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<document id="B8B954CDA73FF8B82850EBD3D33BC395" ID-CLB-Dataset="25602" ID-DOI="10.11646/zootaxa.4664.4.4" ID-GBIF-Dataset="af08637c-3c84-42d2-9b58-a3c88c5b6e28" ID-ISSN="1175-5326" ID-Zenodo-Dep="3386939" ID-ZooBank="8DEC3A3C-193C-4DC4-A968-590F08ED3CD1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1567672545701" checkinUser="plazi" docAuthor="Bezděk, Jan &amp; Lee, Chi-Feng" docDate="2019" docId="D61087F051693232FF1EE8762C72FD89" docLanguage="en" docName="zootaxa.4664.4.4.pdf" docOrigin="Zootaxa 4664 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pyrrhalta schillhammeri Bezděk &amp; Lee 2019, sp. nov." docType="treatment" docUuid="9CB87676-96C3-4CAC-90D7-3622C565D86F" docUuidSource="ZooBank" docVersion="9" lastPageNumber="531" masterDocId="2A29FF885165323FFF89EA61280BFFEA" masterDocTitle="Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra" masterLastPageNumber="534" masterPageNumber="518" pageNumber="530" updateTime="1698754682771" updateUser="ExternalLinkService">
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<mods:title id="00660ADC24643E15F1DE088D1558B393">Revision of Pyrrhalta (Coleoptera: Chrysomelidae: Galerucinae) species with maculate elytra</mods:title>
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<mods:namePart id="8863B08C1D4EE4351D7D642BE9132395">Bezděk, Jan</mods:namePart>
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<mods:namePart id="273C31FDCD26F36C1877A26ED3072D59">Lee, Chi-Feng</mods:namePart>
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<mods:title id="3D41DF726ED23DADF82D24115FFBA83F">Zootaxa</mods:title>
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<mods:date id="A24788CB59977185C307EA72DE9E7D46">2019</mods:date>
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<mods:number id="EE9F97D9AB4656D7EF07783BE2D15F8F">2019-09-05</mods:number>
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<treatment id="D61087F051693232FF1EE8762C72FD89" ID-GBIF-Taxon="159611474" LSID="urn:lsid:zoobank.org:act:9CB87676-96C3-4CAC-90D7-3622C565D86F" httpUri="http://treatment.plazi.org/id/D61087F051693232FF1EE8762C72FD89" lastPageId="13" lastPageNumber="531" pageId="12" pageNumber="530">
<subSubSection id="16A3656D51693233FF1EE8762A2DFDD8" box="[151,550,535,562]" pageId="12" pageNumber="530" type="nomenclature">
<paragraph id="5E0636E651693233FF1EE8762A2DFDD8" blockId="12.[151,550,535,598]" box="[151,550,535,562]" pageId="12" pageNumber="530">
<heading id="054E818A51693233FF1EE8762A2DFDD8" bold="true" box="[151,550,535,562]" fontSize="11" level="1" pageId="12" pageNumber="530" reason="1">
<emphasis id="6CCDEAF451693233FF1EE8762A2DFDD8" bold="true" box="[151,550,535,562]" pageId="12" pageNumber="530">
<taxonomicName id="99B94D6551693233FF1EE87629CBFDD8" authority="Bezděk &amp; Lee, 2019" authorityName="Bezděk &amp; Lee" authorityYear="2019" box="[151,448,535,562]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="530" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
<emphasis id="6CCDEAF451693233FF1EE87629CBFDD8" bold="true" box="[151,448,535,562]" italics="true" pageId="12" pageNumber="530">Pyrrhalta schillhammeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77FE578F51693233FE41E8792A2DFDD8" box="[456,550,536,562]" pageId="12" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="16A3656D51693233FF1EE85D299DFDBC" box="[151,406,571,598]" pageId="12" pageNumber="530" type="description">
<paragraph id="5E0636E651693233FF1EE85D299DFDBC" blockId="12.[151,550,535,598]" box="[151,406,571,598]" pageId="12" pageNumber="530">
(
<figureCitation id="C6822A6351693233FF16E85D28E3FDBF" box="[159,232,571,598]" captionStart="FIGURES 17" captionStartId="2.[151,264,1865,1890]" captionTargetBox="[175,1421,255,1817]" captionTargetId="figure@2.[164,1423,234,1837]" captionTargetPageId="2" captionText="FIGURES 17. Aedeagus. 1—Pyrrhalta maculata (basal half in dorsal view, apical half in dorsal view, lateral view); 2—P. carolusi sp. nov. (dorsal and lateral views); 3—P. kaboureki sp. nov. (dorsal, lateral and ventral views); 4—P. kambaitiensis sp. nov. (basal part in dorsal view, apex in dorsal view, lateral view); 5—P. lucka sp. nov. (dorsal and lateral views); 6—P. schillhammeri sp. nov. (dorsal and lateral views); 7—P. tsoui sp. nov. (basal part in dorsal view, apex in dorsal view, lateral view). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/3386941/files/figure.png" pageId="12" pageNumber="530">Figs 6</figureCitation>
,
<figureCitation id="C6822A6351693233FF7DE85D296EFDBC" box="[244,357,571,598]" captionStart="FIGURES 821" captionStartId="3.[151,264,1668,1693]" captionTargetBox="[188,1421,199,1621]" captionTargetId="figure@3.[151,1436,181,1643]" captionTargetPageId="3" captionText="FIGURES 821. Details of Pyrrhalta males. 8—P. kaboureki (last abdominal ventrite); 9—P. lucka sp. nov. (last abdominal ventrite); 10—P. lucka sp. nov. (protarsus, lateral view); 11—P. schillhammeri sp. nov. (protarsus, lateral view); 12—P. carolusi sp. nov. (mesotarsus, lateral view); 13—P. kaboureki sp. nov. (mesotarsus, lateral view); 14—P. kambaitiensis sp. nov. (mesotarsus, lateral view); 15—P. lucka sp. nov. (mesotarsus, lateral view); 16—P. schillhammeri sp. nov. (mesotarsus, lateral view); 17—P. kaboureki (metatarsus, dorsal view); 18—P. maculata (antenna, dorsal and lateral views); 19—P. carolusi sp. nov. (antenna, dorsal and lateral views); 20—P. lucka sp. nov. (antenna, dorsal view); 21—P. schillhammeri sp. nov. (antenna, dorsal view). Scale bars: 1.0 mm for Figs 89, 0.5 mm for Figs 1021." httpUri="https://zenodo.org/record/3386943/files/figure.png" pageId="12" pageNumber="530">11, 16, 21</figureCitation>
,
<figureCitation id="C6822A6351693233FEF8E85A2985FDBC" box="[369,398,571,598]" captionStart="FIGURES 3136" captionStartId="9.[151,264,1916,1941]" captionTargetBox="[172,1404,345,1835]" captionTargetId="figure@9.[151,1436,309,1891]" captionTargetPageId="9" captionText="FIGURES 3136. Male habitus. 31—Pyrrhalta carolusi sp. nov. (holotype, 4.3 mm); 32—P. kaboureki sp. nov. (holotype, 3.9 mm); 33—P. kambaitiensis sp. nov. (holotype, 4.5 mm); 34—P. lucka sp. nov. (holotype, 3.8 mm); 35—P. schillhammeri sp. nov. (holotype, 3.7 mm); 36—P. tsoui sp. nov. (paratype, 3.9 mm)." httpUri="https://zenodo.org/record/3386949/files/figure.png" pageId="12" pageNumber="530">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="16A3656D51693233FF1EE8E52CCEFCE0" pageId="12" pageNumber="530" type="materials_examined">
<paragraph id="5E0636E651693233FF1EE8E52C95FD74" blockId="12.[151,1437,643,2038]" box="[151,1182,643,670]" pageId="12" pageNumber="530">
<materialsCitation id="EED13CBB51693233FF1EE8E52C91FD74" ID-GBIF-Occurrence="2382692833" box="[151,1178,643,670]" country="Laos" county="Bolavens Plateau" location="Nam Thang" municipality="Muang" pageId="12" pageNumber="530" specimenCount="1" stateProvince="Champasak" typeStatus="holotype">
<emphasis id="6CCDEAF451693233FF1EE8E5293DFD74" bold="true" box="[151,310,643,670]" pageId="12" pageNumber="530">
<typeStatus id="8102884451693233FF1EE8E528D9FD74" box="[151,210,644,670]" pageId="12" pageNumber="530">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="26AE767651693233FEB5E8E5297EFD74" box="[316,373,644,670]" name="Laos" pageId="12" pageNumber="530">Laos</collectingCountry>
,
<collectingRegion id="9C7DF80451693233FE08E8E52A4DFD74" box="[385,582,644,670]" country="Laos" name="Champasak" pageId="12" pageNumber="530">Champasak prov.</collectingRegion>
,
<collectingCounty id="B7674E6A51693233FDD8E8E52B1EFD74" box="[593,789,644,670]" pageId="12" pageNumber="530">Bolavens Plateau</collectingCounty>
,
<collectingMunicipality id="BE62AC9C51693233FCA8E8E52B7BFD74" box="[801,880,644,670]" pageId="12" pageNumber="530">Muang</collectingMunicipality>
Paksong—B.
<location id="5B66603D51693233FB9BE8E52C91FD74" LSID="urn:lsid:plazi:treatment:D61087F051693232FF1EE8762C72FD89:5B66603D51693233FB9BE8E52C91FD74" box="[1042,1178,644,670]" country="Laos" county="Bolavens Plateau" municipality="Muang" name="Nam Thang" pageId="12" pageNumber="530" stateProvince="Champasak">Nam Thang</location>
</materialsCitation>
.
</paragraph>
<paragraph id="5E0636E651693233FF4EE8C92CCEFCE0" blockId="12.[151,1437,643,2038]" pageId="12" pageNumber="530">
<emphasis id="6CCDEAF451693233FF4EE8C92986FD28" bold="true" box="[199,397,679,706]" pageId="12" pageNumber="530">Types examined.</emphasis>
<materialsCitation id="EED13CBB51693233FE1AE8C92CCEFCE0" ID-GBIF-Occurrence="2382692849" collectingDate="1996-05-27" collectionCode="NMW" collectorName="Schillhammer &amp; The" elevation="950" location="Nam Thang" municipality="Bolavens-Plateau" pageId="12" pageNumber="530" specimenCount="1" specimenCount-male="1" stateProvince="Champasak" typeStatus="holotype">
<typeStatus id="8102884451693233FE1AE8C92A0BFD28" box="[403,512,680,706]" pageId="12" pageNumber="530" type="holotype">Holotype</typeStatus>
:
<specimenCount id="48BFFD6F51693233FD80E8C92A2BFD28" box="[521,544,680,706]" pageId="12" pageNumber="530" type="male"></specimenCount>
, “S-LAOS: Prov.
<collectingRegion id="9C7DF80451693233FD61E8C92B60FD28" box="[744,875,680,706]" country="Laos" name="Champasak" pageId="12" pageNumber="530">Champasak</collectingRegion>
/
<collectingMunicipality id="BE62AC9C51693233FCF6E8C92C48FD28" box="[895,1091,680,706]" pageId="12" pageNumber="530">Bolavens-Plateau</collectingMunicipality>
/ M. Paksong—B.
<location id="5B66603D51693233FA90E8C92D96FD28" LSID="urn:lsid:plazi:treatment:D61087F051693232FF1EE8762C72FD89:5B66603D51693233FA90E8C92D96FD28" box="[1305,1437,680,706]" municipality="Bolavens-Plateau" name="Nam Thang" pageId="12" pageNumber="530" stateProvince="Champasak">Nam Thang</location>
/ 26./
<date id="2A07102651693233FF47E8AA294FFD0C" box="[206,324,715,742]" pageId="12" pageNumber="530" value="1996-05-27">
<collectingDate id="3A43E9CE51693233FF47E8AA294FFD0C" box="[206,324,715,742]" pageId="12" pageNumber="530" value="1996-05-27">27.5.1996</collectingDate>
</date>
,
<quantity id="99419B0351693233FEC6E8AD29DFFD0C" box="[335,468,716,742]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="12" pageNumber="530" unit="m" value="950.0" valueMax="1100.0" valueMin="800.0">
<elevation id="D594D1D551693233FEC6E8AD29DFFD0C" box="[335,468,716,742]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="12" pageNumber="530" unit="m" value="950.0" valueMax="1100.0" valueMin="800.0">8001100m</elevation>
</quantity>
/ leg.
<collectorName id="F34C533051693233FD93E8AD2ABCFD0C" box="[538,695,716,742]" pageId="12" pageNumber="530">Schillhammer</collectorName>
(6) [w, p]” (
<collectionCode id="38A8AE2351693233FCCAE8AD2B9AFD0C" box="[835,913,716,742]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="530">NMW</collectionCode>
).
<collectorName id="F34C533051693233FC2AE8AD2BC4FD0C" box="[931,975,716,742]" pageId="12" pageNumber="530">The</collectorName>
<typeStatus id="8102884451693233FC5FE8AD2C33FD0C" box="[982,1080,716,742]" pageId="12" pageNumber="530" type="holotype">holotype</typeStatus>
is provided with one additional printed red label: “
<typeStatus id="8102884451693233FEEEE8912A1AFCE0" box="[359,529,752,778]" pageId="12" pageNumber="530" type="holotype">HOLOTYPUS</typeStatus>
, /
<taxonomicName id="99B94D6551693233FDA4E8912A92FCE0" authorityName="Joannis" authorityYear="1865" box="[557,665,752,778]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="530" phylum="Arthropoda" rank="genus">
<emphasis id="6CCDEAF451693233FDA4E8912A92FCE0" box="[557,665,752,778]" italics="true" pageId="12" pageNumber="530">Pyrrhalta</emphasis>
</taxonomicName>
/
<taxonomicName id="99B94D6551693233FD26E8912B44FCE0" authority="Bezděk &amp; Lee, 2019" authorityName="Bezděk &amp; Lee" authorityYear="2019" box="[687,847,752,778]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="530" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
<emphasis id="6CCDEAF451693233FD26E8912B44FCE0" box="[687,847,752,778]" italics="true" pageId="12" pageNumber="530">schillhammeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77FE578F51693233FCDFE8912BA4FCE0" box="[854,943,752,778]" pageId="12" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
, / Bezděk &amp; Lee, 2019”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="16A3656D51693232FF4EE972299AFF11" lastPageId="13" lastPageNumber="531" pageId="12" pageNumber="530" type="description">
<paragraph id="5E0636E651693233FF4EE9722AB1FCC5" blockId="12.[151,1437,643,2038]" box="[199,698,787,815]" pageId="12" pageNumber="530">
<emphasis id="6CCDEAF451693233FF4EE9722952FCC4" bold="true" box="[199,345,787,814]" pageId="12" pageNumber="530">Description.</emphasis>
Body length:
<specimenCount id="48BFFD6F51693233FE71E9752A06FCC4" box="[504,525,788,814]" pageId="12" pageNumber="530" type="male"></specimenCount>
3.7 (
<typeStatus id="8102884451693233FDCFE9752AA4FCC5" box="[582,687,788,815]" pageId="12" pageNumber="530" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="5E0636E651693233FF4EE9592933FBC0" blockId="12.[151,1437,643,2038]" pageId="12" pageNumber="530">
Male (
<typeStatus id="8102884451693233FE98E959297CFCB8" box="[273,375,824,850]" pageId="12" pageNumber="530" type="holotype">holotype</typeStatus>
,
<figureCitation id="C6822A6351693233FE0DE95929D2FCB8" box="[388,473,823,850]" captionStart="FIGURES 3136" captionStartId="9.[151,264,1916,1941]" captionTargetBox="[172,1404,345,1835]" captionTargetId="figure@9.[151,1436,309,1891]" captionTargetPageId="9" captionText="FIGURES 3136. Male habitus. 31—Pyrrhalta carolusi sp. nov. (holotype, 4.3 mm); 32—P. kaboureki sp. nov. (holotype, 3.9 mm); 33—P. kambaitiensis sp. nov. (holotype, 4.5 mm); 34—P. lucka sp. nov. (holotype, 3.8 mm); 35—P. schillhammeri sp. nov. (holotype, 3.7 mm); 36—P. tsoui sp. nov. (paratype, 3.9 mm)." httpUri="https://zenodo.org/record/3386949/files/figure.png" pageId="12" pageNumber="530">Fig. 35</figureCitation>
). Head pale brown with middle part of vertex black. Pronotum pale brown with one elongate median and two round lateral black spots. Antennae pale brown with darkened antennomeres IV-VII. Scu- tellum black. Elytra pale brown, each elytron with five brownish-black spots: one small round spot between humerus and scutellum, not touching anterior elytral margin; two round spots along lateral elytral margin; two elongate spots along suture. Legs pale brown. Ventral side: prosternum pale brown, mesoventrite black with brown middle part, metaventrite black with brown posterior margin, abdomen pale brown with lateral parts of each abdominal ventrite black.
</paragraph>
<paragraph id="5E0636E651693233FF4EEE552A8EFA84" blockId="12.[151,1437,643,2038]" pageId="12" pageNumber="530">
Head. Labrum transverse with rounded anterior margin, surface with transverse row of several punctures bearing long pale setae. Anterior clypeal margin weakly concave. Anterior part of head semiopaque, sparsely covered with punctures. Interantennal space narrow, 0.6 times as wide as transverse diameter of antennal insertion. Interocular space 2.5 as wide as transverse diameter of eye. Frontal tubercles subtriangular semiopaque, covered with microsculpture, separated each other by thin sulcus, laterally demarcated by thin furrow, posterior margins continuous with vertex. Vertex dull, covered with microsculpture, small punctures and short setae. Antennae (
<figureCitation id="C6822A6351693233FACBEE892D9FFAE8" box="[1346,1428,1256,1282]" captionStart="FIGURES 821" captionStartId="3.[151,264,1668,1693]" captionTargetBox="[188,1421,199,1621]" captionTargetId="figure@3.[151,1436,181,1643]" captionTargetPageId="3" captionText="FIGURES 821. Details of Pyrrhalta males. 8—P. kaboureki (last abdominal ventrite); 9—P. lucka sp. nov. (last abdominal ventrite); 10—P. lucka sp. nov. (protarsus, lateral view); 11—P. schillhammeri sp. nov. (protarsus, lateral view); 12—P. carolusi sp. nov. (mesotarsus, lateral view); 13—P. kaboureki sp. nov. (mesotarsus, lateral view); 14—P. kambaitiensis sp. nov. (mesotarsus, lateral view); 15—P. lucka sp. nov. (mesotarsus, lateral view); 16—P. schillhammeri sp. nov. (mesotarsus, lateral view); 17—P. kaboureki (metatarsus, dorsal view); 18—P. maculata (antenna, dorsal and lateral views); 19—P. carolusi sp. nov. (antenna, dorsal and lateral views); 20—P. lucka sp. nov. (antenna, dorsal view); 21—P. schillhammeri sp. nov. (antenna, dorsal view). Scale bars: 1.0 mm for Figs 89, 0.5 mm for Figs 1021." httpUri="https://zenodo.org/record/3386943/files/figure.png" pageId="12" pageNumber="530">Fig. 21</figureCitation>
) subclavate, 0.62 times as long as body, length ratios of antennomeres IXI equal to
<date id="2A07102651693233FBDAEF6A2CC7FACF" box="[1107,1228,1291,1317]" pageId="12" pageNumber="530" value="2013-06-08" valueMax="2013-08-06" valueMin="2013-06-08">13-6-8-8-6</date>
-6-7-
<date id="2A07102651693233FA8DEF6A2D63FACF" box="[1284,1384,1291,1317]" pageId="12" pageNumber="530" value="2013-06-07" valueMax="2013-06-07" valueMin="2013-06-07">6-7-7-13</date>
, last five antennomeres distinctly dilated, antennomeres VIII-X as wide as long, XI twice as long as wide, antennomeres VIII-XI distinctly flattened on ventral sides.
</paragraph>
<paragraph id="5E0636E651693233FF4EEF192C6FF9AC" blockId="12.[151,1437,643,2038]" pageId="12" pageNumber="530">Pronotum transverse, 1.77 times as wide as long, widest at anterior third. Anterior margin widely shallowly concave, lateral margins obtuse angulate with widely rounded middle part, anteriorly and posteriorly convergent and nearly straight, posterior margin weakly concave, lateral part of posterior margin oblique and concave. All margins thinly bordered. Anterior and posterior angles toothed and bearing long pale seta. Surface with distinct elevation along anterior margin, laterally curved posteriorly, disc transversely impressed. Elevation almost glabrous, with irregular small lustrous parts, disc densely covered with small punctures and short setae.</paragraph>
<paragraph id="5E0636E651693233FF4EEC312988F964" blockId="12.[151,1437,643,2038]" pageId="12" pageNumber="530">Scutellum subtriangular, with widely rounded apex, surface densely covered with microsculpture, small punctures and short setae.</paragraph>
<paragraph id="5E0636E651693233FF4EECF92D3FF8F4" blockId="12.[151,1437,643,2038]" pageId="12" pageNumber="530">Elytra moderately convex, parallel, 0.72 as long as body, 1.62 times as long as wide, dull, covered with small confused punctures with interspaces as wide as or narrower than puncture diameters and dense, short pale setae, surface with weakly impressed subscutellar area, rest of surface even. Humeral calli well developed. Epipleura wide, in apical quarter gradually narrowing towards apex, densely covered with short pale setae. Macropterous.</paragraph>
<paragraph id="5E0636E651693233FF4EED492D88F8A8" blockId="12.[151,1437,643,2038]" box="[199,1411,1832,1858]" pageId="12" pageNumber="530">Ventrally semiopaque, covered with longer pale setae. Last abdominal ventrite with deep U-shaped incision.</paragraph>
<paragraph id="5E0636E651693233FF4EED2D2D03F81C" blockId="12.[151,1437,643,2038]" pageId="12" pageNumber="530">
Legs moderately narrow. Protarsi: protarsomeres I and II subtriangular, short, protarsomere I concave in lateral view, basally with distinct tooth (
<figureCitation id="C6822A6351693233FD9BED112A6FF860" box="[530,612,1904,1930]" captionStart="FIGURES 821" captionStartId="3.[151,264,1668,1693]" captionTargetBox="[188,1421,199,1621]" captionTargetId="figure@3.[151,1436,181,1643]" captionTargetPageId="3" captionText="FIGURES 821. Details of Pyrrhalta males. 8—P. kaboureki (last abdominal ventrite); 9—P. lucka sp. nov. (last abdominal ventrite); 10—P. lucka sp. nov. (protarsus, lateral view); 11—P. schillhammeri sp. nov. (protarsus, lateral view); 12—P. carolusi sp. nov. (mesotarsus, lateral view); 13—P. kaboureki sp. nov. (mesotarsus, lateral view); 14—P. kambaitiensis sp. nov. (mesotarsus, lateral view); 15—P. lucka sp. nov. (mesotarsus, lateral view); 16—P. schillhammeri sp. nov. (mesotarsus, lateral view); 17—P. kaboureki (metatarsus, dorsal view); 18—P. maculata (antenna, dorsal and lateral views); 19—P. carolusi sp. nov. (antenna, dorsal and lateral views); 20—P. lucka sp. nov. (antenna, dorsal view); 21—P. schillhammeri sp. nov. (antenna, dorsal view). Scale bars: 1.0 mm for Figs 89, 0.5 mm for Figs 1021." httpUri="https://zenodo.org/record/3386943/files/figure.png" pageId="12" pageNumber="530">Fig. 11</figureCitation>
), length ratios of protarsomeres IIV equal to 6-6-4-7. Mesotarsi: meso- tarsomere I narrow, subparallel, in lateral view axe-shaped, with narrow basal part and expanded apical two thirds, angles of expanded parts pointed (
<figureCitation id="C6822A6351693233FD9FEDD92A63F838" box="[534,616,1975,2002]" captionStart="FIGURES 821" captionStartId="3.[151,264,1668,1693]" captionTargetBox="[188,1421,199,1621]" captionTargetId="figure@3.[151,1436,181,1643]" captionTargetPageId="3" captionText="FIGURES 821. Details of Pyrrhalta males. 8—P. kaboureki (last abdominal ventrite); 9—P. lucka sp. nov. (last abdominal ventrite); 10—P. lucka sp. nov. (protarsus, lateral view); 11—P. schillhammeri sp. nov. (protarsus, lateral view); 12—P. carolusi sp. nov. (mesotarsus, lateral view); 13—P. kaboureki sp. nov. (mesotarsus, lateral view); 14—P. kambaitiensis sp. nov. (mesotarsus, lateral view); 15—P. lucka sp. nov. (mesotarsus, lateral view); 16—P. schillhammeri sp. nov. (mesotarsus, lateral view); 17—P. kaboureki (metatarsus, dorsal view); 18—P. maculata (antenna, dorsal and lateral views); 19—P. carolusi sp. nov. (antenna, dorsal and lateral views); 20—P. lucka sp. nov. (antenna, dorsal view); 21—P. schillhammeri sp. nov. (antenna, dorsal view). Scale bars: 1.0 mm for Figs 89, 0.5 mm for Figs 1021." httpUri="https://zenodo.org/record/3386943/files/figure.png" pageId="12" pageNumber="530">Fig. 16</figureCitation>
), length ratios of mesotarsomeres IIV equal to 7-6-4-7. Metatarsi: meta- tarsomere I narrow, subtriangular, length ratios of metatarsomeres IIV equal to 7-6-5-8. Claws bifid.
</paragraph>
<paragraph id="5E0636E651683232FF4EEAF82AF0FF3D" blockId="13.[151,1437,153,611]" pageId="13" pageNumber="531">
Aedeagus (
<figureCitation id="C6822A6351683232FECCEAF82982FF5E" box="[325,393,153,180]" captionStart="FIGURES 17" captionStartId="2.[151,264,1865,1890]" captionTargetBox="[175,1421,255,1817]" captionTargetId="figure@2.[164,1423,234,1837]" captionTargetPageId="2" captionText="FIGURES 17. Aedeagus. 1—Pyrrhalta maculata (basal half in dorsal view, apical half in dorsal view, lateral view); 2—P. carolusi sp. nov. (dorsal and lateral views); 3—P. kaboureki sp. nov. (dorsal, lateral and ventral views); 4—P. kambaitiensis sp. nov. (basal part in dorsal view, apex in dorsal view, lateral view); 5—P. lucka sp. nov. (dorsal and lateral views); 6—P. schillhammeri sp. nov. (dorsal and lateral views); 7—P. tsoui sp. nov. (basal part in dorsal view, apex in dorsal view, lateral view). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/3386941/files/figure.png" pageId="13" pageNumber="531">Fig. 6</figureCitation>
) relatively narrow, with apical fifth curved left and tip straight, right side of aedeagus nearly straight, subapical part of left side broadly emarginate.
</paragraph>
<paragraph id="5E0636E651683232FF4EEA80299AFF11" blockId="13.[151,1437,153,611]" box="[199,401,225,251]" pageId="13" pageNumber="531">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="16A3656D51683232FF4EEB652A7EFDF6" pageId="13" pageNumber="531" type="diagnosis">
<paragraph id="5E0636E651683232FF4EEB652A7EFDF6" blockId="13.[151,1437,153,611]" pageId="13" pageNumber="531">
<emphasis id="6CCDEAF451683232FF4EEB6529C4FEF5" bold="true" box="[199,463,260,287]" pageId="13" pageNumber="531">Differential diagnosis.</emphasis>
The habitus of
<taxonomicName id="99B94D6551683232FD0DEB642B92FEF5" authority="Bezděk &amp; Lee, 2019" authorityName="Bezděk &amp; Lee" authorityYear="2019" box="[644,921,261,287]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="531" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
<emphasis id="6CCDEAF451683232FD0DEB642B92FEF5" box="[644,921,261,287]" italics="true" pageId="13" pageNumber="531">Pyrrhalta schillhammeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77FE578F51683232FC28EB642BF2FEF5" box="[929,1017,261,287]" pageId="13" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
is similar to that of
<taxonomicName id="99B94D6551683232FB6AEB672D36FEF5" authority="Bezděk &amp; Lee, 2019" authorityName="Bezděk &amp; Lee" authorityYear="2019" box="[1251,1341,261,287]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="531" phylum="Arthropoda" rank="species" species="lucka" status="sp. nov.">
<emphasis id="6CCDEAF451683232FB6AEB672D36FEF5" box="[1251,1341,261,287]" italics="true" pageId="13" pageNumber="531">P. lucka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77FE578F51683232FACCEB642D96FEF5" box="[1349,1437,261,287]" pageId="13" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
The males of both species possess subclavate antennae and modified ventral sides of pro- and mesotarsomeres I. Both species differ in the structure of aedeagus. The aedeagus of
<taxonomicName id="99B94D6551683232FCF0EB2F2C3EFE8D" authority="Bezděk &amp; Lee, 2019" authorityName="Bezděk &amp; Lee" authorityYear="2019" box="[889,1077,333,359]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="531" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
<emphasis id="6CCDEAF451683232FCF0EB2F2C3EFE8D" box="[889,1077,333,359]" italics="true" pageId="13" pageNumber="531">P. schillhammeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77FE578F51683232FBB4EB2C2C9FFE8D" box="[1085,1172,333,359]" pageId="13" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
has the apical half narrow and slightly curved left while the aedeagus of
<taxonomicName id="99B94D6551683232FD40EB132B2AFE61" authority="Bezděk &amp; Lee, 2019" authorityName="Bezděk &amp; Lee" authorityYear="2019" box="[713,801,369,395]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="531" phylum="Arthropoda" rank="species" species="lucka" status="sp. nov.">
<emphasis id="6CCDEAF451683232FD40EB132B2AFE61" box="[713,801,369,395]" italics="true" pageId="13" pageNumber="531">P. lucka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77FE578F51683232FCAEEB102B75FE61" box="[807,894,369,395]" pageId="13" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
has an elongate, flat, lanceolate apical half (
<figureCitation id="C6822A6351683232FAE3EB1028CEFE45" captionStart="FIGURES 17" captionStartId="2.[151,264,1865,1890]" captionTargetBox="[175,1421,255,1817]" captionTargetId="figure@2.[164,1423,234,1837]" captionTargetPageId="2" captionText="FIGURES 17. Aedeagus. 1—Pyrrhalta maculata (basal half in dorsal view, apical half in dorsal view, lateral view); 2—P. carolusi sp. nov. (dorsal and lateral views); 3—P. kaboureki sp. nov. (dorsal, lateral and ventral views); 4—P. kambaitiensis sp. nov. (basal part in dorsal view, apex in dorsal view, lateral view); 5—P. lucka sp. nov. (dorsal and lateral views); 6—P. schillhammeri sp. nov. (dorsal and lateral views); 7—P. tsoui sp. nov. (basal part in dorsal view, apex in dorsal view, lateral view). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/3386941/files/figure.png" pageId="13" pageNumber="531">Figs 56</figureCitation>
). The additional similar species,
<taxonomicName id="99B94D6551683232FDB0EBF72A80FE45" authority="Bezděk &amp; Lee, 2019" authorityName="Bezděk &amp; Lee" authorityYear="2019" box="[569,651,406,431]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="531" phylum="Arthropoda" rank="species" species="tsoui" status="sp. nov.">
<emphasis id="6CCDEAF451683232FDB0EBF72A80FE45" box="[569,651,406,431]" italics="true" pageId="13" pageNumber="531">P. tsoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77FE578F51683232FD1AEBF42AE0FE45" box="[659,747,405,431]" pageId="13" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
from
<collectingCountry id="26AE767651683232FCB9EBF42B89FE45" box="[816,898,405,431]" name="Taiwan" pageId="13" pageNumber="531">Taiwan</collectingCountry>
and
<collectingCountry id="26AE767651683232FC33EBF42BF8FE45" box="[954,1011,405,431]" name="Laos" pageId="13" pageNumber="531">Laos</collectingCountry>
, has both elytral spots near the suture round (elongate in
<taxonomicName id="99B94D6551683232FEE7EBDB2A21FE39" authority="Bezděk &amp; Lee, 2019" authorityName="Bezděk &amp; Lee" authorityYear="2019" box="[366,554,441,467]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="531" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
<emphasis id="6CCDEAF451683232FEE7EBDB2A21FE39" box="[366,554,441,467]" italics="true" pageId="13" pageNumber="531">P. schillhammeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77FE578F51683232FDBBEBD82A86FE39" box="[562,653,441,467]" pageId="13" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
), antennae without distinctly enlarged apical antennomeres (antennae subclavate in
<taxonomicName id="99B94D6551683232FEBBEBBF29E5FE1D" authority="Bezděk &amp; Lee, 2019" authorityName="Bezděk &amp; Lee" authorityYear="2019" box="[306,494,477,503]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="531" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
<emphasis id="6CCDEAF451683232FEBBEBBF29E5FE1D" box="[306,494,477,503]" italics="true" pageId="13" pageNumber="531">P. schillhammeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77FE578F51683232FE7CEBBC2A5BFE1D" box="[501,592,477,503]" pageId="13" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
), and ventral sides of pro- and mesotarsomeres I not modified (modified in
<taxonomicName id="99B94D6551683232FF1EE8632958FDF1" authority="Bezděk &amp; Lee, 2019" authorityName="Bezděk &amp; Lee" authorityYear="2019" box="[151,339,513,539]" class="Insecta" family="Chrysomelidae" genus="Pyrrhalta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="531" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
<emphasis id="6CCDEAF451683232FF1EE8632958FDF1" box="[151,339,513,539]" italics="true" pageId="13" pageNumber="531">P. schillhammeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="77FE578F51683232FED3E86029B8FDF1" box="[346,435,513,539]" pageId="13" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
, see
<figureCitation id="C6822A6351683232FE60E8602A63FDF6" box="[489,616,513,540]" captionStart="FIGURES 821" captionStartId="3.[151,264,1668,1693]" captionTargetBox="[188,1421,199,1621]" captionTargetId="figure@3.[151,1436,181,1643]" captionTargetPageId="3" captionText="FIGURES 821. Details of Pyrrhalta males. 8—P. kaboureki (last abdominal ventrite); 9—P. lucka sp. nov. (last abdominal ventrite); 10—P. lucka sp. nov. (protarsus, lateral view); 11—P. schillhammeri sp. nov. (protarsus, lateral view); 12—P. carolusi sp. nov. (mesotarsus, lateral view); 13—P. kaboureki sp. nov. (mesotarsus, lateral view); 14—P. kambaitiensis sp. nov. (mesotarsus, lateral view); 15—P. lucka sp. nov. (mesotarsus, lateral view); 16—P. schillhammeri sp. nov. (mesotarsus, lateral view); 17—P. kaboureki (metatarsus, dorsal view); 18—P. maculata (antenna, dorsal and lateral views); 19—P. carolusi sp. nov. (antenna, dorsal and lateral views); 20—P. lucka sp. nov. (antenna, dorsal view); 21—P. schillhammeri sp. nov. (antenna, dorsal view). Scale bars: 1.0 mm for Figs 89, 0.5 mm for Figs 1021." httpUri="https://zenodo.org/record/3386943/files/figure.png" pageId="13" pageNumber="531">Figs 11, 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="16A3656D51683232FF4EE84529AEFDD5" box="[199,421,548,575]" pageId="13" pageNumber="531" type="distribution">
<paragraph id="5E0636E651683232FF4EE84529AEFDD5" blockId="13.[151,1437,153,611]" box="[199,421,548,575]" pageId="13" pageNumber="531">
<emphasis id="6CCDEAF451683232FF4EE845296AFDD5" bold="true" box="[199,353,548,575]" pageId="13" pageNumber="531">Distribution.</emphasis>
<collectingCountry id="26AE767651683232FEEEE84429AAFDD5" box="[359,417,549,575]" name="Laos" pageId="13" pageNumber="531">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="16A3656D51683232FF4EE8282C72FD89" box="[199,1145,585,611]" pageId="13" pageNumber="531" type="etymology">
<paragraph id="5E0636E651683232FF4EE8282C72FD89" blockId="13.[151,1437,153,611]" box="[199,1145,585,611]" pageId="13" pageNumber="531">
<emphasis id="6CCDEAF451683232FF4EE8282945FD89" bold="true" box="[199,334,585,611]" pageId="13" pageNumber="531">Etymology.</emphasis>
Dedicated to Harald Schillhammer (NMW) who collected the
<typeStatus id="8102884451683232FB86E8282C7EFD89" box="[1039,1141,585,611]" pageId="13" pageNumber="531" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>