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<document id="596D04580545B1A2FE6B81FA820722CB" ID-CLB-Dataset="55534" ID-DOI="10.5281/zenodo.5413940" ID-GBIF-Dataset="7efc5233-fc92-4a45-b647-ca79436ae20c" ID-ISSN="0253-116X" ID-Zenodo-Dep="5413940" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630356940101" checkinUser="marcus" docAuthor="Assing, Volker" docDate="2015" docId="165187CC927CFFF0FF6FFC73FD4111DE" docLanguage="en" docName="LinzerbiolBeitr.2015.47.1.101-125.pdf" docOrigin="Linzer biologische Beiträge 47 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Orphnebius formosanus Assing 2015, nov.sp." docType="treatment" docVersion="10" lastPageNumber="114" masterDocId="EA68FFB49276FFFDFFDDFFAFFE1C105D" masterDocTitle="On the Orphnebius fauna of the East Palaearctic region. VI. Six new species from China and Taiwan, and additional records (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="125" masterPageNumber="101" pageNumber="111" updateTime="1699245435375" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="240F81B958FA3259E5E78D143B3CC848" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="67086BEC14231011606DF9EF8AAC2E5A">
<mods:title id="7157D861F1204E549E111112F1185EE0">On the Orphnebius fauna of the East Palaearctic region. VI. Six new species from China and Taiwan, and additional records (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
</mods:titleInfo>
<mods:name id="176C61DFDE90606EE1DBFA31E980D93A" type="personal">
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</mods:role>
<mods:namePart id="ED809511E709F28D3984F7D26A5B67E8">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="26F770228A074B2C97C46AC10AF2CFB7">text</mods:typeOfResource>
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<mods:title id="B892F3ADA2E741F0A26F7F537398F037">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="F84DFFC78220A4DA11FA149C3D246780">
<mods:date id="FE9EE195F58A09559403CDECCDC3371D">2015</mods:date>
<mods:detail id="C4698C758216893D529680CABDD3EED0" type="pubDate">
<mods:number id="F02CE6AAF5D011F76326FD4F6A91B04E">2015-07-31</mods:number>
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<mods:detail id="3D3E6087A074E98F0DE361BCEA1C78D2" type="volume">
<mods:number id="F99F137E4B25BB2695E1AB58E9F42FB5">47</mods:number>
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<mods:number id="457DA36510871830D95B36A66CE888A4">1</mods:number>
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<mods:start id="B44B03983F59ED2D13E3CC0EB020092B">101</mods:start>
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<mods:classification id="531D2E913E08E29F3A3FEFBDA6454FE8">journal article</mods:classification>
<mods:identifier id="2686B26C0EA19F454DB3DDA6DD2E8760" type="CLB-Dataset">55534</mods:identifier>
<mods:identifier id="69D51A28522D97C83E255E6F81A424C5" type="DOI">10.5281/zenodo.5413940</mods:identifier>
<mods:identifier id="97E3AD87E41156914E7D74FFC20621A4" type="GBIF-Dataset">7efc5233-fc92-4a45-b647-ca79436ae20c</mods:identifier>
<mods:identifier id="3D0491AFD6FDB6B44D776AAD6FC9DBE1" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="5982CDFAC1ECE0AD0167C15D5B11C063" type="Zenodo-Dep">5413940</mods:identifier>
</mods:mods>
<treatment id="165187CC927CFFF0FF6FFC73FD4111DE" ID-DOI="http://doi.org/10.5281/zenodo.6547275" ID-GBIF-Taxon="195305768" ID-Zenodo-Dep="6547275" LSID="urn:lsid:plazi:treatment:165187CC927CFFF0FF6FFC73FD4111DE" httpUri="http://treatment.plazi.org/id/165187CC927CFFF0FF6FFC73FD4111DE" lastPageId="13" lastPageNumber="114" pageId="10" pageNumber="111">
<subSubSection id="D6E26551927CFFF7FF6FFC73FC1E13AF" box="[178,514,988,1010]" pageId="10" pageNumber="111" type="nomenclature">
<paragraph id="9E4736DA927CFFF7FF6FFC73FC1E13AF" blockId="10.[178,718,988,1011]" box="[178,514,988,1010]" pageId="10" pageNumber="111">
<heading id="C50F81B6927CFFF7FF6FFC73FC1E13AF" box="[178,514,988,1010]" fontSize="8" level="2" pageId="10" pageNumber="111" reason="2">
<emphasis id="AC8CEAC8927CFFF7FF6FFC73FC1E13AF" bold="true" box="[178,514,988,1010]" pageId="10" pageNumber="111">
<taxonomicName id="59F84D59927CFFF7FF6FFC73FFAC13AF" ID-CoL="8Y7B7" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" box="[178,432,988,1010]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="formosanus" status="sp. nov.">
<emphasis id="AC8CEAC8927CFFF7FF6FFC73FFAC13AF" bold="true" box="[178,432,988,1010]" italics="true" pageId="10" pageNumber="111">Orphnebius formosanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7BF57B3927CFFF7FE6BFC72FC1E13AF" box="[438,514,989,1010]" pageId="10" pageNumber="111" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D6E26551927CFFF7FDD5FC72FCD213AE" box="[520,718,989,1011]" pageId="10" pageNumber="111" type="description">
<paragraph id="9E4736DA927CFFF7FDD5FC72FCD213AE" blockId="10.[178,718,988,1011]" box="[520,718,989,1011]" pageId="10" pageNumber="111">
<heading id="C50F81B6927CFFF7FDD5FC72FCD213AE" box="[520,718,989,1011]" fontSize="8" level="2" pageId="10" pageNumber="111" reason="2">
(
<figureCitation id="06C32A5F927CFFF7FDD2FC72FC9C13AE" box="[527,640,989,1011]" captionStart="Figs 16-21" captionStartId="11.[178,218,1502,1524]" captionTargetBox="[178,1082,212,1476]" captionTargetId="figure-1@11.[178,1082,212,1493]" captionTargetPageId="11" captionText="Figs 16-21: Orphnebius formosanus nov.sp.: forebody (16); antenna (17); abdomen (18); median lobe of aedeagus in lateral view (19); median lobe of aedeagus in ventral view (20); paramere (21). Scale bars: 16-18: 0.5 mm; 19-21: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413948" httpUri="https://zenodo.org/record/5413948/files/figure.png" pageId="10" pageNumber="111">Figs 16-21</figureCitation>
,
<figureCitation id="06C32A5F927CFFF7FD57FC72FCDA13AE" box="[650,710,989,1011]" captionStart="Figs 22-27" captionStartId="13.[178,218,1371,1393]" captionTargetBox="[178,1071,395,1349]" captionTargetId="figure-104@13.[178,1077,395,1361]" captionTargetPageId="13" captionText="Figs 22-27: Orphnebius incrassatus nov.sp. (22-25) and O. formosanus nov.sp. (26-27): forebody (22); antenna (23); abdomen (24); spermatheca (25, 27); tergite VII (26). Scale bars: 22-24: 0.5 mm; 25-27: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413950" httpUri="https://zenodo.org/record/5413950/files/figure.png" pageId="10" pageNumber="111">26-27</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D6E26551927CFFF7FF6FFBADFC131444" box="[178,527,1026,1049]" pageId="10" pageNumber="111" type="reference_group">
<paragraph id="9E4736DA927CFFF7FF6FFBADFC131444" blockId="10.[178,1086,1026,1278]" box="[178,527,1026,1049]" pageId="10" pageNumber="111">
<treatmentCitationGroup id="BEE811F4927CFFF7FF6FFBADFC131444" box="[178,527,1026,1049]" pageId="10" pageNumber="111">
<treatmentCitation id="1F5910CB927CFFF7FF6FFBADFC171444" author="PACE R" box="[178,523,1026,1049]" page=")" pageId="10" pageNumber="111" year="2010">
<taxonomicName id="59F84D59927CFFF7FF6FFBADFC171444" ID-CoL="8Y7BL" authority=": PACE (2010: 23)" authorityName="PACE" authorityPageNumber="23" authorityYear="2010" box="[178,523,1026,1049]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="AC8CEAC8927CFFF7FF6FFBADFF7B144A" box="[178,359,1026,1047]" italics="true" pageId="10" pageNumber="111">Orphnebius hauseri</emphasis>
:
<bibRefCitation id="FA694B2B927CFFF7FEAEFBACFC171444" author="PACE R" box="[371,523,1027,1049]" pageId="10" pageNumber="111" pagination="19 - 54" refId="ref13502" refString="PACE R. (2010): Thamiaraeini, Lomechusini, Oxypodini, Hoplandriini e Aleocharini di Taiwan (Coleoptera, Staphylinidae). - Bollettino del Museo Civico di Storia Naturale di Verona 34: 19 - 54." type="journal article" year="2010">PACE (2010: 23)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D6E26551927CFFF7FF6FFBB3FA1014A3" pageId="10" pageNumber="111" type="materials_examined">
<paragraph id="9E4736DA927CFFF7FF6FFBB3FA1014A3" blockId="10.[178,1086,1026,1278]" pageId="10" pageNumber="111">
Type material:
<materialsCitation id="2E903C87927CFFF7FE77FBB3FF501415" ID-GBIF-Occurrence="4415992301" collectingDate="1998-05-04" collectionCode="cAss" collectorName="A. Smetana" country="Taiwan" determinerName="V. Assing" elevation="2130" location="Meifeng" municipality="Meifeng" pageId="10" pageNumber="111" specimenCount="1" specimenCount-male="1" stateProvince="Nantou Hsien" typeStatus="holotype">
<typeStatus id="41438878927CFFF7FE77FBB3FFE2146F" box="[426,510,1052,1074]" pageId="10" pageNumber="111" type="holotype">Holotype</typeStatus>
<specimenCount id="88FEFD53927CFFF7FDDAFBB3FC06146C" box="[519,538,1052,1073]" pageId="10" pageNumber="111" type="male">♂</specimenCount>
: &quot;
<collectingCountry id="E6EF764A927CFFF7FDEDFBB3FC64146F" box="[560,632,1052,1074]" name="Taiwan" pageId="10" pageNumber="111">Taiwan</collectingCountry>
,
<collectingRegion id="5C3CF838927CFFF7FD59FBB3FD1B146F" box="[644,775,1052,1074]" country="Taiwan" name="Nantou" pageId="10" pageNumber="111">Nantou Hsien</collectingRegion>
,
<collectingMunicipality id="7E23ACA0927CFFF7FCCEFBB3FD7F146F" box="[787,867,1052,1074]" country="Taiwan" name="Meifeng" pageId="10" pageNumber="111" stateProvince="Nantou">Meifeng</collectingMunicipality>
,
<quantity id="59009B3F927CFFF7FCB3FBB3FDA4146C" box="[878,952,1052,1073]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="10" pageNumber="111" unit="m" value="2130.0">
<elevation id="15D5D1E9927CFFF7FCB3FBB3FDA4146C" box="[878,952,1052,1073]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="10" pageNumber="111" unit="m" value="2130.0">2130 m</elevation>
</quantity>
,
<date id="EA46101A927CFFF7FC19FBB3FA07146C" box="[964,1051,1052,1073]" pageId="10" pageNumber="111" value="1998-05-04">
<collectingDate id="FA02E9F2927CFFF7FC19FBB3FA07146C" box="[964,1051,1052,1073]" pageId="10" pageNumber="111" value="1998-05-04">4.V.1998</collectingDate>
</date>
,
<collectorName id="330D530C927CFFF7FBFAFBB3FF1C141A" pageId="10" pageNumber="111">A. Smetana</collectorName>
[T197]
</materialsCitation>
/
<taxonomicName id="59F84D59927CFFF7FEB1FB9DFC7E1415" authority="Epp." authorityName="EPPELSHEIM" authorityYear="1895" box="[364,610,1074,1096]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="AC8CEAC8927CFFF7FEB1FB9DFC34141A" box="[364,552,1074,1095]" italics="true" pageId="10" pageNumber="111">Orphnebius hauseri</emphasis>
Epp.
</taxonomicName>
, det. R. Pace 2005 / Holotypus ♂
<taxonomicName id="59F84D59927CFFF7FC0FFB9DFF001403" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="formosanus" status="sp. nov.">
<emphasis id="AC8CEAC8927CFFF7FC0FFB9DFF001403" italics="true" pageId="10" pageNumber="111">Orphnebius formosanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7BF57B3927CFFF7FEFEFBE6FF4B1403" box="[291,343,1097,1118]" pageId="10" pageNumber="111" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="6B9BDA1E927CFFF7FE57FBE6FFFA1402" box="[394,486,1097,1119]" pageId="10" pageNumber="111">V. Assing</determinerName>
2013&quot; (
<collectionCode id="F8E9AE1F927CFFF7FDECFBE6FC7E1402" box="[561,610,1097,1119]" pageId="10" pageNumber="111">cAss</collectionCode>
).
<materialsCitation id="2E903C87927CFFF7FDAFFBE6FC4714D0" ID-GBIF-Occurrence="3766784338" collectingDate="1992-05-13" collectionCode="cSme" collectorName="A. Smetana" country="Taiwan" determinerName="R. Pace" elevation="2150" location="Taichung" pageId="10" pageNumber="111" specimenCount="1" specimenCount-male="1" stateProvince="Taichung" typeStatus="paratype">
<typeStatus id="41438878927CFFF7FDAFFBE6FCD61403" box="[626,714,1097,1118]" pageId="10" pageNumber="111" type="paratype">Paratypes</typeStatus>
[all dissected prior to present study and labelled &quot;
<taxonomicName id="59F84D59927CFFF7FED4FBCFFFED1428" authority="Epp." authorityName="EPPELSHEIM" authorityYear="1895" box="[265,497,1120,1141]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="AC8CEAC8927CFFF7FED4FBCFFFA21428" box="[265,446,1120,1141]" italics="true" pageId="10" pageNumber="111">Orphnebius hauseri</emphasis>
Epp.
</taxonomicName>
, det.
<determinerName id="6B9BDA1E927CFFF7FDFCFBCFFC7B1428" box="[545,615,1120,1141]" pageId="10" pageNumber="111">R. Pace</determinerName>
2005&quot;]:
<specimenCount id="88FEFD53927CFFF7FD6AFBCFFCC91428" box="[695,725,1120,1141]" pageId="10" pageNumber="111" type="male">1♂</specimenCount>
: &quot;
<collectingCountry id="E6EF764A927CFFF7FD34FBCFFD2D142B" box="[745,817,1120,1142]" name="Taiwan" pageId="10" pageNumber="111">Taiwan</collectingCountry>
,
<collectingRegion id="5C3CF838927CFFF7FCE7FBCFFD8C142B" box="[826,912,1120,1142]" country="Taiwan" name="Taichung" pageId="10" pageNumber="111">Taichung</collectingRegion>
Hsien, Anmashan,
<quantity id="59009B3F927CFFF7FF6FFBD9FEE614D1" box="[178,250,1142,1164]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="10" pageNumber="111" unit="m" value="2150.0">
<elevation id="15D5D1E9927CFFF7FF6FFBD9FEE614D1" box="[178,250,1142,1164]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="10" pageNumber="111" unit="m" value="2150.0">2150 m</elevation>
</quantity>
,
<date id="EA46101A927CFFF7FEDEFBD8FF5214D1" box="[259,334,1143,1164]" pageId="10" pageNumber="111" value="1992-05-13">
<collectingDate id="FA02E9F2927CFFF7FEDEFBD8FF5214D1" box="[259,334,1143,1164]" pageId="10" pageNumber="111" value="1992-05-13">13.V.92</collectingDate>
</date>
,
<collectorName id="330D530C927CFFF7FE85FBD8FFDE14D1" box="[344,450,1143,1164]" pageId="10" pageNumber="111">A. Smetana</collectorName>
[T129]&quot; (
<collectionCode id="F8E9AE1F927CFFF7FDC0FBD8FC4814D0" box="[541,596,1143,1165]" pageId="10" pageNumber="111">cSme</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2E903C87927CFFF7FDB8FBD8FC1014FE" ID-GBIF-Occurrence="3766784308" collectingDate="1992-05-12" collectionCode="cSme" collectorName="A. Smetana" country="Taiwan" elevation="2230" location="Anmashan" municipality="Hsien" pageId="10" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Taichung" typeStatus="paratype">
<specimenCount id="88FEFD53927CFFF7FDB8FBD8FC9F14CC" box="[613,643,1143,1169]" pageId="10" pageNumber="111" type="female">1♀</specimenCount>
: &quot;
<collectingCountry id="E6EF764A927CFFF7FD4AFBD9FCC314D1" box="[663,735,1142,1164]" name="Taiwan" pageId="10" pageNumber="111">Taiwan</collectingCountry>
,
<collectingRegion id="5C3CF838927CFFF7FD34FBD9FD2314D1" box="[745,831,1142,1164]" country="Taiwan" name="Taichung" pageId="10" pageNumber="111">Taichung</collectingRegion>
<collectingMunicipality id="7E23ACA0927CFFF7FC99FBD9FD6714D1" box="[836,891,1142,1164]" pageId="10" pageNumber="111">Hsien</collectingMunicipality>
,
<location id="9B276001927CFFF7FC58FBD9FDF514D1" LSID="urn:lsid:plazi:treatment:165187CC927CFFF0FF6FFC73FD4111DE:9B276001927CFFF7FC58FBD9FDF514D1" box="[901,1001,1142,1164]" country="Taiwan" municipality="Hsien" name="Anmashan" pageId="10" pageNumber="111" stateProvince="Taichung">Anmashan</location>
,
<quantity id="59009B3F927CFFF7FC2FFBD8FA2614D1" box="[1010,1082,1143,1164]" metricMagnitude="3" metricUnit="m" metricValue="2.23" pageId="10" pageNumber="111" unit="m" value="2230.0">
<elevation id="15D5D1E9927CFFF7FC2FFBD8FA2614D1" box="[1010,1082,1143,1164]" metricMagnitude="3" metricUnit="m" metricValue="2.23" pageId="10" pageNumber="111" unit="m" value="2230.0">2230 m</elevation>
</quantity>
,
<date id="EA46101A927CFFF7FF6FFB22FEE114FF" box="[178,253,1165,1186]" pageId="10" pageNumber="111" value="1992-05-12">
<collectingDate id="FA02E9F2927CFFF7FF6FFB22FEE114FF" box="[178,253,1165,1186]" pageId="10" pageNumber="111" value="1992-05-12">12.V.92</collectingDate>
</date>
,
<collectorName id="330D530C927CFFF7FEDAFB22FF6E14FF" box="[263,370,1165,1186]" pageId="10" pageNumber="111">A. Smetana</collectorName>
[T127]&quot; (
<collectionCode id="F8E9AE1F927CFFF7FE13FB22FC1914FE" box="[462,517,1165,1187]" pageId="10" pageNumber="111">cSme</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2E903C87927CFFF7FDC8FB22FC0014E7" ID-GBIF-Occurrence="3766784304" collectingDate="1993-07-04" collectionCode="cSme" collectorName="A. Smetana" country="Taiwan" elevation="2500" location="Peinantashan" municipality="Hsien" pageId="10" pageNumber="111" specimenCount="1" specimenCount-male="1" stateProvince="Kaohsiung" typeStatus="paratype">
<specimenCount id="88FEFD53927CFFF7FDC8FB22FC2814FF" box="[533,564,1165,1186]" pageId="10" pageNumber="111" type="male">1♂</specimenCount>
: &quot;
<collectingCountry id="E6EF764A927CFFF7FD95FB22FC8C14FE" box="[584,656,1165,1187]" name="Taiwan" pageId="10" pageNumber="111">Taiwan</collectingCountry>
,
<collectingRegion id="5C3CF838927CFFF7FD47FB22FCE014FE" box="[666,764,1165,1187]" country="Taiwan" name="Kaohsiung" pageId="10" pageNumber="111">Kaohsiung</collectingRegion>
<collectingMunicipality id="7E23ACA0927CFFF7FCDEFB22FD2614FE" box="[771,826,1165,1187]" pageId="10" pageNumber="111">Hsien</collectingMunicipality>
,
<location id="9B276001927CFFF7FC99FB22FDA014FE" LSID="urn:lsid:plazi:treatment:165187CC927CFFF0FF6FFC73FD4111DE:9B276001927CFFF7FC99FB22FDA014FE" box="[836,956,1165,1187]" country="Taiwan" municipality="Hsien" name="Peinantashan" pageId="10" pageNumber="111" stateProvince="Kaohsiung">Peinantashan</location>
trail,
<quantity id="59009B3F927CFFF7FC2FFB22FA2614FE" box="[1010,1082,1165,1187]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="111" unit="m" value="2500.0">
<elevation id="15D5D1E9927CFFF7FC2FFB22FA2614FE" box="[1010,1082,1165,1187]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="111" unit="m" value="2500.0">2500 m</elevation>
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,
<date id="EA46101A927CFFF7FF6FFB0BFF1C14E4" box="[178,256,1188,1209]" pageId="10" pageNumber="111" value="1993-07-04">
<collectingDate id="FA02E9F2927CFFF7FF6FFB0BFF1C14E4" box="[178,256,1188,1209]" pageId="10" pageNumber="111" value="1993-07-04">4.VII.93</collectingDate>
</date>
,
<collectorName id="330D530C927CFFF7FED3FB0BFF6014E4" box="[270,380,1188,1209]" pageId="10" pageNumber="111">A. Smetana</collectorName>
[T136]&quot; (
<collectionCode id="F8E9AE1F927CFFF7FE00FB0BFC0814E7" box="[477,532,1188,1210]" pageId="10" pageNumber="111">cSme</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2E903C87927CFFF7FDF5FB0BFD2C148C" ID-GBIF-Occurrence="3766784327" collectingDate="1993-07-20" collectionCode="cAss" collectorName="A. Smetana" country="Taiwan" county="Kaohsuing" elevation="2400" location="Kuanshan Trail" municipality="Hsien" pageId="10" pageNumber="111" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="88FEFD53927CFFF7FDF5FB0BFC5A14E4" box="[552,582,1188,1209]" pageId="10" pageNumber="111" type="male">1♂</specimenCount>
: &quot;
<collectingCountry id="E6EF764A927CFFF7FD80FB0BFCB914E7" box="[605,677,1188,1210]" name="Taiwan" pageId="10" pageNumber="111">Taiwan</collectingCountry>
,
<collectingCounty id="77264E56927CFFF7FD6FFB0BFD0814E7" box="[690,788,1188,1210]" pageId="10" pageNumber="111">Kaohsuing</collectingCounty>
[sic]
<collectingMunicipality id="7E23ACA0927CFFF7FC92FB0BFD9A14E7" box="[847,902,1188,1210]" pageId="10" pageNumber="111">Hsien</collectingMunicipality>
,
<location id="9B276001927CFFF7FC4EFB0BFA3F14E7" LSID="urn:lsid:plazi:treatment:165187CC927CFFF0FF6FFC73FD4111DE:9B276001927CFFF7FC4EFB0BFA3F14E7" box="[915,1059,1188,1210]" country="Taiwan" county="Kaohsuing" municipality="Hsien" name="Kuanshan Trail" pageId="10" pageNumber="111">Kuanshan Trail</location>
at
<location id="9B276001927CFFF7FF6FFB14FF40148C" LSID="urn:lsid:plazi:treatment:165187CC927CFFF0FF6FFC73FD4111DE:9B276001927CFFF7FF6FFB14FF40148C" box="[178,348,1211,1233]" country="Taiwan" county="Kaohsuing" municipality="Hsien" name="Kaunshanchi Riv." pageId="10" pageNumber="111">Kaunshanchi Riv.</location>
,
<quantity id="59009B3F927CFFF7FEB4FB14FFA8148D" box="[361,436,1211,1232]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="111" unit="m" value="2400.0">
<elevation id="15D5D1E9927CFFF7FEB4FB14FFA8148D" box="[361,436,1211,1232]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="111" unit="m" value="2400.0">2400 m</elevation>
</quantity>
,
<date id="EA46101A927CFFF7FE1CFB14FC07148D" box="[449,539,1211,1232]" pageId="10" pageNumber="111" value="1993-07-20">
<collectingDate id="FA02E9F2927CFFF7FE1CFB14FC07148D" box="[449,539,1211,1232]" pageId="10" pageNumber="111" value="1993-07-20">20.VII.93</collectingDate>
</date>
,
<collectorName id="330D530C927CFFF7FDF5FB14FC8A148D" box="[552,662,1211,1232]" pageId="10" pageNumber="111">A. Smetana</collectorName>
[T158]&quot; (
<collectionCode id="F8E9AE1F927CFFF7FD25FB14FD34148C" box="[760,808,1211,1233]" pageId="10" pageNumber="111">cAss</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2E903C87927CFFF7FCE0FB14FA1214BA" ID-GBIF-Occurrence="3766784322" collectingDate="1992-04-20" collectionCode="MHNG" collectorName="A. Smetana" country="Taiwan" elevation="2400" location="Kuanshan trail" pageId="10" pageNumber="111" specimenCount="2" specimenCount-female="2" stateProvince="Kaohsiung Hsien" typeStatus="paratype">
<specimenCount id="88FEFD53927CFFF7FCE0FB14FD721488" box="[829,878,1211,1237]" pageId="10" pageNumber="111" type="female">2♀♀</specimenCount>
: &quot;
<collectingCountry id="E6EF764A927CFFF7FC58FB14FDD1148C" box="[901,973,1211,1233]" name="Taiwan" pageId="10" pageNumber="111">Taiwan</collectingCountry>
,
<collectingRegion id="5C3CF838927CFFF7FC07FB14FEF514BA" pageId="10" pageNumber="111">Kaohsiung Hsien</collectingRegion>
,
<location id="9B276001927CFFF7FF2EFB7EFF6514BA" LSID="urn:lsid:plazi:treatment:165187CC927CFFF0FF6FFC73FD4111DE:9B276001927CFFF7FF2EFB7EFF6514BA" box="[243,377,1233,1255]" country="Taiwan" name="Kuanshan trail" pageId="10" pageNumber="111" stateProvince="Kaohsiung Hsien">Kuanshan trail</location>
at
<location id="9B276001927CFFF7FE48FB7EFC2414BA" LSID="urn:lsid:plazi:treatment:165187CC927CFFF0FF6FFC73FD4111DE:9B276001927CFFF7FE48FB7EFC2414BA" box="[405,568,1233,1255]" country="Taiwan" county="Hsien" municipality="Kuanshan" name="Kaunshanchi Riv." pageId="10" pageNumber="111" stateProvince="Kaohsiung Hsien">Kaunshanchi Riv.</location>
,
<quantity id="59009B3F927CFFF7FD99FB7EFC9614BA" box="[580,650,1233,1255]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="111" unit="m" value="2400.0">
<elevation id="15D5D1E9927CFFF7FD99FB7EFC9614BA" box="[580,650,1233,1255]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="111" unit="m" value="2400.0">2400 m</elevation>
</quantity>
<date id="EA46101A927CFFF7FD4DFB7EFCE614BB" box="[656,762,1233,1254]" pageId="10" pageNumber="111" value="1992-04-20">
<collectingDate id="FA02E9F2927CFFF7FD4DFB7EFCE614BB" box="[656,762,1233,1254]" pageId="10" pageNumber="111" value="1992-04-20">20.IV.1992</collectingDate>
</date>
,
<collectorName id="330D530C927CFFF7FCD9FB7EFD7214BB" box="[772,878,1233,1254]" pageId="10" pageNumber="111">A. Smetana</collectorName>
[T94]&quot; (
<collectionCode id="F8E9AE1F927CFFF7FC63FB7EFA1514BA" box="[958,1033,1233,1255]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="111" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2E903C87927CFFF7FBC5FB7EFA1514A3" ID-GBIF-Occurrence="3766784318" collectingDate="1992-05-01" collectionCode="cAss" collectorName="A. Smetana" country="Taiwan" elevation="1500" location="Peitawushan" pageId="10" pageNumber="111" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung Hsien" typeStatus="paratype">
<specimenCount id="88FEFD53927CFFF7FBC5FB7EFA2B14BB" box="[1048,1079,1233,1254]" pageId="10" pageNumber="111" type="male">1♂</specimenCount>
: &quot;
<collectingCountry id="E6EF764A927CFFF7FF67FB47FF1E14A3" box="[186,258,1256,1278]" name="Taiwan" pageId="10" pageNumber="111">Taiwan</collectingCountry>
,
<collectingRegion id="5C3CF838927CFFF7FED6FB47FF8614A3" box="[267,410,1256,1278]" country="Taiwan" name="Pingtung" pageId="10" pageNumber="111">Pingtung Hsien</collectingRegion>
,
<location id="9B276001927CFFF7FE7EFB47FC0A14A3" LSID="urn:lsid:plazi:treatment:165187CC927CFFF0FF6FFC73FD4111DE:9B276001927CFFF7FE7EFB47FC0A14A3" box="[419,534,1256,1278]" country="Taiwan" name="Peitawushan" pageId="10" pageNumber="111" stateProvince="Pingtung Hsien">Peitawushan</location>
trail at
<quantity id="59009B3F927CFFF7FD81FB47FCBF14A0" box="[604,675,1256,1278]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="111" unit="m" value="1500.0">
<elevation id="15D5D1E9927CFFF7FD81FB47FCBF14A0" box="[604,675,1256,1278]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="111" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="EA46101A927CFFF7FD71FB47FD1F14A0" box="[684,771,1256,1277]" pageId="10" pageNumber="111" value="1992-05-01">
<collectingDate id="FA02E9F2927CFFF7FD71FB47FD1F14A0" box="[684,771,1256,1277]" pageId="10" pageNumber="111" value="1992-05-01">1.V.1992</collectingDate>
</date>
,
<collectorName id="330D530C927CFFF7FCD1FB47FD6A14A0" box="[780,886,1256,1277]" pageId="10" pageNumber="111">A. Smetana</collectorName>
[T110]&quot; (
<collectionCode id="F8E9AE1F927CFFF7FC0CFB47FA1E14A3" box="[977,1026,1256,1278]" pageId="10" pageNumber="111">cAss</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D6E26551927CFFF7FF6FFAA3FF421562" pageId="10" pageNumber="111" type="etymology">
<paragraph id="9E4736DA927CFFF7FF6FFAA3FF421562" blockId="10.[178,1085,1292,1343]" pageId="10" pageNumber="111">
Etymology: The specific epithet (adjective) is derived from Formosa, the ancient name of
<collectingCountry id="E6EF764A927CFFF7FED7FA86FF451562" box="[266,345,1321,1343]" name="Taiwan" pageId="10" pageNumber="111">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D6E26551927CFFF1FF6FFAE0FFA7132A" lastPageId="12" lastPageNumber="113" pageId="10" pageNumber="111" type="description">
<paragraph id="9E4736DA927CFFF7FF6FFAE0FFEB1587" blockId="10.[178,1086,1358,1498]" pageId="10" pageNumber="111">
Description: Body length 4.6-5.0 mm; length of forebody
<quantity id="59009B3F927CFFF7FC6AFAE0FA261539" box="[951,1082,1359,1380]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="10" pageNumber="111" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9-2.1 mm</quantity>
. Coloration: forebody blackish; abdomen bright reddish, strongly contrasting with the forebody; legs with dark-brown to blackish-brown femora, brown tibiae, and reddish tarsi; antennae with antennomeres V-XI blackish and antennomeres I-IV of variable coloration (reddish to blackish).
</paragraph>
<paragraph id="9E4736DA927CFFF1FF6FFA46FDDB1176" blockId="10.[178,1085,1513,1593]" lastBlockId="12.[178,1085,219,299]" lastPageId="12" lastPageNumber="113" pageId="10" pageNumber="111">
Head (
<figureCitation id="06C32A5F927CFFF7FF21FA46FF5015A2" box="[252,332,1513,1535]" captionStart="Figs 16-21" captionStartId="11.[178,218,1502,1524]" captionTargetBox="[178,1082,212,1476]" captionTargetId="figure-1@11.[178,1082,212,1493]" captionTargetPageId="11" captionText="Figs 16-21: Orphnebius formosanus nov.sp.: forebody (16); antenna (17); abdomen (18); median lobe of aedeagus in lateral view (19); median lobe of aedeagus in ventral view (20); paramere (21). Scale bars: 16-18: 0.5 mm; 19-21: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413948" httpUri="https://zenodo.org/record/5413948/files/figure.png" pageId="10" pageNumber="111">Fig. 16</figureCitation>
) approximately 1.25 times as broad as long; punctation fine and of somewhat variable density, moderately dense to rather dense. Eyes large and bulging, distinctly longer than distance from posterior margin of eye to posterior constriction in dorsal view. Antenna (
<figureCitation id="06C32A5F927AFFF1FE44FF74FFFF10AC" box="[409,483,219,241]" captionStart="Figs 16-21" captionStartId="11.[178,218,1502,1524]" captionTargetBox="[178,1082,212,1476]" captionTargetId="figure-1@11.[178,1082,212,1493]" captionTargetPageId="11" captionText="Figs 16-21: Orphnebius formosanus nov.sp.: forebody (16); antenna (17); abdomen (18); median lobe of aedeagus in lateral view (19); median lobe of aedeagus in ventral view (20); paramere (21). Scale bars: 16-18: 0.5 mm; 19-21: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413948" httpUri="https://zenodo.org/record/5413948/files/figure.png" pageId="12" pageNumber="113">Fig. 17</figureCitation>
)
<quantity id="59009B3F927AFFF1FE2CFF74FC7B10AD" box="[497,615,219,240]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="12" pageNumber="113" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3-1.4 mm</quantity>
long, slightly asymmetric; antennomeres V-X increasingly transverse and of increasing width; X more than 1.5, but less than 2.0 times as broad as long; XI distinctly elongated, at least as long as VIII-X combined.
</paragraph>
<caption id="CA876652927DFFF6FF6FFA71FC26167C" ID-DOI="http://doi.org/10.5281/zenodo.5413948" ID-Zenodo-Dep="5413948" httpUri="https://zenodo.org/record/5413948/files/figure.png" pageId="11" pageNumber="112" startId="11.[178,218,1502,1524]" targetBox="[178,1082,212,1476]" targetPageId="11">
<paragraph id="9E4736DA927DFFF6FF6FFA71FC26167C" blockId="11.[178,1085,1502,1570]" pageId="11" pageNumber="112">
<emphasis id="AC8CEAC8927DFFF6FF6FFA71FF0415AE" bold="true" box="[178,280,1502,1524]" pageId="11" pageNumber="112">Figs 16-21</emphasis>
:
<taxonomicName id="59F84D59927DFFF6FEFBFA70FC1F15A9" authority="Assing, 2015" authorityName="Assing" authorityYear="2015" box="[294,515,1503,1524]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="formosanus" status="sp. nov.">
<emphasis id="AC8CEAC8927DFFF6FEFBFA70FC1F15A9" box="[294,515,1503,1524]" italics="true" pageId="11" pageNumber="112">Orphnebius formosanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7BF57B3927DFFF6FDD6FA70FC4C15A9" box="[523,592,1503,1524]" pageId="11" pageNumber="112" rank="species">nov.sp.</taxonomicNameLabel>
: forebody (
<emphasis id="AC8CEAC8927DFFF6FD61FA71FCCF15AE" bold="true" box="[700,723,1502,1523]" pageId="11" pageNumber="112">16</emphasis>
); antenna (
<emphasis id="AC8CEAC8927DFFF6FCE2FA71FD4A15AE" bold="true" box="[831,854,1502,1523]" pageId="11" pageNumber="112">17</emphasis>
); abdomen (
<emphasis id="AC8CEAC8927DFFF6FC13FA71FDF915AE" bold="true" box="[974,997,1502,1523]" pageId="11" pageNumber="112">18</emphasis>
); median lobe of aedeagus in lateral view (
<emphasis id="AC8CEAC8927DFFF6FE3FFA5AFFE51657" bold="true" box="[482,505,1525,1546]" pageId="11" pageNumber="112">19</emphasis>
); median lobe of aedeagus in ventral view (
<emphasis id="AC8CEAC8927DFFF6FC51FA5AFDBF1657" bold="true" box="[908,931,1525,1546]" pageId="11" pageNumber="112">20</emphasis>
); paramere (
<emphasis id="AC8CEAC8927DFFF6FBC4FA5AFA2C1657" bold="true" box="[1049,1072,1525,1546]" pageId="11" pageNumber="112">21</emphasis>
). Scale bars: 16-18: 0.5 mm; 19-21: 0.2 mm.
</paragraph>
</caption>
<paragraph id="9E4736DA927AFFF1FF6FFE94FA2011BF" blockId="12.[178,1085,315,483]" pageId="12" pageNumber="113">
Pronotum (
<figureCitation id="06C32A5F927AFFF1FEF4FE94FF6B110C" box="[297,375,315,337]" captionStart="Figs 16-21" captionStartId="11.[178,218,1502,1524]" captionTargetBox="[178,1082,212,1476]" captionTargetId="figure-1@11.[178,1082,212,1493]" captionTargetPageId="11" captionText="Figs 16-21: Orphnebius formosanus nov.sp.: forebody (16); antenna (17); abdomen (18); median lobe of aedeagus in lateral view (19); median lobe of aedeagus in ventral view (20); paramere (21). Scale bars: 16-18: 0.5 mm; 19-21: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413948" httpUri="https://zenodo.org/record/5413948/files/figure.png" pageId="12" pageNumber="113">Fig. 16</figureCitation>
) moderately transverse, 1.17-1.23 times as broad as long; weakly dilated posteriad at most; lateral margins weakly convex in dorsal view; posterior angles weakly marked, rounded; disc with very sparse, minute punctures with long greyish setae; microsculpture absent; lateral margins each with 4-5 setae (often broken off) of moderate length. Elytra (
<figureCitation id="06C32A5F927AFFF1FE6BFE1FFC1E119B" box="[438,514,432,454]" captionStart="Figs 16-21" captionStartId="11.[178,218,1502,1524]" captionTargetBox="[178,1082,212,1476]" captionTargetId="figure-1@11.[178,1082,212,1493]" captionTargetPageId="11" captionText="Figs 16-21: Orphnebius formosanus nov.sp.: forebody (16); antenna (17); abdomen (18); median lobe of aedeagus in lateral view (19); median lobe of aedeagus in ventral view (20); paramere (21). Scale bars: 16-18: 0.5 mm; 19-21: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413948" httpUri="https://zenodo.org/record/5413948/files/figure.png" pageId="12" pageNumber="113">Fig. 16</figureCitation>
) approximately 0.85 times as long as, and at posterior margin distinctly broader than pronotum; punctation fine and sparse. Hind wings present.
</paragraph>
<paragraph id="9E4736DA927AFFF1FF6FFE5DFC911220" blockId="12.[178,1086,498,637]" pageId="12" pageNumber="113">
Abdomen (
<figureCitation id="06C32A5F927AFFF1FEF1FE5DFF621255" box="[300,382,498,520]" captionStart="Figs 16-21" captionStartId="11.[178,218,1502,1524]" captionTargetBox="[178,1082,212,1476]" captionTargetId="figure-1@11.[178,1082,212,1493]" captionTargetPageId="11" captionText="Figs 16-21: Orphnebius formosanus nov.sp.: forebody (16); antenna (17); abdomen (18); median lobe of aedeagus in lateral view (19); median lobe of aedeagus in ventral view (20); paramere (21). Scale bars: 16-18: 0.5 mm; 19-21: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413948" httpUri="https://zenodo.org/record/5413948/files/figure.png" pageId="12" pageNumber="113">Fig. 17</figureCitation>
) wedge-shaped, widest at base; tergites III-VI with sharp and pronounced paratergites gradually decreasing in height; tergites III-VI impunctate except for some minute punctures at posterior margins; tergite VII (
<figureCitation id="06C32A5F927AFFF1FCF5FD82FD69121E" box="[808,885,557,579]" captionStart="Figs 22-27" captionStartId="13.[178,218,1371,1393]" captionTargetBox="[178,1071,395,1349]" captionTargetId="figure-104@13.[178,1077,395,1361]" captionTargetPageId="13" captionText="Figs 22-27: Orphnebius incrassatus nov.sp. (22-25) and O. formosanus nov.sp. (26-27): forebody (22); antenna (23); abdomen (24); spermatheca (25, 27); tergite VII (26). Scale bars: 22-24: 0.5 mm; 25-27: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413950" httpUri="https://zenodo.org/record/5413950/files/figure.png" pageId="12" pageNumber="113">Fig. 26</figureCitation>
) with rather coarse non-setiferous punctures in posterior half, but without striate sculpture; posterior margin of tergite VII with pronounced palisade fringe.
</paragraph>
<paragraph id="9E4736DA927AFFF1FF6FFD22FCEE1368" blockId="12.[178,1085,653,821]" pageId="12" pageNumber="113">
♂: tergite VIII posteriorly with two transverse rows of black setae, a submarginal row composed of 6 long setae and a marginal row composed of more numerous and shorter setae; sclerites of segments IX and X modified, with dense and moderately long pubescence; median lobe of aedeagus (
<figureCitation id="06C32A5F927AFFF1FD9AFD4BFCA512A6" box="[583,697,740,763]" captionStart="Figs 16-21" captionStartId="11.[178,218,1502,1524]" captionTargetBox="[178,1082,212,1476]" captionTargetId="figure-1@11.[178,1082,212,1493]" captionTargetPageId="11" captionText="Figs 16-21: Orphnebius formosanus nov.sp.: forebody (16); antenna (17); abdomen (18); median lobe of aedeagus in lateral view (19); median lobe of aedeagus in ventral view (20); paramere (21). Scale bars: 16-18: 0.5 mm; 19-21: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413948" httpUri="https://zenodo.org/record/5413948/files/figure.png" pageId="12" pageNumber="113">Figs 19-20</figureCitation>
) approximately
<quantity id="59009B3F927AFFF1FCBEFD4AFDAE12A7" box="[867,946,741,762]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="113" unit="mm" value="0.7">0.7 mm</quantity>
long; ventral process rather short; paramere (
<figureCitation id="06C32A5F927AFFF1FE2AFCADFC5D1345" box="[503,577,770,792]" captionStart="Figs 16-21" captionStartId="11.[178,218,1502,1524]" captionTargetBox="[178,1082,212,1476]" captionTargetId="figure-1@11.[178,1082,212,1493]" captionTargetPageId="11" captionText="Figs 16-21: Orphnebius formosanus nov.sp.: forebody (16); antenna (17); abdomen (18); median lobe of aedeagus in lateral view (19); median lobe of aedeagus in ventral view (20); paramere (21). Scale bars: 16-18: 0.5 mm; 19-21: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413948" httpUri="https://zenodo.org/record/5413948/files/figure.png" pageId="12" pageNumber="113">Fig. 21</figureCitation>
)
<quantity id="59009B3F927AFFF1FD8CFCADFCFC134A" box="[593,736,770,792]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="12" pageNumber="113" unit="mm" value="0.625" valueMax="0.65" valueMin="0.6">0.60-0.65 mm</quantity>
long; paramerite distinctly longer than condylite and with four long setae at base of velum.
</paragraph>
<paragraph id="9E4736DA927AFFF1FF6FFCE5FFA7132A" blockId="12.[178,1085,836,888]" pageId="12" pageNumber="113">
<specimenCount id="88FEFD53927AFFF1FF6FFCE5FEDB1302" box="[178,199,842,863]" pageId="12" pageNumber="113" type="female">♀</specimenCount>
: sclerites of segments IX and X with very long and dense dark pubescence; spermatheca as in
<figureCitation id="06C32A5F927AFFF1FEB1FCCDFFA7132A" box="[364,443,866,888]" captionStart="Figs 22-27" captionStartId="13.[178,218,1371,1393]" captionTargetBox="[178,1071,395,1349]" captionTargetId="figure-104@13.[178,1077,395,1361]" captionTargetPageId="13" captionText="Figs 22-27: Orphnebius incrassatus nov.sp. (22-25) and O. formosanus nov.sp. (26-27): forebody (22); antenna (23); abdomen (24); spermatheca (25, 27); tergite VII (26). Scale bars: 22-24: 0.5 mm; 25-27: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413950" httpUri="https://zenodo.org/record/5413950/files/figure.png" pageId="12" pageNumber="113">Fig. 27.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D6E26551927AFFF1FF6FFC28FF471666" pageId="12" pageNumber="113" type="discussion">
<paragraph id="9E4736DA927AFFF1FF6FFC28FCDD13A9" blockId="12.[178,1085,903,1013]" pageId="12" pageNumber="113">
Comment: This species was erroneously recorded from
<collectingCountry id="E6EF764A927AFFF1FCE7FC28FD9A13C0" box="[826,902,903,925]" name="Taiwan" pageId="12" pageNumber="113">Taiwan</collectingCountry>
as
<taxonomicName id="59F84D59927AFFF1FC70FC28FA0713C1" authorityName="EPPELSHEIM" authorityYear="1895" box="[941,1051,903,924]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="AC8CEAC8927AFFF1FC70FC28FA0713C1" box="[941,1051,903,924]" italics="true" pageId="12" pageNumber="113">O. hauseri</emphasis>
</taxonomicName>
by
<bibRefCitation id="FA694B2B927AFFF1FF6FFC0AFF2713E7" author="PACE R" box="[178,315,932,954]" pageId="12" pageNumber="113" pagination="19 - 54" refId="ref13502" refString="PACE R. (2010): Thamiaraeini, Lomechusini, Oxypodini, Hoplandriini e Aleocharini di Taiwan (Coleoptera, Staphylinidae). - Bollettino del Museo Civico di Storia Naturale di Verona 34: 19 - 54." type="journal article" year="2010">PACE (2010)</bibRefCitation>
. The distribution of
<taxonomicName id="59F84D59927AFFF1FDF9FC0AFC8A13E4" authorityName="EPPELSHEIM" authorityYear="1895" box="[548,662,932,954]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="AC8CEAC8927AFFF1FDF9FC0AFC8A13E4" box="[548,662,932,954]" italics="true" pageId="12" pageNumber="113">O. hauseri</emphasis>
</taxonomicName>
is confined to North
<collectingCountry id="E6EF764A927AFFF1FC53FC0BFDDE13E7" box="[910,962,932,954]" name="India" pageId="12" pageNumber="113">India</collectingCountry>
and
<collectingCountry id="E6EF764A927AFFF1FBDCFC0BFA2113E7" box="[1025,1085,932,954]" name="Nepal" pageId="12" pageNumber="113">Nepal</collectingCountry>
(
<bibRefCitation id="FA694B2B927AFFF1FF64FC6DFF4D138A" author="ASSING V" box="[185,337,961,983]" pageId="12" pageNumber="113" pagination="1 - 26" refId="ref13078" refString="ASSING V. (2006 a): A revision of the Palaearctic species of Orphnebius MOTSCHULSKY (Insecta: Coleoptera: Staphylinidae: Aleocharinae). - Entomological Problems 36 (2): 1 - 26." type="journal article" year="2006">ASSING 2006a</bibRefCitation>
). The records from mainland
<collectingCountry id="E6EF764A927AFFF1FD54FC6EFCD9138A" box="[649,709,961,983]" name="China" pageId="12" pageNumber="113">China</collectingCountry>
(
<collectingRegion id="5C3CF838927AFFF1FD08FC6EFD36138A" box="[725,810,961,983]" country="China" name="Sichuan" pageId="12" pageNumber="113">Sichuan</collectingRegion>
,
<collectingRegion id="5C3CF838927AFFF1FCEAFC6DFD95138A" box="[823,905,962,983]" country="China" name="Yunnan" pageId="12" pageNumber="113">Yunnan</collectingRegion>
) by
<bibRefCitation id="FA694B2B927AFFF1FC66FC6DFA211385" author="PACE R" box="[955,1085,962,984]" pageId="12" pageNumber="113" pagination="77 - 102" refId="ref13542" refString="PACE R. (2012): Biodiversita delle Aleocharinae della Cina: Lomechusini e Thamiaraeini (Coleoptera, Staphylinidae). - Beitrage zur Entomologie, Keltern 62 (1): 77 - 102." type="journal article" year="2012">PACE (2012)</bibRefCitation>
are most likely based on misidentified material, too.
</paragraph>
<paragraph id="9E4736DA927AFFF1FF6FFBABFDAF14F1" blockId="12.[178,1085,1028,1196]" pageId="12" pageNumber="113">The female from Anmashan is distinguished from the remaining material by extremely sparse punctation of the head and by a distinctly shorter antennomere XI. A clarification of whether these differences are an expression intra- or of interspecific variation is currently not possible. The paramere of the male from Peinantawushan has the apex of the condylite nearly reaching that of the paramerite. Since no additional distinguishing characters were found, this difference is attributed to intraspecific variation.</paragraph>
<paragraph id="9E4736DA927AFFF1FF6FFB13FF471666" blockId="12.[178,1085,1212,1595]" pageId="12" pageNumber="113">
Comparative notes: Based on the external (coloration, body proportions, punctation, chaetotaxy) and the sexual characters (morphology of the aedeagus and the spermatheca),
<taxonomicName id="59F84D59927AFFF1FE9AFB56FFC21553" authorityName="Assing" authorityYear="2015" box="[327,478,1273,1294]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="AC8CEAC8927AFFF1FE9AFB56FFC21553" box="[327,478,1273,1294]" italics="true" pageId="12" pageNumber="113">O. formosanus</emphasis>
</taxonomicName>
undoubtedly belongs to the
<taxonomicName id="59F84D59927AFFF1FCDAFB56FD681550" authorityName="EPPELSHEIM" authorityYear="1895" box="[775,884,1272,1294]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="AC8CEAC8927AFFF1FCDAFB56FD681550" box="[775,884,1272,1294]" italics="true" pageId="12" pageNumber="113">O. hauseri</emphasis>
</taxonomicName>
group (see
<bibRefCitation id="FA694B2B927AFFF1FC2CFB56FEEE1570" author="ASSING V" pageId="12" pageNumber="113" pagination="1 - 26" refId="ref13078" refString="ASSING V. (2006 a): A revision of the Palaearctic species of Orphnebius MOTSCHULSKY (Insecta: Coleoptera: Staphylinidae: Aleocharinae). - Entomological Problems 36 (2): 1 - 26." type="journal article" year="2006">ASSING 2006a</bibRefCitation>
,
<bibRefCitation id="FA694B2B927AFFF1FF20FAB8FF5D1570" author="ASSING V" box="[253,321,1303,1325]" pageId="12" pageNumber="113" pagination="75 - 84" refId="ref13117" refString="ASSING V. (2006 b): On the Orphnebius species of China (Insecta: Coleoptera: Staphylinidae: Aleocharinae). - Entomological Problems 36 (2): 75 - 84." type="journal article" year="2006">2006b</bibRefCitation>
). The male sexual characters are most similar to those of
<taxonomicName id="59F84D59927AFFF1FC55FAB8FA191571" authorityName="ASSING" authorityYear="2006" box="[904,1029,1303,1324]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="AC8CEAC8927AFFF1FC55FAB8FA191571" box="[904,1029,1303,1324]" italics="true" pageId="12" pageNumber="113">O. mutabilis</emphasis>
</taxonomicName>
from
<collectingCountry id="E6EF764A927AFFF1FF6FFA9AFEED1516" box="[178,241,1333,1355]" name="Nepal" pageId="12" pageNumber="113">Nepal</collectingCountry>
, from which
<taxonomicName id="59F84D59927AFFF1FE5DFA9AFC041517" authorityName="Assing" authorityYear="2015" box="[384,536,1333,1354]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="AC8CEAC8927AFFF1FE5DFA9AFC041517" box="[384,536,1333,1354]" italics="true" pageId="12" pageNumber="113">O. formosanus</emphasis>
</taxonomicName>
is distinguished by the much more elongated apical antennomere (
<taxonomicName id="59F84D59927AFFF1FE97FAFCFFD21535" authorityName="ASSING" authorityYear="2006" box="[330,462,1363,1384]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="AC8CEAC8927AFFF1FE97FAFCFFD21535" box="[330,462,1363,1384]" italics="true" pageId="12" pageNumber="113">O. mutabilis</emphasis>
</taxonomicName>
: only approximately as long as antennomeres IX and X combined), longer antennae (
<taxonomicName id="59F84D59927AFFF1FE3EFADEFC7F15DB" authorityName="ASSING" authorityYear="2006" box="[483,611,1393,1414]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="AC8CEAC8927AFFF1FE3EFADEFC7F15DB" box="[483,611,1393,1414]" italics="true" pageId="12" pageNumber="113">O. mutabilis</emphasis>
</taxonomicName>
:
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), more transverse antennomeres VI-X, the denser punctation and pubescence of the head, the less sparse punctation of the elytra, the less extensive and sparser punctation of the abdominal tergite VII, the different shape of the crista apicalis of the median lobe of the aedeagus, the distinctly larger and differently shaped paramere (
<taxonomicName id="59F84D59927AFFF1FD93FA45FCD015A3" authorityName="ASSING" authorityYear="2006" box="[590,716,1513,1535]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="AC8CEAC8927AFFF1FD93FA45FCD015A3" box="[590,716,1513,1535]" italics="true" pageId="12" pageNumber="113">O. mutabilis</emphasis>
</taxonomicName>
: approximately
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long), and the slightly different shape of the spermatheca. For illustrations of
<taxonomicName id="59F84D59927AFFF1FC53F9A7FA0D1641" authorityName="ASSING" authorityYear="2006" box="[910,1041,1543,1565]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="mutabilis">
<emphasis id="AC8CEAC8927AFFF1FC53F9A7FA0D1641" box="[910,1041,1543,1565]" italics="true" pageId="12" pageNumber="113">O. mutabilis</emphasis>
</taxonomicName>
see
<bibRefCitation id="FA694B2B927AFFF1FF6FF989FF4A1666" author="ASSING V" box="[178,342,1573,1595]" pageId="12" pageNumber="113" pagination="1 - 26" refId="ref13078" refString="ASSING V. (2006 a): A revision of the Palaearctic species of Orphnebius MOTSCHULSKY (Insecta: Coleoptera: Staphylinidae: Aleocharinae). - Entomological Problems 36 (2): 1 - 26." type="journal article" year="2006">ASSING (2006a)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D6E26551927BFFF0FF6FFF74FD4111DE" pageId="13" pageNumber="114" type="distribution">
<paragraph id="9E4736DA927BFFF0FF6FFF74FD4111DE" blockId="13.[178,1086,219,387]" pageId="13" pageNumber="114">
Distribution and natural history: The species is known only from
<collectingCountry id="E6EF764A927BFFF0FF30FF57FF271153" box="[237,315,248,270]" name="Taiwan" pageId="13" pageNumber="114">Taiwan</collectingCountry>
, where it was found in several localities in
<collectingRegion id="5C3CF838927BFFF0FCCCFF57FD421150" box="[785,862,248,269]" country="Taiwan" name="Nantou" pageId="13" pageNumber="114">Nantou</collectingRegion>
,
<collectingRegion id="5C3CF838927BFFF0FCB6FF57FDD11153" box="[875,973,248,270]" country="Taiwan" name="Taichung" pageId="13" pageNumber="114">Taichung</collectingRegion>
,
<collectingRegion id="5C3CF838927BFFF0FC06FF57FA251153" box="[987,1081,248,270]" country="Taiwan" name="Pingtung" pageId="13" pageNumber="114">Pingtung</collectingRegion>
, and
<collectingRegion id="5C3CF838927BFFF0FF02FEBAFF511176" box="[223,333,277,299]" country="Taiwan" name="Kaohsiung" pageId="13" pageNumber="114">Kaohsiung</collectingRegion>
provinces and where it is currently the sole representative of the genus. The specimens were sifted from wet moss and debris along a river, from leaf litter under broadleaved bushes, and from leaf litter in primary broadleaved evergreen and in primary mixed forests at altitudes of
<quantity id="59009B3F927BFFF0FE0CFEC2FC4411DE" box="[465,600,365,387]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="13" pageNumber="114" unit="m" value="2000.0" valueMax="2500.0" valueMin="1500.0">1500-2500 m</quantity>
(SMETANA pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
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