPageNumber="81" masterPageNumber="1" pageNumber="1" updateTime="1732478763716" updateUser="ExternalLinkService">
|
||
<mods:mods id="8B6C412FE75596BC650637FD663335AC" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="07340BE5565354B7B91EA7C86B2B5D56">
|
||
<mods:title id="C143C51AB5C812231802CFC82D772554">A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="257E01FEA563670C27E757B35A7F6762" type="personal">
|
||
<mods:role id="E25A7BB22344F47059E7003ADE88B332">
|
||
<mods:roleTerm id="0E61904702D73FFA6B163ABA82236991">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="498929929899AE80B9099C52778FAF2E">Balkenohl, Michael</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="C2A6A0C41864A75D453B07E44DD1A6A6">text</mods:typeOfResource>
|
||
<mods:relatedItem id="018A68EB76366FC77EFD91CDF581FAAC" type="host">
|
||
<mods:titleInfo id="16A5A5C1F3753AA96FEDEF604190425F">
|
||
<mods:title id="59405781E5343EB6B31EBB659489333D">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="2D906F9FAA0661B3B36CF839390DE52D">
|
||
<mods:date id="644CF59438422D2C099CFF3F3A90A6A0">2020</mods:date>
|
||
<mods:detail id="A5875E2227E54E1477547CC57195FFC0" type="volume">
|
||
<mods:number id="00E25AC49D6ED604E8042E95DCEF8D1D">901</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="2D71B229054ED21B1546DAC54FC14B69" unit="page">
|
||
<mods:start id="3BDD25E821DF90AFE8555B8FF247C727">1</mods:start>
|
||
<mods:end id="7D9404DD87F31C55769EAC3811B0644F">81</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="2C21CCD2486907EA83919A5241729B64">
|
||
<mods:url id="EBF429582F63A4AAC62A1DE5DEFB9A76">http://dx.doi.org/10.3897/zookeys.901.39432</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="9E37B110B406207FCCDF62429EA431FC">journal article</mods:classification>
|
||
<mods:identifier id="1576FDB56DD9D80B3EE93896E73D569D" type="DOI">http://dx.doi.org/10.3897/zookeys.901.39432</mods:identifier>
|
||
<mods:identifier id="29361E9FA7385D23661F047867A8CA3E" type="Pensoft-Pub">1313-2970-901-1</mods:identifier>
|
||
<mods:identifier id="54C7F0A56E6C4FB760C532098049AFF0" type="Pensoft-UUID">D8C38C1C3844549F8BEC7C1D884CDF8E</mods:identifier>
|
||
<mods:identifier id="85E8998C594E2C621713EC6BD04AE79E" type="ZooBank">843F6065-D00E-48E6-9A26-9CEB044BF102</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="2539ACAEEF2E5F4DA7DF755F030D1F55" ID-GBIF-Taxon="161671761" LSID="urn:lsid:plazi:treatment:2539ACAEEF2E5F4DA7DF755F030D1F55" httpUri="http://treatment.plazi.org/id/2539ACAEEF2E5F4DA7DF755F030D1F55" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Scaritinae" scope_tribe="Salcediini">
|
||
<subSubSection id="EAE50459455E31F2DFA020507C9E121B" pageId="0" pageNumber="1" type="nomenclature">
|
||
<paragraph id="3274CEF889D5DE07478E406A91C769BD" pageId="0" pageNumber="1">
|
||
<taxonomicName id="BF7CA18876CE7834D2A836AFAD26ED2D" ID-CoL="6XBSD" LSID="2539ACAE-EF2E-5F4D-A7DF-755F030D1F55" authority="Fairmaire, 1899" authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia perrieri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="perrieri">Salcedia perrieri Fairmaire, 1899</taxonomicName>
|
||
<figureCitation id="5CE570E8E2E1EC7BDD1BF6B2D6E52760" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Semi-schematic dorsal view of Salcedia perrieri Fairmaire highlighting some of the features and discriminating characters used in the descriptions and the identification key. Interneurs 7 and 8 visible in ventral view (see Fig. 2)." figureDoi="10.3897/zookeys.901.39432.figure1" httpUri="https://binary.pensoft.net/fig/370602" pageId="0" pageNumber="1">Figs 1</figureCitation>
|
||
<figureCitation id="B82716D1F89F3CBC997F2AE325CC6C4C" captionStart="Figures 3, 4" captionStartId="F3" captionText="Figures 3, 4. Identical Salcedia specimens in uncleaned (a) and cleaned (b) condition 3 S. perrieri Fairmaire 4 S. lukulua sp. nov." figureDoi="10.3897/zookeys.901.39432.figures3-4" httpUri="https://binary.pensoft.net/fig/370604" pageId="0" pageNumber="1">, 3</figureCitation>
|
||
<figureCitation id="4890B7D5EC4083C4E5F99AB430B15C94" captionStart="Figures 8–13" captionStartId="F7" captionText="Figures 8 - 13. Salcedia species, habitus, dorsal view, holotype 8 S. perrieri Fairmaire 9 S. unifoveata sp. nov. 10 S. faillei sp. nov. 11 S. coquilhati Alluaud 12 S. elongata Alluaud 13 S. africana (Britton)." figureDoi="10.3897/zookeys.901.39432.figures8-13" httpUri="https://binary.pensoft.net/fig/370608" pageId="0" pageNumber="1">, 8</figureCitation>
|
||
<figureCitation id="3A2F17550924D43EB4507C27DE378FB6" captionStart="Figures 26–35" captionStartId="F10" captionText="Figures 26 - 35. Salcedia species, pronotum, dorsal view 26 S. perrieri Fairmaire 27 S. unifoveata sp. nov. 28 S. faillei sp. nov. 29 S. coquilhati Alluaud 30 S. elongata Alluaud 31 S. africana (Britton) 32 S. schoutedeni Alluaud 33 S. nigeriensis Alluaud 34 S. putzeysi (Oberthuer) 35 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures26-35" httpUri="https://binary.pensoft.net/fig/370611" pageId="0" pageNumber="1">, 26</figureCitation>
|
||
<figureCitation id="FC2FF1AD952315111FBA866B17D51A80" captionStart="Figures 44–51" captionStartId="F12" captionText="Figures 44 - 51. Salcedia species, left elytron, basal half, dorsal view 44 S. perrieri Fairmaire 45 S. unifoveata sp. nov. 46 S. elongata Alluaud 47 S. africana (Britton) 48 S. schoutedeni Alluaud 49 S. nigeriensis Alluaud 50 S. baroensis sp. nov. 51 S. miranda (Andrewes)." figureDoi="10.3897/zookeys.901.39432.figures44-51" httpUri="https://binary.pensoft.net/fig/370613" pageId="0" pageNumber="1">, 44</figureCitation>
|
||
<figureCitation id="5E83857C1FD6BBE770161716FF6E2415" captionStart="Figures 52–57" captionStartId="F13" captionText="Figures 52 - 57. Salcedia species, male genitalia, dorsal view of aedeagus (a) and parameres (b, c), lateral view (d). Small sketches in black symbolising the apex in cross section 52 S. perrieri Fairmaire 53 S. unifoveata sp. nov. 54 S. faillei sp. nov. 55 S. coquilhati Alluaud 56 S. elongata Alluaud 57 S. africana (Britton)." figureDoi="10.3897/zookeys.901.39432.figures52-57" httpUri="https://binary.pensoft.net/fig/370614" pageId="0" pageNumber="1">, 52</figureCitation>
|
||
<figureCitation id="37FF42E9015E098F1E0FE53D4E69BB89" captionStart="Figures 68–76" captionStartId="F16" captionText="Figures 68 - 76. Salcedia species, female coxostyli (gonopod IX) and laterotergite IX (epipleutite IX) 68 S. perrieri Fairmaire 69 S. unifoveata sp. nov. 70 S. coquilhati Alluaud 71 S. elongata Alluaud 72 S. africana (Britton) 73 S. schoutedeni Alluaud 74 S. nigeriensis Alluaud 75 S. putzeysi (Oberthuer) 76 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures68-76" httpUri="https://binary.pensoft.net/fig/370617" pageId="0" pageNumber="1">, 68</figureCitation>
|
||
<figureCitation id="78177C927F8FD83474F4E727B7C7BC24" captionStart="Figure 84" captionStartId="F18" captionText="Figure 84. Map of tropical Africa providing an overview of the occurrence of all African Salcedia species described to date (recorded localities plotted). 1 (black S. perrieri Fairmaire; 2 (white) S. unifoveata sp. nov.; 3 (black circle) S. faillei sp. nov.; 4 (red) S. coquilhati Alluaud; 5 (yellow) S. elongata Alluaud; 6 (blue) S. africana (Britton); 7 (orange) S. utetea sp. nov.; 8 (grey) S. procera sp. nov.; 9 (light green) S. robusta sp. nov.; 10 (turquoise) S. schoutedeni Alluaud; 11 (purple) S. lukulua sp. nov.; 12 (dark green) S. tuberculata sp. nov.; 13 (brown) S. baroensis sp. nov.; 14 (light blue) S. nigeriensis Alluaud; 16 (dark blue) S. putzeysi (Oberthuer); 17 (black) S. matsumotoi sp. nov.;? in a circle (twice): collection locality of a single female Salcedia specimen of uncertain species. Basic schematic map of Africa taken from SimpleMappr. net." figureDoi="10.3897/zookeys.901.39432.figure84" httpUri="https://binary.pensoft.net/fig/370619" pageId="0" pageNumber="1">, 84</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F4BBFA2B5F7449AD706E17AC29CA70BD" pageId="0" pageNumber="1" type="reference_group">
|
||
<paragraph id="1A1298DFA07E6A625D5A98A9F73490C3" pageId="0" pageNumber="1">
|
||
<taxonomicName id="DF07BA08BDF02B99967347FF432523DF" authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia perrieri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="perrieri">Salcedia perrieri</taxonomicName>
|
||
Fairmaire, 1899: 512;
|
||
<bibRefCitation id="5065F3AA770EFA2B000CA073E4F89167" author="Csiki, E" journalOrPublisher="Entomologist's monthly magazine, May" pageId="0" pageNumber="1" refId="B20" refString="Csiki, E, 1927. Coleopterorum Catalogus auspiciis et auxilio. W. Junk editus a S. Schenkling. 91: Carabidae: Carabinae I. W. Junk, Berlin, 1-553." title="Coleopterorum Catalogus auspiciis et auxilio. W. Junk editus a S. Schenkling. 91: Carabidae: Carabinae I. W. Junk, Berlin, 1 - 553." year="1927">Csiki 1927</bibRefCitation>
|
||
: 546;
|
||
<bibRefCitation id="C6018C5C1A779CA89A582E63E18A01FB" author="Alluaud, C" journalOrPublisher="Afra" pageId="0" pageNumber="1" pagination="10 - 23" refId="B2" refString="Alluaud, C, 1930. Note sur les Scaritides de Madagascar. Afra 1: 10 - 23" title="Note sur les Scaritides de Madagascar." volume="1" year="1930">Alluaud 1930</bibRefCitation>
|
||
: 22;
|
||
<bibRefCitation id="1628FAB0A56235AECD094B2FF8A34B10" author="Jeannel, R" journalOrPublisher="Office de la Recherche Scientifique Coloniale, Paris" pageId="0" pageNumber="1" refId="B31" refString="Jeannel, R, 1946. Faune de l'empire francais. VI. Coleopteres carabiques de la region Malgache (premiere partie). Office de la Recherche Scientifique Coloniale, Paris" title="Faune de l'empire francais. VI. Coleopteres carabiques de la region Malgache (premiere partie)." year="1946">Jeannel 1946</bibRefCitation>
|
||
: 233;
|
||
<bibRefCitation id="486983EBB1B6CA201ACBF3E1A251B0DC" author="Basilewsky, P" journalOrPublisher="Faune de Madagascar" pageId="0" pageNumber="1" pagination="1 - 322" refId="B14" refString="Basilewsky, P, 1973. Insectes Coleopteres Carabidae Scaritinae. Faune de Madagascar 37: 1 - 322" title="Insectes Coleopteres Carabidae Scaritinae." volume="37" year="1973">Basilewsky 1973</bibRefCitation>
|
||
: 293;
|
||
<bibRefCitation id="8EB739E5FC7FB1ECAFD46582CA199C3C" author="Reichardt, H" journalOrPublisher="Revista Brasileira Entomologia" pageId="0" pageNumber="1" pagination="99 - 110" refId="B38" refString="Reichardt, H, 1975. The genera of Salcediina (Coleoptera, Carabidae, Scaritini). Revista Brasileira Entomologia 19: 99 - 110" title="The genera of Salcediina (Coleoptera, Carabidae, Scaritini)." volume="19" year="1975">Reichardt 1975</bibRefCitation>
|
||
: 103;
|
||
<bibRefCitation id="11B839EE12632E8398837D5CB85260CA" author="Peyrieras, A" journalOrPublisher="Faune de Madagascar" pageId="0" pageNumber="1" pagination="1 - 161" refId="B37" refString="Peyrieras, A, 1976. Insectes Coleopteres Carabidae Scaritinae. II. Biologie. Faune de Madagascar 41: 1 - 161" title="Insectes Coleopteres Carabidae Scaritinae. II. Biologie." volume="41" year="1976">Peyrieras 1976</bibRefCitation>
|
||
: 157;
|
||
<bibRefCitation id="4904F54009653AB7816CF93BDCF49AB9" author="Dostal, A" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="0" pageNumber="1" pagination="117 - 121" refId="B24" refString="Dostal, A, 1993. Beschreibung einer neuen Gattung aus Vietnam: Androzelma gen. nov. (Coleoptera, Carabidae, Scaritinae: Salcediini). Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen 45: 117 - 121" title="Beschreibung einer neuen Gattung aus Vietnam: Androzelma gen. nov. (Coleoptera, Carabidae, Scaritinae: Salcediini)." volume="45" year="1993">Dostal 1993</bibRefCitation>
|
||
: 121;
|
||
<bibRefCitation id="FEAD85EAF4FE3D4C5DF297AFAF72DFDF" author="Deuve, T" journalOrPublisher="Memoires du Museum national d'Histoire naturell" pageId="0" pageNumber="1" pagination="1 - 184" refId="B23" refString="Deuve, T, 1993. L'abdomen et les genitalia des femelles de Coleopteres Adephaga. Memoires du Museum national d'Histoire naturell 155: 1 - 184" title="L'abdomen et les genitalia des femelles de Coleopteres Adephaga." volume="155" year="1993">Deuve 1993</bibRefCitation>
|
||
: 141;
|
||
<bibRefCitation id="D2E3BF6898E16DBF1BD348B1E2A0C983" author="Lorenz, W" journalOrPublisher="Printed by the author, Tutzing" pageId="0" pageNumber="1" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant Ground Beetles of the World (InsectaColeoptera "Geadephaga": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae), Second Edition. Printed by the author, Tutzing" title="Systematic list of extant Ground Beetles of the World (InsectaColeoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae), Second Edition." year="2005">Lorenz 2005</bibRefCitation>
|
||
: 155.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="319E9E575DE369003B2397365EB8AEE2" pageId="0" pageNumber="1" type="type material">
|
||
<paragraph id="3193F6F9B3DDAA1CBBB23485CEFBC278" pageId="0" pageNumber="1">Type material.</paragraph>
|
||
<paragraph id="C0E9C681DA92592B51E0165C138BA689" pageId="0" pageNumber="1">
|
||
<emphasis id="D964574882DB28F13E40E93A936CF76F" bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||
: ♂, with labels and data: brownish, printed, black framed "Madagr. Suberblle. H.Perrier" (MNHN).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="33087C72D8A7C9F974ABB0673DDCBCA8" pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph id="116B85AA4DA5D03BF88A4D7E05642BB7" pageId="0" pageNumber="1">Remark.</paragraph>
|
||
<paragraph id="552A2FDFEB84C671758B5193C24AAA40" pageId="0" pageNumber="1">The holotype is complete. The specimen was located in the collection Fairmaire in MNHN.</paragraph>
|
||
<caption id="396BCB1115B32E5A01ACF5645734736F" doi="10.3897/zookeys.901.39432.figures8-13" httpUri="https://binary.pensoft.net/fig/370608" pageId="0" pageNumber="1" start="Figures 8–13" startId="F7">
|
||
<paragraph id="F04BC4480C3990C12504EA48B55E86FB" pageId="0" pageNumber="1">
|
||
<emphasis id="5F1533693DDF6026247B45AFBB4D6281" bold="true" pageId="0" pageNumber="1">Figures 8-13.</emphasis>
|
||
<taxonomicName id="CF384A4FFB7820250F4FE989BD8031D7" authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="699896E16AF9B8BEE6D5EC4591C6124B" italics="true" pageId="0" pageNumber="1">Salcedia</emphasis>
|
||
</taxonomicName>
|
||
species, habitus, dorsal view, holotype
|
||
<emphasis id="9088708B7A0203E854088ECEB6F727E0" bold="true" pageId="0" pageNumber="1">8</emphasis>
|
||
<taxonomicName id="F458E0DA724D448C603926BD42317942" lsidName="S. perrieri" pageId="0" pageNumber="1" rank="species" species="perrieri">
|
||
<emphasis id="743019475D74852143E919FB2C65C473" italics="true" pageId="0" pageNumber="1">S. perrieri</emphasis>
|
||
</taxonomicName>
|
||
Fairmaire
|
||
<emphasis id="F77B7CDEFD21A19C418D7F283FCB9AA5" bold="true" pageId="0" pageNumber="1">9</emphasis>
|
||
<taxonomicName id="01617018E1CABBE545E805A8EE8C62CB" lsidName="S. unifoveata" pageId="0" pageNumber="1" rank="species" species="unifoveata">
|
||
<emphasis id="065D866AE12F0B9DE6555C4E5D4D2DF2" italics="true" pageId="0" pageNumber="1">S. unifoveata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="BB7634F887E3D9FBF84E48DA6B5990F0" bold="true" pageId="0" pageNumber="1">10</emphasis>
|
||
<taxonomicName id="36BEC1C9604A547582F25BD169B2B303" lsidName="S. faillei" pageId="0" pageNumber="1" rank="species" species="faillei">
|
||
<emphasis id="C05FE86044D58F3BA1B4F9CD5E354F1D" italics="true" pageId="0" pageNumber="1">S. faillei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="0F3CF0EB8977D988966BD1874AB088AF" bold="true" pageId="0" pageNumber="1">11</emphasis>
|
||
<taxonomicName id="DA946D0BCC03D37282694861CCB6690A" lsidName="S. coquilhati" pageId="0" pageNumber="1" rank="species" species="coquilhati">
|
||
<emphasis id="4E48E866CAA7E84170CCF493DA825CA0" italics="true" pageId="0" pageNumber="1">S. coquilhati</emphasis>
|
||
</taxonomicName>
|
||
Alluaud
|
||
<emphasis id="1B900F92228C099009CCAECFEE5F4E81" bold="true" pageId="0" pageNumber="1">12</emphasis>
|
||
<taxonomicName id="32089FC1657371FA7635AEFAA69A5DC2" lsidName="S. elongata" pageId="0" pageNumber="1" rank="species" species="elongata">
|
||
<emphasis id="D2A36CBC557C45F7FCE62D628E3E3289" italics="true" pageId="0" pageNumber="1">S. elongata</emphasis>
|
||
</taxonomicName>
|
||
Alluaud
|
||
<emphasis id="8FDE4D8B8801E984FF846F04753B1AE9" bold="true" pageId="0" pageNumber="1">13</emphasis>
|
||
<taxonomicName id="36BFB87C6237915C1509BDC216741569" lsidName="S. africana" pageId="0" pageNumber="1" rank="species" species="africana">
|
||
<emphasis id="73677F0606010C283D4FC51FCD79C204" italics="true" pageId="0" pageNumber="1">S. africana</emphasis>
|
||
</taxonomicName>
|
||
(Britton).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="C16676CD639A301E621CBFA92E7903BC" pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph id="E131EE55EFDCD639C4308B4DBC637C54" pageId="0" pageNumber="1">Additional material.</paragraph>
|
||
<paragraph id="A1A25B4752F59B13C773B1C5A907A397" pageId="0" pageNumber="1">
|
||
1 spec., Madagascar Ron Maroantsetra V 35 Vadon! (MNHN, CBB); 29 specs., Madagascar Ron Maroantsetra IV 38 Vadon! / Andranofotsy (MNHN); 2 ♂, 2 specs., same data but with label COLL. MUS. Congo Madagascar: Col.P.Basilewsky / one of them with additional blue label:
|
||
<normalizedToken id="D17BEF0582C164EEC24F2ADAE7C91CE3" originalValue="MUSÉUM">MUSEUM</normalizedToken>
|
||
PARIS 1938 J. VADON et E. LEBIS (MRACT); 4 specs., Madagascar Ron Maroantsetra IV 38 Vadon! / Antakotako (MNHN); 1 spec., same data but Mahakiry (MNHN); 1 ♀, 3 specs., Madagascar, Fampanambo XII.1958 86 (Inondations) / MUS. ROY. CENTR. Madagascar Est: Bale
|
||
<normalizedToken id="381468D26277F1440247185673ECD583" originalValue="d’Antongli">d'Antongli</normalizedToken>
|
||
J. Vadon (MRACT); 1 spec., same data but Maroantsetra Fampanambo III. 56 (MRACT); 1 spec., Madagascar: Fampanambo II.1960 J. Vadon /
|
||
<taxonomicName id="39B0AB3BFD2242525C797785BC833083" authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia perrieri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="perrieri">
|
||
<emphasis id="78090645B0A1028343D419F57420B414" italics="true" pageId="0" pageNumber="1">Salcedia perrieri</emphasis>
|
||
</taxonomicName>
|
||
Frm. P. Basilewsky det., 1970 / COMP. TYP. BASILEWSKY (MRACT); 1 spec., same data but III 65 (Vadon) (MNHN); 1 spec., Maroantsetra Ambodivoangy J.Vadon / INSTITUT SCIENTIFIQUE MADAGASCAR (MNHN); 11 specs., Ankofa VII 56 n. 7 / INSTITUT SCIENTIFIQUE MADAGASCAR (MNHN, CBB); 1 ♂, 3 ♀, 1 spec., Ankofa XII. 55 n. 7 / INSTITUT SCIENTIFIQUE MADAGASCAR / COLL. MUS: TERVUREN (ex
|
||
<normalizedToken id="18EB36AAAFF00BC2F7910DC67222B0D6" originalValue="Muséum">Museum</normalizedToken>
|
||
Paris) Coll. P. Basilewsky (MRACT); 1 spec., Tananarive Tsimbazaza / INSTITUT SCIENTIFIQUE MADAGASCAR /
|
||
<taxonomicName id="E644870C1091F9814093E8D9134C6876" authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia perrieri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="perrieri">
|
||
<emphasis id="0EE5DB5A2E73DE9C14F10A606E4F3B4D" italics="true" pageId="0" pageNumber="1">Salcedia perrieri</emphasis>
|
||
</taxonomicName>
|
||
Fairm. P. Basilewsky det. 1971 (MNHN); 1 spec., same data but with add. label COLL. MUS: TERVUREN (ex
|
||
<normalizedToken id="02276986D225C1E4D0886B7FC7B6AFA2" originalValue="Muséum">Museum</normalizedToken>
|
||
Paris) Coll. P. Basilewsky (MRACT); 1 ♂, 2 ♀, MADAGASCAR c. Antsiranana prov., 2002 Ambanja, Ambodidimaka r. 15-16.12., Ivo
|
||
<normalizedToken id="EC2FCD0BFC4D68970A7C0B332C0952E3" originalValue="Jeniš">Jenis</normalizedToken>
|
||
leg. (CBP); 1 spec., Madagascar Ron Maroantsetra IV 38 Vadon! / Andranofotsy /
|
||
<taxonomicName id="CC72B803837B5D78837E44911E84C719" authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia perrieri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="perrieri">
|
||
<emphasis id="0139DB430A54310F3EB6C7473D51028B" italics="true" pageId="0" pageNumber="1">Salcedia perrieri</emphasis>
|
||
</taxonomicName>
|
||
Fm. / COLLECTIO KAREL KULT COLL.A.DOSTAL, 1999 / Coll. P. Bulirsch Prague Czech Republic (CBP); 1 ♀, MADAGASCAR ?? Perinet 1989 (CBP); 1 spec., right antenna, right hind tibia and right intermediate tarsomere missing, MADAGASCAR, Ambanja Antsiranana pr. XII. 2002 / BMNH (E) 2008-52 P.Bulirsch/ SALCEDIA
|
||
<taxonomicName id="5F152C40BEEA5A39B17EABE0AF666EEB" authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="perrieri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="perrieri">perrieri</taxonomicName>
|
||
Fairmaire, 1899 P.Bulirsch det. 2011 (BMNH); 2 specs., Madagascar Ron Maroantsetra IV 38 Vadon! / Andranofotsy / with additional blue label:
|
||
<normalizedToken id="16909306AC32391B14FD9E01AA46D34D" originalValue="MUSÉUM">MUSEUM</normalizedToken>
|
||
PARIS 1938 J. VADON et E. LEBIS / with additional white label: BRIT.MUS. 1947-246. (BMNH); 1 ♀, Madag Perrier (MNHN); 1 ♂, Morondava
|
||
<normalizedToken id="AF146FBAB518DF47528CAA5339E44A03" originalValue="fôret">foret</normalizedToken>
|
||
sud de Befasy I-56 R.P. / INSTITUT SCIENTIFIQUE MADAGASCAR (MNHN).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B93937524A393BFA051107695193CF79" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="73CA2EE87A7ED2965ACC651F1F313481" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="4316F5C715B3E7FE941E89CF541559A7" pageId="0" pageNumber="1">
|
||
A medium sized species, with sub-elongate oval outline of the elytra with maximum width at middle and the pronotum with three additional lateral carinae. The pseudohumerus is nearly rectangular and distinctly dentate. The antennomeres are elongate. Distinguished most clearly from the similar species
|
||
<taxonomicName id="02E20EF582A6CD63AC28FAACA0FA4963" lsidName="S. faillei" pageId="0" pageNumber="1" rank="species" species="faillei">
|
||
<emphasis id="403A714AB25E17AD4D42C73606CDF3C7" italics="true" pageId="0" pageNumber="1">S. faillei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by the tubercle on the frons of the head, the twelve tubercles at the lateral margin of the pronotum, and the pseudohumerus of the elytron which shows a laterally projecting tooth. Moreover, the median lobe of the male genitalia is different and the dorsal paramere has a pilus at the apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8DC4524D626A858F3235A409418F6FCB" pageId="0" pageNumber="1" type="redescription">
|
||
<paragraph id="6115D4E27C42E2FEE0AA9CE7D4BF8E55" pageId="0" pageNumber="1">Redescription.</paragraph>
|
||
<paragraph id="0094733FAA8D0CCE20EB6EDBBBE158AD" pageId="0" pageNumber="1">
|
||
Measurements in Table
|
||
<tableCitation id="260C9A94A8BCBA9DF7D9635F384D165D" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and ratios of Salcedia species. N = number of specimens measured; x ̄ = arithmetic mean; " term used " refers to the shape of the elytra." httpUri="http://table.plazi.org/id/FDDA58A1B347C9F0593A738CF707345F" pageId="0" pageNumber="1" tableUuid="FDDA58A1B347C9F0593A738CF707345F">1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="55BF99C1062D0C018E66D131C5736F33" pageId="0" pageNumber="1">
|
||
<emphasis id="27EAADC33258C38CECE1E1EF4BD1109B" bold="true" italics="true" pageId="0" pageNumber="1">Colour and surface:</emphasis>
|
||
Anthracite grey, opaque; top of carinae on head, pronotum and elytra shiny; mandibles and legs piceous, antennae and palpi leoninous.
|
||
</paragraph>
|
||
<paragraph id="8E83949F35C053760541B65CF090AAF9" pageId="0" pageNumber="1">
|
||
<emphasis id="95A81241B06170695F78C211907686BF" bold="true" italics="true" pageId="0" pageNumber="1">Head:</emphasis>
|
||
Two-thirds of width of pronotum. Outline semi-circular. Clypeus wide, straight anteriorly, separated from convex clypeal wings by indistinct obtuse notches, clypeal wings separated from supra-antennal plates by distinct notches, with raised pentagon-shaped field at middle, separated from frons by deep transverse furrow, frons with two raised paramedian carinae, joining anteriorly, prolonged anteriorly into a central distinctly erected tubercle, with two small glossy teeth bilaterally anterior to central tubercle, with two parallel carinae paralaterally near base; frons and clypeus separated from supra-antennal and supraorbital plates by deep broad furrows, each furrow with deep circular pit between supra-antennal plate and clypeus; supra-antennal and supraorbital plates distinctly margined, margin raised, carina-like, indistinctly crenulated, supra-antennal plates vaulted. Base emarginated at middle, obtuse angled laterally (angle 120°). Eyes convex, genae slightly convex, both of them partly visible from above, with pentagonal shape in lateral view. Antenna with segments five to ten elongate (L/W 1.26), densely pubescent, segments two to four scarcely pubescent, scapus with longitudinal reticulation. Labrum excised anteriorly. Mandible moderately short, wide, slightly arcuate at apex. Mentum small, epilobes wide, projecting and obtuse angled anteriorly, completely margined, surface coriaciate-like with flat impressions.
|
||
</paragraph>
|
||
<paragraph id="5FC8F845E0CC4FE5A8B90CE17CDC9C42" pageId="0" pageNumber="1">
|
||
<emphasis id="CA91A026C8155CEFC8CFDF0C93B6637B" bold="true" italics="true" pageId="0" pageNumber="1">Pronotum</emphasis>
|
||
(Fig.
|
||
<figureCitation id="3CEF41AF9E7D89642275EF32F3DA6F71" captionStart="Figures 26–35" captionStartId="F10" captionText="Figures 26 - 35. Salcedia species, pronotum, dorsal view 26 S. perrieri Fairmaire 27 S. unifoveata sp. nov. 28 S. faillei sp. nov. 29 S. coquilhati Alluaud 30 S. elongata Alluaud 31 S. africana (Britton) 32 S. schoutedeni Alluaud 33 S. nigeriensis Alluaud 34 S. putzeysi (Oberthuer) 35 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures26-35" httpUri="https://binary.pensoft.net/fig/370611" pageId="0" pageNumber="1">26</figureCitation>
|
||
): Outline rectangular, transverse, a third wider than long. Lateral margin convex, maximum width at middle, converging anteriorly and posteriorly. Lateral margin distinctly crenulated, with 12 tubercles, with distinct emargination at posterior angles. Base posteriorly moderately wing-like produced, with distinct notch between wing and central part of base, straight posteriorly. Declining flat keel at middle of base pointing posteriorly. Disc slightly convex (lateral view), with two raised paramedian carinae parallel to median line, slightly diverging posteriorly, with median line small, short, ending in pits anteriorly and posteriorly, with four additional shorter carinae bilaterally, the paralateral one joining with the paramedian carinae at base and forming a tooth-like tubercle pointing posteriorly, extended anteriorly as less raised paralateral carina. Two much shorter and less raised inner and outer lateral carinae, the inner one joining with the anterior extension of paralateral carina, all carinae sub-crenulate. Lateral margin and space between lateral margin and paralateral carina wing-like bent up, with six large transverse pits, the basal ones partly separated into two small circular pits.
|
||
</paragraph>
|
||
<paragraph id="414848D0FDB0874821C01459777DB81B" pageId="0" pageNumber="1">
|
||
<emphasis id="9876C6A32BBCC92D3115EC115F6E8A17" bold="true" italics="true" pageId="0" pageNumber="1">Elytron</emphasis>
|
||
(Fig.
|
||
<figureCitation id="8C0CB9BC8EE48F1FF5C9C3070527EE15" captionStart="Figures 44–51" captionStartId="F12" captionText="Figures 44 - 51. Salcedia species, left elytron, basal half, dorsal view 44 S. perrieri Fairmaire 45 S. unifoveata sp. nov. 46 S. elongata Alluaud 47 S. africana (Britton) 48 S. schoutedeni Alluaud 49 S. nigeriensis Alluaud 50 S. baroensis sp. nov. 51 S. miranda (Andrewes)." figureDoi="10.3897/zookeys.901.39432.figures44-51" httpUri="https://binary.pensoft.net/fig/370613" pageId="0" pageNumber="1">44</figureCitation>
|
||
): Flattened in anterior half (lateral view), convex in frontal view. Sub-elongate, margin moderately convex, maximum width at middle. Pseudohumerus nearly rectangular, distinctly dentate. Apex rounded, acutely denticulate at suture. Disc with interneur six sub-crenulate, interneur two running up to apex as slightly convex line, conspicuously raised; interneur four running in parallel to interneur six, not reaching base, shortened at apex. Interneur five and six with two rows of serial pits, the latter ones partly doubled at base and transversally connected apically.
|
||
</paragraph>
|
||
<paragraph id="BCA02ACF547B4D030A3F05B3402EF279" pageId="0" pageNumber="1">
|
||
<emphasis id="305A24F7C04CE3EDEC3FEB6AA72A3DEA" bold="true" italics="true" pageId="0" pageNumber="1">Hind wings:</emphasis>
|
||
Fully developed.
|
||
</paragraph>
|
||
<paragraph id="D91CBF2AB30CB1AECB7A738232448B7D" pageId="0" pageNumber="1">
|
||
<emphasis id="7D7AD0975C4B726DAB5A073B0CAB8239" bold="true" italics="true" pageId="0" pageNumber="1">Lower surface:</emphasis>
|
||
Antennal channel of pronotum with isodiametric reticulation. Pseudoepipleura with one row of serial pits, lateral margin of elytron smooth. Metasternum and last five abdominal sternites with numerous irregularly situated larger and smaller pits, regularly arranged at carinae and margins.
|
||
</paragraph>
|
||
<paragraph id="4EDC40DEF43BD01ED8F169E49666C945" pageId="0" pageNumber="1">
|
||
<emphasis id="8B50EB7B51BFFDCC8751805644164F3D" bold="true" italics="true" pageId="0" pageNumber="1">Legs:</emphasis>
|
||
Profemora with irregular pits on dorsal surface. Protibia laterally with two larger and one smaller tooth, dorsally with one and ventrally with two carinae. Metafemora with small, short grooves at dorsal surface.
|
||
</paragraph>
|
||
<paragraph id="1DA785BC7F1545C7E903F4CDF7DDB8ED" pageId="0" pageNumber="1">
|
||
<emphasis id="FEE259F02325B07A93119177B605110C" bold="true" italics="true" pageId="0" pageNumber="1">External sexual dimorphism:</emphasis>
|
||
Not observed.
|
||
</paragraph>
|
||
<paragraph id="88A3AB7ADA07458D8106A5F733A79DFF" pageId="0" pageNumber="1">
|
||
<emphasis id="1273A2A1FBF1DECA211D918405D1B883" bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
|
||
(Fig.
|
||
<figureCitation id="73A06BC92B1FF0C328B973D222F7147D" captionStart="Figures 52–57" captionStartId="F13" captionText="Figures 52 - 57. Salcedia species, male genitalia, dorsal view of aedeagus (a) and parameres (b, c), lateral view (d). Small sketches in black symbolising the apex in cross section 52 S. perrieri Fairmaire 53 S. unifoveata sp. nov. 54 S. faillei sp. nov. 55 S. coquilhati Alluaud 56 S. elongata Alluaud 57 S. africana (Britton)." figureDoi="10.3897/zookeys.901.39432.figures52-57" httpUri="https://binary.pensoft.net/fig/370614" pageId="0" pageNumber="1">52</figureCitation>
|
||
): Median lobe large, moderately slender, in lateral view nearly straight, in dorsal view slightly arcuate in basal half and at middle, distinctly arcuate in apical third, with few fine pili at beginning of apical third, apex stick-like. Endophallus with two longitudinal groups of densely situated microtrichia, groups as long as one third of median lobe. Dorsal paramere moderately wide at base, bisinuate, with pilus at apex, lateral and basal apophysis blunt; ventral one of moderate size; both parameres somewhat distorted.
|
||
</paragraph>
|
||
<paragraph id="53FBF5B7ECE32A9775415EECF949AF94" pageId="0" pageNumber="1">
|
||
<emphasis id="467A93A62C8E1A2242CD3F8B3C9D8261" bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
|
||
(Fig.
|
||
<figureCitation id="B686EBD60DE4CD34418BB755EC9F8CF3" captionStart="Figures 68–76" captionStartId="F16" captionText="Figures 68 - 76. Salcedia species, female coxostyli (gonopod IX) and laterotergite IX (epipleutite IX) 68 S. perrieri Fairmaire 69 S. unifoveata sp. nov. 70 S. coquilhati Alluaud 71 S. elongata Alluaud 72 S. africana (Britton) 73 S. schoutedeni Alluaud 74 S. nigeriensis Alluaud 75 S. putzeysi (Oberthuer) 76 S. matsumotoi sp. nov." figureDoi="10.3897/zookeys.901.39432.figures68-76" httpUri="https://binary.pensoft.net/fig/370617" pageId="0" pageNumber="1">68</figureCitation>
|
||
): Coxostylus moderately broad at base, obtusely curved, moderately broad in apical third, rounded at apex, at end of basal third with one broader and five slender nematiform setae laterally, with one long slender nematiform seta ventrally, SSO with one microtrichium.
|
||
</paragraph>
|
||
<paragraph id="9CDD5EE693B9D37CC29F3AB53F536B6D" pageId="0" pageNumber="1">
|
||
<emphasis id="C4538C8D9CB3EE0A9385707B3646949D" bold="true" italics="true" pageId="0" pageNumber="1">Variation:</emphasis>
|
||
The carinae on the frons of the head are tuberculated to a different degree. The number and shape of the pits on the ventral surface varies to a small degree. Slight differences could also be observed with the flat impressions on the lower surface. The female specimen from Perinet (label indicating uncertain location) is slightly larger than the other ones but in all other characters matches
|
||
<taxonomicName id="DA835C7FEF9686C24343C0352E29FD87" lsidName="S. perrieri" pageId="0" pageNumber="1" rank="species" species="perrieri">
|
||
<emphasis id="FD96F54EFFFE696B8FB9DD8A0E0070F0" italics="true" pageId="0" pageNumber="1">S. perrieri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7EC439EA30756F521CD52986284A35A2" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="628267DB41D7455A9CD18439F0E70F74" pageId="0" pageNumber="1">Distribution</paragraph>
|
||
<paragraph id="16EA4C00F48C800F15B33739B87D38A1" pageId="0" pageNumber="1">
|
||
(Fig.
|
||
<figureCitation id="974441EFF75AE6CFC1A66066F47FAD06" captionStart="Figure 84" captionStartId="F18" captionText="Figure 84. Map of tropical Africa providing an overview of the occurrence of all African Salcedia species described to date (recorded localities plotted). 1 (black S. perrieri Fairmaire; 2 (white) S. unifoveata sp. nov.; 3 (black circle) S. faillei sp. nov.; 4 (red) S. coquilhati Alluaud; 5 (yellow) S. elongata Alluaud; 6 (blue) S. africana (Britton); 7 (orange) S. utetea sp. nov.; 8 (grey) S. procera sp. nov.; 9 (light green) S. robusta sp. nov.; 10 (turquoise) S. schoutedeni Alluaud; 11 (purple) S. lukulua sp. nov.; 12 (dark green) S. tuberculata sp. nov.; 13 (brown) S. baroensis sp. nov.; 14 (light blue) S. nigeriensis Alluaud; 16 (dark blue) S. putzeysi (Oberthuer); 17 (black) S. matsumotoi sp. nov.;? in a circle (twice): collection locality of a single female Salcedia specimen of uncertain species. Basic schematic map of Africa taken from SimpleMappr. net." figureDoi="10.3897/zookeys.901.39432.figure84" httpUri="https://binary.pensoft.net/fig/370619" pageId="0" pageNumber="1">84</figureCitation>
|
||
): Known from several localities in north-east and Central Madagascar. As far as the label information allows it can be stated the specimens were collected at the margins of inland waters. Some of the specimens were collected after flooding. The type locality Suberbieville is located on the River Ikopa near Maevatana in the north-west of Madagascar.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |