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<document id="591FA73E2AAF713D0DEB79718E86C3E5" ID-DOI="10.11646/zootaxa.2952.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5281673" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629734144550" checkinUser="felipe" docAuthor="Nielson, M. W." docDate="2011" docId="E70787A4FF9DE176FF5DE25ECE648F73" docLanguage="en" docName="zt02952p086.pdf" docOrigin="Zootaxa 2952 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2952.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Docalidia acutula Nielson 2011, sp. nov." docType="treatment" docVersion="5" lastPageNumber="17" masterDocId="1B3EFFDCFF8DE166FFCAE141CF7F8839" masterDocTitle="New species in the Neotropical genus Docalidia with a key to known species, notes on distribution, taxonomy and a synoptic catalogue of the genus (Hemiptera: Cicadellidae: Coelidiinae: Teruliini) 2952" masterLastPageNumber="86" masterPageNumber="1" pageNumber="17" updateTime="1699103888837" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="17EADC9CE3DBA669E8D9D236C3A8624E">New species in the Neotropical genus Docalidia with a key to known species, notes on distribution, taxonomy and a synoptic catalogue of the genus (Hemiptera: Cicadellidae: Coelidiinae: Teruliini) 2952</mods:title>
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<mods:namePart id="D9A708D3DE205B4AF272C64F65E35061">Nielson, M. W.</mods:namePart>
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<mods:date id="D52106291E718375E24CF4F109ABA3FC">2011</mods:date>
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<treatment id="E70787A4FF9DE176FF5DE25ECE648F73" ID-DOI="http://doi.org/10.5281/zenodo.5286310" ID-Zenodo-Dep="5286310" LSID="urn:lsid:plazi:treatment:E70787A4FF9DE176FF5DE25ECE648F73" httpUri="http://treatment.plazi.org/id/E70787A4FF9DE176FF5DE25ECE648F73" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="27B46539FF9DE176FF5DE25ECEC78B63" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="6F1136B2FF9DE176FF5DE25ECEA48B00" blockId="16.[151,475,799,858]" box="[151,475,799,825]" pageId="16" pageNumber="17">
<heading id="345981DEFF9DE176FF5DE25ECEA48B00" bold="true" box="[151,475,799,825]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="5DDAEAA0FF9DE176FF5DE25ECEA48B00" bold="true" box="[151,475,799,825]" pageId="16" pageNumber="17">
<emphasis id="5DDAEAA0FF9DE176FF5DE25ECE0A8B00" bold="true" box="[151,373,799,825]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="A8AE4D31FF9DE176FF5DE25ECE108B00" ID-CoL="8JRRR" authority="Nielson, 2011" authorityName="Nielson" authorityYear="2011" box="[151,367,799,825]" class="Insecta" family="Cicadellidae" genus="Docalidia" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="acutula" status="sp. nov.">Docalidia acutula</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="46E957DBFF9DE176FEB7E25ECEA48B00" box="[381,475,799,825]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6F1136B2FF9DE176FF5DE203CEC78B63" blockId="16.[151,475,799,858]" box="[151,440,834,858]" pageId="16" pageNumber="17">
(Plate 1B,
<figureCitation id="F7952A37FF9DE176FEC4E203CECF8B63" box="[270,432,834,858]" captionStart="FIGURES 1017" captionStartId="17.[151,264,1918,1941]" captionTargetBox="[152,1408,204,1878]" captionTargetId="figure-66@17.[151,1436,65,1955]" captionTargetPageId="17" captionText="FIGURES 1017. Docalidia acutula, sp. nov. (10) pygofer, lateral view; (11) right subgenital plate, ventral view; (12) right style, lateral view (Fig. 13); right style, dorsal view (14); aedeagus and dorsal connective, lateral view (15); aedeagus, ventral view (16); connective, caudal view (Fig. 17); dorsal connective (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5281677" httpUri="https://zenodo.org/record/5281677/files/figure.png" pageId="16" pageNumber="17">Figures 1017</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="27B46539FF9DE176FF5DE2CACB1B8DDB" pageId="16" pageNumber="17" type="description">
<paragraph id="6F1136B2FF9DE176FF5DE2CACD098B9B" blockId="16.[151,1436,906,1866]" box="[151,630,906,931]" pageId="16" pageNumber="17">
<emphasis id="5DDAEAA0FF9DE176FF5DE2CACF8C8B9A" bold="true" box="[151,243,907,931]" inLineHeading="true" pageId="16" pageNumber="17" reason="1">Length.</emphasis>
Male 6.80 mm., female unknown.
</paragraph>
<paragraph id="6F1136B2FF9DE176FF0DE2EECA2D8C36" blockId="16.[151,1436,906,1866]" pageId="16" pageNumber="17">
<emphasis id="5DDAEAA0FF9DE176FF0DE2EECEB28BFE" bold="true" box="[199,461,943,967]" inLineHeading="true" pageId="16" pageNumber="17" reason="1">External morphology.</emphasis>
Moderately large, robust species. General color of dorsum concolorous yellow except for black anterior half of mesonotum; forewings with veins of clavus ivory; eyes translucent (Plate 1B); face yellow; clypeus with row of short, black, transverse stripes on lateral margins, large black spot below each eye.
</paragraph>
<paragraph id="6F1136B2FF9DE176FF0DE55BCE4A8CDF" blockId="16.[151,1436,906,1866]" pageId="16" pageNumber="17">Head distinctly narrower than pronotum, anterior margin broadly rounded; crown broad, about as wide as eye, produced distally about ¼ of entire length, disk foveate of each side of middle; eyes large, semiglobular; pronotum slightly shorter than crown, surface rugulose; mesonotum large, about twice as long as pronotum; forewings long, very broad, venation typical of genus; clypeus long, broad, lateral margins broadly convex, median longitudinal carina very prominent; clypellus about 1/3 as long as clypeus, narrow, lateral margins expanded apically, median ridge inflated.</paragraph>
<paragraph id="6F1136B2FF9DE176FF0DE5B2CB1B8DDB" blockId="16.[151,1436,906,1866]" pageId="16" pageNumber="17">
<emphasis id="5DDAEAA0FF9DE176FF0DE5B2CE038D32" bold="true" box="[199,380,1266,1291]" inLineHeading="true" pageId="16" pageNumber="17" reason="1">Male genitalia.</emphasis>
Pygofer in lateral view triangulate, with moderately long, stout caudodorsal process, caudoventral process absent (
<figureCitation id="F7952A37FF9DE176FE74E457CD6C8D17" box="[446,531,1302,1326]" captionStart="FIGURES 1017" captionStartId="17.[151,264,1918,1941]" captionTargetBox="[152,1408,204,1878]" captionTargetId="figure-66@17.[151,1436,65,1955]" captionTargetPageId="17" captionText="FIGURES 1017. Docalidia acutula, sp. nov. (10) pygofer, lateral view; (11) right subgenital plate, ventral view; (12) right style, lateral view (Fig. 13); right style, dorsal view (14); aedeagus and dorsal connective, lateral view (15); aedeagus, ventral view (16); connective, caudal view (Fig. 17); dorsal connective (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5281677" httpUri="https://zenodo.org/record/5281677/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
); right subgenital plate long, narrow, with few short setae in distal ¼ (
<figureCitation id="F7952A37FF9DE176FAF0E457CAF28D17" box="[1338,1421,1302,1326]" captionStart="FIGURES 1017" captionStartId="17.[151,264,1918,1941]" captionTargetBox="[152,1408,204,1878]" captionTargetId="figure-66@17.[151,1436,65,1955]" captionTargetPageId="17" captionText="FIGURES 1017. Docalidia acutula, sp. nov. (10) pygofer, lateral view; (11) right subgenital plate, ventral view; (12) right style, lateral view (Fig. 13); right style, dorsal view (14); aedeagus and dorsal connective, lateral view (15); aedeagus, ventral view (16); connective, caudal view (Fig. 17); dorsal connective (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5281677" httpUri="https://zenodo.org/record/5281677/files/figure.png" pageId="16" pageNumber="17">Fig. 11</figureCitation>
); right style long, shorter than aedeagus, numerous setae along margin of distal half and apically, dense spongy cover in distal 1/
<quantity id="A8569B57FF9DE176FEDBE41FCE408D4F" box="[273,319,1374,1398]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="16" pageNumber="17" unit="in" value="3.0">3 in</quantity>
lateral view (
<figureCitation id="F7952A37FF9DE176FE15E41FCD148D4F" box="[479,619,1374,1398]" captionStart="FIGURES 1017" captionStartId="17.[151,264,1918,1941]" captionTargetBox="[152,1408,204,1878]" captionTargetId="figure-66@17.[151,1436,65,1955]" captionTargetPageId="17" captionText="FIGURES 1017. Docalidia acutula, sp. nov. (10) pygofer, lateral view; (11) right subgenital plate, ventral view; (12) right style, lateral view (Fig. 13); right style, dorsal view (14); aedeagus and dorsal connective, lateral view (15); aedeagus, ventral view (16); connective, caudal view (Fig. 17); dorsal connective (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5281677" httpUri="https://zenodo.org/record/5281677/files/figure.png" pageId="16" pageNumber="17">Figs. 12, 13</figureCitation>
); aedeagus asymmetrical, in lateral view broadly sinuate, shaft tubular, preatrium exceptionally long, very long, needle-like subapical process closely appressed to shaft, almost hidden, directed basally, gonopore medial on ventral margin (
<figureCitation id="F7952A37FF9DE176FD36E4E7CCF78D87" box="[764,904,1446,1470]" captionStart="FIGURES 1017" captionStartId="17.[151,264,1918,1941]" captionTargetBox="[152,1408,204,1878]" captionTargetId="figure-66@17.[151,1436,65,1955]" captionTargetPageId="17" captionText="FIGURES 1017. Docalidia acutula, sp. nov. (10) pygofer, lateral view; (11) right subgenital plate, ventral view; (12) right style, lateral view (Fig. 13); right style, dorsal view (14); aedeagus and dorsal connective, lateral view (15); aedeagus, ventral view (16); connective, caudal view (Fig. 17); dorsal connective (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5281677" httpUri="https://zenodo.org/record/5281677/files/figure.png" pageId="16" pageNumber="17">Figs. 14, 15</figureCitation>
); connective T-shaped, arms stout, stem short, membrane absent (
<figureCitation id="F7952A37FF9DE176FEA1E48BCEC18DDB" box="[363,446,1482,1506]" captionStart="FIGURES 1017" captionStartId="17.[151,264,1918,1941]" captionTargetBox="[152,1408,204,1878]" captionTargetId="figure-66@17.[151,1436,65,1955]" captionTargetPageId="17" captionText="FIGURES 1017. Docalidia acutula, sp. nov. (10) pygofer, lateral view; (11) right subgenital plate, ventral view; (12) right style, lateral view (Fig. 13); right style, dorsal view (14); aedeagus and dorsal connective, lateral view (15); aedeagus, ventral view (16); connective, caudal view (Fig. 17); dorsal connective (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5281677" httpUri="https://zenodo.org/record/5281677/files/figure.png" pageId="16" pageNumber="17">Fig. 16</figureCitation>
); dorsal connective very narrow, expanded basally (
<figureCitation id="F7952A37FF9DE176FBCEE48BCB288DDB" box="[1028,1111,1482,1506]" captionStart="FIGURES 1017" captionStartId="17.[151,264,1918,1941]" captionTargetBox="[152,1408,204,1878]" captionTargetId="figure-66@17.[151,1436,65,1955]" captionTargetPageId="17" captionText="FIGURES 1017. Docalidia acutula, sp. nov. (10) pygofer, lateral view; (11) right subgenital plate, ventral view; (12) right style, lateral view (Fig. 13); right style, dorsal view (14); aedeagus and dorsal connective, lateral view (15); aedeagus, ventral view (16); connective, caudal view (Fig. 17); dorsal connective (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.5281677" httpUri="https://zenodo.org/record/5281677/files/figure.png" pageId="16" pageNumber="17">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="27B46539FF9DE176FF0DE4AFCBE18E13" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="6F1136B2FF9DE176FF0DE4AFCBE18E13" blockId="16.[151,1436,906,1866]" pageId="16" pageNumber="17">
<emphasis id="5DDAEAA0FF9DE176FF0DE4AFCEBE8E3E" bold="true" box="[199,449,1518,1543]" inLineHeading="true" pageId="16" pageNumber="17" reason="1">Material examined.</emphasis>
<materialsCitation id="DFC63CEFFF9DE176FE12E4AFCBE18E13" collectingDate="2004-05-29" collectingDateMax="2004-06-19" collectingDateMin="2004-05-29" collectorName="Malaise &amp; H. Vargas &amp; Leg., M." country="Colombia" elevation="710" latitude="3.8" location="Cabana Las Mirlas" longLatPrecision="1307" longitude="-73.86667" municipality="Sumapare" pageId="16" pageNumber="17" specimenCount="710" stateProvince="Meta" typeStatus="holotype">
<typeStatus id="B0158810FF9DE176FE12E4AFCD3F8E3F" box="[472,576,1518,1542]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="17B97622FF9DE176FD67E4AFCC388E3F" box="[685,839,1518,1542]" name="Colombia" pageId="16" pageNumber="17">COLOMBIA</collectingCountry>
:
<collectingRegion id="AD6AF850FF9DE176FCA8E4AECCE18E3F" box="[866,926,1519,1542]" country="Colombia" name="Meta" pageId="16" pageNumber="17">Meta</collectingRegion>
, PNN,
<collectingMunicipality id="8F75ACC8FF9DE176FBDBE4AFCBFB8E3F" box="[1041,1156,1518,1542]" pageId="16" pageNumber="17">Sumapare</collectingMunicipality>
,
<location id="6A716069FF9DE176FB55E4AFCAE88E3F" LSID="urn:lsid:plazi:treatment:E70787A4FF9DE176FF5DE25ECE648F73:6A716069FF9DE176FB55E4AFCAE88E3F" box="[1183,1431,1518,1542]" country="Colombia" latitude="3.8" longLatPrecision="1307" longitude="-73.86667" municipality="Sumapare" name="Cabana Las Mirlas" pageId="16" pageNumber="17" stateProvince="Meta">Cabaña Las Mirlas</location>
,
<geoCoordinate id="0A9A5075FF9DE176FF5DE753CF978E13" box="[151,232,1554,1578]" degrees="3" direction="north" minutes="48" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="3.8">3º48N</geoCoordinate>
.
<geoCoordinate id="0A9A5075FF9DE176FF21E753CE298E13" box="[235,342,1554,1578]" degrees="73" direction="west" minutes="52" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="-73.86667">73º52W</geoCoordinate>
.,
<specimenCount id="79A8FD3BFF9DE176FEAFE753CECA8E13" box="[357,437,1554,1578]" pageId="16" pageNumber="17" type="generic">
<quantity id="A8569B57FF9DE176FEAFE753CECE8E13" box="[357,433,1554,1578]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="16" pageNumber="17" unit="m" value="710.0">
<elevation id="E483D181FF9DE176FEAFE753CECE8E13" box="[357,433,1554,1578]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="16" pageNumber="17" unit="m" value="710.0">710 m</elevation>
</quantity>
.
</specimenCount>
,
<collectorName id="C25B5364FF9DE176FE0BE753CD638E13" box="[449,540,1554,1578]" pageId="16" pageNumber="17">Malaise</collectorName>
,
<date id="1B101072FF9DE176FDEDE753CDD78E13" box="[551,680,1554,1578]" pageId="16" pageNumber="17" value="2004-05-29">
<collectingDate id="0B54E99AFF9DE176FDEDE753CDD78E13" box="[551,680,1554,1578]" pageId="16" pageNumber="17" value="2004-05-29">29/05/2004</collectingDate>
</date>
<date id="1B101072FF9DE176FD7DE753CC498E13" box="[695,822,1554,1578]" pageId="16" pageNumber="17" value="2004-06-19">
<collectingDate id="0B54E99AFF9DE176FD7DE753CC498E13" box="[695,822,1554,1578]" pageId="16" pageNumber="17" value="2004-06-19">19/06/2004</collectingDate>
</date>
,
<collectorName id="C25B5364FF9DE176FC88E752CCCC8E12" box="[834,947,1555,1579]" pageId="16" pageNumber="17">H. Vargas</collectorName>
,
<collectorName id="C25B5364FF9DE176FC74E752CB638E13" box="[958,1052,1554,1579]" pageId="16" pageNumber="17">Leg., M.</collectorName>
4688 (HB).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="27B46539FF9DE176FF0DE776CFB28E4B" pageId="16" pageNumber="17" type="etymology">
<paragraph id="6F1136B2FF9DE176FF0DE776CFB28E4B" blockId="16.[151,1436,906,1866]" pageId="16" pageNumber="17">
<emphasis id="5DDAEAA0FF9DE176FF0DE776CE328E76" bold="true" box="[199,333,1591,1615]" inLineHeading="true" pageId="16" pageNumber="17" reason="1">Etymology.</emphasis>
The name is descriptive for the subtle, almost hidden, very fine, needle-like aedeagal ventral process.
</paragraph>
</subSubSection>
<subSubSection id="27B46539FF9DE176FF0DE73ECE648F73" pageId="16" pageNumber="17" type="discussion">
<paragraph id="6F1136B2FF9DE176FF0DE73ECE648F73" blockId="16.[151,1436,906,1866]" pageId="16" pageNumber="17">
<emphasis id="5DDAEAA0FF9DE176FF0DE73ECE448EAE" bold="true" box="[199,315,1663,1687]" inLineHeading="true" pageId="16" pageNumber="17" reason="1">Remarks.</emphasis>
From
<taxonomicName id="A8AE4D31FF9DE176FE4DE73ECD248EAF" authority="Nielson" authorityName="Nielson" box="[391,603,1662,1686]" class="Insecta" family="Cicadellidae" genus="Docalidia" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="oriellyi">
<emphasis id="5DDAEAA0FF9DE176FE4DE73ECEE48EAF" box="[391,411,1663,1686]" italics="true" pageId="16" pageNumber="17">D</emphasis>
.
<emphasis id="5DDAEAA0FF9DE176FE61E73FCE838EAF" box="[427,508,1662,1686]" italics="true" pageId="16" pageNumber="17">oriellyi</emphasis>
Nielson
</taxonomicName>
to which it is similar in preatrial and aedeagal configurations,
<taxonomicName id="A8AE4D31FF9DE176FAEEE73ECAE38EAF" authorityName="Nielson" authorityYear="2011" box="[1316,1436,1662,1686]" class="Insecta" family="Cicadellidae" genus="Docalidia" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="acutula">
<emphasis id="5DDAEAA0FF9DE176FAEEE73ECA478EAF" box="[1316,1336,1663,1686]" italics="true" pageId="16" pageNumber="17">D</emphasis>
.
<emphasis id="5DDAEAA0FF9DE176FA82E73FCAE38EAF" box="[1352,1436,1662,1686]" italics="true" pageId="16" pageNumber="17">acutula</emphasis>
</taxonomicName>
can be distinguished by the pygofer which lacks the basal flange on the caudodorsal process, absence of caudoventral process, dense spongy, marginal covering in the apical 1/3 of the style and the longer preatrium and ventral process of the aedeagus. The former species is known from
<collectingCountry id="17B97622FF9DE176FCD4E7ABCC0A8F3B" box="[798,885,1770,1794]" name="Guyana" pageId="16" pageNumber="17">Guyana</collectingCountry>
and
<collectingCountry id="17B97622FF9DE176FC64E7ABCB728F3B" box="[942,1037,1770,1794]" name="Ecuador" pageId="16" pageNumber="17">Ecuador</collectingCountry>
, the latter locality represented by a male specimen depicting variation in genitalia features. (For comparison, see
<bibRefCitation id="0B3F4B43FF9DE176FB8FE64FCB928F1F" author="Nielson" box="[1093,1261,1806,1830]" firstAuthor="Nielson" pageId="16" pageNumber="17" pagination="1 - 329" refId="ref55314" refString="Nielson, M. W. (1979 b) A revision of the subfamily Coelidiinae (Homoptera: Cicadadellidae). III. Tribe Teruliini. Pacific Insects - Monograph, 35, 1 - 329." type="journal article" year="1979">Nielson, 1979</bibRefCitation>
e:247;
<bibRefCitation id="0B3F4B43FF9DE176FAF4E64FCE708F73" author="Nielson" firstAuthor="Nielson" page="303" pageId="16" pageNumber="17" pagination="297 - 305" refId="ref55534" refString="Nielson, M. W. (1992 f) New species of Neotropical leafhoppers in the tribe Teruliini with a revised key to some species (Homoptera: Cicadellidae: Coelidiinae). Entomologica Scandinavica, 23 (3), 297 - 305." type="journal article" year="1992" yearSuffix="f">Nielson, 1992f:303</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>